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Handbook on Common
BUTTERFLIES
of U ttarakhand
PARMOD KUMAR
Zoological Survey of India, Northern Regional Station,
218 Kaulagarh Road, Dehra Dun 248 195
;;nA
ISBN 97~-81-8171-198-4
PRICE
Indian Rs. 750.00
Foreign : $ 40; £ 30
Published at the Publication Division by the Director Zoological Survey of India, 2341
4, AJe Bose Road, 2nd MSO Building, 13th floor, Nizam Palace, Kolkata 700020 and
printed at Mis Shiva Offset Press, Dehra Dun 248 001.
FOREWORD
The Lepidoptera (Butterflies) is an important group of insects whose members are
economically important for mankind from the time immmemorial. They play a key role in
nature as an important component of the food chain of birds, spiders and other predatory
insects. They also act as pollinators. The old relationship between the human and butterflies is
today going through a period of great stress as humans by their activities are directly or
indirectly threatening the existence of butterflies. The presence of butterfl ies depend upon the
variety of factors including the climate, suitable food, appropriate adult nectar sources of other
food, suitable areas for flight and courtship and in some instances, the presence of certain
symbiotic species, notably ants.
The actual number of known butterflies from Uttarakhand is 323 but the present pictorial
handbook contains a descriptive and illustrated account of 11 0 species of common and fascinating
butterflies belonging to nine families of the suborder Rhopalocera, order Lepidoptera that are
most common in Uttarakhand. The handbook contains the scientific names of each butferfly,
followed by the common name, general ecology, diagnostic characters, wingspan and distribution
in the State vis-a-vis in India.
It is expectedthat this volume will assist scientific workers in identification of these important
group of insects.
Dr. Ramakrishna
Director
Zoological Survey of India
Dated: 5st August, 2008
Kolkata - 700 053
CONTENTS
INTRODUCTION .............................................................................................................. 1
CLASSIFICATION ............................................................................................................. 3
PHySIOGRAPHy ............................................................................................................... 5
TOPOGRAPHY .................................................................................................................. 5
VEGETATION .................................................................................................................... 6
The descriptions are based on the collections made by survey parties of the Northern
Regional Station, Dehra Dun, since its inception in the year 1960, as also collected by author
during the last decade. The Handbook deals with 110 species representing nine families viz.,
Papilionidae, Pieridae, Danaidae, Satyridae, Nymphalidae, Acraeidae, Erycinidae, Lycaenidae
and Hesperiidae.
The handbook has been prepared keeping in mind its worthiness for the amateurs, naturalists
and students as an aid in field identification of the butterflies that fascinates the onlooker.
The colouration and patterns on the wings are very useful characters in identifying butterflies
an.d their classification. Characters like shape of wings, their margin and markings are important
morphological characters of taxonomic value. The figure of wings, showing different areas on
them would help readers to understand the text in a more meaningful way.
It is expected that the present attempt of the author would cater to the needs of the next
generation in understanding Butterflies.
Parmod Kumar
Zoological Survey of India,
Northern Regional Station,
Dehra Dun - 248 195
ACKNOWLEDGEMENTS
The aythor is thankful to Dr. Ramakrishna, Director, Zoological Survey of India, for his
kind pennission to undertake the study, to Dr. Arun Kumar, fonner Additional Director and
Shri P.T. Bhutia, Scientist-E and Officer-in-Charge, Northern Regional Station, Zoological
Survey of India, Dehra Dun for providing necessary facilities and encouragement throughout
the preparation of the document. I would not let the opportunity to go without expressing my
sincere thanks to Dr. G.S. Arora, an eminent Lepidopterist and fonner Joint Director and
Officer-in-Charge of the Station, for his guidance, help and literature provided, without which
the work would not have been completed. It is my pleasure to place on record my sincere
thanks to my senior colleagues namely, Dr. H.S. Mehta, Dr. P.C. Tak and Shri J.P Sati for
rendering manifold help, to Dr. S.K.Gupta for critically going through the manuscript and
providing useful and constructive suggestions.
My sincere thanks are also due to Scientific, Technical and Administrative Staff of Northern
Regional Station, for extending help during the completion of work and also thanks to Shri M.
Vijay, Senior Research Fellow for assistance in computer inputs.
Thanks are also due to Shri Rakesh Kumar, Photographer, Northern Regional Station, ZSI,
Dehra Dun for photographing the beautiful digital images ofbutterflies that have been included
in the document.
Last, but not the least, my grateful thanks are due to Dr. Vinod Khanna for guidance.
My sincere thanks are also to all of my family members who encouraged and supported me
during the writing of this handbook and for sparing the time, which was rightfully their's.
IN RODUCTION
Butterflies play a key fo) ,e in nature. They are important component in the food chain
of birds, spiders and others pr,edatory insects. They also act as poHinators. The old relationship
between the man and buttertlies is today going through a period of great stress as humans
by their activities are ditectly or indirectly threatening the existence of butterflies. The
pres'ence of butterflies depends upon the a vari,ety of factors including the climate., suitable
food, appropriate adult nectar sources, suitable areas for flight and courtship and in some
instances, the presence of certain symbiotic sp,ecies, notably ants.
The butterflies reach adulthood through ,a series of stages, viz., egg; larvae; chrysalis, and
finally the imago or the sexually mature stage. Thus, the life history of a butterfly shows a
compi'ete metamorphosis from larval stage to adult stag,e, ,and this period is variable in
different species. Human activities are disturbing and destroying the delicate balance that
sustains these facto~s, leading to the decrease in butterfly populations as well as their divers'tty.
Some of the threats faced by the butterflies today are habitat destrulction, use of pesticides,
illegal collection of butterflies and their trading.
Like birds, some butterflies also migrate but the butterflYlnigration often takes the form
of persistent flight in one direction over long distances.
,Many species of butterfli,es show mimicry. They may camouflage thernselves with leaf
like paUerns and shapes, like the leaf butterflies, of the genus Kallima.
D ~B TT RF
2 Zoological Survey of ["dia
V.9
V.7 _ _
V.6 ,.,...---~
V.5
V.4
FOREWING
HIND WING
The actual number of known butterflies from Uttarakhand is 323 but the present pictorial
handbook contains a descriptive and illustrated account of 110 species of butterflies that are
most cOlnmon in Uttarakhand.
The scientific names of each butterfly is folIowed by the common name, general ecology,
diagnostic characters, size of wingspan (in mIn) and their distribution.
The tenninologx used in the text and figure of wings of butterflies (Fig. 1) are as follows :
CLASSIFICATION
Butterflies are insects belonging to the order Lepidoptera. The order is divided into two
suborders, Heterocera and Rhopalocera. The Heterocera constitutes the Moths while
Rhopalocera constitute the Butterflies. Some authorities distinguish a third division, the
Grypocera, which comprise.s the family Hesperiidae (the Skippers).
The suborder Rhopalocera includes nine families i.e. Papilionidae, Pieridae, Danaidae,
Satyridae, Nymphalidae, Amathusiidae, Acraeidae, Erycinidae and Lycaenidae under a single
super-family Papilionoidea and Hesperioidea that includes only one family, the Hesperiidae.
The family Amathusiidae does not occur in the Uttarakhand.
HISTORICAL RESUME
The butterflies of the Indian Region, including those from western Himalayas, were
named and described by Linnaeus and Fabricius for the first time during the eighteenth
century. Doherty (1886) published a list of butterflies, exclusively from Kumaon, for the
first time, followed by Hannyngton (1910-11) who published an account of 373 butterflies.
Mackinon and de Niceville (1897-98) published a list of 323 butterflies from Mussoorie
and neighbouring region from the western Himalayas followed by Ollenbach (1930-31) who
collected, identified and published a list of 143 butterflies from Mussoorie.
The other important works referable to the study of Garhwal Himalayas are by Stemptfer
(1952); Bernardi (1952) and Lesse (1952).
Wynter-Blyth (1957) recorded as many as 415 species from western Himalayas out of
which 323, species were listed from Uttarakhand. Subsequently Arora and Mandai (1977)
recorded 45 species from Garhwal; MandaI and Chaudhary (1977) recorded 10 species froln
Va] ley of flowers, Chaturvedi (1981) and MandaI (1984) recorded 17 species froln Garhwal
Himalayas and adjoining slopes of Tons valley forest. Arora (1994) recorded 68 species froln
4 Zoological Survey of India
t
N
...
I.-
•
Rajaji National Park; Singh (1994) recorded 148 species froln New forest campus, Dehra
Dun; Arora, Ghosh and Chaudhary (1995) recorded 223 species from western Himalayas.
Arora (1997) identified 80 species of butterflies collected from Nanda Devi Biosphere.
Singh and Bhandari (2003) recorded 183 species from lower western Himalayan forests of
Doon Valley; Kumar and Kumar (1998) identified 21 species from Panchchuli (Dist.
Pithoragarh); Kumar and Gupta (2004) recorded 48 species from Govind Pashu Vihar,
Uttarkashi district.
Combining the lists as above, the present inventory of valid species of butterflies belonging
to nine families is somewhere nearly 323 species, of which the present work is restricted to
110 species the most common ones.
PHYSIOGRAPHY
Uttarakhand lies between 28 0 44' and 31 0 28' N latitude and 77 0 35' and 81 0 01 'E longitude.
It was carved out from the state of Uttar Pradesh by separating the hill region with a geographical
area of 53,483 sq km constituting 1.630/0 of country land area, (Forest Survey of India, 1999).
The State has 13 districts, namely Chamoli, Dehra Dun, Haridwar, Pauri, Rudra Prayag ,Tehri
and Uttarkashi in the Garhwal Division; and Almora, Bageshwar, Champawat, Nainital,
Pithoragarh and Udham Singh Nagar in the Kumaon division. (Fig. 2).
Boundary
Three districts of the state viz. Uttarkashi, Chamoli and Pithoragarh share an international
boundary in the North with Tibet (China) while in the East, Pithoragarh, Champawat and
Udham Singh Nagar share a boundary with Nepal.
Uttarkashi and Dehra Dun share its boundary with Himachal Pradesh in the Northwest
while in the South, Dehra Dun, Haridwar and a part of Nainital and Udham Singh Nagar
join the boundaries of Uttar Pradesh
Drainage
Ganges, Yamuna, Ramganga and Sharada are the four major rivers that originate frOln the
Uttar~khand Himalayas.
TOPOGRAPHY
1. Plains of Haridwar, Udham Singh Nagar, Dehra Dun district Bhabar and Terai area of
Dehra Dun, Garhwal and Nainital.
6 Zoological Survey of India
VEGETATION
The altitude of the state varies from 300 m to 3,500 In and above. Seven out of 16 forest
types existing in India can be found in Uttarakhand (based on Champion and Seth, 1968).
These are:
1. Moist Alpine scrub : At tree line around an altitude of 3,500 m. The major species are
Betula uti/is and Rhododendron campanulatum.
2. Sub-alpine forests: At altitudes of 2,900 m to 3,500 m above sea level in the middle and
upper Himalayas. The forests are characterized by patches of Abeis-Betula interspersed
with shrubby growth and grassy patches or alpine grasslands called bugya/s
3. Himalayan Dry temperate forests: In the inner dry Trans Hilnalayan valleys of the
state. Major species occurring in this forest are Cedrus deodara, Pinus wallichiana and
Juniperus species.
5. Sub-tropical pine forests: In lower regions of the Himalayas with pines as the dominant
specIes.
6. Tropical Dry deciduous forests : In the dry southern faces of Shiwaliks and adjoining
plains. These forests are open and mixed with major species as Shorea robusta, Anogeissus
latifolia, Terminalia tomentosa etc.
7. Littoral and Swamp forests : Occurs at a few locations in the valleys in the foothills
and are characterized by the presence of moisture loving species such as Syzigium cumini,
Ficus glomerata, Pterospermum acerifolium and Diospyros embrioptyris. The undergrowth
is characterized by the presence of cane Calamus ten ius.
The State of Uttarakhand is bestowed with varied vegetation types ranging from
tropical deciduous to alpine vegetation. In the lower altitude region up to 800 m, moist
tropical and dry deciduous forest of Sal, teak or mixed/pure forest of syzygium, Aegle,
Haldina, Acacia and Terminalia are found. At places, scrub forests having evergreen
species are also found.
