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DE GENERE TURBINICARPUS F. BUXB. ET BACKEB. (CACTACEA)

A. B. Doweld, National Institute of Catpology (Gaertnerian Institution), Moscow, Russia

The species delimitation in the genus Turbinicarpus remains mostly unresolved, although extensive field and morphological data have been provided (Anderson, 1986). The creation of a distinct genus Kadenicarpus Doweld (1998) to accommodate anomalous southern species, T pseudomacrochele (Backeb.) F. Buxb. & Backeb .. questions the taxonomic position of another southern species, T laui Glass & Foster, which has a similar type of gross seed morphology anp spination (cf Anderson. 1986: 22). However, the fine structure of its seed cuticular coats is reported by Glass and Foster (1977) as papillate and radially-striate (cf fig. 45 in Glass & Foster, I. e.). Such a type of cuticular sculpturing is similar to that of sect. Normanbokea (T. valdezianus. T pseudopectinatus), and is different from that in sect. Turbinicarpus. The somewhat structure of the seed cuticular sculpturing is noticed within the genus Gymnocactus treated as a distinct sister generic entity. A detailed seed morphological and taxonomic survey would be presented elsewhere. The species, T pailanus Halda & Panarotto, on the basis of quite another seed-coat morphology has been placed into Neol/oydia. This is an additional step in the amplification of the genus Neo//oydia (Doweld, 1999,2000). The species of the genus are re-classified as follows (genera Bravocactus Doweld (1998), Kadenicarpus Doweld (1998) and Gymnocactus Backeb. (1938) are accepted as distinct): Thrbinicarpus F. Buxb. & Backeb. 1937, Cactaceae (Berlin), I : 27. 1969, in Krainz, Kakteen, 40-41 : sine pag. [CVIIIb]. -

Normanbokea Kladiwa & F. Buxb.

Pseudosolisia Y. Ito, 1981, Cactaeeae: 477, nom.

il/eg. T y pus: T schmiedickeanus (Bod.) F. Buxb. & Baekeb. 9 species:

Sect. I. Normanbokea (Kladiwa & F. Buxb.) Doweld, 1999, Sukkulenty, I : 27. T Y pus: T valdezianus (Moll.) Glass & Foster. 3 species: Ser. I. Valdeziani Doweld, ser. nov. - Plantae habitu Solisiae, aculeis centralibus nul/is, radialibus pluribus (25-

50), semina ornamenlatione cutciularis radialihus-striatibus. T y pus: T valdezianus (Moll.) Glass & Foster. l. T valdezianus (Moll.) Glass & Foster, 1977, Caet. Succ.J. (Los Angeles), 49. 4: 174. 2. T pseudopectinatus (Backeb.) Glass & Foster, 1977, Cact. Succ. 1. (Los Angeles), 49,4: 175.
3. T. x mombergeri Riha, 1996, Kaktusy, 32, 2: 54. Ser. 2. Lauani Doweld, ser. nov. - Plantae habitu Kadenicarpi, solitariae, aculeis radialibus 6-8, centralibus

absentis, seminibus ornamentatio cuticularis radialihus-striatibus.

Ty pus: T laui Glass & Foster. 4. T laui Glass & Foster, 1975, Caet. Suee. J. (Los Angeles), 47, 3: 116. Sect. 2. Turbinicarpus T y pus (as for genus): T schmiedickeanus (Bod.) F. Buxb. & Backeb. 5 species:

1. T gracilis Glass & Foster, 1976, Cact. Suce. J. (Los Angeles), 48, 4: 176.
1 a. ssp.gracilis lb. ssp. dickisoniae (Glass & Foster) Panarotto, 1999, Piante Grasse, 18,3 : 120.

2. T schmiedickeanus (Bod.) F. Buxb. & Backeb. 3. T macrochele (Werderm.) F. Buxb. & Backeb.
3a. ssp. macrochele - Neotypus (Anderson, 1986: 20): E. F. Anderson 4940 (US). 3b. ssp.flaviflorus (G. R. W. Frank et Lau) Doweld, stat. nov.

= Turbinicarpusflaviflorus G. R. W. Frank et Lau,

1979, Kakteen Sukk. 30,1: 6-

7. -

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Holotypus,- Lau 1185 (ZSS). 3c. ssp. schwarz;; (Shurly) Doweld, comb. et stat. nov. == Stromhocactus schwarzii Shuriy, 1948, Cact. Succ. J. Grt. Brit. 10 : 93. - Holotypus: Shurly in 1948 (K). 3d. ssp. klinker;anus (Backeb. et H. Jacobs.) Doweld, stat. nov. = Turhinicarpus klinkerianus Backeb. et H. Jacobs. 1948, Sukkulentenk. 2: 27-28. Lectotypus (Anderson, 1986: 20): fig. in Backeb. et H. Jacobs. I.e. 3e. ssp. anderson;; (Mosco) Doweld, comb. nov. = Turhinicarpus schmiedickeanus ssp. andersonii Mosco, 1999, Cactaceae Cons. Init. 7 : 18. - Holotypus: E. F. Anderson 4932 (US).
4. T. a/onsoi Glass & S. Arias, 1996, Kakteen Sukk. 47, 2: 25. 5. T. holeri Liithy & Lau, 1991, Kakteen Sukk. 42, 2 : 34. Species ex/udendae: Neolloydia pai/ana (Halda & Panarotto) Doweld, comb. nov. = Turhinicarpus pai/anus Halda & Panarotto, 1998, Acta Mus. Richnov. 5,4: 162. - Holotypus: Panarotto 850477 (PR).

Literature Cited I. Anderson E.F.A revision of the genus Neolloydia B. & R. (Cactaceae) II Bradleya. 1986. Vol. 4. P. 1-28. 2. Doweld A. B. Amplification of the genus Neol/oydia Britton & Rose II Cactaceae Cons. Init. 1999. N 7. P. 1117. 3. Doweld A. B. Amplification of the genus Neolloydia Briton & Rose: addenda et corrigenda II Cactaceae Syst. Init. 2000. N 9. P. 19-23. 4. Glass C.. Foster R.A revision of the genus Turhinicarpus (Backbg.) Buxb. & Backbg. II Cact. Succ. J. (Los Angeles). 1977. Vol. 49. N 4. P. 161-176.

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