Giant Panda

You might also like

Download as pdf or txt
Download as pdf or txt
You are on page 1of 19

1/23/2019 Giant panda - Wikipedia

Giant panda
The giant panda (Ailuropoda melanoleuca, literally "black and white cat-
foot"; Chinese: 大熊猫; pinyin: dà xióng māo, literally "big bear cat"),[4] also
Giant panda
known as panda bear or simply panda, is a bear[5] native to south central
China.[1] It is easily recognized by the large, distinctive black patches around
its eyes, over the ears, and across its round body. The name "giant panda" is
sometimes used to distinguish it from the unrelated red panda. Though it
belongs to the order Carnivora, the giant panda's diet is over 99%
bamboo.[6] Giant pandas in the wild will occasionally eat other grasses, wild
tubers, or even meat in the form of birds, rodents, or carrion. In captivity,
they may receive honey, eggs, fish, yams, shrub leaves, oranges, or bananas Giant panda at the Ocean Park Hong
along with specially prepared food.[7][8]
Kong
The giant panda lives in a few mountain ranges in central China, mainly in Conservation status
Sichuan, but also in neighbouring Shaanxi and Gansu.[9] As a result of
farming, deforestation, and other development, the giant panda has been
driven out of the lowland areas where it once lived.

The giant panda is a conservation-reliant vulnerable species.[10][11] A 2007 Vulnerable (IUCN 3.1)[1]
report showed 239 pandas living in captivity inside China and another 27 Scientific classification
outside the country.[12] As of December 2014, 49 giant pandas lived in
Kingdom: Animalia
captivity outside China, living in 18 zoos in 13 different countries.[13] Wild
population estimates vary; one estimate shows that there are about 1,590 Phylum: Chordata
individuals living in the wild,[12] while a 2006 study via DNA analysis Class: Mammalia
estimated that this figure could be as high as 2,000 to 3,000.[14] Some
reports also show that the number of giant pandas in the wild is on the
Order: Carnivora
rise.[15] In March 2015, Mongabay stated that the wild giant panda Family: Ursidae
population had increased by 268, or 16.8%, to 1,864.[16] In 2016, the IUCN
Genus: Ailuropoda
reclassified the species from "endangered" to "vulnerable".[11]
Species: A. melanoleuca
While the dragon has often served as China's national symbol,
Binomial name
internationally the giant panda appears at least as commonly. As such, it is
becoming widely used within China in international contexts, for example Ailuropoda melanoleuca
since 1982 issuing gold panda bullion coins or as one of the five Fuwa David, 1869[2]
mascots of the Beijing Olympics.
Subspecies

†A. m. hastorni
Contents A. m. melanoleuca
Taxonomy A. m. qinlingensis
Classification
Etymology
Subspecies

https://en.wikipedia.org/wiki/Giant_panda 1/19
1/23/2019 Giant panda - Wikipedia

Description
Pathology
Genomics
Ecology
Diet
Predators
Behavior
Reproduction
Uses and human interaction
Early references Giant panda range
Western discovery
Panda diplomacy Giant panda
Biofuel
Conservation
In zoos
Population chart
Reference in medicine
In cryptozoology
"Panda" in Traditional (top) and
See also
Simplified (bottom) Chinese
References
characters[3]
External links
Traditional Chinese 熊貓
Simplified Chinese 熊猫
Taxonomy Literal meaning "bear cat"
Transcriptions
Classification Standard Mandarin
For many decades, the precise taxonomic classification of the giant panda was Hanyu Pinyin xióngmāo
under debate because it shares characteristics with both bears and raccoons.[17] Wade–Giles hsiung2-mao1
However, molecular studies indicate the giant panda is a true bear, part of the IPA [ɕjʊ̌ŋ.máu]
family Ursidae.[5][18] These studies show it differentiated early (about 19
Wu
million years ago)[19] from the main ursine stock; since it is the most basal
Suzhounese yón-mau
member of the group, it is equidistant from all other extant ursids.[20][19] The
giant panda has been referred to as a living fossil.[21] Yue: Cantonese
Yale Romanization hùhng-māau
Despite the shared name, habitat type, and diet, as well as a unique enlarged
IPA [hȍŋ.máːu]
bone called the pseudo thumb (which helps them grip the bamboo shoots they
eat) the giant panda and red panda are only distantly related. Jyutping hung4-maau1
Alternative Chinese name

Etymology Traditional Chinese 貓熊

The word panda was borrowed into English from French, but no conclusive
Simplified Chinese 猫熊
explanation of the origin of the French word panda has been found.[22] The Literal meaning "cat bear"
closest candidate is the Nepali word ponya, possibly referring to the adapted Transcriptions
wrist bone of the red panda, which is native to Nepal. The Western world
Hakka
Romanization meu-yùng
Southern Min
https://en.wikipedia.org/wiki/Giant_panda 2/19
1/23/2019 Giant panda - Wikipedia

originally applied this name to the red panda. Until 1901, when it was Hokkien POJ niau-hîm
erroneously stated to be related to the red panda, the giant panda was known
as "black and white cat-footed animal" (Ailuropus melanoleucus).[23]

In many older encyclopedic sources, the name "panda" or "common panda"


originally referred to the lesser-known red panda,[24] thus necessitating the
inclusion of "giant" and "lesser/red" prefixes in front of the names. Even in
2013, the Encyclopædia Britannica still used "giant panda" or "panda bear" for
the bear,[25] and simply "panda" for the Ailuridae,[26] despite the popular
usage of the word "panda".

Since the earliest collection of Chinese writings, the Chinese language has
given the bear 20 different names, such as huāxióng (花熊 "spotted bear") and
zhúxióng (竹熊 "bamboo bear").[27] The most popular names in China today is Panda cubs
dàxióngmāo (大熊貓 literally "giant bear cat"), or simply xióngmāo (熊貓 "bear
cat"). The name xióngmāo (熊貓 "bear cat") was originally used to describe the
red panda (Ailurus fulgens), but since giant panda was thought to be closely related to red panda, dàxióngmāo (大熊貓)
was named relatively.[27]

In Taiwan, another popular name for panda is the inverted dàmāoxióng ( 大 貓 熊 "giant cat bear"), though many
encyclopediae and dictionaries in Taiwan still use the "bear cat" form as the correct name. Some linguists argue, in this
construction, "bear" instead of "cat" is the base noun, making this name more grammatically and logically correct, which
may have led to the popular choice despite official writings.[27] This name did not gain its popularity until 1988, when a
private zoo in Tainan painted a sun bear black and white and created the Tainan fake panda incident.[28][29]

Subspecies
Two subspecies of giant panda have been recognized on the basis of distinct
cranial measurements, color patterns, and population genetics.[30]

The nominate subspecies Ailuropoda m. melanoleuca consists of most


extant populations of panda. These animals are principally found in
Sichuan and display the typical stark black and white contrasting colors.
The Qinling panda, A. m. qinlingensis[31] is restricted to the Qinling
Mountains in Shaanxi at elevations of 1,300–3,000 m. The typical black
and white pattern of Sichuan giant pandas is replaced with a dark brown
versus light brown pattern.[30] The skull of A. m. qinlingensis is smaller The Qinling panda has a light-brown
than its relatives, and it has larger molars. and white pattern
A detailed study of the giant panda's genetic history from 2012[32] confirms
that the separation of the Qinlin population occurred about 300,000 years ago,
and reveals that the non-Qinlin population further diverged into two groups, named the Minshan and the Qionglai-
Daxiangling-Xiaoxiangling-Liangshan group respectively, about 2,800 years ago.[33]

Description
The giant panda has luxuriant black-and-white fur. Adults measure around 1.2 to 1.9 m (4 to 6 ft) long, including a tail of
about 10–15 cm (3.9–5.9 in), and 60 to 90 cm (2.0 to 3.0 ft) tall at the shoulder.[34][35] Males can weigh up to 160 kg
(350 lb).[36] Females (generally 10–20% smaller than males)[37] can weigh as little as 70 kg (150 lb), but can also weigh up
to 125 kg (276 lb).[10][34][38] Average adult weight is 100 to 115 kg (220 to 254 lb).[39]

https://en.wikipedia.org/wiki/Giant_panda 3/19
1/23/2019 Giant panda - Wikipedia

The giant panda has a body shape typical of bears. It has black fur on its ears,
eye patches, muzzle, legs, arms and shoulders. The rest of the animal's coat is
white. Although scientists do not know why these unusual bears are black and
white, speculation suggests that the bold coloring provides effective
camouflage in their shade-dappled snowy and rocky habitat.[40] The giant
panda's thick, wooly coat keeps it warm in the cool forests of its habitat.[40]
The panda's skull shape is typical of durophagous carnivorans. It has evolved
from previous ancestors to exhibit larger molars with increased complexity and
expanded temporal fossa.[41][42] A 110.45 kg (243.5 lb) giant panda has a 3D
The skull of giant panda at the
canine teeth bite force of 2603.47 newtons and bite force quotient of 292. Smithsonian Museum of Natural
Another study had a 117.5 kg (259 lb) giant panda bite of 1298.9 newtons (BFQ History
151.4) at canine teeth and 1815.9 newtons (BFQ 141.8) at carnassial teeth.[43]