KUMAR: H,an,dbook ,on Common Buft,erj1ies of Uttarakhan.d 7
8 ZOQlogical Survey of India
KUMAR: Handbook 011 Common Blitlel/lies of Utlarakhand 9
1. Family PAPILIONIDAE
Swallowtails
The species of the family Papilionidae consists of the largest and most beautiful of the
Indian Butterflies and cOlnmonly known as Swallow-tails. The Birdwings have a Wingspan
up to 250 mm. The members of this family are predominantly black or black-brown, except
Apollos, (Parnassius), which are white, spotted with red, blackened or blue spots and are
found in high altitudes in the Himalayas. The tail-less form are also abundant.
They are active, fast fliers and if disturbed immediately retract to the forest areas. Some
of the species of this falnily are fond of visiting and feeding on flowers and some settle on
damp ground, either alone or in congregation. Papilionid butterflies love basking in the sun.
They are mainly found in the forest and hilly regions but some of them are common in
the plains. They are characterized by their forewings being fully developed; the hind wings
with the pre-costal spur present, vein 1 b absent; claws are generally simple.
They are distributed up to 18,000 feet in Himalayan reaches. The caterpillars of most of
the species feed on pungent and evil tasting food plants including citrus, orange, lime, bel etc.
The average life span of an adult Swallow-tail butterfly may be as long as 4 months, but
generally up to 20-30 days. The eggs are spherical, smooth and white, green and blotched
The larvae have fleshy spines, but are never hairy. They have a pair of structure called
"osmeterium", a forked scent gland. This gland produces powerful chelnicals, which help in
repelling predators.
Some of the common species are Common Mormon (Papilio polytes) , Lime Butterfly
(Papilio demoleus), Common Peacock (Papillo polyctor) and Yellow Swallow-tails (Papilio
machaon).
10 Zo%gictl/ Sun1ey of India
Gener.,1 Ecology : The COlntnOn Ros'e is a red ~ bodied Swallowtail huuerlly of lower
elevations and is very cotnmon in the plains.
It is common during the rainy season. Male
emits a strong and slightly pungent odour. It
is migratory and roosts in large numbers
together with the Crimson Rose. The flight
is slow and del'iberate. Male forms of PapUio
polytes romulus Cralner form stich ius is an
excellentmimic of this butterfly. It is a
COlnmon visitor of garden flowers of Lantana,
Cosmos, and Zinnia etc. and visits wet soil.
It can be seen basking near treetops with it
wings spread out. nder s'l de Male
Diagnostic Characters: Antennae, thorax and ,abdomen above black~ h,ead and the whole
of thorax and abdolnen beneath vermiii on··red.
,Male and female: Tailed. Upper side: Wings black. Forewing area paler with black~ fold
stripes and well lnarked pale vein-stripes. Hindwing with a d'iscal row of elongate white
spots from 2 to 5 ,and a row of sub... marginal fled lunules; tail black. Under side' Discal spot
in the anal area 1 usua11y red. In male, scent "wool absent. Feluale similar, but wings broader
than in male.
Wingspan : 80-110 lnm.
Distribution : This specie is common throughout in India and recorded upto 3,000 feet
in North-west Hhnalayas. Also occurs in Sri Lanka and MyanInar. Uttarakhand : Recorded
from DehraDun, Haridwar, Pauri, Tehri and Pithoragarh districts andCOmlTIOn in the State.
1939. Cfrilosa ely/fa c/ylia (Linn.aeus) : Talbot, Fauna of British Jl.1dj.a (Butte,I:/lies) 1 : ) 16, figs . 28 .. 29.
General Ecology: This butterfly is found in hilly regions usually .at low elevations. They
congregate in large numbers on sandy banks of streaJns. It is plentiful in the pre-monsoon
and monsoon season. It feeds mainly frOlTI flowers of shrubs and small trees. It basks with
its wings spread flat or closed.
KUMAR: Halldbook on Common Bllu<' ,:/lies (~l U/tarakhand 11
This tailless s~'allowtail has two forms 'in northern India: elytia and dissil11ilis, both the
fonns are available in the state. The clylia milnics the Common Crow, Euploea core Cralner
and dissimilis Inirnics, The Blue Tiger, Tirultutl" Iilnniace (era.ner).
Diagnostics ,Char(lct,e~' : Form elytfa : Male and.felnale : Upper side: Wings rich dark
brown with cre,aITI-coioured markings" Forewing with sub-Jnarginal white spots .outwardly
truncat'e or ,elnarginated; the spot in ar'ea 4 shifted in\vards out 'Of lines~ those in areas 6" 7.
and 8 oblique to the ,costa, the low,est and the upper two spots elongate,~ a eri,. s of slnal1er
marginal white spots, two in area 1, one
above theother, and two in area 8; a single
sub~apical spot betw,e,en thelnarginal and sub-
marginal series" Hindwing with a discal series
of long arrow-shap,ed streaks in area I to 5,
a sub-marginal series of white crescents in
broadly divided across the middle by a bar
of tbe ground colour. Underside: Pale to
rich d.ark brown. Forewing markings as
above. Hindwing with a series of large yellow
terminal spots .
.Wingspan : 90-100 mIn. Upper side M.lIe form (/isSillij/t\
Form dissimilis : Male and felnale : Upper side: Wings black. Forewing with four 'crealny
streaks, joined at the base, a long streak in area I b, two streaks in area 1 with two spots"
streaks in are·a 21uuch broader with an outwardly emarginaled spot beyond' in area 3 a spot
at the base and one beyond, in area 4 a single similar spot, in area 5 and 6 ,elongate streaks,
and in 8 ;and 9 much smaller elongate spots; sub-ma~ginal and Jna~ginal Jnarkings ,as in tonn
c!ytia, Underside: Markings as upper side. Hindwing with large y,e llowlnarginal spots.
Wing.~pan : 90... 100 1nln.
Di~tribulion.: The subspeci,es is not very cornmon in its range of distribution. Oc'casionally
up to 7,500 feet in the HiJnalayas, usually 3,000 feet in the southern India and 5,000 feet
in Srianka. Uttarakhand .: Both the forms are recorded from Dehra Dun, Haridwar, Pauri
Nainital and Tehri districts.
12 Zoological Surv(,'J' (~llndia
1939. Papilla fJolytes fomulus Cramer: Talbot Fauna of British India (Bullel:!liefi) 1 : 178 .
Upper side l\lale form romulus Upper side FCluale rtJl1Ut/US form ~..tichius
Diagnostic Characters : Head, thorax and abdolnen black with white spots. Male and
.female : Upper side.: Wings black. For'ewing withrnarginaI se.ries of white or pale yellow
spots decreasing in size towards the apex. Hindwing: Upper side with complete discal band
of elongate white spots, lTIOre prominent in female, ending in red lunu l,es in the toroal region
and sub~marginal series of crimson lunules in female . Under side DuH coloured. The InaJ,es
are slnaller in size.
1909a. PapiUo pro/enol' protenor (Cramer) : Jordan, In Seitz, Macrolep . Fauna Indo-A listral., '9 : 75-76.
Distribu.tion : The subsp·ecies is not uncommon within its range of distribution from
Kashmir to Kumaon., rather rar'e west of Mussoorie. The Spangle fl i'es in the North-West
Himalayas upto an altitude of 8,,500 f:eet. Also occurs in Nepal. Uttarakhand : Almora,
Bageshwar, Chamoli, DehraDun, Haridwar, Pauri, Rudra Prayag, Pithoragarh, Tehri and
UUarkashi districts.
14 Z,o%gi,ca.l Survey (~r I"dia
1895. Papilio po/yetor polyetor (Boisduval) : Rothschild, A f,evision of the Papilios 'Of the Eastern Hemishere.
Nov. Zool., 2 : 381.
General Ecology: This beautiful butterfly flies throughout the Himalayas and is comrnon
in wann valleys. Dry season fonn may be seen settling in large numbers on damp patches
of mud and the bank of streams, and w,et season form within gardens. It visits the flowers
of Lantana Jasmine etc., and generally hovers for a long time over flower.
Diagnostic Characters : Antennae, head., thorax and abdomen brownish..bla,ck; the head,
thorax and abdomen above thinly irrorated w'th green scales above.
Male and female: Similar. Upper sid,e : Wings dark black and tailed. Male with woolly
scent stripes on forewing. Forewing with diffuse golden green irroration, becoming bluish
,anteriorly. Hindwing with less dense golden-gre,enirroration, turning to blue on the ,anterior
portions of the wing; a broad bright blue upper discal patch, broad between areas 4 and 1,
and has its outer edge uneven, occupying the base of area 4 and tbe outer portions of areas
5,6, and 7; posteriorly this patch is continued in ar'eas I to 3 by much smaller diffuse
quadrat,e spots of brilliant golden-green
scales; tomus with three or four 'crarets-
red sub-marginal crescents. Underside :
Chocolate brown, somewhat thin .l y
irrorated with yellowish scales, which
are, however, absent from a more or less
triangular p,atch in the middle of the
forewing post'eriorly. Hindwing without
discal patch; sub-marginal series of
claret-red ,c rescents, ea'c h crescent
transversed inwardly by a line of
purplish blue, follow,ed by velvety black
spots and broad white m,a rginal
crescents. Upper Side Male
Distribution : The subspecies is common froln Chitral (Pakistan) to Kumaon from low
,elevation to about 7,500 feet. Uttarakhand: Dehra Dun, Haridwar, Pauri, Nainital, Pithor,agarh,
Udham Singh Nagar, Champawat, Tehri and Uttarkashi districts.
KUMAR : Ha 1.1 db O,ok ,on Common Butterflies of Uttarakhand 15
1939. P,apilio demoleus demoleus . innaeus, Talbot, Fauna 0/ British India (Buttet:/lies) 1 : ) 89.
General Ecology: It is also known as the Lemon Butterfly or ime Swallowtail. The
most common butterfly occuring throughout the country. Visits garden flow'ers and flowers
of all Citrus, Radish and Curry leaf plants during and after monsoons. It has a very quick
& straight flight. It is an op'en plain species but also found in smaller numbers in forested
country. They aggr,egate in hundreds and mud puddle with ,closed wings. As it grows old,
the yellow markings on the wings frequently tum to deep orange.
Diagnostic Characters: Antennae dark reddish-brown; head, thorax and abdomen dusky
black. Palpi, thorax and ,abdomen beneathcrealn yellow.
1895. Papillo mac/lOon punjabcllsis imler, Arb. Ve,,: Bei Schmell. , 2 : 104.
1939. Papilio machaoJl pLmjabellsis Eimer: Talbot, Fauna of British India (Buller/lies)., 1 :1 96 .
•
General Ecology. It is a tailed, black and yellow butterfly. A fast flier but frequently
interrupted by visits to low growing flowers over which it rests. It is fond of hill tops
and visits garden flowers. It inhabits the open, grassy, flower-,c ov,e red tneadows of the
hills.
Diagno51ic Characters: Antennae black; head., thorax and abdomen with crealny colour
with black lines. Abdomen beneath with narrow lateral ill-defined black lining.
Male and felnale : Upper side Forew ing with the ground colour bla,ck, dusted with
yellow-slcales, rest of cell yellow with a black bar and another bar at its end. Discal area
yellow with bla,ck veins. Marginal area black with a cOlnplet'e series of regular slnaH
yellow spots near the terlnen.
Hindwing basal half creamy yellow
'with bl.ack veins, 'Outer half black with
a series of diffused blue spots and
marginal series of creamy-yellow large
crescent spots. Blue .. topped red spot
at tornus. Under side : Pale-cream
colour. Forewing with two transverse
short bands across the cell. Hindwing
veins prominently black. P. machaon
punjabensis Eimer is ,close to Papi/io
machaon asiatica Menetries that
occurs from Chitral (Pakistan) to
Kashmir and Tibet.
Upper 'Side Male
'939. Popillo paris paris Linnaeus : Talbot, Fauna ql British India (Bultel1Ues) 1 : 150.
General Ecology : This beautiful and large Butterfly is very cOlnrnon in the wooded
regions .of evergr,e,en forests.. (t is abundant during the southw'est Inonsoon Inonths. Both
sexes visit damp patches, sOlnetimes settling with their wings held flat against the ubstratulTI,
It visits flowers .of Lantana and Citrus spp.
