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Biology
Biology is the natural science that studies life and living
organisms, including their physical structure, chemical
processes, molecular interactions, physiological
[1]
mechanisms, development and evolution. Despite the
complexity of the science, certain unifying concepts
consolidate it into a single, coherent field. Biology
recognizes the cell as the basic unit of life, genes as the
basic unit of heredity, and evolution as the engine that
propels the creation and extinction of species. Living
organisms are open systems that survive by transforming
energy and decreasing their local entropy[2] to maintain a
stable and vital condition defined as homeostasis.[3]
Sub-disciplines of biology are defined by the research Biology deals with the study of life and organisms.
methods employed and the kind of system studied:
theoretical biology uses mathematical methods to top: E. coli bacteria and gazelle
formulate quantitative models while experimental biology
bottom: Goliath beetle and tree fern
performs empirical experiments to test the validity of
proposed theories and understand the mechanisms
underlying life and how it appeared and evolved from
non-living matter about 4 billion years ago through a gradual increase in the complexity of the
system.[4][5][6] See branches of biology.
Contents
Etymology
History
Foundations of modern biology
Cell theory
Evolution
Genetics
Homeostasis
Energy
Study and research
Structural
Physiological
Evolutionary
Systematic
Kingdoms
Ecological and environmental
Basic unresolved problems in biology
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Etymology
Biology derives from the Ancient Greek words of βίος; romanized bíos meaning "life" and -λογία;
romanized logía (-logy) meaning "branch of study" or "to speak". [7][8] Those combined make the Greek
word βιολογία; romanized biología meaning biology. Despite this, the term βιολογία as a whole didn't
exist in Ancient Greek. The first to borrow it was the English and French (biologie). Since the advent of
the scientific era, reanalyzable as a compound using the combining forms bio + logy.
The Latin-language form of the term first appeared in 1736 when Swedish scientist Carl Linnaeus (Carl
von Linné) used biologi in his Bibliotheca Botanica. It was used again in 1766 in a work entitled
Philosophiae naturalis sive physicae: tomus III, continens geologian, biologian, phytologian generalis,
by Michael Christoph Hanov, a disciple of Christian Wolff. The first German use, Biologie, was in a 1771
translation of Linnaeus' work. In 1797, Theodor Georg August Roose used the term in the preface of a
book, Grundzüge der Lehre van der Lebenskraft. Karl Friedrich Burdach used the term in 1800 in a
more restricted sense of the study of human beings from a morphological, physiological and
psychological perspective (Propädeutik zum Studien der gesammten Heilkunst). The term came into its
modern usage with the six-volume treatise Biologie, oder Philosophie der lebenden Natur (1802–22) by
Gottfried Reinhold Treviranus, who announced:[9]
The objects of our research will be the different forms and manifestations of life, the conditions and
laws under which these phenomena occur, and the causes through which they have been effected.
The science that concerns itself with these objects we will indicate by the name biology [Biologie]
or the doctrine of life [Lebenslehre].
History
Although modern biology is a relatively recent development, sciences related to and included within it
have been studied since ancient times. Natural philosophy was studied as early as the ancient
civilizations of Mesopotamia, Egypt, the Indian subcontinent, and China. However, the origins of
modern biology and its approach to the study of nature are most often traced back to ancient
Greece.[10][11] While the formal study of medicine dates back to Pharaonic Egypt, it was Aristotle (384–
322 BC) who contributed most extensively to the development of biology. Especially important are his
History of Animals and other works where he showed naturalist leanings, and later more empirical
works that focused on biological causation and the diversity of life. Aristotle's successor at the Lyceum,
Theophrastus, wrote a series of books on botany that survived as the most important contribution of
antiquity to the plant sciences, even into the Middle Ages.[12]
Scholars of the medieval Islamic world who wrote on biology included al-Jahiz (781–869), Al-Dīnawarī
(828–896), who wrote on botany,[13] and Rhazes (865–925) who wrote on anatomy and physiology.