The giant panda's paw has a


"thumb" and five fingers; the
"thumb" – actually a modified
sesamoid bone – helps it to hold
bamboo while eating.[44] Stephen The skeleton (left) and taxidermy
Jay Gould discusses this feature in model (right) of "Tong Tong", once
his book of essays on evolution and bred in Ueno Zoo at the National
biology, The Panda's Thumb. Museum of Nature and Science,
Tokyo
The giant panda's tail, measuring
10 to 15 cm (4 to 6 in), is the
second-longest in the bear family. (The longest belongs to the sloth bear.)[37]

The giant panda typically lives around 20 years in the wild and up to 30 years
in captivity.[45] A female named Jia Jia was the oldest giant panda ever in
captivity, born in 1978 and died at an age of 38 on 16 October 2016.[46]
Bones of the left forelimb

Pathology
A seven-year-old female named Jin Yi died in 2014 in a zoo in Zhengzhou,
China, after showing symptoms of gastroenteritis and respiratory disease. It
was found that the cause of death was toxoplasmosis, a disease caused by
Toxoplasma gondii and infecting most warm-blooded animals, including
humans.[47]

Genomics
The giant panda genome was sequenced in 2009 using Illumina dye
Toxoplasma gondii (arrow) in
sequencing.[48] Its genome contains 20 pairs of autosomes and one pair of sex macrophages in the lung of a giant
chromosomes. panda[47]

Ecology

https://en.wikipedia.org/wiki/Giant_panda 4/19
1/23/2019 Giant panda - Wikipedia

Diet
Despite its taxonomic classification as a carnivoran, the giant panda's diet is
primarily herbivorous, consisting almost exclusively of bamboo.[45] However,
the giant panda still has the digestive system of a carnivore, as well as
carnivore-specific genes,[49] and thus derives little energy and little protein
from consumption of bamboo. Its ability to digest cellulose is ascribed to the
microbes in its gut.[50][51] Pandas are born with sterile intestines and require
bacteria obtained from their mother's feces to digest vegetation.[52] The giant
panda is a "highly specialized" animal with "unique adaptations", and has lived
in bamboo forests for millions of years.[53] The average giant panda eats as Pandas eating bamboo.
much as 9 to 14 kg (20 to 30 lb) of bamboo shoots a day to compensate for the
limited energy content of its diet. Ingestion of such a large quantity of material
is possible because of the rapid passage of large amounts of indigestible plant
material through the short, straight digestive tract.[54][55] It is also noted,
however, that such rapid passage of digesta limits the potential of microbial
digestion in the gastrointestinal tract,[54] limiting alternative forms of
digestion. Given this voluminous diet, the giant panda defecates up to 40 times
a day.[56] The limited energy input imposed on it by its diet has affected the
panda's behavior. The giant panda tends to limit its social interactions and
avoids steeply sloping terrain to limit its energy expenditures.[57]
Panda eating, standing, playing
Two of the panda's most distinctive features, its large size and round face, are
adaptations to its bamboo diet. Anthropologist Russell Ciochon observed: "
[much] like the vegetarian gorilla, the low body surface area to body volume [of the giant panda] is indicative of a lower
metabolic rate. This lower metabolic rate and a more sedentary lifestyle allows the giant panda to subsist on nutrient poor
resources such as bamboo."[57] Similarly, the giant panda's round face is the result of powerful jaw muscles, which attach
from the top of the head to the jaw.[57] Large molars crush and grind fibrous plant material.

The morphological characteristics of extinct relatives of the giant panda suggest that while the ancient giant panda was
omnivorous 7 million years ago (mya), it only became herbivorous some 2-2.4 mya with the emergence of A.
microta.[58][59] Genome sequencing of the giant panda suggests that the dietary switch could have initiated from the loss
of the sole T1R1/T1R3 umami taste receptor, resulting from two frameshift mutations within the T1R1 exons.[60] Umami
taste corresponds to high levels of glutamate as found in meat and may have thus altered the food choice of the giant
panda.[61] Although the pseudogenization of the umami taste receptor in Ailuropoda coincides with the dietary switch to
herbivory, it is likely a result of, and not the reason for, the dietary change.[59][60][61] The mutation time for the T1R1 gene
in the giant panda is estimated to 4.2 mya[59] while fossil evidence indicates bamboo consumption in the giant panda
species at least 7 mya,[58] signifying that although complete herbivory occurred around 2 mya, the dietary switch was
initiated prior to T1R1 loss-of-function.

Pandas eat any of 25 bamboo species in the wild, such as Fargesia dracocephala[62] and Fargesia rufa.[63] Only a few
bamboo species are widespread at the high altitudes pandas now inhabit. Bamboo leaves contain the highest protein
levels; stems have less.[64]

Because of the synchronous flowering, death, and regeneration of all bamboo within a species, the giant panda must have
at least two different species available in its range to avoid starvation. While primarily herbivorous, the giant panda still
retains decidedly ursine teeth and will eat meat, fish, and eggs when available. In captivity, zoos typically maintain the
giant panda's bamboo diet, though some will provide specially formulated biscuits or other dietary supplements.[65]

https://en.wikipedia.org/wiki/Giant_panda 5/19
1/23/2019 Giant panda - Wikipedia

Pandas will travel between different habitats if they need to, so they can get the nutrients that they need and to balance
their diet for reproduction. For six years, scientists studied six pandas tagged with GPS collars at the Foping Reserve in the
Qinling Mountains. They took note of their foraging and mating habits and analyzed samples of their food and feces. The
pandas would move from the valleys into the Qinling Mountains and would only return to the valleys in autumn. During
the summer months bamboo shoots rich in protein are only available at higher altitudes which causes low calcium rates in
the pandas and during breeding season the pandas would trek back down to eat bamboo leaves rich in calcium.[66]

Predators
Although adult giant pandas have few natural predators other than humans, young cubs are vulnerable to attacks by snow
leopards, yellow-throated martens,[67] eagles, feral dogs, and the Asian black bear. Sub-adults weighing up to 50 kg
(110 lb) may be vulnerable to predation by leopards.[68]

Behavior
The giant panda is a terrestrial animal and primarily spends its life roaming and feeding in the bamboo forests of the
Qinling Mountains and in the hilly province of Sichuan.[69] Giant pandas are generally solitary.[53] Each adult has a
defined territory and a female is not tolerant of other females in her range. Social encounters occur primarily during the
brief breeding season in which pandas in proximity to one another will gather.[70] After mating, the male leaves the female
alone to raise the cub.[71]

Pandas were thought to fall into the crepuscular category, those who are active twice a day, at dawn and dusk; however,
Jindong Zhang found that pandas may belong to a category all of their own, with activity peaks in the morning, afternoon
and midnight. Due to their sheer size, pandas do not need to fear predators like other herbivores. They can therefore be
active at any time of the day.[72]

Pandas communicate through vocalization and scent marking such as clawing trees or spraying urine.[10] They are able to
climb and take shelter in hollow trees or rock crevices, but do not establish permanent dens. For this reason, pandas do
not hibernate, which is similar to other subtropical mammals, and will instead move to elevations with warmer
temperatures.[73] Pandas rely primarily on spatial memory rather than visual memory.[74]

Though the panda is often assumed to be docile, it has been known to attack humans, presumably out of irritation rather
than aggression.[75][76][77]

Reproduction
Initially, the primary method of breeding giant pandas in captivity was by artificial insemination, as they seemed to lose
their interest in mating once they were captured.[79] This led some scientists to try extreme methods, such as showing
them videos of giant pandas mating[80] and giving the males sildenafil (commonly known by name Viagra).[81] Only
recently have researchers started having success with captive breeding programs, and they have now determined giant
pandas have comparable breeding to some populations of the American black bear, a thriving bear species. The normal
reproductive rate is considered to be one young every two years.[15][69]

Giant pandas reach sexual maturity between the ages of four and eight, and may be reproductive until age 20.[82] The
mating season is between March and May, when a female goes into estrus, which lasts for two or three days and only
occurs once a year.[83] When mating, the female is in a crouching, head-down position as the male mounts her from
behind. Copulation time is short, ranging from 30 seconds to five minutes, but the male may mount her repeatedly to
ensure successful fertilization. The gestation period ranges from 95 to 160 days.[83]

https://en.wikipedia.org/wiki/Giant_panda 6/19
1/23/2019 Giant panda - Wikipedia

Giant pandas give birth to twins in about half of pregnancies.[84] If twins are
born, usually only one survives in the wild. The mother will select the stronger
of the cubs, and the weaker cub will die due to starvation. The mother is
thought to be unable to produce enough milk for two cubs since she does not
store fat.[85] The father has no part in helping raise the cub.

When the cub is first born, it is pink, blind, and toothless,[86] weighing only 90
1
to 130 grams (3.2 to 4.6 ounces), or about 800 th of the mother's weight,[17]
proportionally the smallest baby of any placental mammal.[87] It nurses from
its mother's breast six to 14 times a day for up to 30 minutes at a time. For A giant panda cub. At birth, the giant
panda typically weighs 100 to 200
three to four hours, the mother may leave the den to feed, which leaves the cub
grams (31⁄2 to 7 oz) and measures
defenseless. One to two weeks after birth, the cub's skin turns gray where its
15 to 17 centimeters (6 to 7 in)
hair will eventually become black. Slight pink color may appear on cub's fur, as
long.[78]
a result of a chemical reaction between the fur and its mother's saliva. A month
after birth, the color pattern of the cub's fur is fully developed. Its fur is very
soft and coarsens with age. The cub begins to crawl at 75 to 80 days;[17]
mothers play with their cubs by rolling and wrestling with them. The cubs can
eat small quantities of bamboo after six months,[88] though mother's milk
remains the primary food source for most of the first year. Giant panda cubs
weigh 45 kg (100 pounds) at one year and live with their mothers until they are
18 months to two years old. The interval between births in the wild is generally
two years.