Diagnostic Characters : Antennae, head, thorax and abdolnen black sprinkled with
green scales above.
Distl"ibution : The subspecies ranges from KUlnaon eastwards to Orissa, and is COIHlnon
fr.om 5.,000 to 7,000 feet in Hitnalayas. Also oc'curs in Myanlnar. Uttarakhand : Dehra Dun .
Haridwar, Nainital and Uttarkashi distdcts,
1939 . ,Graphium Clool1thus cloan(Iu,15 (Westwood) : Talbot. Fauna oj British Il.ldi,a (BlIllel:/lies) 1 : 229.
Gene~al Ecology : The Glassy Blue Bottle occurs in the hilly areas. It visits da'inp
patches, where m,ales can be seen ,congregating. It has a rapid 'flight and is fond of cir,cling
i8 Zoological Sun'ey of Indic,
round trees~ ,especially those on hi II lOpS. It is attracted to flowers of Buddleia and Horse
chest-nut.
Diagnostic ,Characters Antennae, head, thorax, and abdolnen dark brownish ~ black,
thorax with lateral dark grey pubescence, underside of palpi, thorax and abdomen touched
with dingy white, the abdomen ith three
lateral \fhitish stripes.
Distribution : The subspecies is common at low el,evation upto 4,500 feet, but also
recorded as high as 9,.000 fe,et in the Shilnla hills. It ranges from Kashmir to Assam , Also
occurs in Myanmar.Uttarakhand : Dehra Dun, Haridwar, Pauri, Nainital, Tehri Rudra Prayag,
Chamoli, Almor;a, Pithoragarh, and Bageshwar districts.
General Ecology : This is an extremely local butterfly. These butterflies are found in
open places in fore, ted areas between 3000 feet to 8000 feet Males fly high around some
select,ed tree, often a number of them together, settling high up now and then, though
occasionally sweeping down low enough to be taken in the net. The females are not so Ii vely
and fly c'lose to the ground and often settle while searching for host plant on which they lay
their eggs.
Diagnostic Characters Male and fe.male : Upper side: Whitish ,abov,e, head, thorax and
abdom'en creamy whit,e. Fore'wing semi-transpar,ent with ten bla,ck bars or bands of 'which
KUMAR: H,a ndbook on Common Bulle rfUe~' of IU uarakhand 19
the first, second and third marginal bands more or less reach the dorsum. Hindwing with
black tornal patch dusted with greyish blue and a double yellow spot in 1 and 2 above tbe
area. Under side : Hindwing with Inarginal black bands, a complete row of ,central pale
yellow spots of which tornal ones are yellow. Male with no scent wool in dorsal fo ld.
Wingspan : 60-70Inln.
Distribution : The species is distributed from Kashmir to Sikki'TI and orth-east India.
Also occurs in Nepal, Bhutan and Myanmar. Uttarakhand : Nainital and Uttarkashi districts.
1932. Pathysa nomius nomius (Esper) : Evans, Idellt{{icalion of Indian Butfe'ilies, J. Bombay nal. Hisl.
Soc., 29 : 238, pi: 5, fig. A., 1932a. p, 54.
Male and female ' . Upper side : Whitish above. Forewing with five dark bands and with
a broad dark rnarg in bearing a row of white rounded spots. Hindwing with broad outer
band containing white crescents and a patch of grey suffusion at the tomus.Unders'ide :
Whitish markings brownish nearly as on upper side. Hindwing with central red spots. Males
with cottony scent wool in dorsal fold. 'Tail long and black,
General Ecology: Tbe Common Blue Bottle is the strong,est flier of this group, settling
seldom and only momentarily and often difficult to catch. It feeds on nectar and visit flowers
hurriedly. The males often congregate in large numbers for mud-puddling. It occurs in the
evergreen and semi-evergreen forests. It is found mainly at low elevations.
Diagnostic Charact.ers : Male and female : Upper side: Wings brownish-black with a
short tail. Forewing with pale blue or greenish blue discal band running from inner margin,
narrowing anteriorly towards apex and ending in a sman spot just before the apex . Hindwing
with the band scaled with white, continue
from below the inner margin of for'ewing to
basal part of hiridwing, narrowing
postleriorly and with sub-marginal semi-
crescents shaped blue spoOts; costal part of
band whitish; scent fold on the inner margin
greyish. Underside : Brownish-black with
hand 'm or,e bluish than the upper side.
Forew' ng with slight indications of sub...
marginal spots before the tornus. Hindwing
with a red spot :at the base and the disc
with five red spots.
Upper side Male
Wingspan : 80 90 mm.
Distribution: The su species is common from Kashmir to Sri Lanka and commonest at
low elevations, but in South Indian hills and Himalayas,it has been recorded upto 9,000 fe,et.
Uttarakhand: Dehra Dun, Haridwar and Tehri dis·tricts.
KUMAR: Handbook 011 Common Butte/~t7ies of U((arakhalld 21
1932. Parnassius fl,ar.dwickei hardwickei (Gray) : Evans, Identific.atioJl of Indian Buttel/lies., J. Bombay nut.
Hist. Soc., 29 : 59.
General Ecology: The 'Colnmon Blue Apollos are h'igh altitude butterflies and rarely
found below 7,500 feet These butterflies are hardy and tough. They are strong fliers and fly
,close to the ground in the sunshine. They are found usually in open alpine cQuntry or low
growing shrubs and junipers. They visit flowers of Asters, Senecio. and Tll~axaCUln spp.
When the weather is cloudy or cold, and towards evening, they become very sluggish. ~the
femal'e is seen much less often than the male.
Diagnostic Characters: Antennae bl,ack, with only a f w white specks; head with brownish-
yellow pubes,cence; thorax and abdomen densely cov'ered with long whit,e hairs, Inore or less
narrowly, ,also clothes, the inner margin of hind-wings. Male : Upper side : Wings crearny
white. Forewing with base and costal area d nsely irroratedwith black scales; a broad, short,
velvety black bar across lniddle of cell, another at its apex and a third beyond the apex, the
last mentioned bearing a spot of crimson; a crilnson-'c'entered black spot in middle of area
1; curved prominent post.. basal series of dusky black spots. Hindwingwith base and inner
margin broadly dusky black:; a crilnson centered black spot near base of area 5, and another
at middle of area 7, followed by a curved marginal series of blue spots ringed with black
and centred with white. Under side: Silnilar as upper side with a glossy appearance. Forewing
with markings visible froln the upper side. Hindwing with a brQad basal band above the
tomal angle; ,aU the crimson spots encircled by black rin,g sand following centred with white
spots in area 2 and 7 .and the basal andlnedian spots in area 7. Felnaie : Upper side: Shnilar
to male, forewing with the dusky black irroration more extensiv,e. The critnson spots larger,
with additional spots in area 6 of forewing and a pre-tornal sp'ot on the hindwing. Under side:
Similar to male but all the red spots much larger and centered with white.
Wingspan ,: 50 .. 65 min .
Distribution: The species ranges froln Chitral (Pakistan) to, KU(l1aOIl., and found between
7 500 feet to 13,000 feet. Uttarakhand: Ahnora, ChaJnoli, Nainital and Pithoragarh districts.
Zoological Slln'e.~ ollndi,(I
1939. Graphillm agamemnon ( innaeus) Talbot, Fauna (?f British India (Bulle/~fljes) I : 229.
Ge';era/ Ecology: This is an extretnely restless black butterfly with yellow-green spots.
It starts it ~ activi,ty in the tTIorning and ,ends late in the afternoon. It vibrates it5wings
constantly while feeding . It regularly visits garden flowers such as Lantana, red 'flowers,
ixora, Mussaenda etc. It is not fond of visiting wet soil.
Diagnostic Characters : Antennae, head, thorax and abdomen black; thorax above ,and
abdolnen on sides streaked with greenish grey; the thorax touched with pink beneath.
Male and,female : Upper side Wings blackish-brown with yellow-green spots. Forewing
with a spot at extrelne base of costa, a tr.ansverse short bar near bas,e of cell, and seven spots
beyond , two spots beyond apex of cell; a spot at base of areas 1a and 1, fol owed by two
oblique short macular bands; .a discal series 'Of spots decreasing in size towards the costa,
and a post-discal series of smaner spots that begins with two in area 1., the spots in ar1ea 7.
Hindwing with a basal stripe, ,ending in .a pO'int in area I, a discal stripe interrupted at lower
dge of cell and not reaching
beyond the middle of area 2; two
post-discsl series of spots; these
Inarkings are white in area 7.
Hindwing with tail, which is long r
in the fe'male than in the male.
Underside: Paler, the green spots
partly covered with white or
brownish scales, both wings
clouded with violet gr,ey. Hind
wing between costa and cell has a
black cresClent, which is basically
margined with red between vein 8
and cell. Male with scent wool 'I n
the dorsal fold.
Upper . de Male
Wingspan : 85-100 mIn.
Distribution The species is comlnon from Kumaon to Assam. Also occurs in Myanmar.
Uttar,akhand : Dehra Dun H,aridwar, P.auri and Pithoragarh distri1cts.
KUMAR: Handbook on CommoJ1 Bllfle/~/7ies of UlfaraklIand 23
2. Family PIERIDAE
White and Yellows
The Inelnbers of family Pieridae are slnall to Inedium sized. They have a predolninance
of white and yellow colour, hence they are called 'Whites and Yellows' This family is
alnong the slnaller families of butterflies. 109 species are recorded in India.
The family is divided into two sub-falnilies; the Whites belong to Pierinae and the
Yellows, to Coliadinae.
These are sun loving butterflies especially males and sOlnetilnes found in large
congregations on river or strealn beds or damp patches during the hot weather, where they
are seen sucking Inoisture and salts. Females are Inostly solitary, preferring shady habitats.
The flight is usually slow, sluggish and close to the ground. They visit various types of
flowers and some species bask with their wings fully open and others partly open.
They are characterized by the presence of fully developed fore legs in both the sexes, the
inner Inargin of hind wing is channeled to receive the abdomen. Vein 1 b is present.
The seasonal and sexual variations are quite comlnon and well Inarked in some species
(Eurema, Catopsilia etc.).
They are distributed throughout the peninsular India and occur up to 15,000 feet in
Hilnalayas. The caterpillars are cylindrical, green or brown with stripes and pubescence.
They feed on plants of Brassiacae, Fabaceae, and Cappraridaceae.
Some common species of this family are Leptosia nina (Fabricius), Valeria valeria
(Fabricius), Delias eucharis (Drury), Pieris brassicae Doubleday etc.
24 Zoological Survey ollndill
Subfamily PI RINAE
1939. Leplosia nil1,a nina (Fabricius) : Talbot, Fauna of British India (Buttct:lhes) I : 305.
General Ecology : This is a little, white butterfly with black marking on the wing tips.
The under side of the wings is faintly marked with green.. It can be identifi,ed from any
distance because of its typica'\ flight. It is a very weak flier and flies near the ground in the
under growth and bushes. When it settles, it sInks the forewings back into the hindwings.
Diagnostic Characters : Head br.ownish; antennae dark br.own, spotted with white; thorax
and ;a bdomen whitish.
General Ecology: The Great Blackvein has a heavy floating flight and prefers open and
wooded country. They are very fond of Horse-ch stnut blossotns.
Diagnostic Characters : Antennae, head, thorax and abdomen black; abdomen beneath
white.
Male and female .: Upper side: Both the sexes ar,e black with white or dusky discal and
sub-marginal stripes on wings, cell of the hindwing white. Under side: Similar, the hindwing
at the base of costa bright yellow.
Distribution : 'T he species ranges from Kashmir eastwards through North-West and western
Himalayas . Wynter Blyth (1940) states that both the subspecies are common up to Garhwal
above 5,000 feet.
1910. Aporia agathon pJuyxe (Boisduval) : Fruhstorfer,ln : Seitz, .Macro/ep. Fauna ()llnd() ~Aust"" 9 ~ 139.
Diagnostic Characters : Male andfelnale With ground colour black and darker than in
phryxe (Boisduval).Upper ~ ide : Forewing heavily dusted with black. Both wings with tbe
e,en whitish. Under side Paler, markings as on upper side' sub apical spots yellowish.
