BIOX.

CN 2006-9-5

()

[5]

8600
300
[3]

4
2.1
Wynne-Edwards(1967)

[1]
2.2 [5]
(George Williams)
[8](Maynard Smith)[9]
[10]

l2 12
l 14

2.3 [5]
(Robert Axelrod)[11]

2.4 [5]

(docility)

()

3 4

[1] 2001
[2] 1992
[3] 2000 9
[4] 1999 6
[5] 2000 3
[6] 1991,10(2)4956
[7] () 1992 4
[8] Williams,G.C. Adapration and Nature Selection [M].Princeton:Princeton University
Press,1966
[9] Smith,Maynard J and G R Price.The Logic of Animal Confliers[J].Nature,Vol.246(1973).15-
18
[10] Hamilton,W H The Generical Evolution of Social Behaviour [J].Journal of Theoretical
Biology,Vol.7(1964).1-52
[11] Axelrod,Rolert,and William D Hamilion.The Evolution of Cooperation
[J].Science,Vol.211(1981).1890-1896