Panda Research and Breeding


In July 2009, Chinese scientists confirmed the birth of the first cub to be
Center in Chengdu.
successfully conceived through artificial insemination using frozen sperm.[89]
The cub was born at 07:41 on 23 July that year in Sichuan as the third cub of
You You, an 11-year-old.[89][90][91] The technique for freezing the sperm in liquid nitrogen was first developed in 1980 and
the first birth was hailed as a solution to the problem of lessening giant panda semen availability, which had led to
inbreeding.[91][92] Panda semen, which can be frozen for decades, could be shared between different zoos to save the
species.[89][90] It is expected that zoos in destinations such as San Diego in the United States and Mexico City will now be
able to provide their own semen to inseminate more giant pandas.[92] In August 2014, a rare birth of panda triplets was
announced in China; it was the fourth of such births ever reported.[93]

Attempts have also been made to reproduce giant pandas by interspecific pregnancy by implanting cloned panda embryos
into the uterus of an animal of another species. This has resulted in panda fetuses, but no live births.[94]

Uses and human interaction

Early references
In the past, pandas were thought to be rare and noble creatures – the Empress Dowager Bo was buried with a panda skull
in her vault. The grandson of Emperor Taizong of Tang is said to have given Japan two pandas and a sheet of panda skin
as a sign of goodwill. Unlike many other animals in Ancient China, pandas were rarely thought to have medical uses. The
few known uses include the Sichuan tribal peoples' use of panda urine to melt accidentally swallowed needles, and the use
of panda pelts to control menses as described in the Qin Dynasty encyclopedia Erya.[95]

https://en.wikipedia.org/wiki/Giant_panda 7/19
1/23/2019 Giant panda - Wikipedia

The creature named mo ( 貘 ) mentioned in some ancient books has been interpreted as giant panda.[95] The dictionary
Shuowen Jiezi (Eastern Han Dynasty) says that the mo, from Shu (Sichuan), is bear-like, but yellow-and-black,[96]
although the older Erya describes mo simply as a "white leopard".[97] The interpretation of the legendary fierce creature
pixiu (貔貅) as referring to the giant panda is also common.[98]

During the reign of the Yongle Emperor (early 15th century), his relative from Kaifeng sent him a captured zouyu (騶虞),
and another zouyu was sighted in Shandong. Zouyu is a legendary "righteous" animal, which, similarly to a qilin, only
appears during the rule of a benevolent and sincere monarch. It is said to be fierce as a tiger, but gentle and strictly
vegetarian, and described in some books as a white tiger with black spots. Puzzled about the real zoological identity of the
creature captured during the Yongle era, J.J.L. Duyvendak exclaims, "Can it possibly have been a Pandah?"[99]

The comparative obscurity of the giant panda throughout most of China's history is illustrated by the fact that, despite
there being a number of depictions of bears in Chinese art starting from its most ancient times, and the bamboo being one
of the favorite subjects for Chinese painters, there are no known pre-20th-century artistic representations of giant pandas.

Western discovery
The West first learned of the giant panda on 11 March 1869, when the French missionary Armand David[17] received a skin
from a hunter. The first Westerner known to have seen a living giant panda is the German zoologist Hugo Weigold, who
purchased a cub in 1916. Kermit and Theodore Roosevelt, Jr., became the first Westerners to shoot a panda, on an
expedition funded by the Field Museum of Natural History in the 1920s. In 1936, Ruth Harkness became the first
Westerner to bring back a live giant panda, a cub named Su Lin[100] which went to live at the Brookfield Zoo in Chicago. In
1938, five giant pandas were sent to London.[101][102]

Panda diplomacy
Gifts of giant pandas to American and Japanese zoos formed an important part
of the diplomacy of the People's Republic of China (PRC) in the 1970s, as it
marked some of the first cultural exchanges between the PRC and the West.
This practice has been termed "panda diplomacy".

By 1984, however, pandas were no longer given as gifts. Instead, the PRC
began to offer pandas to other nations only on 10-year loans, under terms Adult male giant panda
including a fee of up to US$1,000,000 per year and a provision that any cubs
born during the loan are the property of the PRC. Since 1998, because of a
WWF lawsuit, the United States Fish and Wildlife Service only allows a US zoo to import a panda if the zoo can ensure the
PRC will channel more than half of its loan fee into conservation efforts for the giant panda and its habitat.

In May 2005, the PRC offered a breeding pair to Taiwan. The issue became embroiled in cross-Strait relations – both over
the underlying symbolism, and over technical issues such as whether the transfer would be considered "domestic" or
"international", or whether any true conservation purpose would be served by the exchange.[103] A contest in 2006 to
name the pandas was held in the mainland, resulting in the politically charged names Tuan Tuan and Yuan Yuan (from
tuanyuan, meaning "reunion", i.e. "reunification"). PRC's offer was initially rejected by Chen Shui-bian, then President of
Taiwan. However, when Ma Ying-jeou assumed the presidency in 2008, the offer was accepted, and the pandas arrived in
December of that year.[104]

Biofuel

https://en.wikipedia.org/wiki/Giant_panda 8/19
1/23/2019 Giant panda - Wikipedia

Microbes in panda waste are being investigated for their use in creating biofuels from bamboo and other plant
materials.[105]

Conservation
The giant panda is a vulnerable species, threatened by continued habitat loss and habitat fragmentation,[106] and by a very
low birthrate, both in the wild and in captivity.[45] Its range is currently confined to a small portion on the western edge of
its historical range, which stretched through southern and eastern China, northern Myanmar, and northern Vietnam.[1]

The giant panda has been a target of poaching by locals since ancient times and by foreigners since it was introduced to
the West. Starting in the 1930s, foreigners were unable to poach giant pandas in China because of the Second Sino-
Japanese War and the Chinese Civil War, but pandas remained a source of soft furs for the locals. The population boom in
China after 1949 created stress on the pandas' habitat and the subsequent famines led to the increased hunting of wildlife,
including pandas. During the Cultural Revolution, all studies and conservation activities on the pandas were stopped.
After the Chinese economic reform, demand for panda skins from Hong Kong and Japan led to illegal poaching for the
black market, acts generally ignored by the local officials at the time.

Though the Wolong National Nature Reserve was set up by the PRC
government in 1958 to save the declining panda population, few advances in
the conservation of pandas were made, owing to inexperience and insufficient
knowledge of ecology. Many believed the best way to save the pandas was to
cage them. As a result, pandas were caged at any sign of decline and suffered
from terrible conditions. Because of pollution and destruction of their natural
habitat, along with segregation caused by caging, reproduction of wild pandas
was severely limited. In the 1990s, however, several laws (including gun
Closeup of a seven-month-old
control and the removal of resident humans from the reserves) helped their
panda cub
chances of survival. With these renewed efforts and improved conservation
methods, wild pandas have started to increase in numbers in some areas,
though they still are classified as a rare species.

In 2006, scientists reported that the number of pandas living in the wild may have been underestimated at about 1,000.
Previous population surveys had used conventional methods to estimate the size of the wild panda population, but using a
new method that analyzes DNA from panda droppings, scientists believe the wild population may be as large as 3,000.[45]
In 2006, there were 40 panda reserves in China, compared to just 13 reserves in 1998.[14] As the species has been
reclassified to "vulnerable" since 2016, the conservation efforts are thought to be working. Furthermore, in response to
this reclassification, the State Forestry Administration of China announced that they would not accordingly lower the
conservation level for panda, and would instead reinforce the conservation efforts.[107]

The giant panda is among the world's most adored and protected rare animals, and is one of the few in the world whose
natural inhabitant status was able to gain a UNESCO World Heritage Site designation. The Sichuan Giant Panda
Sanctuaries, located in the southwest province of Sichuan and covering seven natural reserves, were inscribed onto the
World Heritage List in 2006.[108][109]

Not all conservationists agree that the money spent on conserving pandas is well spent. Chris Packham has argued that the
breeding of pandas in captivity is "pointless" because "there is not enough habitat left to sustain them".[110] Packham
argues that the money spent on pandas would be better spent elsewhere,[110] and has said he would "eat the last panda if I
could have all the money we have spent on panda conservation put back on the table for me to do more sensible things
with",[111] though he has apologized for upsetting people who like pandas.[112] He points out, "The panda is possibly one of

https://en.wikipedia.org/wiki/Giant_panda 9/19
1/23/2019 Giant panda - Wikipedia

the grossest wastes of conservation money in the last half century."[111] However, a 2015 paper found that the giant panda
can serve as an umbrella species as the preservation of their habitat also helps other endemic species in China, including
70% of the country's forest birds, 70% of mammals and 31% of amphibians.[113]

In 2012, Earthwatch Institute, a global nonprofit that teams volunteers with scientists to conduct important
environmental research, launched a program called "On the Trail of Giant Panda". This program, based in the Wolong
National Nature Reserve, allows volunteers to work up close with pandas cared for in captivity, and help them adapt to life
in the wild, so that they may breed, and live longer and healthier lives.[114]

In zoos
Pandas have been kept in zoos as early as the Western Han Dynasty in China, where the writer Sima Xiangru noted that
the panda was the most treasured animal in the emperor's garden of exotic animals in the capital Chang'an (present
Xi'an). Not until the 1950s were pandas again recorded to have been exhibited in China's zoos.[115]

Chi Chi at the London Zoo became very popular. This influenced the World Wildlife Fund to use a panda as its symbol.[116]

A 2006 New York Times article[117] outlined the economics of keeping pandas, which costs five times more than that of
the next most expensive animal, an elephant. American zoos generally pay the Chinese government $1 million a year in
fees, as part of a typical ten-year contract. San Diego's contract with China was to expire in 2008, but got a five-year
extension at about half of the previous yearly cost.[118] The last contract, with the Memphis Zoo in Memphis, Tennessee,
ended in 2013.[117]

Population chart

Year Wild[119] Change Captivity[53] Change Total Change

1976 1,000 n/a n/a n/a n/a n/a


1985 800–1,200 n/a n/a n/a n/a n/a
1987 >1,000 n/a n/a n/a n/a n/a
1994 1,200 n/a n/a n/a n/a n/a
1995 1,000 −200 n/a n/a n/a n/a

2003 1,596 +596 164[120] n/a 1,760 n/a

2012 n/a n/a 341[121] +178 n/a n/a

2013 1,864[122] +268 375[120][123] +34[123] 2,239 +479

Reference in medicine
The Face of the Giant Panda Sign is an MRI sign in patients with Wilson's disease, named for the midbrain's resemblance
to a giant panda's face.