J857,. Aporia soractaMoore, In: Horsfield and Moore's Cat. Mus. E.l.C., 1 : 83.
Male and female .: Upp,er side: W'ings Inilky white. Forewing with black veins. Forewing
with a post-discal black band, which is usually obsolete in area 3; a he,a vy disco-cellular bar;
a thin marginal black Hne from base of costa to base of inner margin, more or less interrupted
at area 1. Hindwing veins bla,c kand unmark,ed.. Under side : Forewing white; ,a pex and costal
edge paJ'e yellowish with black markings as on upper side but narrower. Hindwing p,a le
yellow., the pre",costalarea chrome yellow but the vein heavily marked.
1881. DeJias eucharis (Drury) : Moore, Lep. Ceylon, 1 : 140, PI. 54, fig. 1., la.
General Ecology; A very beautiful butterfly, seen commonly ,around gardens and forest
edges. It is very fond of visiting flowers of Lantana camara, Anacarditan occidentale.,
Tridax procumbens, mango inflorescence and wild flowers. Females are known to fly higber
than the males in sear-ch of food plants to lay their eggs. Males frequently are found on
flowers or occasionally drinking from wet patches.
Wingspan : 66 . . 85mm.
Distribution : It is the only widely distributed species of Delias in India, found everywhere
up to 7,000 feet in Himalayas and hjlls of Assam. Also occurs in Sri Lanka and Myanmar.
Uttarakhand : Dehra Dun., Haridwar, Almor,a, Bageshwar, Champaw,at, Nainital, Pauri, Tehri
and Uttarkashi districts.
28 ZOO/OgiC(I/ Survey (~( India
1925.De/ias belladonna horsfieldi (Gr,ay) : Jorden "On Delias belladonna H()r.~/leldi and a/Ued species
"Nov. Zool.. 32 : 285.
General Ecology It is a fairly l.arge, striking butterfly and v,ery common in the lower
v.alleys froln June to November. It is found in open areas, around fields, forest edges and
disturbed human habitations. They are fond of visiting dalnp patches and flowers of Lantana
Budd/eta, Horse Cbestnut, and Aesculu ' indicum. It is easily identified by the extensive
black, red and yeUowlnarkings.. Its flight is slow and lazy and can be rapid and irregular
on disturbance. Males fly close to the ground.
Diagnostic ,Characters : Male and jeJnaie ~ Ground colour of wings black with white to
pale cream coloured spots. Upper side: Forewing with cell spot at tbe end white; post. . disca\
and sub-marginal series of white spots. Hindwing 'with a large basal y How spot; ·a tarnal
yellow' patch and white inner area, post discal and sub marginal series of white spots.
Und,er sid'e : Forewing : Markings larger and more sharply defined than upperside, the
cell stripes on both wIngs prominent. Forewing with a bar across the discal end of the cell.
Hindwing : Basal spot in .area 7, cell end spot ,and inner area along with the tornal patch
entirely yeHow; post-discal series of spots, from above tornal patch to area 6., whitish.sub ..
Inarginal series of spots, yellowish, slnal1 and rounded froln beyond tornal patch to area 4,
and elong,ated in ar'ea 5 and 6.
Distribution : The subspecies is fairly common. Its r.anges from Hilnalayas to Kulu
(Himachal Pradesh) eastwards and common at 2;000 to 10,000 feet in Himalayas. Uttarakhand :
Dehra Dun, Haridwar, Pauri, Tehri, Chamoli, Nainita'l, Ahnora, Pithoragarh, Bag,eshwar,
Champawat and Udham Singh Nagar districts.
KUMAR: Handbook 011 Common Buttel:flies ,of Uuarakhand 29
1932 . Anap.haeis auro,ta (Fabricius) : Hemming " The Butterfl i,es of Transjorden" Trans. Ellt. Soc
Lon.d.. 80(2) : 283.
General Ecology : 'This species is quite conunon, a strong flier and fond of sunshine.
Both sexes visit flowers, males found on damp patches.
Diagnostic ,Characters : Antennae black, sparsely dotted with whi te; head, thor,ax and
abdolnen white above. SOlnetimes thorax bluish-grey.
Male and f emale : Upper side: Wings white. Forewing with costal area to base of vein
11 dusky-black; a prominent curved jet-black disco-cellular bar.; a broad black api'cal area,
narrowing to vein 2.. Hindwing with a marginal black border froln vein 2 to 6, bearing four
round white spots in areas 2 to 5. Under side: Forewing similar to the upper side, the
markings more clearly defined and the apical spots larger. Hindwing yeHowish~white, aU
the veins very bro.adly bordered with bla'c k;areas 1,,2;6, and 7 with black cross bars:~m,arginal
bla,ck border with white spots as on upper side. Female: Reselnbles the male, but the black
Inarkings broader, white apical spots of forewing often partly obsolete on up'per side. Upper
side of hindwing varying from white (Dry season form) to yeUow (W,et season form), with
black veins.
Wingspan : 40-55 mm.
Distribution : 'This sp1ecies is common throughout India, except ASSalTI, and ascends
to about 8,500 feet in Himalayas. Uttarakhand: Dehra Dun, Haridwar, Bageshwar,
Champawat, Pauri, Pithoragarh., Tehri, Uttarkashi, Nainital AI'mora and Udhaln Singh
Nagar districts.
30 Zoological Survey of '",d.ia
1'907. Ceporia neriss.a plllyne (F.abricius) = Bingham, Fauna of British India (Butte,ilies)" 2 : 18S.
General Ecology: It isa very activ,e butterfly, It tli,es among the bushes, grass and
dry dec'iduous forests of the plains and the low lying hills of India .It spends much titne on
flowers in search of nectar. In sum:mer, the males gather in large numbers on patches of wet
soil. It is most common between the pre-monsoon and post-monsoon months, the numbers
get decT,eased gradually towards the end of monsoon as well as in the dry season.
Diagnostic Characters: Male: Upper side: Wings white. Forewlng with apical black area
bearing prominent white spots in the middle of area 2. Hindwing with outer marginal black
border dentate on the inner edge. Underside : yellowish-whit . Felnale : Upper side Wings
in male, with extensive blackish.. brown
markings. Forewing with a sub-marginal black
spot in ,area I b. Hindwing with sub-marginal
white spots within the blackish .. brown
marginal border; upper margin of cell
blackened, the outer vein less darkened than
on the for1ewing. Under side : Fof,ewing as .on
upper side. Hlndwing much paler. In Dry
season form, the black markings are reduced
in both the sexes. Wet season forms ar,e
heavily marked.
Wingspan : 40.-65 mm. Under side Female
1939. Pieris canidia indica Evans, Talbot, Fauna of British India (Butterflies) 2 : 425-426.
General Ecology: This is ,a predominantly white butterfly, very 'c ommon at the edges of
cultivated fi,elds, grasslands, on roads, tea estat'es and vicinity ,a nd visits flowers of various
species.
Diagnostic Characters : Male and female: Upper side: Wings white. he apex of
forewing on the upper side is black with a few terminal black spots, a prominent black spot
in area 3, and one in 1b moOre oOr less
distinct. Hindwing with bla,c k marginal
spots and a costal spot. A large black
spot present in the outer half of the
forewing. Under side: Forewing
without marginal black; spots in areas 3
and 1b large. Hindwing with dirty white
and greenish scales. The dry season
forms are mor,e dusted with such scales.
The females have an additional discal
black spot on the upper side of the
forewing.
Upper s.l de Ma e
Wingspan : 45-60 mm.
Distribution : 'T his subspecies ranges from North Baluchistan and Chitral (Pakist.an)
eastwards to Myanmar and is widely distributed in India and does not show much s,e asonal
variation. It flies from 2,500 to 12,500 feet in Himal,ayas. Uttarakhand : Dehra Dun Haridwar,
Tehri, N,ainital, Pauri, Pithor,a garh, Rudra Prayag, Chamoli, Almora, Bageshwar, Champaw,at,
Udham Singh Nagar and Uttarkashi districts.
1939. Pieris brassicae nepalensis Doubled.ay: Talbot, Faun,a of British India (BlIlI(u:/lies) 1 : 427-428.
Diagno.stic CI,aractels : Antennae bla'ek, the club tipped with white~ he,ad, thorax and
abdomen black, with SOlne white hairs; underside whitish.
Distribution: This sub . . species ranges froln Baluchistan and Chitral (pakistan) eastwards
to Assaln. It is very COlnlnon in Hilnalayas and plains adjoining Hilnalayas, ascending to
about 12,000 feet in Himalayas. Uttarakhand : R,ecorded from De~raDun, aridwar, Pauri,
TehrL, Rudra Prayag, Chamoli, Nainital., Pithoragarh andUttarkashi districts.
KUMAR: Handbook 01.1 Common BUlle/lUes o/Utlar.akltal1d 33
J'904, P0l11ia d(lplidiceMoore, Lep. Indica. 4 ~ ] 37, pI. 521, fig. L l.a, J b.
1907. Lellcochloe daplidice moore; Rober, In : Seitz. Macro/ep Fallna Pa/eaearClicQ, 1 : 49.
Gene/~al Ecology : The Bath White is prilnarily a butterfly of the higher hills but Jnay
be found in the plains and is COlnlnon in the valleys of western H:imalayas. It is sun~Joving
and visits flowers of various sp ci s. It flies close to the ground and rests with the wings
closed. -
Diagl10stic Character'; : Antennae dusky black; head, thorax., and abdolnen fusco'Us-
black' under side of bead, thorax, ,and abdolnen white.
Male Upper side : Forewing white with basal half of costa dusted with black~ a large
and quadrate black spot at the apex of the cell; api·cal and Jnarginal area above vein 3
broadly black with a series of white spots continued as fine lines to the outer lnargin.
Hindwing unifonn., the Inarkings of under side show.ing faintly through; base of both wings
black. Under side : Forewing white, markings as on upper side, but the apical area dusted
with grleen, the sub-Inarginal spot obscure; a black spot in the outer half of ar1ea J b. HindwJng
green with thr·ee curved series of white spots .in area 1 to 6; inner margin white; a round
white spot in lniddle of the cell, and other spot above it in area 7.
Female: Upper side: Forewing white with basal area dusted with black; a quadrate black
spot in the outer half of area 1b,with sometimes a short ll-defined black lines below it~
apical and lnarginal black broader than in the male, the sub-Inarginal spots blun'·_d and
obscure. Hindwing with a iargecostal black spot before the ap·ex, a broad, black~ sub-
Jnarginal inwardly curved band and Inarginal series of black spots connected by thin :lnarkil1gs
to the sub .. marginal band. Under side : SaIne as in the tnale but Jnarkings broader.
Wingspan : Male and female : 45 .. 50 Inln.
Dh;lrihuliol1 : The subspecies is distributed froln Baluchistan and Chitral (Pakistan)
through Kashlnir to west,ern H'ilnalayas, ascending to about 7,000 feet. Uttarakhand: Dehra
Duo, H,aridwar, Almora, Tehn, Bag1e shwarand Uttarkashi districts.
34 Zoologkul Survey of India
Male ,: Upper side : Wings chalk white. Forewing with broad apical half black; ,a broad
apical patch of orange extending f~om costa and narrowed posteriorly to a little below vein
2" Hindwingwith broad terminal black border.
1939. Ixias pyl'ene kausala Moor'e : Talbot~ Fauna of British India (Butf,erjUes), I : 446.
General Ecology : h is a very ,common butterfly, flies in the bushes and shrubs at the
edge of forested ar,e as,with a fast and straight flight. It basks in the Inorning with its wings
open. It feeds on the nector of various spec'l~s .of flowers, also visits darnp pat1ches, str,eaJns
and shaded naUahs.
Diagnostic Characters: Male : Upper s:ide : Wings yellow, for'e wingwith costal and
api,cal area black and ,a large .orange band nearer the Icell than to apex, ,e.ntering the end of
the cell. Hindwing narrow with black
outer border. Under side: Pale yellow,
speckl,ed with fuscous-brown; row of
blackish-brown spots, more or less in
the form of ocelli. Forewing usually
with a black tornal spot.