Medicine was especially well studied by Islamic scholars working in Greek philosopher traditions, while
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The discovery of the physical representation of heredity came along with evolutionary principles and
population genetics. In the 1940s and early 1950s, experiments pointed to DNA as the component of
chromosomes that held the trait-carrying units that had become known as genes. A focus on new kinds of
model organisms such as viruses and bacteria, along with the discovery of the double-helical structure of
DNA in 1953, marked the transition to the era of molecular genetics. From the 1950s to the present
times, biology has been vastly extended in the molecular domain. The genetic code was cracked by Har
Gobind Khorana, Robert W. Holley and Marshall Warren Nirenberg after DNA was understood to
contain codons. Finally, the Human Genome Project was launched in 1990 with the goal of mapping the
general human genome. This project was essentially completed in 2003,[23] with further analysis still
being published. The Human Genome Project was the first step in a globalized effort to incorporate
accumulated knowledge of biology into a functional, molecular definition of the human body and the
bodies of other organisms.
Cell theory
Cell theory states that the cell is the fundamental unit of life, that
all living things are composed of one or more cells, and that all
cells arise from pre-existing cells through cell division. In
multicellular organisms, every cell in the organism's body
derives ultimately from a single cell in a fertilized egg. The cell is
also considered to be the basic unit in many pathological
processes.[24] In addition, the phenomenon of energy flow
occurs in cells in processes that are part of the function known
as metabolism. Finally, cells contain hereditary information
(DNA), which is passed from cell to cell during cell division.
Research into the origin of life, abiogenesis, amounts to an HeLa cells with nuclei (specifically the
attempt to discover the origin of the first cells. DNA) stained blue. The central and
rightmost cell are in interphase, so the
entire nuclei are labeled. The cell on the
Evolution left is going through mitosis and its DNA
has condensed.
A central organizing concept in biology is that life changes and
develops through evolution, and that all life-forms known have a
common origin. The theory of evolution postulates that all organisms on the Earth, both living and
extinct, have descended from a common ancestor or an ancestral gene pool. This universal common
ancestor of all organisms is believed to have appeared about 3.5 billion years ago.[25] Biologists regard
the ubiquity of the genetic code as definitive evidence in favor of the theory of universal common descent
for all bacteria, archaea, and eukaryotes (see: origin of life).[26]
The term "evolution" was introduced into the scientific lexicon by Jean-Baptiste de Lamarck in 1809,[27]
and fifty years later Charles Darwin posited a scientific model of natural selection as evolution's driving
force.[28][29][30] (Alfred Russel Wallace is recognized as the co-discoverer of this concept as he helped
research and experiment with the concept of evolution.)[31] Evolution is now used to explain the great
variations of life found on Earth.
Darwin theorized that species flourish or die when subjected to the processes of natural selection or
selective breeding.[32] Genetic drift was embraced as an additional mechanism of evolutionary
development in the modern synthesis of the theory.[33]
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Genetics
DNA is found as linear chromosomes in eukaryotes, and circular A Punnett square depicting a cross
chromosomes in prokaryotes. A chromosome is an organized between two pea plants heterozygous for
structure consisting of DNA and histones. The set of purple (B) and white (b) blossoms
chromosomes in a cell and any other hereditary information
found in the mitochondria, chloroplasts, or other locations is
collectively known as a cell's genome. In eukaryotes, genomic DNA is localized in the cell nucleus, or with
small amounts in mitochondria and chloroplasts. In prokaryotes, the DNA is held within an irregularly
shaped body in the cytoplasm called the nucleoid.[37] The genetic information in a genome is held within
genes, and the complete assemblage of this information in an organism is called its genotype.[38]
Homeostasis
Homeostasis is the ability of an open system to regulate its internal environment to maintain stable
conditions by means of multiple dynamic equilibrium adjustments that are controlled by interrelated
regulation mechanisms. All living organisms, whether unicellular or multicellular, exhibit
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homeostasis.[40]
Energy
The organisms responsible for the introduction of energy into an ecosystem are known as producers or
autotrophs. Nearly all such organisms originally draw their energy from the sun.[41] Plants and other
phototrophs use solar energy via a process known as photosynthesis to convert raw materials into
organic molecules, such as ATP, whose bonds can be broken to release energy.[42] A few ecosystems,
however, depend entirely on energy extracted by chemotrophs from methane, sulfides, or other non-
luminal energy sources.[43]
Some of the energy thus captured produces biomass and energy that is available for growth and
development of other life forms. The majority of the rest of this biomass and energy are lost as waste
molecules and heat. The most important processes for converting the energy trapped in chemical
substances into energy useful to sustain life are metabolism[44] and cellular respiration.[45]
Structural
Molecular biology is the study of biology at the molecular level.[46] This field overlaps with other areas of
biology, particularly those of genetics and biochemistry. Molecular biology is a study of the interactions
of the various systems within a cell, including the interrelationships of DNA, RNA, and protein synthesis
and how those interactions are regulated.
