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List of Human Evolution Fossils
List of Human Evolution Fossils
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The following tables give a brief overview of several notable hominin fossil
finds relating to human evolution beginning with the formation of the
Hominini tribe in the late Miocene (roughly 6 million years ago).
Most of the fossils shown are not considered direct ancestors to Homo
sapiens but are closely related to direct ancestors and are therefore
important to the study of the lineage.
Contents
1 Undated
2 Late Miocene
2.1 7 million - 5.3 million years old
3 Pliocene
3.1 5.3 million - 2.58 million years old
4 Pleistocene
4.1 Lower Paleolithic: 2.58 million - 300,000
4.2 Middle Paleolithic: 300,000 - 50,000 years old
4.3 Upper Paleolithic: 50,000 - 10,000 years old
5 Holocene
5.1 Mesolithic / Neolithic: 10,000 - 5000 years old
6 Abbreviations used in fossil catalog name
7 Timelines
8 See also
9 Further reading
10 References
10.1 Bibliography
11 External links
Undated
Year Discov Now
Specie Countr
Name Age discove ered located
s y
red by at
Universi
Dinaledi Rick
ty of the
Chambe Hunter
Homo South Witwate
r Undated 2013 and
naledi Africa rsrand
hominin Steven
(South
s Tucker
Africa)
Late Miocene
7 million - 5.3 million years old
Pliocene
5.3 million - 2.58 million years old
s
Australo
KSD-VP-
pithecu Yohanne
1/1
3.58 Ma s 2005 Ethiopia s Haile-
(Kadanu
afarensi Selassie
umuu)
s
Mamelb
Australo
aye N'Djame
KT- pithecu
Tomalta na
12/H1 3.5 Ma s 1995 Chad
and (Chad),
(Abel) bahrelg
Michel BEAC
hazali
Brunet
Justus
KNM-WT
Kenyant Erus
40000 Lake
hropus and
(Flat 3.5 Ma 1999 Turkana,
platyop Meave
Faced Kenya
s Leakey[1
Man)[13] 4]
Australo Yohann
pithecu es
BRT-VP- 3.3 - 3.5
s 2015 Ethiopia Haile-
3/14 Ma
deyirem Selassie[
eda 15]
Pleistocene
Lower Paleolithic: 2.58 million - 300,000
OH 24
Homo Tanzani Peter
(Twiggy) 1.8 Ma 1968
[29]
habilis a Nzube
Homo Tanzani
OH 8[30] 1.8 Ma 1960
habilis a
OH 5 1.8 Ma Paranth 1959 Tanzani Mary
(Zinj or ropus a Leakey
nutcrac boisei
ker
man)
David
Lordkip
D2700
anidze
(Dmanis Homo
1.8 Ma 2001 Georgia and
i Skull erectus
Abesalo
3)
m
Vekua
D3444
David
(Dmanis Homo
1.8 Ma 2003 Georgia Lordkip
i Skull erectus
anidze
4)
D4500 2005
David
(Dmanis Homo (publish
1.8 Ma Georgia Lordkip
i Skull erectus ed in
anidze
5) 2013)
1.78 Homo Koobi Meave
KNM-ER
-1.9 rudolfen 2012 Fora, Leakey'
62000[31]
Mya sis Kenya s team
KNM-ER
62003
1.78 Homo Koobi Meave
(mandib
-1.9 rudolfen 2012 Fora, Leakey'
ular
Mya sis Kenya s team
fragmen
t)[31]
KNM-ER
1.78 Homo Koobi Meave
60000
-1.9 rudolfen 2012 Fora, Leakey'
(mandib
Mya sis Kenya s team
le)[31]
Jonatha
Homo Tanzani
OH 7 1.75 Ma 1960 n
habilis a
Leakey
Homo
ergaste
KNM ER Bernard
1.75 Ma r[32] (aka 1975 Kenya
3733 Ngeneo
Homo
erectus)
KNM ER Homo Paul
1.74 Ma 1973/4 Kenya
1805 habilis Abell
1.7 Ma /
0.5-0.6
Yuanmo Homo Fang
Ma 1965 China
u Man erectus Qian
(dispute
d)[33]
Paranth
KNM ER Richard
1.7 Ma ropus 1969 Kenya
406 Leakey
boisei
Paranth
KNM ER Richard
1.7 Ma ropus 1970 Kenya
732[34] Leakey
boisei
Paranth Koobi
KNM ER Benson
1.7 Ma ropus 1990 Fora,
23000[35] Kyongo
boisei Kenya
National
Museum
KNM WT Paranth
Not Not s of
17400[36] 1.7 Ma ropus Kenya
know[38] know[38] Kenya,
[37]
boisei
Nairobi
(Kenia)
Homo
KNM WT
ergaste Kenya
15000 Kamoya
1.6 Ma r (aka 1984 Kenya National
(Turkan Kimeu
Homo Museum
a Boy)
erectus)
Homo
1.5 -2 South A. R.