Distribution: The subspecies ranges from Baluchistan and Chitral (Pakistan) to KUlnaon,
ascending to about 7.,000 fe'et in HimalayasUttarakhand : Dehra Dun, Haridwar, Ahnora,
Bageshwar, Nainital, Tehri and Uttarkashi districts.
1939. J.{aleria valeria hippia (Fabricius) : Talbot. The Fauna of British India (Bune~flies) I : 486.
neith r fast nor slow, but when dislurbed it can be errati,c and fast. The felnales are nluch
less COlTIlnonly seen than the 1l1aies.
Diagno'tic ,C hal'acters
Antennae black~ head, thor,ax and
abdo.nen fuscous; the thorax
clothed with long bluish hairs;
under side of palpi, thorax and
abdolnen pale silvery bluish-white
Male Upper side Wings Shining
sky~blue. Forewing with costa
broadly and apex and outer rnargin
v'ery broad'ly black. Hindwi ng with
costal and inner Inargins broadly
whitish; ,o uter Inargin broadly
Upper ,side Male
black especiat1y at the apex. AU
veins blackened,. Under sid,e :
Forew ing pal le blue, the outer
lnargins obscurely fuscous; a suh..
Inarginal series '0f very indistinct
whitish [unulated spots. Forewing
\vith the vein broadly, bordered
\vith black. Hindwing with veins
6, 7and 8 broadly, the rest of veins
very narrowly, edged with black.
Fenu.t/e : Upper side: Bla'ek with
bluish white tnarkings. For,ewing
with two parallel streaks in the cen~
a s,eries of post-cleUular stripes,
vlery irr,egular in length.Hind wing
Under side Male b1.a ck; costa and inner Inargin
l
Wingspan 65-80 m :m .
Distribution: The subspecies is comlnon throughout India, Myanmar and other parts of
the Oriental region. Uttanlkhand Dehra Duo) Haridwar, Tehri, Almora, Bageshwar and
Pauri districts.
KUMAR : Handbook on Common Butt(!I:/lies of Uttarakhand 37
Subfamily COLIADINAE
28. Catop~..ili(,
cro,cai,e (Cram'er)
The Common Emigrant
1775. Papilio crocale Cramer~ Pap.. EXOI., I : 87.
1990. Catopsilia croca/e crocale (Cramer) : Ghosh, Mondal& Chaudhary. Rec zool. SUf1~ Indi,a, 86( I) : 20.
General Ecology : A very cOlnmon butterfly found throughout the country, including
drier parts. Som,e tilnes swanns of this butterfly can be seen just ,after the monsoon. This is
a fast flier, rises high in the air, and cover long distances. The flight is straight w ,ith up and
down curves. His also known to Inigrate,. It is extrernely fond of settling on various species
of flowers like Lantana, Fiddle- leaves, Jatroph.a, POinsettia, 'C rape luyrtle, Pomegranate,
Bougainvillea and Sunflower etc. It also visits damp patches., sometilnes a large congregation
of these butterflies can be seen lnud puddling.
Diagnostic Characters : Male : Upper side: Wings chalky white, yellow or green ish
yellow. Fore wing with costa narrowly black to ' the base" wider at the apex. Iindwing
unlnarked rarely with marginal
b1.a'c k vein dots. Under side
without markings., yellow or
yellowish white,. Forewing often
whit,e in the post'e rior distal area.
Female : Upper side : Creamy
white to yellow. For,e wing with a
black disco-c,e llular spot, usually
forming a bar to the costa; costal
Inargin black froln base to apex;
an outer lnarginal black border,
'w ider at the apex. Hindwing with
outer Inarginal black border.
Under side: Similar to the lna le,
v,a rying to ochraceous buff.
Upper side Mil e
Antennae black.
Wingspan : 55 80 mIn.
Distribution: The species is very throughout India and ascends up to 8,000 feet
COITIlnOn
in Himalayas. Also occurs in Sri Lanka, Myanmar to Southern Asia . Utta~ak hand: Dehra
Duo, Haridwar, Nainital, Pauri, Rudra Prayag, Chamoli, Pithoragarh, Bageshwar, Chall1paw,at,
TehriUttarkashi ,andUdhalTI Singh Nagar districts.
38 Zo%gical Survey of [ndill
1881a. Catopsilia pyron/he (Linnaeus) Moore, Lep.. Ceylon, 1 : 124, pI. 47, fig. 2,. 2a.
Geper:ai Ecology : This is an energetic and very common butterfly, which rarely rests
during its activity period. It is most active during morning and ,eady afternoon. It covers long
dist.anc,es in a single flight. On hot days the males gather in large numbers on wet soil. When
resting., the wings are closed over the back but the forewings are not drawn between the
hindwings . It is a r,egular migr.ant in the plains of peninsular India. It feeds on the flowers
of herbs and shrubs e.g. Lantana camara, Pomegranate, red gram, bitter gourd, Medicago
s'ativa and several other plants.
Diagnostic Characters : Antennae reddish, head and thorax brown; abdolnen white;
under side of paJpi, thorax and abdomen white,.
Male: Upper side: Wings chalky white 'or greenish. Forewing with apical and Inarginal
narrow black border; a hlack disco-cellular spot, which tnay be absent. Hindwing with
marginal black dots or spots with a continuous narrow border. Under side : Greenish, w'ith
reddish-brown strigae and disco-cellular spots.
Female: Upper side: Wings greenish-white to greenish . . yellow. Forewing with black
border, and a black disco-cellular spot. Under side : Similar to male with proll1inent disco. .
ceHul,ar ring spots. Both wings with post-discal series of reddish brown spots.
Wingspan : 50-7'0 mm .
Wynter~B 'lyth(1957), Varshney (1997) and Larsen (1987) believe that Catopsilia jIore/la
is only a dry season form of pyranthe and hence there seems to be no justification to treat
Catopsi!ia. jlorella separately.
Distribution: The speci1es is very common throughout India, ascending upto 9,000 feet
in Hhnalayas. Also occurs in Sri Lanka, Myanmar and South-east Asia. Uttarakhand : Dehra
Dun, Handw,ar, Ahnora, Chamoli, Bageshwar, Nainital, Paurl, Pithoragarh, Tehri, Champawat,
Uttarkashi and Udham Singh Nagar districts.
KUMAR: HO'tdbQok on Common Bulterflies of Uuarakhalld 39
1907. Gonepteryxr rhamni nepalensis (Doubleday) : Rober, 111 ,: Seitz.. Macro/.ep .• Fauna Palaearctica. I
: 6 , t. 246.
D,i stribution: This subspecies is distributed from Chitral (Pakistan) to northern Myanlnar
and occurs from 2,000 to 10,500 feet in the NorthWest Hinlalayas.Uttarakhand . Dehra
Duo, Haridwar, Almora, Chamoli, Pauri, Nainital, ~ehri, Rudra Prayag and Uttarkashi
districts.
4'0 Zoological Sun'c) of India
1990. Eurem,a la,ela facIa (Boisduval) : Ghosh Mondal '& Chaudhary, Rec. zool. Sur". India. 86( I ) : 22.
General Ecology: A very common butterfly .of drier regions. It flies close to the ground
for very short distances and visits various flowers like Smithia sensitivex, Tridax, Vernonia,
Sida sp. and many more lowly growing flowers.
Diagnostic Characters : Antennae yellow, with scattered dusky scales; head with pinkish
pubescence; thorax and abdolnen black with scattered yellow hairs ,and scales; under side of
pal pi, thorax and abdomen pale yellowish-white.
Male and female: Upp,e r side: Wings yellowish. Wet season fonn: Forewing apex and
tennen rounded and broadly black, but is not excavated between vein 3 and does not run
on the inner edge being unevenly rounded. Hindwing with narrow terminal black border,
Under side: DuUer ground colour. Both
wings evenly but sparsely dusted with
bIalck. Forewing paler yleUow.
Hindwing with black dots.
1867. Terias fimbriata Wallace, Trans. ell I. Soc. Lond., (3) 4(3) : 323.
1932. Eurem,a hecabe jimbri.at,a (Wallace) : Corbet and Penl,ebury, Bull. Ra.D/'es Mus. Sing .. 1 : 160, pI. 5.
General Ecology : It is found aU over India and abundant in many places. It occurs in
large open patches in the evergreen forests, semi ~evergreen and deciduous forests and also
near the human habitations. It has a weak fluttering flight. Although it is named HGrass
Yellow", it is not dependent on grasses. It feeds mostly on small, low growing flowers of
plant species of the gener,a Cassl~a, Wagatea, Acacia, Serbania, A /hizzia, Caesalpinia etc ..
The males also visit da'mp patches and are a regular member of snud-puddhng assemblage.
It is most active in the morning and at noon.
Diagnostic Characters: Head with antennae greyish yellow, club black~ head, thorax and
,abdomen yellow, shaded 'with fuscous; under side of palpi, thorax and abdomen yellowish
white.
Male and female : Upper side: Wings bright yellow. Wet season form : Forewing with
black broad border ,and has an ,excavated between veins 2 and 4. Under side: Pal'er than
upper side. Forewing with a
prominent reddish-brown patch .
Hindwing with a slightly curved
sub-basal series of three spots. Both
wings with minut,e reddish-brown
costal and outer marginal t(~rminal
specks at vein ends.
Distribution : The subspecies ranges from Punjab to KUlnaon, ascending to about 9,000
feet in North- West H'imalayas. Also oc,curs in Pakistan. Uttarakhand : Dehra Dun, Haridwar,
Nainital, Paun, Alrnora, Pithoragarh., Tehri., Bageshwar, Chalupawat, UUarkashi and Udham
Singh Nagar districts.
42 Zoological SUI'l'C!.v of India
1867 , Terias rubella WaJJaee, r~ans. ell!. Soc. Lond" (3) 4(3) : 323 .
1939. Euy,ema brigilla rubella (Walla,ee) : Talbot, Fauna of British India (Buuer./lies). 1 : 515-517.
General Ecology : This is a very comlnon butterfly. t flies close to ground, around
bushes. It is a weak flier. Its congregation can be seen in the undergrowth . It vis;its low
growing flowers.
Diagnostic ,Characters : Antenna,e, head, thorax and abdolnen dusky black, the thorax
with some yellow hairs; under side of pa~lpi, thorax and abdolnen whitish-yellow.
Male and female : ypper side : Wings bright yellow. Forewing with apex and ternlen
broadly black with inner edge evenly rounded. Hindwing with the tennen broadly or narrowly
black. Under side : Forewing with costa narrowly edge w'th. pink; two well Inarked sman
black spots on the end cell. Hindwing with a spot at the base, foUow,e d by thr,e e transv1ers,eJy
placed spots and an elongate,
delicate, loop-like disco~cellular
spot; black above and below the
disco-cellular spot are slnall
diffuse bl,ack spots. Both the wings
with black marginal vein dots.
This species shows seasona I
variations. Wet season form has
broad black areas. The name
rubella . (Wallace) applies to the
dry season form. The dry season
tonn has more dusting and is
paler. The males have no sex
bands on the hindwing. Upper side Male
1939. Colias eJecto field; M'enetries : Talbot,f',auna a/British India (Buller/lies), I : 562.. 563,
Diagnostic Characters
Antenna,e, head and thorax
interiorly sa'imon pink, antennae
club darkening to brown; thorax and l PP"'r side Male
,abdomen dusky greenish-black.
Male : Upper side: Forewing
deep orange" inner margin broad
pale yeUow; outer margin broadly
black; disco-cellular spot pale
yellow in colour. Under side : Pal f,
the costal margin narrowly.
Hindwing overlaid with pale dull
gr,een and disco-,cel1ular spot large.
Fem,a Je: Upper side : Sam,e as
male, but borders spotted with
ground colour. H'indwing heavily
dusted with black scaling. Under
side.: As in the male. Unde side Male
1880. 1(b), Colias erale erate (Esper) : Butler, uOn a collection of Lepidoptera fronl Kandahar" P,ne.
Zool. Soc. Lond.. : 409.
(kneral Ecology: It is a very busy butterfly, keeps const,antly on Inove, fluttering r,apidly
within a meter from the ground, It visits flowers. Both the sexes frequently settle to feed on
low growing flowers.