In cryptozoology
Skeptical cryptozoologist Joe Nickell, notes that since Giant Pandas were known to local people, they qualify as
cryptids.[124] However, unlike Bigfoot, pandas specimens were quickly obtained after Armand David learned of them.
Also, fossil evidence shows that pandas were once widespread, including the two million year old skull of Ailuropoda
https://en.wikipedia.org/wiki/Giant_panda 10/19
1/23/2019 Giant panda - Wikipedia

microta.[125]

See also
Red panda Pygmy giant panda
List of giant pandas Wildlife of China
Panda tea List of endangered and protected species of China

References
Notes

1. Swaisgood, R.; Wang, D.; Wei, F. (2016). "Ailuropoda melanoleuca" (http://oldredlist.iucnredlist.org/details/712/0).


IUCN Red List of Threatened Species. IUCN. 2016: e.T712A45033386. Retrieved 5 September 2016.
2. David, Armand (1869). "Voyage en Chine" (https://www.biodiversitylibrary.org/page/13953952). Bulletin des Nouvelles
Archives du Muséum. 5: 13. "Ursus melanoleucus"
3. Like the English "giant", the term dà ("large") is technically prefixed to the name "panda" in Chinese, but is not
generally in everyday use.
4. Scheff, Duncan (2002). Giant Pandas. Animals of the rain forest (illustrated ed.). Heinemann-Raintree Library. p. 7.
ISBN 0-7398-5529-8.
5. Lindburg, Donald G.; Baragona, Karen (2004). Giant Pandas: Biology and Conservation. University of California
Press. ISBN 0-520-23867-2.
6. Quote: "Bamboo forms 99 percent of a panda's diet", "more than 99 percent of their diet is bamboo": p. 63 of Lumpkin
& Seidensticker 2007 (as seen in the 2002 edition).
7. "Giant Panda" (https://web.archive.org/web/20100107032418/http://animal.discovery.com/mammals/giant-panda/).
Discovery Communications, LLC. Archived from the original (http://animal.discovery.com/mammals/giant-panda/) on 7
January 2010. Retrieved 8 August 2010.
8. "Giant Pandas" (http://nationalzoo.si.edu/Animals/GiantPandas/PandaFacts/default.cfm). National Zoological Park.
Retrieved 7 November 2010.
9. Scheff, Duncan (2002). Giant Pandas. Animals of the rain forest (illustrated ed.). Heinemann-Raintree Library. p. 8.
ISBN 0-7398-5529-8.
10. "Global Species Programme – Giant panda" (https://web.archive.org/web/20080704204350/http://www.panda.org/abo
ut_wwf/what_we_do/species/our_solutions/endangered_species/giant_panda/index.cfm). World Wildlife Fund. 14
November 2007. Archived from the original (http://panda.org/about_wwf/what_we_do/species/our_solutions/endanger
ed_species/giant_panda/index.cfm) on 4 July 2008. Retrieved 22 July 2008.
11. "Four out of six great apes one step away from extinction – IUCN Red List" (http://iucnworldconservationcongress.or
g/news/20160904/article/four-out-six-great-apes-one-step-away-extinction-iucn-red-list). 4 September 2016.
12. "Number of pandas successfully bred in China down from last year" (http://news.xinhuanet.com/english/2007-11/08/c
ontent_7034856.htm). Xinhua. 8 November 2007. Retrieved 22 July 2008.
13. "Panda Zoos Around The World" (https://web.archive.org/web/20160102213245/http://www.giantpandazoo.com/pand
a/panda-zoos/). www.GiantPandaZoo.com. Archived from the original (http://www.giantpandazoo.com/panda/panda-z
oos/) on 2 January 2016.
14. Briggs, Helen (20 June 2006). "Hope for future of giant panda" (http://news.bbc.co.uk/2/hi/science/nature/5085006.st
m). BBC News. Retrieved 14 February 2007.
15. Warren, Lynne (July 2006). "Pandas, Inc" (http://ngm.nationalgeographic.com/ngm/0607/feature1/?fs=animals-panthe
r.nationalgeographic.com). National Geographic. Retrieved 10 April 2008.
16. "Giant panda population rises by nearly 17 percent" (http://news.mongabay.com/2015/0302-hance-giant-panda-popul
ation.html). Mongabay Environmental News.

https://en.wikipedia.org/wiki/Giant_panda 11/19
1/23/2019 Giant panda - Wikipedia

17. "Giant Panda" (http://www.britannica.com/EBchecked/topic/441032/giant-panda). Encyclopædia Britannica Online.


2010. Retrieved 9 August 2010.
18. O'Brien, Nash, Wildt, Bush & Benveniste, A molecular solution to the riddle of the giant panda's phylogeny (http://ww
w.nature.com/nature/journal/v317/n6033/abs/317140a0.html), Nature Page 317, and pages 140 – 144 (12 September
1985)
19. Krause, J.; Unger, T.; Noçon, A.; Malaspinas, A.; Kolokotronis, S.; Stiller, M.; Soibelzon, L.; Spriggs, H.; Dear, P. H.;
Briggs, A. W.; Bray, S. C. E.; O'Brien, S. J.; Rabeder, G.; Matheus, P.; Cooper, A.; Slatkin, M.; Pääbo, S.; Hofreiter, M.
(2008). "Mitochondrial genomes reveal an explosive radiation of extinct and extant bears near the Miocene-Pliocene
boundary" (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2518930). BMC Evolutionary Biology. 8 (220): 220.
doi:10.1186/1471-2148-8-220 (https://doi.org/10.1186%2F1471-2148-8-220). PMC 2518930 (https://www.ncbi.nlm.ni
h.gov/pmc/articles/PMC2518930). PMID 18662376 (https://www.ncbi.nlm.nih.gov/pubmed/18662376).
20. Yu, Li; Li, Yi-Wei; Ryder, Oliver A.; Zhang, Ya-Ping (2007). "Analysis of complete mitochondrial genome sequences
increases phylogenetic resolution of bears (Ursidae), a mammalian family that experienced rapid speciation" (http://b
mcevolbiol.biomedcentral.com/articles/10.1186/1471-2148-7-198). BMC Evolutionary Biology. 7 (198): 198.
doi:10.1186/1471-2148-7-198 (https://doi.org/10.1186%2F1471-2148-7-198).
21. "Behind the News – Panda Granny" (https://web.archive.org/web/20080504213933/http://www.abc.net.au/tv/btn/storie
s/s1947589.htm). Australian Broadcasting Corporation. 12 June 2007. Archived from the original (http://www.abc.net.
au/tv/btn/stories/s1947589.htm) on 4 May 2008. Retrieved 22 July 2008.
22. Oxford English Dictionary, s.v. panda n. 1.
23. New York Zoological Society (1987). Animal Kingdom, Volumes 90–91. p. 14.
24. "Animal Info – Red Panda" (http://www.animalinfo.org/species/carnivor/ailufulg.htm).
25. "giant panda (mammal) -- Encyclopedia Britannica" (https://web.archive.org/web/20130515123427/http://www.britanni
ca.com/EBchecked/topic/441032/giant-panda). Britannica.com. Archived from the original (http://www.britannica.com/
EBchecked/topic/441032/giant-panda) on 2013-05-15. Retrieved 2017-01-23.
26. "panda (mammal, Ailurus species) -- Encyclopedia Britannica" (https://web.archive.org/web/20130515120439/http://w
ww.britannica.com/EBchecked/topic/441037/panda). Britannica.com. Archived from the original (http://www.britannica.
com/EBchecked/topic/441037/panda) on 2013-05-15. Retrieved 2017-01-23.
27. "Discussion about the Chinese name for giant panda (in Chinese)" (http://huayuqiao.org/articles/yaodehuai/yaodh08.h
tm).
28. "Government Information Office will now use dàxióngmāo as the proper name (in Chinese)" (http://udndata.com/ndap
p/Story2007?no=31&page=4&udndbid=udndata&SearchString=v9%2B6tSu6tb%2FfK6TptME%2BPTE5NTEwMTAxK
6TptME8PTIwMDAwMTAx&sharepage=10&select=0&kind=2&article_date=1990-08-09&news_id=100993201&show
UserSearch=+%3Cstrong%3E%3Cfont+color%3D%23333333+class%3Dtitle03%3E%B1%7A%A5%48%3C%2Ffon
t%3E+%3Cfont+color%3D%23FF6600+class%3Dtitle04%3E%BF%DF%BA%B5%2B%BA%B5%BF%DF%2B%A4%
E9%B4%C1%3E%3D19510101%2B%A4%E9%B4%C1%3C%3D20000101%3C%2Ffont%3E+%3Cfont+color%3D%
23333333+class%3Dtitle03%3E%A6%40%B7%6A%B4%4D%A8%EC%3C%2Ffont%3E+%3Cfont+color%3D%23FF
6600+class%3Dtitle04%3E68%3C%2Ffont%3E+%3Cfont+color%3D%23333333+class%3Dtitle03%3E%B5%A7%B
8%EA%AE%C6%3C%2Ffont%3E%3C%2Fstrong%3E&firstFatherCateID=&collectCateNewsPage=1). 聯合報
. 1990-
08-09.
29. " "bear cat" or "cat bear" (in Chinese)" (http://udndata.com/ndapp/Story2007?no=307&page=31&udndbid=udndata&S
earchString=tsKl1bq1L7q1v98vv9%2B6tSuk6bTBPj0xOTg3MTIwMSuk6bTBPD0xOTg4MTIzMQ%3D%3D&sharepag
e=10&select=1&kind=2&article_date=1987-12-29&news_id=103310434&showUserSearch=+%3Cstrong%3E%3Cfon
t+color%3D%23333333+class%3Dtitle03%3E%B1%7A%A5%48%3C%2Ffont%3E+%3Cfont+color%3D%23FF6600+
class%3Dtitle04%3E%B6%C2%A5%D5%BA%B5%2F%BA%B5%BF%DF%2F%BF%DF%BA%B5%2B%A4%E9%B
4%C1%3E%3D19871201%2B%A4%E9%B4%C1%3C%3D19881231%3C%2Ffont%3E+%3Cfont+color%3D%23333
333+class%3Dtitle03%3E%A6%40%B7%6A%B4%4D%A8%EC%3C%2Ffont%3E+%3Cfont+color%3D%23FF6600+
class%3Dtitle04%3E317%3C%2Ffont%3E+%3Cfont+color%3D%23333333+class%3Dtitle03%3E%B5%A7%B8%E
A%AE%C6%3C%2Ffont%3E%3C%2Fstrong%3E&firstFatherCateID=&collectCateNewsPage=1). 聯合報
. 1987-12-
29.
30. Wan, Wu & Fang 2005.