[39]
StW 53 gauteng 1976
Ma Africa Hughes
ensis
SK 1.5 - 2 Homo South
1949
847[40] Ma habilis Africa
Paranth
DNH 7
1.5 - 2 ropus South Andr
(Eurydic 1994
Ma robustu Africa Keyser
e)[41]
s
Paranth
1.5 - 1.8 ropus South Robert
SK 46[42] 1949
Ma robustu Africa Broom
s
Peninj Paranth
Tanzani Richard
Mandibl 1.5 Ma ropus 1964
a Leakey
e boisei
OH 9
(Chellea Homo Tanzani Louis
1.5 Ma 1960
n Man) erectus a Leakey
[43]
Homo
ergaste
KNM ER Richard
1.5 Ma r (aka 1971 Kenya
992 Leakey
Homo
erectus)
Mojoker Andojo,
to 1 G.H.R.
1.43 - Homo Indonesi
(Mojoke 1936 von
1.49 Ma erectus a
rto Koenigs
child) wald
KNM ER 1.4 - 1.6 Homo Richard
1976 Kenya
3883 Ma erectus Leakey
Paranth
KGA 10- A.
1.4 Ma ropus 1993 Ethiopia
525[44][45] Amzaye
boisei
Museo
de la
Atapuer
Eudald Evoluci
ca Homo
1.2 Ma 2008 Spain Carbone n
Jawbone sp.
[46]
ll Humana
, Burgos
(Spain)
Homo
1.1 Ma M. Cihat
Kocabas [47] erectus 2002 Turkey
[48]
Aliek
Homo Henry
Daka 1.0 Ma 1997 Ethiopia
erectus Gilbert
G.H.R.
Sangira Homo Indonesi von
1 Ma 1939
n4 erectus a Koenigs
wald
ATD6-15 Museo
and de la
ATD6-69 Bermd Evoluci
Nio de 780k - Homo
1994 Spain ez & n
la Gran 858k erectus
Arsuaga Humana
Dolina3 , Burgos
42 (Spain)
G.H.R.
Sangira 0.7 - 1.6 Homo Indonesi von
1937
n2 Ma erectus a Koenigs
wald
Trinil 2
Pitheca
nthropu
0.7 -1 Homo Indonesi Eugne
s-1 1891
Ma erectus a Dubois
or
Java
Man[49]
C.
Ternifine
Arambo
2-3 now Homo
700k 1954 Algeria urg & B.
Tighenn erectus
Hoffstet
if[50]
ter
Sangira Homo Indonesi S.
700k 1969
n 17[51] erectus a Sartono
Peking 680k - Homo 1921 China Davidso Lost/stol
Man 780k erectus n Black en
Homo
heidelb National
Madam 600k - ergensi Ernesto Museum
1997 Eritrea
Buya[52] 1.4 Ma s or Abbate of
Homo Eritrea
erectus
Homo
heidelb
ergensi A.
Bodo[53] 600k 1976 Ethiopia
s or Asfaw
Homo
erectus
Mauer 1 Homo
Daniel
(Heidelb heidelb German
500k 1907 Hartma
erg ergensi y
nn
Man) s
Homo
Saldanh South
500k rhodesi 1953
a man[54] Africa
ensis
Homo
Boxgrov 478k - heidelb
1994 UK
e Man 524k ergensi
s
Arago
Henry
21 Homo
450k 1971 France de
(Tautav erectus
Lumley
el Man)
Hexian[56 400k - Homo
[55]
1980 China
]
500k erectus
Homo
ceprane
nsis?
Homo
Argil
400 - anteces Ceprano Italo
Ceprano 1994
500k sor? , Italia Biddittu
Man
Homo
heidelb
ergensi
s?