Diagnostic Characters : Male : Upper side : Wings lemon yellow. Forewing with a
small, oval., disco-cellular black spot; apex ,and outer margin broadly black, broadest at the
apex and narrowing to tornus; with or without yellow sub-tnarginal black spot. Hindwing
de1ep orange-yellow with disco ..cellular and broken lnarginal black spots.
Distribution: This subspec·ies ranges from Baluchistan to ChitraJ (Pakistan) and Kumaon.
It has been recorded betwe'en 5,000 to 11,0.00 feet in the Him,alayas between March and July
before rain and then between November and March ·in the dry season. Uttarakhand : Dehra
Dun, Haridwar, ~ehri andUttarkashi districts.
KUMAR : Handbook on Comlnon Butterflies of Uttar,a khand 45
3. Family DANAI AE
Milkweed Butterflies
The members of this family are commonly known as Tigers, Crows and 'Tree Nymphs.
They ar-e large sized ,a nd tough, and leathery Butterflies possessing .an unpl,e asant smell and
unp,aletable juic1es. 'T he odour and unpleasant taste has been evolved to protect them from
their natural enemies like birds and lizards.
All the speCl'es are fond of visiting and feeding on flowers, most of them like resting on
damp patches, and ov,e r ripe fruits. They are sun-loving. They have a slow flight, feign death
when ,caught.
The males have special scent organs and also posses a pair of abdominal hair pencils. The
forelegs being imperfect unfitted for the walking in both the sexes ,c haracterize this family.
The cens are closed on both the wings No seasonal variation is observed in this group of
butterflies.
Talbot (1947) recorded only three genera viz. Danaus Kluk, Euploea Fabricjus, and Idea
Fabricius from the ndian region. The present study is re~erable to first two gener,a, the genus
Idea is not known from Western Himalayas.
Some common species of this family are EupJoea core core, D,anaus crysippus, Danaus
genutia etc.
1947. ,Dana us cllrysippus cluysippus (Linnaeus): Talbot~ Fauna of British India (ButteJ:/lies) , 2 : 21.
,and variable numbers of white spots in the costa and apex; api,cal half black, four white
spots from costa to vein 4; smaller white spots also inare,a 3 and at the eel end, on costa
and ,at the outer margin; veins not blackened. Hindwing: Ground colour paler; outer margin
narrowly black, with an incomplete series of white spots; with four small black spots
around the cen in mal,e ,and three in female and ,a pouch in m,ale, Under side: Dun orange.
Forewings dark brown in the upper half with spots in the black area. Hindwing with six
black tipots.
The fem,ale of Danaid Egg~ny (Hypolim.nas ,misippus) 'mitnics the Plain Tiger and it is
very interesting that mimic and the model are found together.
Wingspan: 70 .. 80 mm.
Distribution: The subspecies is very common throughout India, ascending to a height of
about 9,000 feet in H'imalayas. Uttarakhand : Chamoli" Dehra Dun, Haridwar,Ahnora,
Bageshw,ar, NainitaJ, Pauri, Pithoragarh, Tehri andUttarkashi districts.
'General Ecology: It is popularly known ,as the Monarch Butterfly all ov,e r the world.
Also known as uThe Common Tiger" It is ,a wen known migrant and found throughout India
in the areas of heavy rainfall, forest edges ,and scrub jungles. It visits flowers of Indian Chest
nut, Feather Cock's Comb, Zinnia, Cosmos and CadHla etc. It is known to tnigrate in large
numbers in the plains and hiUs of South India.
Diagnostic ,Characters: Antennae black; head
and thorax black with white spots and streaks;
abdomen orange brown, with white markings.
Male and female: Upper side: Forewing
reddish brown with bold black veins and white
apic,al spots cell end. Hindwing paler than
forewing bearing two compl,ete series of white
spots. Under side: Paler, Hindwing with more
prominent black lines along the veins.
Upper side Male
,Wingspan : 75 100 mm.
Distribution: The subspeci'es is available throughout the year, though less common in
,cold weather. When occurring in drier region, the orange brown p.art of the hindwing tends
to becom,e pal,er or whitish. It is found up to 8,000 feet in Himalayas. Uttarakhand : Chamoli,
Dehra Dun, Haridwar, Almora, Nainital Bageshwar, Rudra Prayag, Pauri, Tehri, Pithoragarh"
Champawat, Uttarkashi and Udham Singh Nag,ar districts,
KUMAR :H,a ndbook ,o n Common Butter,/lies of Uttarakhand 47
1984. Parontjea sila sila (Kollar) : Ackery & Vane-Wright, MUkwe,e d Buuerjlies = ) 78.
General Ecology: It is fairly common along most of the Himalayas, and in the hills of
Assam. It is found in all sort of country. It has a high and soaring flight often well above
the tree level. It visits damp patches and also se,en on the flowers of chestnut tree, Cedrella
etc.
Diagnostic Characters : Antennae black; head and thorax black, spotted with white~
abdomen from brown to bright ochraceous, whitish beneath.
Male ,and female: Upper side: For,ewing black or fuJiginous black with the bluish white
sub-hyaline markings . The whole of cell and thr,ee-fourths of the basal area is filled with
broad bluish-wh'ite streak;ftve very large quadrate discal spots; two long sub-apical streaks;
three short streaks beyond cell; a sub-marginal series of rounded spots, decreasing in size
anteriorly, and curved inwards opposite the apex; an incomplet,e sub-marginal series of
smaller spots. Hindwing : Chestnut-red,with sub-hyaline streaks ,and spots, streak from the
base, in ar'e,a la, 1 b, not reaching the margin, two broad streaks in area 1 c,' united to near
their apex; a stripe filling the celL Male with two scent pouches, the largest on the area 'I
b and the smaller one in the I a. Under side: Markings similar, clearer ,and more complete.
Forewing with the ,apical area also chestnut brown .
.Wingspan : 85 ~ 11 °mm.
Distribution : The species is common from Kashmir to Sikk!m and AssaJn, ascending to
about 8,000 feet in Himalayas. Also occurs in Bhutan and Myanmar. Uttarakhand : ChalnoJi,
Dehr.a Dun., Haridwar, Pauri and Pithoragarh districts.
48 Zoological Survey of Illdia
1'866 . Dallais Iinmiace var. leopardus Buder, Proc. .zool. Soc. LOl1d., : 2, fig. lO (Female).
1984. 1iru.mala limniac.e leQpardus (Butler) Ackery & Vane-Wright. Milkwe.ed Blilfe/~flies : 1'9 8.
General Ecology: It is a large sized and very com:mon butterfly found throughout the
p1:ainsand hills of lndi;a. It flies slowly, but faster during migration Qr when disturbed. It
settles frequently on wet places. t is mainly an inhabitant of woodlands, gardens, edges of
forests etc.
Diagnostic Characters : Antennae black; head and thora.x blackish with white spots and
streaks; abdomen dusky black.
Distribution : The subspecies ranges from Baluchistan and ,Chitral to Kum,aon, West
Bengal and Sikkim, ascending to 5,000 feet in Himalayas. Also .occurs in Sri Lanka,
Bangladesh, Bhutan, China, Nepal, Myanmar Sumatra, Thailand and Vietnam. Uttarakhand
: Dehr.a Dun, Haridwar, Pauri, Nainital and 'Tehri districts.
2006. nrumala hamata septentrio.nis (Butler) Gupta .& MauUk, Fauna .o f N~galand. Slate Fauna Series,
12 : 283.
KUMAR : Handbo,ok ,oil Common Buuerflies of Uttarak/l.and 49
General Ecology: It is ,a very 'common butterfly throughout the hilly and forest region.
It is found throughout the year but commonest at low elevations during wet weather. It is
a slow flier and visits lnoist places. It visits flowers and can be seen feed'ing on flowers of
'Cirsium species.
It closely reselnbles Tirumala liminiace (Cramer), but is always sufficiently distinct to be
easily recognized, even on the wings.
Diagnostic Characters: Male and female: Upper side: The back ground colour of wings
is dark and semi-hyaline markings narrower and 'Ofa bluish~whit,e It int. Forewing with two
narrow streaks In area 1 b, nev'e r
coalesc,ent, the upper one fonning an oval
detached spot; the short streaks above vein
5 are outwardly never truncate, ,always
acute. Hindwing with two cell streaks
united at the base but s'eparate distally.
Distribution: 'The subspecies is fairly comlnon in Himalayas, ranges froln Kulu to Assaln,
ascend'jng to about 8,000 feet ,and Nagaiand, Arunachal Pradesh, Manipur, Meghalaya, Jamlnu
'& Kashmir, Sikkim, West Beng.al ,and Darjeeling. Also occurs in Bhu.tan, Chjna,Malaya,
Myanmar, Nepal, Sumatra, Taiwan and Thailand. Uttarakhand :Almora, Dehra Dun, Haridwar
Nainital, Pithoragarh, Tehri and Uttarkashi districts.
•'984. Parontlea ag/ea melan.oides Moore: A1ckery & VaneeWr.ight, Milkweed BUlle/file.,., : J 75.
G,enerai Ecology ~ A transparent blue-brown buttertly of hilly regions, mostly :in areas of
heavy rainfalL It is a weak flier as compared to other Tigers and com,es out of for,ests 'Only
to visittlowers o{Lantana, Ageratum etc. It is a known migrant and frequents dalnp patches
occasional (y.
Diagnostic ,Characters : Antennae black; head ,and thorax black, spotted with white.;
abdomen bla,ckish-brown, ochraceous beneath.
50 Zoological Survey of J"dia
Male and female : Upper side Wings dark brown. Forewing with bluish-white semi-
hyaline streaks and spots; cell streak divided lengthwise into two portions, and united at
base~wjth black lines traversing throughout; most spots progressively decreasing in size; a
marginal series of spots much smaHer and
placed in pairs between Veins. Hindwing
streaks long and broad; the ceHwith two
broad streaks which are united at base; str~ak
'in inner area and below the cell, long and
narrow; other spots larger in post-diseal area
but decreasing towards margin. Males with
a larger scent pouch on vein 2. Under side
: Dark brown, the streaks are often Inuch
blurred.
Upper sideM,a le
Distribution: The subspecies is fairly common from Kashmir to Sikkim and Assam,
asc,e nding to about 8,000 fe·e t in Himalayas. Also occurs in Myanmar. UUarakhand : Dehra
Dun, Haridwar, Tehri, N,a inital, Champawat, Udham Singh Nag,a r and Uttarkashi districts.
General Ecology: This glossy brown butterfly is very common. It Jnay be seen sailing
lazily in aU types of country and flying about shrubs and bushes in s,e arch of its foodstuff.
It visits low growing flow,e r plants, i.e. Lantana, ,C ommon Zinnia, MeyenlQ /axijlora, marigold
etc. Even if disturbed, it does not usually flyaway and even if takes off, soon returns to the
same spot It generally flies during early morning and evening and rests in groups during
afternoon.
'This butterfly is mimi'cked by Chilasa clytia Linnaeus, and fetnale of Hypolilnnas bo/inu
Linnaeus.
Distribution : The subspecies is very common throughout India at altitude below 8,000
feet.. Uttarakhand : Almora, Chalnoli,Dehra Dun, Nainital, P,auri, Pithorag,arh, Bagesh'war,
Tehri, Champawat, Uttarkashi andUdhaln Singh Nagar districts.
1911. Eup/oe,a mulciber mulc.iber (Cramer) Tytler, J. Bombay nal. l-lis.t. Soc., 21 : 4<) ,
Diagnostic ,C haracters
Antenna'e , head, thor,ax and the
abdomen dark brown .; abdom1en
glossed with greenish blue above;
head, thorax and ,abdom,en beneath
with white spots and transversle'ly
banded.
Female : Upper side : Pale and similar to lnaie, except on the hindwing narrow white
discal streaks, spots silnilar but white. Forewing blue glossed areas are similar. Under side
: Markings as on the upper side but clear, distinct and broad.
4. ami y ACRAEIDAE
'Members of family Acraeidae are smaU to medium-sized butterfli'es. They ar,e tawny or
yellow in colour. They are mostly distributed in Africa. Only two species are known to occur
in India viz Acraea issoria (Hubner) and Acraea vioJae (Fabricius). Both of them .occur in
Uttaranchal.