https://en.wikipedia.org/wiki/Giant_panda 12/19
1/23/2019 Giant panda - Wikipedia

31. Hammond, Paula (2010). The Atlas of Endangered Animals: Wildlife Under Threat Around the World. Marshall
Cavendish. p. 58. ISBN 0-7614-7872-8.
32. Shancen Zhao; Pingping Zheng; Shanshan Dong; Xiangjiang Zhan; Qi Wu (16 December 2012). "Whole-genome
sequencing of giant pandas provides insights into demographic history and local adaptation" (http://www.nature.com/n
g/journal/vaop/ncurrent/full/ng.2494.html). Nature Genetics. 45 (1): 67–71. doi:10.1038/ng.2494 (https://doi.org/10.10
38%2Fng.2494). PMID 23242367 (https://www.ncbi.nlm.nih.gov/pubmed/23242367). Retrieved 17 December 2012.
33. "Scientists Discover Evidence of Giant Panda's Population History and Local Adaptation" (https://www.sciencedaily.co
m/releases/2012/12/121216132511.htm). 16 December 2012. Retrieved 17 December 2012.
34. Giant Panda (http://www.arkive.org/giant-panda/ailuropoda-melanoleuca/#text=Facts), Arkive
35. "Physical Description" (https://web.archive.org/web/20111204035805/http://www.giantpandaonline.org/naturalhistory/d
escription.htm). Giant Panda Species Survival Plan. Archived from the original (http://www.giantpandaonline.org/natur
alhistory/description.htm) on 4 December 2011. Retrieved 26 October 2011.
36. Boitani, Luigi, Simon & Schuster's Guide to Mammals. Simon & Schuster/Touchstone Books (1984), ISBN 978-0-671-
42805-1
37. Brown, Gary (1996). Great Bear Almanac. p. 340. ISBN 1-55821-474-7.
38. "Archived copy" (https://web.archive.org/web/20150924121859/http://www.science.smith.edu/msi/pdf/i0076-3519-110-
01-0001.pdf) (PDF). Archived from the original (http://www.science.smith.edu/msi/pdf/i0076-3519-110-01-0001.pdf)
(PDF) on 24 September 2015. Retrieved 5 February 2016. (2011).
39. [1] (http://www.animalfactguide.com/animalfacts/giant-panda/) (2011).
40. Dudley, Karen (1997). Giant Pandas. Untamed world (illustrated ed.). Weigl Educational Publishers. p. 9. ISBN 0-
919879-87-X.
41. Ellis, Richard (2004). No turning back: the life and death of animal species (illustrated ed.). HarperCollins. p. 315.
ISBN 0-06-055803-2.
42. Figueirido, Borja; Zhijie Jack Tseng, Alberto Mart ́ın-Serra (July 2013). "Skull shape evolution in durophagous
carnivorans". Evolution; international journal of organic evolution. 67 (7): 1975–93. doi:10.1111/evo.12059 (https://doi.
org/10.1111%2Fevo.12059). PMID 23815654 (https://www.ncbi.nlm.nih.gov/pubmed/23815654).
43. Stephen Wroe. "Bite forces and evolutionary adaptations to feeding ecology in carnivores (Ecology)" (https://www.aca
demia.edu/239888/Bite_forces_and_evolutionary_adaptations_to_feeding_ecology_in_carnivores_Ecology_).
academia.edu.
44. Morris, Paul; Susan F. Morris. "The Panda's Thumb" (http://www.athro.com/evo/pthumb.html). Athro Limited.
Retrieved 7 August 2010.
45. Earth's Changing Environment. Learn & Explore. Encyclopædia Britannica, Inc. 2010. p. 49. ISBN 1-61535-339-9.
46. " 'Oldest' panda in captivity Jia Jia dies at the age of 38" (https://www.bbc.com/news/world-asia-37672804). BBC.
2016-10-16. Retrieved 2016-10-16.
47. Ma, Hongyu; Wang, Zedong; Wang, Chengdong; Li, Caiwu; Wei, Feng; Liu, Quan (2015). "Fatal Toxoplasma gondii
infection in the giant panda" (http://www.parasite-journal.org/articles/parasite/full_html/2015/01/parasite150075/parasit
e150075.html). Parasite. 22: 30. doi:10.1051/parasite/2015030 (https://doi.org/10.1051%2Fparasite%2F2015030).
ISSN 1776-1042 (https://www.worldcat.org/issn/1776-1042). PMC 4626621 (https://www.ncbi.nlm.nih.gov/pmc/article
s/PMC4626621). PMID 26514595 (https://www.ncbi.nlm.nih.gov/pubmed/26514595).
48. Li, R.; Fan, W.; Tian, G.; Zhu, H.; He, L.; Cai, J.; Huang, Q.; Cai, Q.; Li, B.; Bai, Y.; Zhang, Z.; Zhang, Y.; Wang, W.; Li,
J.; Wei, F.; Li, H.; Jian, M.; Li, J.; Zhang, Z.; Nielsen, R.; Li, D.; Gu, W.; Yang, Z.; Xuan, Z.; Ryder, O. A.; Leung, F. C.
C.; Zhou, Y.; Cao, J.; Sun, X.; Fu, Y. (2009). "The sequence and de novo assembly of the giant panda genome" (http
s://www.ncbi.nlm.nih.gov/pmc/articles/PMC3951497). Nature. 463 (7279): 311–317. Bibcode:2010Natur.463..311L (htt
p://adsabs.harvard.edu/abs/2010Natur.463..311L). doi:10.1038/nature08696 (https://doi.org/10.1038%2Fnature0869
6). PMC 3951497 (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3951497). PMID 20010809 (https://www.ncbi.nlm.ni
h.gov/pubmed/20010809).

https://en.wikipedia.org/wiki/Giant_panda 13/19
1/23/2019 Giant panda - Wikipedia