Museo
Bermd
de la
Homo ez,
Skull 5 Evoluci
heidelb Arsuaga
(Miguel 400k 1992 Spain n
ergensi &
n) Humana
s Carbone
, Burgos
ll
(Spain)
Swansc 400k Homo 1935 UK Alvan T
heidelb
ombe
ergensi Marston
Man
s
Homo A
Sal[57][58]
[59]
400k rhodesi 1971 Morocco quarry
ensis worker
Homo
Tanzani A.A.
Ndutu[60] 350k rhodesi 1973
a Mturi
ensis
Homo
Steinhei heidelb German
350k 1933
m Skull ergensi y
s
Homo Mrton
Samu 350k 1964 Hungary
erectus Pcsi
Holocene
Mesolithic / Neolithic: 10,000 - 5000 years old
Timelines
Life timeline
view discuss edit
-4500
-4000
-3500
-3000
-2500
-2000
-1500
-1000
-500
water
Single-celled
life
photosynthesis
Eukaryotes
Multicellular
life
Land life
Dinosaurs
Mammals
Flowers
Earliest Earth (4540)
Earliest water
Earliest life
LHB meteorites
Earliest oxygen
Atmospheric oxygen
Oxygen crisis
Earliest sexual reproduction
Ediacara biota
Cambrian explosion
Earliest humans
P
h
a
n
e
r
o
z
o
i
c
P
r
o
t
e
r
o
z
o
i
c
A
r
c
h
e
a
n
H
a
d
e
a
n
Pongola
Huronian
Cryogenian
Andean
Karoo
Quaternary
-9
-8
-7
-6
-5
-4
-3
-2
-1
Human-like
apes
Nakalipithecus
Ouranopithecus
Sahelanthropus
Orrorin
Ardipithecus
Australopithecus
Homo habilis
Homo erectus
Neanderthal
Homo sapiens
Earlier apes
Possibly bipedal
Earliest bipedal
Earliest stone tools
Earliest exit
from Africa
Earliest fire use
Earliest cooking
Earliest clothes
Modern humans
P
l
e
i
s
t
o
c
e
n
e
P
l
i
o
c
e
n
e
M
i
o
c
e
n
e
s
Axis scale: millions of years.
Also see: Life timeline and Nature timeline
See also
Dawn of Humanity (2015 PBS film)
Human evolution
List of countries and islands by first human settlement
List of fossil primates
List of fossil sites
List of notable fossils
List of transitional fossils
Most recent common ancestor
Nature timeline
Paleoanthropology
Timeline of human evolution
Further reading
Gibbons, Ann. The First Human: The Race to Discover our Earliest
Ancestor. Anchor Books (2007). ISBN 978-1-4000-7696-3
Hartwig, Walter (ed.). The Primate Fossil Record. Cambridge University
Press (2002). Reprinted 2004. ISBN 978-0-521-08141-2.
Johanson, Donald & Wong, Kate. Lucy's Legacy: The Quest for Human
Origins. Three Rivers Press (2009). ISBN 978-0-307-39640-2
Jones, Steve; Martin, Robert D.; Pilbeam, David R (Editors). (1994). The
Cambridge Encyclopedia of Human evolution. Cambridge University
Press. ISBN 978-0-521-46786-5 (Note: this book contains very
useful, information dense chapters on primate evolution in general,
and human evolution in particular, including fossil history).
Leakey, Richard & Lewin, Roger. Origins Reconsidered: In Search of
What Makes us Human. Little, Brown and Company (1992). ISBN 0-
316-90298-5
Lewin, Roger. Bones of Contention: Controversies in the Search for
Human Origins. Penguin Books (1987). ISBN 0-14-022638-9
Morwood, Mike & van Oosterzee, Penny. A New Human: The Startling
Discovery and Strange Story of the 'Hobbits' of Flores, Indonesia.
Smithsonian Books (2007). ISBN 978-0-06-089908-0
Oppenheimer, Stephen. Out of Eden: The Peopling of the World.
Constable (2003). ISBN 1-84119-697-5
Roberts, Alice. The Incredible Human Journey: The Story of how we
Colonised the Planet. Bloomsbury (2009). ISBN 978-0-7475-9839-8
Shreeve, James. The Neanderthal Enigma: Solving the Mystery of
Modern Human Origins. Viking (1996). ISBN 0-670-86638-5
Stringer, Chris. The Origin of Our Species. Allen Lane (2011). ISBN 978-
1-84614-140-9
Stringer, Chris & Andrews, Peter. The Complete World of Human
Evolution. Thames & Hudson (2005). ISBN 0-500-05132-1
Stringer, Chris & McKie, Robin. African Exodus: The Origins of Modern
Humanity. Jonathan Cape (1996). ISBN 0-224-03771-4
van Oosterzee, Penny. The Story of Peking Man. Allen & Unwin (1999).