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Genetics is the science of genes, heredity, and the variation of organisms.[48][49] Genes encode the
information needed by cells for the synthesis of proteins, which in turn play a central role in influencing
the final phenotype of the organism. Genetics provides research tools used in the investigation of the
function of a particular gene, or the analysis of genetic interactions. Within organisms, genetic
information is physically represented as chromosomes, within which it is represented by a particular
sequence of amino acids in particular DNA molecules.
Developmental biology studies the process by which organisms grow and develop. Developmental
biology, originated from embryology, studies the genetic control of cell growth, cellular differentiation,
and "cellular morphogenesis," which is the process that progressively gives rise to tissues, organs, and
anatomy. Model organisms for developmental biology include the round worm Caenorhabditis
elegans,[50] the fruit fly Drosophila melanogaster,[51] the zebrafish Danio rerio,[52] the mouse Mus
musculus,[53] and the weed Arabidopsis thaliana.[54][55] (A model organism is a species that is
extensively studied to understand particular biological phenomena, with the expectation that discoveries
made in that organism provide insight into the workings of other organisms.)[56]
Physiological
Physiology is the study of the mechanical, physical, and biochemical processes of living organisms
function as a whole. The theme of "structure to function" is central to biology. Physiological studies have
traditionally been divided into plant physiology and animal physiology, but some principles of physiology
are universal, no matter what particular organism is being studied. For example, what is learned about
the physiology of yeast cells can also apply to human cells. The field of animal physiology extends the
tools and methods of human physiology to non-human species. Plant physiology borrows techniques
from both research fields.
Physiology is the study the interaction of how, for example, the nervous, immune, endocrine, respiratory,
and circulatory systems, function and interact. The study of these systems is shared with such medically
oriented disciplines as neurology and immunology.
Evolutionary
Evolutionary research is concerned with the origin and descent of species, and their change over time. It
employs scientists from many taxonomically oriented disciplines, for example, those with special training
in particular organisms such as mammalogy, ornithology, botany, or herpetology, but are of use in
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Evolutionary biology is partly based on paleontology, which uses the fossil record to answer questions
about the mode and tempo of evolution,[57] and partly on the developments in areas such as population
genetics.[58] In the 1980s, developmental biology re-entered evolutionary biology after its initial
exclusion from the modern synthesis through the study of evolutionary developmental biology.[59]
Phylogenetics, systematics, and taxonomy are related fields often considered part of evolutionary
biology.
Systematic
A phylogenetic tree of all living things, based on rRNA gene data, showing
the separation of the three domains bacteria, archaea, and eukaryotes as
described initially by Carl Woese. Trees constructed with other genes are
generally similar, although they may place some early-branching groups
very differently, presumably owing to rapid rRNA evolution. The exact
relationships of the three domains are still being debated.
Further, each kingdom is broken down recursively until each species is separately classified. The order is:
Domain; Kingdom; Phylum; Class; Order; Family; Genus; Species.
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Outside of these categories, there are obligate intracellular parasites that are "on the edge of life"[64] in
terms of metabolic activity, meaning that many scientists do not actually classify such structures as alive,
due to their lack of at least one or more of the fundamental functions or characteristics that define life.
They are classified as viruses, viroids, prions, or satellites.
The scientific name of an organism is generated from its genus and species. For example, humans are
listed as Homo sapiens. Homo is the genus, and sapiens the species. When writing the scientific name of
an organism, it is proper to capitalize the first letter in the genus and put all of the species in
lowercase.[65] Additionally, the entire term may be italicized or underlined.[66]
The dominant classification system is called the Linnaean taxonomy. It includes ranks and binomial
nomenclature. How organisms are named is governed by international agreements such as the
International Code of Nomenclature for algae, fungi, and plants (ICN), the International Code of
Zoological Nomenclature (ICZN), and the International Code of Nomenclature of Bacteria (ICNB). The
classification of viruses, viroids, prions, and all other sub-viral agents that demonstrate biological
characteristics is conducted by the International Committee on Taxonomy of Viruses (ICTV) and is
known as the International Code of Viral Classification and Nomenclature (ICVCN).[67][68][69][70]
However, several other viral classification systems do exist.