49. "(...)indicating that the panda probably has all the necessary components for a carnivorous digestive system."
Ruiqiang Li; Tian, Geng; Zhu, Hongmei; He, Lin; Cai, Jing; Huang, Quanfei; Cai, Qingle; Li, Bo; Bai, Yinqi; Zhang,
Zhihe; Zhang, Yaping; Wang, Wen; Li, Jun; Wei, Fuwen; Li, Heng; Jian, Min; Li, Jianwen; Zhang, Zhaolei; Nielsen,
Rasmus; Li, Dawei; Gu, Wanjun; Yang, Zhentao; Xuan, Zhaoling; Ryder, Oliver A.; Leung, Frederick Chi-Ching; Zhou,
Yan; Cao, Jianjun; Sun, Xiao; et al. (2010). "The sequence and de novo assembly of the giant panda genome" (http
s://www.ncbi.nlm.nih.gov/pmc/articles/PMC3951497). Nature. 463 (21): 311–317. Bibcode:2010Natur.463..311L (htt
p://adsabs.harvard.edu/abs/2010Natur.463..311L). doi:10.1038/nature08696 (https://doi.org/10.1038%2Fnature0869
6). PMC 3951497 (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3951497). PMID 20010809 (https://www.ncbi.nlm.ni
h.gov/pubmed/20010809).
50. "We did not find any homologues of digestive cellulase genes, including endoglucanase, exoglucanase and beta-
glucosidase, indicating that the bamboo diet of the panda is unlikely to be dictated by its own genetic composition,
and may instead be more dependent on its gut microbiome." Ruiqiang Li; Tian, Geng; Zhu, Hongmei; He, Lin; Cai,
Jing; Huang, Quanfei; Cai, Qingle; Li, Bo; Bai, Yinqi; Zhang, Zhihe; Zhang, Yaping; Wang, Wen; Li, Jun; Wei, Fuwen;
Li, Heng; Jian, Min; Li, Jianwen; Zhang, Zhaolei; Nielsen, Rasmus; Li, Dawei; Gu, Wanjun; Yang, Zhentao; Xuan,
Zhaoling; Ryder, Oliver A.; Leung, Frederick Chi-Ching; Zhou, Yan; Cao, Jianjun; Sun, Xiao; et al. (2010). "The
sequence and de novo assembly of the giant panda genome" (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC395149
7). Nature. 463 (21): 311–317. Bibcode:2010Natur.463..311L (http://adsabs.harvard.edu/abs/2010Natur.463..311L).
doi:10.1038/nature08696 (https://doi.org/10.1038%2Fnature08696). PMC 3951497 (https://www.ncbi.nlm.nih.gov/pm
c/articles/PMC3951497). PMID 20010809 (https://www.ncbi.nlm.nih.gov/pubmed/20010809).
51. Zhu, L.; Wu, Q., Dai, J., Zhang, S., Wei, F.; Dai, J.; Zhang, S.; Wei, F. (17 October 2011). "Evidence of cellulose
metabolism by the giant panda gut microbiome" (http://www.pnas.org/content/early/2011/10/11/1017956108).
Proceedings of the National Academy of Sciences. 108 (43): 17714–17719. Bibcode:2011PNAS..10817714Z (http://a
dsabs.harvard.edu/abs/2011PNAS..10817714Z). doi:10.1073/pnas.1017956108 (https://doi.org/10.1073%2Fpnas.101
7956108). PMC 3203778 (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3203778).
52. "BBC Nature — Dung eater videos, news and facts" (http://www.bbc.co.uk/nature/adaptations/Coprophagia).
Bbc.co.uk. Retrieved 2011-11-27.
53. "Giant Panda Facts" (http://nationalzoo.si.edu/animals/giantpandas/pandafacts/default.cfm). nationalzoo.si.edu.
National Zoological Park. Retrieved 8 June 2012.
54. Dierenfeld, E. S.; Hintz, H. F.; Robertson, J. B.; Van Soest, P. J.; Oftedal, O. T. (1982). "Utilization of bamboo by the
giant panda". Journal of Nutrition. 112 (4): 636–641. PMID 6279804 (https://www.ncbi.nlm.nih.gov/pubmed/6279804).
55. Finley, T. G.; Sikes, Robert S.; Parsons, Jennifer L.; Rude, Brian J.; Bissell, Heidi A.; Ouellette, John R. (2011).
"Energy digestibility of giant pandas on bamboo-only and on supplemented diet". Zoo Biology. 30 (2): 121–133.
doi:10.1002/zoo.20340 (https://doi.org/10.1002%2Fzoo.20340). PMID 20814990 (https://www.ncbi.nlm.nih.gov/pubm
ed/20814990).
56. "Panda tests bring population hope" (http://news.bbc.co.uk/2/hi/uk_news/wales/5095448.stm). BBC. 20 June 2006.
Retrieved 28 October 2010.
57. Ciochon, Russell L.; Eaves-Johnson, K. Lindsay (20 July 2007). "Bamboozled! The Curious Natural History of the
Giant Panda Family" (https://web.archive.org/web/20070721092645/http://www.scitizen.com/screens/blogPage/viewB
log/sw_viewBlog.php?idTheme=27&idContribution=855). Scitizen. Archived from the original (http://scitizen.com/scre
ens/blogPage/viewBlog/sw_viewBlog.php?idTheme=27&idContribution=855) on 21 July 2007. Retrieved 22 July
2008.
58. Jin, C; Ciochon, R. L.; Dong, W; Hunt Jr, R. M.; Liu, J; Jaeger, M; Zhu, Q (2007). "The first skull of the earliest giant
panda" (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1904166). Proceedings of the National Academy of Sciences
of the United States of America. 104 (26): 10932–10937. Bibcode:2007PNAS..10410932J (http://adsabs.harvard.edu/
abs/2007PNAS..10410932J). doi:10.1073/pnas.0704198104 (https://doi.org/10.1073%2Fpnas.0704198104).
PMC 1904166 (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1904166). PMID 17578912 (https://www.ncbi.nlm.nih.g
ov/pubmed/17578912).
59. Zhao, S; Zheng, P; Dong, S; Zhan, X; Wu, Q; Guo, X; Hu, Y; He, W; Zhang, S; Fan, W; Zhu, L; Li, D; Zhang, X; Chen,
Q; Zhang, H; Zhang, Z; Jin, X; Zhang, J; Yang, H; Wang, J; Wang, J; Wei, F (2013). "Whole-genome sequencing of
giant pandas provides insights into demographic history and local adaptation". Nature Genetics. 45 (1): 67–71.
doi:10.1038/ng.2494 (https://doi.org/10.1038%2Fng.2494). PMID 23242367 (https://www.ncbi.nlm.nih.gov/pubmed/23
242367).
https://en.wikipedia.org/wiki/Giant_panda 14/19
1/23/2019 Giant panda - Wikipedia

60. Li, R; Fan, W; Tian, G; Zhu, H; He, L; Cai, J; Huang, Q; Cai, Q; Li, B; Bai, Y; Zhang, Z; Zhang, Y; Wang, W; Li, J; Wei,
F; Li, H; Jian, M; Li, J; Zhang, Z; Nielsen, R; Li, D; Gu, W; Yang, Z; Xuan, Z; Ryder, O. A.; Leung, F. C.; Zhou, Y; Cao,
J; Sun, X; et al. (2010). "The sequence and de novo assembly of the giant panda genome" (https://www.ncbi.nlm.nih.
gov/pmc/articles/PMC3951497). Nature. 463 (7279): 311–317. Bibcode:2010Natur.463..311L (http://adsabs.harvard.e
du/abs/2010Natur.463..311L). doi:10.1038/nature08696 (https://doi.org/10.1038%2Fnature08696). PMC 3951497 (htt
ps://www.ncbi.nlm.nih.gov/pmc/articles/PMC3951497). PMID 20010809 (https://www.ncbi.nlm.nih.gov/pubmed/20010
809).
61. Jin, K; Xue, Chenyi; Wu, Xiaoli; Qian, Jinyi; Zhu, Yong; Yang, Zhen; Yonezawa, Takahiro; Crabbe, M. James C.; Cao,
Ying; Hasegawa, Masami; Zhong, Yang; Zheng, Yufang (2011). "Why does the giant panda eat bamboo? A
comparative analysis of appetite-reward-related genes among mammals" (https://www.ncbi.nlm.nih.gov/pmc/articles/
PMC3144909). PLoS ONE. 6 (7): 22602. Bibcode:2011PLoSO...622602J (http://adsabs.harvard.edu/abs/2011PLoS
O...622602J). doi:10.1371/journal.pone.0022602 (https://doi.org/10.1371%2Fjournal.pone.0022602). PMC 3144909
(https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3144909). PMID 21818345 (https://www.ncbi.nlm.nih.gov/pubmed/21
818345).
62. Li, De-Zhu; Guo, Zhenhua; Stapleton, Chris (2007). "Fargesia dracocephala" (http://www.efloras.org/florataxon.aspx?f
lora_id=2&taxon_id=200025413). In Wu, Z. Y.; Raven, P.H.; Hong, D.Y. Flora of China. 22. Beijing: Science Press; St.
Louis: Missouri Botanical Garden Press. p. 93.
63. Li, De-Zhu; Guo, Zhenhua; Stapleton, Chris (2007). "Fargesia rufa" (http://www.efloras.org/florataxon.aspx?flora_id=2
&taxon_id=200025444). In Wu, Z. Y.; Raven, P.H.; Hong, D.Y. Flora of China. 22. Beijing: Science Press; St. Louis:
Missouri Botanical Garden Press. p. 81.
64. Dolberg, Frands (1 August 1992). "Progress in the utilization of urea-ammonia treated crop residues: biological and
socio-economic aspects of animal production and application of the technology on small farms" (http://lrrd.cipav.org.c
o/lrrd4/2/dolberg.htm). University of Arhus. Retrieved 10 August 2010.
65. Lumpkin & Seidensticker 2007, pp. 63–64 (page numbers as per the 2002 edition)
66. "Pandas roam to find better bamboo" (http://www.australiangeographic.com.au/news/2014/07/pandas-roam-to-find-be
tter-bamboo). Australian Geographic. 25 July 2014. Retrieved 17 November 2014.
67. "Predator of giant panda" (http://wwf.panda.org/what_we_do/endangered_species/giant_panda/panda/kung_fu_pand
a_enemies_defences/). WWF.
68. Schaller, G.B., Jinchu, H., Wenshi, P., and Jing, Z. (1985). The giant pandas of Wolong. Chicago and London:
University of Chicago Press.
69. "Panda behavior & habitat" (https://web.archive.org/web/20080607110445/http://www.wwfchina.org/english/pandacen
tral/htm/learn_about_giant_panda/panda_q_a/panda_behavior_habitat.htm). World Wildlife Federation China.
Archived from the original (http://www.wwfchina.org/english/pandacentral/htm/learn_about_giant_panda/panda_q_a/p
anda_behavior_habitat.htm) on 7 June 2008. Retrieved 16 June 2008.
70. "Giant Panda" (http://nationalzoo.si.edu/Animals/GiantPandas/PandaFacts/default.cfm). National Zoological Park.
Retrieved 17 July 2008.
71. Dudley, Karen (1997). Giant Pandas. Untamed world (illustrated ed.). Weigl Educational Publishers. p. 23. ISBN 0-
919879-87-X.
72. Pandas Live by a Different Rhythm (http://www.laboratoryequipment.com/news/2015/08/pandas-live-different-rhythm)
published on 08/07/2015 by Michigan State Univ.
73. Paul Massicot (13 February 2007). "Animal Info – Giant Panda" (http://www.animalinfo.org/species/carnivor/ailumela.
htm). Animal Info. Retrieved 17 June 2008.
74. Deborah Smith Bailey (January 2004). "Understanding the giant panda" (http://www.apa.org/monitor/jan04/pandas.ht
ml). American Psychological Association.
75. "Teenager hospitalized after panda attack in Chinese zoo" (http://www.foxnews.com/story/0,2933,304249,00.html).
Fox News/Associated Press. 23 October 2007.
76. "Panda attacks man in Chinese zoo" (http://news.bbc.co.uk/2/hi/asia-pacific/7743748.stm). BBC News. 22 November
2008.
77. "Giant panda in China bites third victim" (http://www.cnn.com/2009/WORLD/asiapcf/01/09/panda.attack/index.html).
CNN News. 10 January 2009.