ISBN 1-86508-632-0
Walker, Allan & Shipman, Pat. The Wisdom of the Bones: In Search of
Human Origins. Weidenfeld and Nicholson (1996). ISBN 0-297-
81670-5
Wade, Nicholas. Before the Dawn: Recovering the Lost History of our
Ancestors. Penguin Press (2006). ISBN 978-0-7156-3658-9
Weiss, M.L., & Mann, A.E. (1985). "'Human Biology and Behaviour: An
anthropological perspective" (4th ed.). Boston: Little Brown.
ISBN 978-0-673-39013-4 (Note: this book contains very accessible
descriptions of human and non-human primates, their evolution,
and fossil history).
Wells, Spencer (2004). The Journey of Man : A Genetic Odyssey. New
York, NY: Random House Trade Paperbacks. ISBN 978-0-8129-7146-
0.
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Smithsonian
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Inverted strata
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Smithsonian
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Saldanha man[self-published source?] Archived July 11, 2007, at the Wayback
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Smithsonian
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Smithsonian's Human Origins Program
http://www.thelocal.it/20151104/italy-dates-its-first-neanderthal-to-250k-
years-ago
Chang, C.-H.; Kaifu, M.; Kona, R. T.; Grn, R.; Matsu'ura, S.; Kinsley, L.; Lin,
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Choi, Charles Q. (January 27, 2015). "Ancient Human Fossil Could Be New
Primitive Species". Live Science.
Smith TM, Tafforeau P, Reid DJ, et al. (April 2007). "Earliest evidence of
modern human life history in North African early Homo sapiens".
Proceedings of the National Academy of Sciences of the United States of
America. 104 (15): 612833. doi:10.1073/pnas.0700747104.
PMC 1828706. PMID 17372199. Lay summary Phys.org (March 13,
2007).
Smith TM, Tafforeau P, Reid DJ, et al. (December 2010). "Dental evidence
for ontogenetic differences between modern humans and Neanderthals".
Proceedings of the National Academy of Sciences of the United States of
America. 107 (49): 209238. doi:10.1073/pnas.1010906107.
PMC 3000267. PMID 21078988.
Zimmer, Carl (16 November 2015). "In a Tooth, DNA From Some Very Old
Cousins, the Denisovans". New York Times. Retrieved 16 November 2015.
Johanson, Donald; Edgar, Blake (2006). From Lucy to Language. Simon &
Schuster. ISBN 978-0-7432-8064-8.
Demeter, F.; Shackelford, L. L.; Bacon, A.-M.; Duringer, P.; Westaway, K.;
Sayavongkhamdy, T.; Braga, J.; Sichanthongtip, P.; Khamdalavong, P.;
Ponche, J.-L.; Wang, H.; Lundstrom, C.; Patole-Edoumba, E.; Karpoff, A.-M.
(2012). "Anatomically modern human in Southeast Asia (Laos) by 46 ka".
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http://www.modernhumanorigins.net/neanderthal1.html Neanderthal 1
images at Modern Human Origins
http://news.softpedia.com/newsImage/Experts-Recreate-the-Face-of-the-
First-European-2.jpg/ Images of Petera cu Oase[self-published source?]
Timeline at Bone & Stone
http://www.modernhumanorigins.net/minatogawa1.html Minatogawa 1
images at Modern Human
Template:Cita magazine
"Lake Tandou Skull". Australia: The Land Where Time Began. Retrieved
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McKie, Robin (June 20, 2010). "Bones from a Cheddar Gorge cave show
that cannibalism helped Britain's earliest settlers survive the ice age". The
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"Mystery of a West African skull from 13,000 years ago". Natural History
Museum, London, UK. Retrieved July 27, 2012.
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Retrieved August 21, 2013.
Storm, Paul; Nelson, Andrew (1992). "The many faces of Wadjak man".
Archaeology in Oceania. 27 (1): 3746. doi:10.1002/j.1834-
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Origin of Native Americans (Google Books). Cambridge University Press.
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1. Angel, J.L.; Phenice, T.W.; Robbins, L.H.; Lynch, B.M. (1980). Late stone
age fishermen of Lothagam, Kenya. National Anthropological
Archives, Sithsonian Institution, Part 3.
Bibliography
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