A merging draft, BioCode, was published in 1997 in an attempt to standardize nomenclature in these
three areas, but has yet to be formally adopted.[71] The BioCode draft has received little attention since
1997; its originally planned implementation date of January 1, 2000, has passed unnoticed. A revised
BioCode that, instead of replacing the existing codes, would provide a unified context for them, was
proposed in 2011.[72][73][74] However, the International Botanical Congress of 2011 declined to consider
the BioCode proposal. The ICVCN remains outside the BioCode, which does not include viral
classification.
Kingdoms
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population ecology, although population biology is more frequently used in the case of diseases, viruses,
and microbes, while the term population ecology is more commonly applied to the study of plants and
animals. Ecology draws on many subdisciplines.
Ethology is the study of animal behavior (particularly that of social animals such as primates and canids),
and is sometimes considered a branch of zoology. Ethologists have been particularly concerned with the
evolution of behavior and the understanding of behavior in terms of the theory of natural selection. In
one sense, the first modern ethologist was Charles Darwin, whose book, The Expression of the Emotions
in Man and Animals, influenced many ethologists to come.[77]
Biogeography studies the spatial distribution of organisms on the Earth, focusing on such topics as plate
tectonics, climate change, dispersal and migration, and cladistics.
Origin of life. While there is very good evidence for the abiotic origin of biological compounds such as
amino acids, nucleotides and lipids, it is largely unclear how these molecules came together to form the
first cells. Related is the question of extra-terrestrial life. If we understand how life originated on earth,
we can predict more reliably which conditions are required to generate life on other planets.
Aging. At present, there is no consensus view on the underlying cause of aging. Various competing
theories are outlined in Ageing Theories.
Pattern formation. We have a good understanding of pattern formation in some systems, such as the
early insect embryo, but the generation of many patterns in nature cannot be explained easily, e.g. the
stripes in zebras or many snakes, such as coral snakes. While we know that the patterns are generated by
selective activation or repression of genes, many of these genes and their regulatory mechanisms remain
unknown.
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Bioinformatics – the use of information technology for the study, collection, and storage of genomic
and other biological data
Biolinguistics – the study of the biology and evolution of language
Biomechanics – the study of the mechanics of living beings
Biomedical research – the study of health and disease
Biophysics – the study of biological processes by applying the theories and methods traditionally
employed in the physical sciences
Biotechnology – the study of the manipulation of living matter, including genetic modification and
synthetic biology
Synthetic biology – research integrating biology and engineering; construction of biological
functions not found in nature
Botany – the study of plants
Phycology – scientific study of algae
Plant physiology – concerned with the functioning, or physiology, of plants
Astrobotany - the study of plants in space
Cell biology – the study of the cell as a complete unit, and the molecular and chemical interactions
that occur within a living cell
Chronobiology – the study of periodic events in living systems
Cognitive biology – the study of cognition
Conservation biology – the study of the preservation, protection, or restoration of the natural
environment, natural ecosystems, vegetation, and wildlife
Cryobiology – the study of the effects of lower than normally preferred temperatures on living beings
Developmental biology – the study of the processes through which an organism forms, from zygote
to full structure
Embryology – the study of the development of embryo (from fecundation to birth)
Gerontology – study of ageing processes
Ecology – the study of the interactions of living organisms with one another and with the non-living
elements of their environment
Evolutionary biology – the study of the origin and descent of species over time
Genetics – the study of genes and heredity
Genomics – the study of genomes
Epigenetics – the study of heritable changes in gene expression or cellular phenotype caused by
mechanisms other than changes in the underlying DNA sequence
Immunology – the study of the immune system
Marine biology (or biological oceanography) – the study of ocean ecosystems, plants, animals, and
other living beings