https://en.wikipedia.org/wiki/Giant_panda 15/19
1/23/2019 Giant panda - Wikipedia

78. "Animal Info – Giant Panda" (http://www.animalinfo.org/species/carnivor/ailumela.htm#Early). Retrieved 29 May 2009.


79. "National Zoo's Giant Panda Undergoes Artificial Insemination" (http://www.nbc4.com/news/15643501/detail.html).
NBC. Associated Press. 19 March 2008. Retrieved 13 April 2008.
80. Prapanya, Narunart (25 January 2006). " 'Panda porn' to encourage mating" (http://edition.cnn.com/2006/WORLD/asi
apcf/01/25/panda.passion/index.html). Time Warner. Retrieved 13 April 2008.
81. "Pandas unexcited by Viagra" (http://news.bbc.co.uk/2/hi/asia-pacific/2246588.stm). BBC News. 9 September 2002.
Retrieved 13 April 2008.
82. "Giant Panda Reproduction" (https://web.archive.org/web/20080527204441/http://newsdesk.si.edu/kits/pandas/nzp_p
anda_reproduction.pdf) (PDF). National Zoological Park. Archived from the original (http://newsdesk.si.edu/kits/panda
s/nzp_panda_reproduction.pdf) (PDF) on 27 May 2008. Retrieved 13 April 2008.
83. Kleiman, Devra G. "Giant Panda Reproduction" (https://web.archive.org/web/20080807161537/http://www.4panda.co
m/panda/pandatips/reproduction.htm). Archived from the original (http://www.4panda.com/panda/pandatips/reproducti
on.htm) on 7 August 2008. Retrieved 14 April 2008.
84. Ruane, Michael E.; Koh, Elizabeth; Weil, Martin (23 August 2015). "National Zoo's giant panda Mei Xiang gives birth
to two cubs hours apart" (https://www.washingtonpost.com/local/giant-panda-preparing-to-give-birth-national-zoo-say
s/2015/08/22/bba0fd2c-4804-11e5-8ab4-c73967a143d3_story.html). The Washington Post. Retrieved 24 August
2015.
85. "Panda Facts" (https://web.archive.org/web/20150924063658/http://www.pandasinternational.org/wptemp/education-
2/panda-facts/). Pandas International. Archived from the original (http://www.pandasinternational.org/wptemp/educatio
n-2/panda-facts/) on 24 September 2015. Retrieved 26 August 2015.
86. Dudley, Karen (1997). Giant Pandas. Untamed world (Illustrated ed.). Weigl Educational Publishers. p. 26. ISBN 0-
919879-87-X.
87. Guinness World Records 2013, Page 050, hardcover edition. ISBN 9781904994879
88. "Panda Update: September Cub Exam" (http://animal.discovery.com/videos/panda-update-september-cub-
exam.html). Discovery Communications, LLC. 4 May 2006. Retrieved 9 August 2010.
89. "Baby panda born from frozen sperm" (http://news.bbc.co.uk/2/hi/asia-pacific/8168491.stm). BBC. 25 July 2009.
Retrieved 26 July 2009.
90. "World's 1st giant panda born from frozen sperm in SW China" (https://web.archive.org/web/20091226013440/http://n
ews.xinhuanet.com/english/2009-07/24/content_11766726.htm). Xinhua News Agency. 24 July 2009. Archived from
the original (http://news.xinhuanet.com/english/2009-07/24/content_11766726.htm) on 26 December 2009. Retrieved
26 July 2009.
91. "First panda cub born using frozen sperm" (http://www.irishtimes.com/newspaper/world/2009/0725/1224251307368.ht
ml). The Irish Times. 25 July 2009. Retrieved 26 July 2009.
92. Tran, Tini (24 July 2009). "China announces first panda from frozen sperm" (https://www.usatoday.com/tech/science/2
009-07-24-panda_N.htm). USA TODAY. Retrieved 24 January 2011.
93. "Rare panda triplets born in China" (http://www.cbc.ca/news/technology/rare-panda-triplets-born-in-china-1.2734071).
cbc.ca. 12 August 2014.
94. Chen, D. Y.; Wen, D. C.; Zhang, Y. P.; Sun, Q. Y.; Han, Z. M.; Liu, Z. H.; Shi, P.; Li, J. S.; Xiangyu, J. G.; Lian, L.; Kou,
Z. H.; Wu, Y. Q.; Chen, Y. C.; Wang, P. Y.; Zhang, H. M. (2002). "Interspecies implantation and mitochondria fate of
panda-rabbit cloned embryos". Biology of Reproduction. 67 (2): 637–642. doi:10.1095/biolreprod67.2.637 (https://doi.
org/10.1095%2Fbiolreprod67.2.637). PMID 12135908 (https://www.ncbi.nlm.nih.gov/pubmed/12135908).
95. Schaller 1993, p. 61
96. Shuowen Jiezi, Chapter 10, radical 豸 (http://chinese.dsturgeon.net/text.pl?node=26160&if=en&searchu=%E8%B2%
98#n32614): "貘:似熊而黃黑色,出蜀中" ("Mo: like bear, but yellow-and-black, comes from Shu").
97. Erya, Chapter "釋獸" ("About animals") (http://chinese.dsturgeon.net/text.pl?node=39144&if=en): "貘,白豹" (Mo,
white leopard).

https://en.wikipedia.org/wiki/Giant_panda 16/19
1/23/2019 Giant panda - Wikipedia