The forewings are long and hindwings rounded, fre,e from the abdomen along their inner
margins, not channeled to receive it.
They love sunshine and visit various flowers. They are weak fliers. They fly close to the
ground and often seen basking early in the mornings. Like Danaids, Acraeids are also well
protected, and when in captivity, the species ,eject a sort of protectiv,e, oily, yellowish evH-
smelling fluid, which is obnoxious to predators.
In both the sexes; the forelegs are unsuitable for walking; the cells are closed in both the
wings. Anal veins are free in forewing..
t 844. Acraea anomala Kollar, In : Hugel's .Kaschmir und das Reich de,. Siek, 4(2) : 4'5, pI. 3, flgs.
3, 4.
1910. Acraea issoria ,anolnala Kollar, The Butterflies of Kumaon, Part~ l, J, Bombay nat. Hist. Soc., 20
; 141.
General Ecology :A localised rapidtlier but when at r,est can easily be captured, It is
fond of sett ing on leaves and flowers. It inhabits open and scrubby country. Sun loy' ng, can
be seen basking early in the morning.
DiagnosticCh.aracters : Antennae, head., thorax and abdolnen black and thorax vvilh
ochraceous yellow pubescene,e anteriorly~ under side of thorax and abdonlcn black and
spotted with a very pale ochraceous spots.
Underside of forewing yellow, becoming paler towards the apex, the veins much darker,
and the black disco-cellular mark showing through. Hindwingwith ground colour a delicate
pinkish-white" the veins ,conspicuously black; sub-marginal line lunul,ate, edg,ed with black
on inner as well as outer margIn. The Inarkings of female are usually darker than the males.
According to Peile (t 937) the species is abundant at Mussoorie in the month of June at
3,000 feet. .
Wingspan : 45 .. 65 mm,.
Distribution : The subspecies ranges from Kulu to KUlnaon between 3,000 to 6,000 feet.
Uttarakhand : Ahnora, 'C hamoh,Dehra Dun, Haridw,ar, Bageshwar, Champawat, Nainital,
Rudra Pr,ayag., Tehri and Udham Singh Nagar districts.
177.5. Papilio violae Fabricius, Sysl. Enl.• (3) I : 460 no. 74.
1947. ACl'aea viQ/ae (Fabricius) : Talbot, The fauna of British India (Buttc.1:flies),. 2 : 466 ~469 .
General EcoJ~gy : This yel1owish .. brown butterfly seen during the latter part of June
becomes most abundant in July between 2,000 to 7,000 feet. A localized butterfly, seldom
straying far from its food plants. Its flight is very slow) fluttering and close to the ground.
Often found basking in the early morning hours,. It is Fond of settl ing on leaves and flowers
of Land-Caltrops, Lantana, Tall chaste, Monk's Pepper tree, etc. It inhabits open and scrubby
country but not forest. It excudes ,a yeHowish, fou)..slnelling, oily liquid when disturbed.
Diagnostic Characters : Antennae black; head and thorax black, spotted with ochraceous
,and white; abdomen ,anteriorly black, posteriorly ochraceous yellow with black lines. Under
side of palpi, thorax and ,abdomen ochraceous; thorax spotted 'with ochraceous, abdomen
with basal longitudinal black lines.
Male : Upper side : Wings tawny_
Forewing with a transverse black cell
spot; an irregular disco cellular black
bar; post.. discal black spots in areas I b,
3, 4, 5, 6, ,and 10, those in 4 to 6
parallel to end of cell; apex and outer
margin narrowly black, narrowing
posteriorly, and with short projections
on the veins. Hindwing with a basal
lines of four or five black spots, a
similar spot in the middle of cell, and
a sub-costal black spot above it· ,a discal
seri'es of ObSCUf'e blackish spots; a Upper side Male
KUMAR: Handbook on Common Blltte/~f1ies of Ullarakhand 55
minute post-discal black dot in areas 4 and 6; a broad black marginal border, bearing a
sub-marginal series of small spots of the ground colour. Under side : Ochraceous-yellow.
Forewing: Paling to whitish on the apex, with black markings as on upper side, but more
clearly defined; sub-marginal spots much larger, white, not tawny; base of wing black,
separated from the basal black spots by two or three larger whitish spots.
Female : Silnilar to male. Upper side : Ground colour duller, the black spots on both
wings larger, upper post-discal spots on forewing often coalescing, and fonning an irregular
oblique short band; marginal black border on both wings proportionally broader. Hindwing
: Sub-marginal spot larger and whitish. Under side: Paler and duller, markings as on upper
side, spot on hindwing well defined.
Distribution : Peninsular region to Punjab and the lower altitudes of Himalayas upto
7,000 feet, West Bengal, Bihar and Sikkim. Also occurs in Sri Lanka. Uttarakhand : Dehra
Dun, Haridwar, Almora and Bageshwar districts.
56 Zoological Survey ollndia
5. Fami y SATYRIDAE
Browns
Melnbers of the family Satyridae are usually dun brown or blackish in colour, occasionally
with yellowish or white patches on wings. Under side often ocel1ated and beauti Cully
vari,egated. The flight is weak, slow, and Jerky or bouncing, rather close to the ground
littered with dead fallen leaves.
Some species particularly in hilly regions, prefer open meadows and grasslands with
slnall herbs, shrubs and bushes. They are 'less dependant on sunlight than the gaudy coloured
butterflies" often seen on the wings even on cloudy and rainy days when no other butterfly
usually venture to corne out.
The species mostly feed on grasses including bamboos. Most of the species do not visit
flowers but are attracted to over-ripe fruits, sap and salts like Com'moo Evening Brown, even
get attra,cted to light.
The forelegs in both the sexes are very small and unfitted for walking and cell closed in
both the wings.
Seasonal variations are best known in this family, particularly in Evening Brown where
no two specimens in the dry season look alike and appear like dead leaves" often with fungus
like markings,looking all the more natura].
Some common species of this family occurring in the State are Mycalesis perseus
(Fabricius.), Mycafesis mineus mineus (Linnaeus.), Lethe maitrya (de Niceville), Lethe confusa
con/usa AuriviUius, Lethe verma verma (Kollar).
Diagnostic Cha~acters: Antennae brown above, greyish white below the ,Club tipped
with ochraceous; head, thorax and abdomen brown, paler beneath.
Mal,e and female : Upper side For'ewing with an ocellus in area 2. Ground colour of
both wings dark to pale brown. Both
wings with a pale sub .. rnarginaJ and a
marginal line, placed ,close tog _ther, and
obscure on forewing. Underside = Both
wings with ground colour as on upper
side and ,crossed by a narrow, dis-caL
bluish white line, its inner side edge
with darker brown; sub-marginal and
marginal lines paler.
Distribution: The subspecies ranges fr'Om Kangra (HiJnachal Pradesh) to Myanlnar and
ascends to 5,000 feet in Himalayas. Uttarakhand : Dehra Dun, Haridwar and Udham Singh
N,agar districts .
Male and .fe.1nale Upper side of wings dark hro\vn; boOth wings with sub-Inarginal and
luargina\.,lender, pale lines. Fore wing with a single 'white centred fulvous ring. black
oceJlus in area 2; SOIne times a siJnHar slnal'ler ocellus in area 5, without any pale surrounding
area. Hindwing unmarked, sometimes\vith one or t\\"o obscure ocelli.
Distribution : The subspecies is generally COlnmon froln Kulu to Myanmar and ascends
to about 6,000 feet in Himalayas. Uttarakhand : 'DehraDun~ Haridwar Alnlora, Bageshwar,
Nainital and Pauri districts.
1932a. Lethe C01~rUsa C01~rUsa (AuriviUiu') : Evans, Ident~fic,alioll 0/ Indian BUllelllies. 2 : lOS.
General Ecology : Very COJurnon species, fond of visiting moist pat,ches and salt
encrustations. It has territorial habit very aggressive towards intruders. R,e sts with its wings
cJos _d and hardly seen basking.
Diagnostic ,Characters : Antennae, head,
thorax and ,abdomen du 11 brown.; ant,ennae
tipped with ochraceous.
marginal sinuous lalaeine white lines. Forewing with discal white bar as on upper side, a
sub-apical lilac patch bearing three ocelli in straight line and wi th a very short white bar
joining it with the costa. Hindwing with two medial lilac Jines meeting on the dorsal margin
,and a strongly ar'ched seri'es of bla,ck 'Ocelli with white specks in centre, an inner ochraceous
ring and outer lilac ring, the api,eal ocellus very large.
Wingspan : 50-55 Inm .
Male : Upper side : Wings brown to dark brown. Forewing with discal band showing
through from below; two sub-api< c al, small ochraceous spots. Hindwing uniform, a post..
discal, curved series of four round black spots, the apical spots sometianes elongate; sub-
marginal and marginal slender black Hnes. Under side : Paler. For< ewing with basal h:a1lf
darkest; a disca1, broad, oblique pale bar, a sub-apical, short, narrow band bearing three
white centered obscure, small ocelli. Hindwing with ,a sub-basal, discal and sub-tnarginal
slender brown lines; a curved post discaI series of black ocelli 'w ith white-ce:nter~d ix ocelli,
each ocellus with inner ochra< ceous and outer paL nngs.
1911. Lethejalauridajalaurida (de Niceville) : Fruhstorfer: In Sellz.Grass-Sclunett. der Erde. Fauna lndo-
Austral., 9 : 313.
Diagnostic Characters: Antenna< e, be,ad, thorax and abdomen brown, pale ochraceous
beneath. Male and .felnale : Upper side: Wings reddish-brown with a silky gloss. Forewing
KUMAR: Handbook on Common BUlleJ:lhes(~l Ullarakltal1(/ 61
Distribution: The subspec'ies ranges from Kulu to Kumaon and ascends to 7,000 to
11,000 feet in the North . . West Himalay.as. Wynter. . Blyth ([940) records it ,as oe,eurring sparingly
in the Narkanda~Bhagi forest and in Kulu, above 6,000 feet, from June to September.
Uttarakhand : Almora, Chamoli, Nainital, Bag,eshwar, Champawat andUtt.arkashi districts.
1865. Zophoessa baladeva Moore" Proc. zool. Soc. Lond., 769, pI. 41, fig. 5 .
1911.Lethe bala,deva baladeva Fruhstorfer: In Seitz.Gross- Schmelt. der Erde. Fauna Indo-Austra/.., 9 :
314, t. 97d ..
'General Ecology : It is very shy, an uncom'mon species; flies in the forested areas and
along paths and streams.
Diagnostic Characters : Antenna'e, head, thorax and ,abdolnen brown; antennae tipped
with ochraceous.
62 Zoological SllIT(~Y o{ India
Distribution: The subspecies ranges from KUlnaon to Sikkim, .at ,an altitud ranging froln
5,000 to 9,000 feet. Uttarakhand : Almora, Bageshwar, Champawat and Nainital dist.
General Ecology: Inhabit low elevations wbere bamboos grow in abundance. A v,ery shy
insect and weak flier; hardly ever visit flowers and seldo:m settle on the damp patches,
undergrowth or tree trunk.
Diagnostic Characters: Antennae brown, annulated with 'white; head, thorax and abdomen
brown deep shining.
Male and female : Upper side: Wings deep shining brown. Forewing crossed beyond the
ceH by a pale ochraceous. outwardly curved band, narrow,esl and more prominent at the
costa, almost obsolete on reaching the third median nervule. Hindwing with the ocelli below
showing through indistinctly in black spots, and a sub-marginal series of black lunules.
KUMAR: Handbook all Common Blillerjli(, ,\' (~r U((arakhand 63
1947. Lethe rohria rohria Talbot, Parma of British India (Bulterflies), 2 : 200-201.
General Ecology: A butterfly of hills with thick growth of bamboo forests, Relnain "'
a,c tiv,e during early morning ,and late evening, spending rest of the day at rest.1t is ,a weak
flier and never flies far, even if disturbed; gets attracted towards animal dung., over ripe fruits
and tree sap.
Diagnostic ,Character : Antennae black with ochraceous tip; head, thorax and abdolnen
brown .