Microbiology – the study of microscopic organisms (microorganisms) and their interactions with other
living things
Bacteriology – the study of bacteria
Mycology – the study of fungi
Parasitology – the study of parasites and parasitism
Virology – the study of viruses and some other virus-like agents
Molecular biology – the study of biology and biological functions at the molecular level, some cross
over with biochemistry
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Nanobiology – the application of nanotechnology in biological research, and the study of living
organisms and parts on the nanoscale level of organization
Neuroscience – the study of the nervous system
Paleontology – the study of fossils and sometimes geographic evidence of prehistoric life
Pathobiology or pathology – the study of diseases, and the causes, processes, nature, and
development of disease
Pharmacology – the study of the interactions between drugs and organisms
Phycology – the study of seaweeds and other algae
Physiology – the study of the functions and mechanisms occurring in living organisms
Phytopathology – the study of plant diseases (also called Plant Pathology)
Psychobiology – the application of methods traditionally used in biology to study human and non-
human animals behaviour
Quantum biology – the study of the role of quantum phenomena in biological processes
Sociobiology - the study of social behavior in terms of evolution
Systems biology – the study of complex interactions within biological systems through a holistic
approach
Structural biology – a branch of molecular biology, biochemistry, and biophysics concerned with the
molecular structure of biological macromolecules
Theoretical biology – the branch of biology that employs abstractions and mathematical models to
explain biological phenomena
Zoology – the study of animals, including classification, physiology, development, evolution and
behaviour, including:
Ethology – the study of animal behaviour
Entomology – the study of insects
Herpetology – the study of reptiles and amphibians
Ichthyology – the study of fish
Mammalogy – the study of mammals
Ornithology – the study of birds
See also
Biology in fiction National Association of Biology Teachers
Glossary of biology Outline of biology
List of biological websites Periodic table of life sciences in Tinbergen's
List of biologists four questions
List of biology journals Reproduction
List of biology topics Science tourism
List of omics topics in biology Terminology of biology
Notes
a. For a more detailed list, see Outline of biology.
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Further reading
Alberts B, Johnson A, Lewis J, Raff M, Roberts K, Walter P (2002). Molecular Biology of the Cell (http
s://archive.org/details/molecularbiolog000wils) (4th ed.). Garland. ISBN 978-0-8153-3218-3.
OCLC 145080076 (https://www.worldcat.org/oclc/145080076).
Begon M, Townsend CR, Harper JL (2005). Ecology: From Individuals to Ecosystems (4th ed.).
Blackwell Publishing Limited. ISBN 978-1-4051-1117-1. OCLC 57639896 (https://www.worldcat.org/o
clc/57639896).
Campbell N (2004). Biology (7th ed.). Benjamin-Cummings Publishing Company. ISBN 978-0-8053-
7146-8. OCLC 71890442 (https://www.worldcat.org/oclc/71890442).
Colinvaux P (1979). Why Big Fierce Animals are Rare: An Ecologist's Perspective (https://archive.or
g/details/whybigfierceanim00paul) (reissue ed.). Princeton University Press. ISBN 978-0-691-02364-
9. OCLC 10081738 (https://www.worldcat.org/oclc/10081738).
Mayr, Ernst (1982). The Growth of Biological Thought: Diversity, Evolution, and Inheritance (https://b
ooks.google.com/books?id=pHThtE2R0UQC). Harvard University Press. ISBN 978-0-674-36446-2.
Hoagland M (2001). The Way Life Works (reprint ed.). Jones and Bartlett Publishers inc. ISBN 978-0-
7637-1688-2. OCLC 223090105 (https://www.worldcat.org/oclc/223090105).
Janovy, John (2004). On Becoming a Biologist (2nd ed.). Bison Books. ISBN 978-0-8032-7620-8.
OCLC 55138571 (https://www.worldcat.org/oclc/55138571).
Johnson, George B. (2005). Biology, Visualizing Life (https://archive.org/details/holtbiologyvisua00joh
n). Holt, Rinehart, and Winston. ISBN 978-0-03-016723-2. OCLC 36306648 (https://www.worldcat.or
g/oclc/36306648).
Tobin, Allan; Dusheck, Jennie (2005). Asking About Life (3rd ed.). Belmont, CA: Wadsworth.
ISBN 978-0-534-40653-0.
External links
Biology (https://curlie.org/Science/Biology) at Curlie
OSU's Phylocode (http://www.ohio.edu/phylocode/index.html)
Biology Online – Wiki Dictionary (http://www.biology-online.org/dictionary/Main_Page)
MIT video lecture series on biology (http://ocw.mit.edu/courses/biology/7-012-introduction-to-biology-f
all-2004/)
The Tree of Life (http://tolweb.org/tree/phylogeny.html): A multi-authored, distributed Internet project
containing information about phylogeny and biodiversity.
Journal links
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