98. China Giant Panda Museum: Historical Records in Ancient China (http://www.kepu.net.cn/english/giantpanda/giantpa
nda_know/200409230028.html). Supposed Chinese historical terminology appears in the Chinese version of this
article, 我国古代的历史记载 (http://www.kepu.net.cn/gb/lives/panda/know/now001.html) Archived (https://web.archive.
org/web/20120706193346/http://www.kepu.net.cn/english/giantpanda/giantpanda_know/200409230028.html) 6 July
2012 at the Wayback Machine
99. Duyvendak, J.J.L. (1939). "The True Dates of the Chinese Maritime Expeditions in the Early Fifteenth Century The
True Dates of the Chinese Maritime Expeditions in the Early Fifteenth Century". T'oung Pao, Second Series. 34 (5):
402. JSTOR 4527170 (https://www.jstor.org/stable/4527170)
100. Watson, DA. "The Panda Lady: Ruth Harkness (Part 1)" (http://femexplorers.com/full_article.php?article_id=17).
Female explorers. Retrieved 1 February 2007.
101. "Giant Pandas Through Singapore. Rare Animals from Wilds of China. Will be First to Reach Europe in Captivity" (htt
p://newspapers.nl.sg/Digitised/Article.aspx?articleid=straitstimes19381127.2.109&sessionid=9fb04643596f4181a3eba
55881450408&keyword=antenor+panda). Straits Times. 27 November 1938. Retrieved 2 February 2010.
102. Austin, A. B. (8 January 1939). "How Giant Pandas Arrived in London" (http://newspapers.nl.sg/Digitised/Article.aspx?
articleid=straitstimes19390108.2.79&sessionid=feffb0825aa74ad486527dce35b920cc&keyword=antenor+panda&sort
=&page=1&sort=). Straits Times. Retrieved 2 February 2010.
103. China's Panda Politics (http://www.newsweek.com/id/45901/). Newsweek. 15 October 2007. Retrieved 23 May 2008.
Archived (https://web.archive.org/web/20121010223523/http://www.newsweek.com/id/45901/) 10 October 2012 at the
Wayback Machine
104. China sends panda peace offering (https://www.theguardian.com/world/2008/dec/23/china-taiwan-pandas). The
Guardian. 28 December 2008.
105. "Panda Poop Might Help Turn Plants Into Fuel" (http://news.nationalgeographic.com/news/energy/2013/09/130910-pa
nda-poop-might-help-turn-plants-into-fuel). News.nationalgeographic.com. 2013-09-10. Retrieved 2013-10-02.
106. Li, Renqiang; Xu, Ming; Wong, Michelle Hang Gi (February 2015). "Climate change threatens giant panda protection
in the 21st century". Biological Conservation. 182: 93–101. doi:10.1016/j.biocon.2014.11.037 (https://doi.org/10.101
6%2Fj.biocon.2014.11.037).
107. The Panda is still endangered species, and the conservation efforts still need to be reinforced (http://www.forestry.gov.
cn/main/195/content-902929.html) State Forestry Administration of the People's Republic of China (in Chinese)
108. Pandas gain world heritage status (http://news.bbc.co.uk/2/hi/asia-pacific/5174854.stm) BBC News
109. Panda sanctuaries now World Heritage sites (http://www.upi.com/NewsTrack/view.php?StoryID=20060718-072442-2
033r) United Press International
110. Chris Packham: 'Giant pandas should be allowed to die out' (https://www.telegraph.co.uk/earth/wildlife/6216775/Chris
-Packham-Giant-pandas-should-be-allowed-to-die-out.html). Telegraph.co.uk. 22 September 2009.
111. Beyond cute and cuddly (http://www.theaustralian.news.com.au/story/0,25197,22730796-30417,00.html) Archived (htt
ps://web.archive.org/web/20090116040347/http://www.theaustralian.news.com.au/story/0,25197,22730796-30417,00.
html) 16 January 2009 at the Wayback Machine. The Australian. 10 November 2007.
112. "TV Packham says sorry for 'ditch pandas' blast" (https://web.archive.org/web/20091229234817/http://www.mirror.co.
uk/news/top-stories/2009/09/23/bear-raid-115875-21693846/). Daily Mirror. UK. 23 September 2009. Archived from
the original (https://www.mirror.co.uk/news/top-stories/2009/09/23/bear-raid-115875-21693846/) on 29 December
2009. Retrieved 24 April 2010.
113. Pimm, Stuart L.; Li, Binbin V. (2015). "China's endemic vertebrates sheltering under the protective umbrella of the
giant panda". Conservation Biology. 30 (2): 329–339. doi:10.1111/cobi.12618 (https://doi.org/10.1111%2Fcobi.12618).
114. "Earthwatch: On the Trail of Giant Panda" (http://www.earthwatch.org/exped/zhang.html).
115. Schaller pg.62.
116. "Giant Panda: Overview" (http://www.worldwildlife.org/species/finder/giantpanda/panda.html). World Wildlife Fund.
Retrieved 26 October 2011.
117. Goodman, Brenda (12 February 2006). "Eats Shoots, Leaves and Much of Zoos' Budgets" (https://www.nytimes.com/
2006/02/12/national/12panda.html). The New York Times. Atlanta. Retrieved 9 August 2010.

https://en.wikipedia.org/wiki/Giant_panda 17/19
1/23/2019 Giant panda - Wikipedia

118. "Zoo negotiates lower price to rent bears from China" (http://www.signonsandiego.com/news/metro/20081213-9999-1
n13panda.html). SignOnSanDiego.com. 13 December 2008.
119. "Animal Info – Giant Panda" (http://www.animalinfo.org/species/carnivor/ailumela.htm#Population).
www.animalinfo.org. Retrieved 2015-09-02.
120. "China's panda population increases by 17 per cent, major census finds" (https://www.telegraph.co.uk/news/worldnew
s/asia/china/11441960/Chinas-panda-population-increases-by-17-per-cent-major-census-finds.html). Retrieved
2015-09-02.
121. Brooks, Melody. "Summary – Giant Panda (Ailuropoda melanoleuca) Fact Sheet, 2001 – ResearchGuides at
International Environment Library Consortium" (http://ielc.libguides.com/sdzg/factsheets/giantpanda).
ielc.libguides.com. Retrieved 2015-09-02.
122. "How many are left in the wild?" (http://wwf.panda.org/what_we_do/endangered_species/giant_panda/panda/how_ma
ny_are_left_in_the_wild_population/). wwf.panda.org. Retrieved 2015-09-02.
123. "Panda Census 2013 | Wild Panda Population Increases to 1,864 | Pandas International" (http://www.pandasinternati
onal.org/wptemp/panda-census-2013-wild-panda-population-increases-to-1864/). www.pandasinternational.org.
Retrieved 2015-09-02.
124. Nickell, Joe (January 2018). "The Giant Panda: Discovered in the Land of Myth" (https://www.csicop.org/si). Skeptical
Inquirer. 42 (1): 12–14.
125. "Remains Of Earliest Giant Panda Discovered" (https://web.archive.org/web/20180311182107/https://www.sciencedail
y.com/releases/2007/06/070618174710.htm). ScienceDaily. ScienceDaily. June 19, 2007. Archived from the original
(https://www.sciencedaily.com/releases/2007/06/070618174710.htm) on 11 March 2018. Retrieved 11 March 2018.

Bibliography

AFP (via Discovery Channel) (2006, 20 June). Panda Numbers Exceed Expectations (https://web.archive.org/web/20
070325105323/http://dsc.discovery.com/news/2006/06/20/panda_ani.html?category=animals&guid=20060620103030
&dcitc=w19-502-ak-0000).
Associated Press (via CNN) (2006). Article link (https://web.archive.org/web/20060111121511/http://www.cnn.com/20
06/WORLD/asiapcf/01/06/china.taiwan.ap/index.html).
Catton, Chris (1990). Pandas. Christopher Helm.
Friends of the National Zoo (2006). Panda Cam: A Nation Watches Tai Shan the Panda Cub Grow. New York:
Fireside Books.
Goodman, Brenda (2006, 12 February). Pandas Eat Up Much of Zoos' Budgets (https://www.nytimes.com/2006/02/1
2/national/12panda.html). The New York Times.
Lumpkin, Susan; Seidensticker, John (2007). Giant Pandas. London: Collins. ISBN 0-06-120578-8. (An earlier edition
is available as The Smithsonian Book of Giant Pandas, Smithsonian Institution Press, 2002, ISBN 1-58834-013-9.)
Panda Facts At a Glance (https://web.archive.org/web/20061219071409/http://www.wwfchina.org/english/pandacentr
al/htm/wwf_at_work/panda_survey/q%26a.htm) (N.d.). www.wwfchina.org. WWF China.
Ryder, Joanne (2001). Little panda: The World Welcomes Hua Mei at the San Diego Zoo. New York: Simon &
Schuster.
Schaller, George B. (1993). The Last Panda (https://books.google.com/books?id=BkU6IwfjmYAC). Chicago:
University of Chicago Press. ISBN 0-226-73628-8. (There are also several later reprints)
Wan, Qiu-Hong; Wu, Hua; Fang, Sheng-Guo (2005). "A New Subspecies of Giant Panda (Ailuropoda melanoleuca)
from Shaanxi, China". Journal of Mammalogy. 86 (2): 397–402. doi:10.1644/BRB-226.1 (https://doi.org/10.1644%2FB
RB-226.1). JSTOR 4094359 (https://www.jstor.org/stable/4094359).
Warren, Lynne (July 2006). "Panda, Inc." National Geographic. (About Mei Xiang, Tai Shan and the Wolong Panda
Research Facility in Chengdu China).

External links
BBC Nature: (http://www.bbc.co.uk/nature/life/Giant_Panda) Giant panda news, and video clips from BBC
programmes past and present.
Panda Pioneer: the release of the first captive-bred panda 'Xiang Xiang' in 2006 (http://www.abc.net.au/catalyst/storie
s/s1755621.htm)

https://en.wikipedia.org/wiki/Giant_panda 18/19
1/23/2019 Giant panda - Wikipedia

WWF (https://web.archive.org/web/20080704204350/http://www.panda.org/about_wwf/what_we_do/species/our_solut
ions/endangered_species/giant_panda/index.cfm) – environmental conservation organization
Pandas International (http://www.pandasinternational.org/) – panda conservation group
National Zoo Live Panda Cams (http://nationalzoo.si.edu/Animals/GiantPandas/) – Baby Panda Tai Shan and mother
Mei Xiang
Information from Animal Diversity (http://animaldiversity.ummz.umich.edu/site/accounts/information/Ailuropoda_melan
oleuca.html)
NPR News 2007/08/20 – Panda Romance Stems From Bamboo (https://www.npr.org/templates/story/story.php?storyI
d=13746175)
View the panda genome (http://www.ensembl.org/Ailuropoda_melanoleuca/Info/Index/) on Ensembl.
Texts and pictures of the Panda exhibition at the Museum für Naturkunde Berlin (http://biowikifarm.net/v-mfn/panda/e
n)

Retrieved from "https://en.wikipedia.org/w/index.php?title=Giant_panda&oldid=877792646"

This page was last edited on 10 January 2019, at 22:51 (UTC).

Text is available under the Creative Commons Attribution-ShareAlike License; additional terms may apply. By using this
site, you agree to the Terms of Use and Privacy Policy. Wikipedia® is a registered trademark of the Wikimedia
Foundation, Inc., a non-profit organization.

https://en.wikipedia.org/wiki/Giant_panda 19/19

You might also like