Male and female: Upper side : Dark brown. Male: Upp,er side: Fore'wing with a costal
and two sub-apica.l white spots. Hindw.ing with oceJJi of undersid,e showing through, fOfllling
64 Zoological Survcy (~r India
Distribution: The subspecies ranges from South India to Panchmarhi and Kashmir to
Sikkim, found between 3,000 to 6.,000 feet in western Himalayas. Also occurs in Myanm,ar.
Uttarakhand: Dehra Dun, Almora, Bageshwar and Tehri districts.
1844. Satyrus verma Kollar, In : Hugel's Kaschmir und dol' Reich der Seik, 4 (2) : 447, pI. 10 fig. 23.
]'9( L Lethe v,erma verma (Kollar) : Fruhs,t orfer; ,( n Se:itz.Gross-Schmett. Erde Fauna Indo-AuslraL, 9: 324.
General Ecology : Abundant species especially in North west Himalayas betw,een 4,000
to 9,000 feet from spring to autumn. The flight is weak and of very short distance. It is fond
of settling on the bark of trees and earthen banks along roadsides.
Diagnostic Characters: Antennae, head, thorax and abdomen brown, antennae tipped
with ochraceous.
KUMAR : Handbook on Common ButlC!l:/lies of Utlarakhalld 6S
1947. Pararge schakra schakra (Ko11ar) : Talbot, Fauna of British India (Butle~/li(?s) , 2 : 242~243.
G.enera/ Ecology: The Common Wall is one of the comlnonest butterflies of the western
Himalayas, extending ·eastwards as far ·as Sikkim. It can be seen in all s,easons around the
rocky roadside, and pitching on the rocks or sunny banks. The flight is lively and fairly close
to the ground.
Diagnostic Characters : Antennae brown; head and thorax dark grey and abdolnen pale
brown. Both sexes differing slightly.
66 Zoological Slirvey ,o f India
1947.Rhaphicera m.oore; moore; (Butler) : Talbot, Faun.a .of British India (Butterflies), 2 ; 250, fig. 76.
,General Ecolo,gy : The Small Tawny \Va'il is a local species of temperat,e regions. It is
found in the Himalayas between 7,500 to 14,000 feet during August. Some times it visits
the How,ers and rests in shady undergrowths or under rocks with wings closed.
Diagnostic ,C haracters: Antennae bla,ck; head, thorax and abdom'en dusky brown.
KUMAR: Handbook on Common Buue.l}7ies (~r Uu,araklralld 67
Distribution : The subspecies ranges froln ShiinJa to S'ikkitn, found froln 7,000 to ll ,000
feet in Kumaon hiUs.Uttarakhand: Almora, Chamoli, Dehra Dun, Bageshwar and Pithoragarh
districts.
1932. Au/oceta Brahminus Blanchard : Evans. Idellt(/icalion of Indian BUffelilier;, 2nd ed. : 116.
..•-......
~
--
....... -
-.. / "....-
...
•- ,
\;_\
l
r4
~
Male and fe/nale Upper side Wings bla'ck with a bronzy she,en; both wings crossed
by a post-discal oblique band of white spots, ex curved on hindwing. Forewing with band-
interrupted sub-apically by a large round black spot., enclosed by a white spot on outer, upp,er
·and inner edges. Hindwing band narrow, not reaching inner margin, cilia alternately black
and white. Under side Pale brown, markings as on upper side but irrorated with minute
specks and striae of white; for1ewing irroration less prolninent at basal, discaJ and inner
areas. Hindwing darker in lower basal half,with a post discal, lunular, irregularly curved
black band.
Distribution : The spe,cies is not common and occurs at high elevation to ,above 13,5.00
f; et in Himalayas. Uttarakhand : Chamoli, Dehra Dun, Almora, Bag,eshwar, Pithoragarh and
Uttarkashi districts.
1932 . Aulocera padm,a padma (Kollar) : Ev.ans., Identification of Indian Butlelllies" 2nd ,ed. : 116.
'General Ecology : It is the largest species of the Aulocera genus. It has a powerful and
graceful flight, very difficuh to catch. It is fond of settling on roads, hilltops and ridges,
onen sits on the ground with closed wings. The male is very aggressive and chases intruders,.
Male and female: It is larger than Aulocera brahminus. Upper side : Wings dark brown
to black with out bronzy sheen. Forewing with broad discal band, straight not tinged,
Inarginal three white spots w,ell separated, conical and pointed outwardly; sub-api,cal black
spot not promi nent; white spot in inner side of sub-apical spot continued to costa in female.
Hindwing with broad band., g'enerally touching inner margin.
Under sid,e : Silky brown; dark brown discal bands sharply defined on inner edges,
diffused on outer side. Forewing with a sub-margina , broad dark brown band; spots on ar'ea
three and four of disc,al band separate. Basal area of hindwing very dark, the greyish white,
very delicate and fine striae and ting'ed with purplish. Post.. discal black band broad, having
a s,eries of irregular triangu ar white blotches.
Distribution: It is found in aU sort of country above 4.,000 feet, and ranges from Kashlllir
to Assam and ascends to about 9., 000 feet in Himalayas. Uttarakhand : ChaJTIoli, Dehra Dun .
Rudra Prayag and Uttarkashi districts.
1932. Aulocera s'Waha swaha (Kollar) : Evans, Ident~/icafio1.l of Indi,all BUllel:/lies 2"d ed. : 117.
Distribution The subspecies is common frotn Chitral (Pakistan) to Sikkim and ascends
t'0 about 10,000 feet in Hilna ayas. Peile (1937) recorded it frotTI August to October at 6,000
to 10,000 feet [rotn hillsid,es and very COlTIlnOn at Nainital in Septernber. A'iso oocurs in
Nepa1. Uttarakhand Ahnora., 'Chalnoli and Rudra Prayag districts.
l
1867. Aulocera saras~va(i (Kollar) : Buder,. Enl. Month. Mag .., 4: ]2 1, no. 2.
General Ecology The Striated Satyr greatly resembles the Common Satyr. Unlike the
Comlnon Satyr, it shows a decided preference for open, grassy ditch sides, where it swarms
in late August.
Male and Female : Upper side : Wings dark brown, gloss,ed greenish,with a whitish
band, the white band very broad and nearly even throughout and reaching inner Inargin on
hind wing; apex ,and tomusmuch more rounded. A large blind 'Ocellus present in s'Ub ~apical
area. In the female forewing" the whitish spot on the inner edge of the b ~ ack sub ~apical spot
is large and elongate" in the male minut'e.
1844. Erebia 5canda KoUar1 In .' Hugel's Kaschmir und das Reich der Siek, 4(2) ; 452, pI. 171 figs, 3,
4.
1867 Ca/.l,e~ehia sctlllda Kollar: Butler. Ann. and Mag. of Nat, Hisl., Third serles, 20 : 217,. n. 1,
General Ecology ,: A very common butterfly seen along roadside in the month of August.
It has a typical Brown's flight. When disturbed., it flies dodging out of reach of the collector.
It can be seen basking for a very short titne.
Distribution : The subspecies has been recorded froln Kashlni r to KUJnaon and uplO an
,altitude of 8,000 fe,et in Hhnalayas . Uttarakhand : D,ehra Dun, Ahnora Bagesl1\var, ~l\;h ri~
Pithoragarh, Uttarkashi and Chalnoli districts.
1932. Yplltima sakra nikaea Moore Evans, The Identification of Indian BUllel:flies : 123.
General Ecology : This butterfly is COlnfnon in the rainy seasons but found" throughout
the year, .on grassy slopes and edg,es of forest. It fe'eds on flowers of various speci1es and flies
close to the ground. Its flight is weak. It also visits damp pat'ch'es. This butterfly does not
show any s,easonal forms.
Diagnostic Characlerf) :Antennae~ head, thorax and abdolnen alnber-brown, pa ler beneath.
72 Zoolog.ical Survey of India
Distribution . The subspecies is common from Murree (P,akistan) to Kumaon (India) and
ascends to about 9,000 feet in westemHimalayas from May to October. Uttarakhand :
Almora, ChamoIi, Tehri, Bageshwar and Uttarkashi districts.
1947. Yph.tima c.eylonica hubneri (Kirby) : Talbot, TheF:auna of British India (Butterflies), 2 : 329~330.
General Ecology : A v'ery common butterfly of faidy open hill country. The flight is
short, constantly or frequently settling on eaves or on grass along roadsides.
Diagnostic Characters : Antennae, bead, thorax and abdomen greyish-brown, ,abdomen
paler beneath. .
Male and fe.male : 'Wet s'eason fonn Upper side Wings greyish-brown. Forewing with
a sub-apical1arge ocellus, black, bi-pupiHed, with yellow-ring. Hindwing with two, sometimes
three, uni-pupilled post discat ocelli. Under side: Greyish white, not v'ery densely covered
with transverse brown striae. For,ewing with sub-api'c,al oc,ellus as on upper side. Hindwing
with one sub-apical and three contiguous posterior oc'el1L Both wings ,crossed by dark bands,
faint on the hindwing. In dry season form both wings somewhat paller on both sides, discal
and sub.. marginal bands more pr-onounc'ed. Hindwing under side with ocelli 'minute or absent.
Wingspan : 30-35 mm.
Distribution : This subspecies is common from 4,00.0 to 5,,000 fe,et in the hiBs and
throughout Peninsular India to ,Assam. Also OQcurs in Myanmar. Uttarakhand : Dehra Dun,
Haridw,ar, 'Tehri and Pauri districts.
1932,. Ypthima nare,da nareda (KolJar) : Evans, IdentificatIOn allndian Butterflies (2nd ed.) : 120.
Gene~al Ecol()gy : It is a hill flying species found in open, grassy 'khud' sides and in
lightly wooded country, s,een throughout the year.
Diagnostic Characters : Antenna,e, head, thorax and abdomen, brown, abdomen pale
beneath. Male and female: Upper side : Wings pale brown.; cilia of both the wings whitish-
brown; the broad sub-marginal band of under sid,e shows through. Forewing with sub-apical
ocellus. Hindwing with a sub-tornal and a Ininute tornal spot. Und,er side: Pale ochrac,eous;
striations well marked and uniformly spread; ocelli as on under side more or less in line;
hindwing with a large sub-apilcal oc'ellu . Both \vings with sub-'marginal, dark band" that in
the for'ewing broadening posteriorly.
1947. Ypthima baldus baldus (Fabricius) : Talbot, The Fauna of British India (Buttel.'!lies). 2 :337-338,
Gellerai Ecology : It is a v,ery comrnon butterfly of dry and lnoist deciduous forests. It
Oie' in open country and ,evergreen forest areas. It i' active throughout the year. It visits wet
soil e pecially in the sumrner .or on hot days. It is a frequent visitor of flowers, unlike rnost
other Browns' Fond of slnaU herbs and hrubs., especially Adeloclyslll11, Tridax etc.
Th _ Dry season fonn and wet ' ason fOfln tnay be obtained froln the suitable localities.
Diagno lie Characters Male andfelnale:Wet Season fonn Upper side Wings brown;
both fore and hindwings with the outer margins much darker, with sub basal and dis'cal
bands. Forewing with a sub-apical large ocellus, oval, black, hipupiHed, and yellow-ringed.
Hindwingwith two' smaller post disca) ocelli and two~minute tornal ocelli. Under side :
Brownish . .white, both wings With prominent sub .. basal, discal and sub"'marginal brown bands.
Hind wing with six smaller ocelli in pairs" two tomal, two medians and two in sub apical
area.. Female: Upper side of both the wings with the area surrounding the ocelli paler, closely
striated with brown. Under side is paler than the male and the transverse dark bands more
clearly defined. Dry season form in paler and the ocelli on the upper side of hindwlng reduce
to specks.
Wingspan : 32-48 mm.
Distribution ': The subspecies is common from North-West Himalayas (Chalnba) to Assa'm
upto 5,000 feet in Himalayas. Also occurs in Myanmar. Uttarakhand : Mussoorie in the
month of March and April, and from July to October in Dehra Dun, Haridwar ,and Tehri
districts.,.
1947, Melanitis leda ismene (Cramer) : Talbot, Fauna of British India. Bulter/lies. 2 : 366 369.
KUMAR: Handbook 011 Common Buttelilies of Utlarakltand 75
1773. Papilio undularis Drury, II/ustf: Exat. Ins ., ,2 : pI. 10, figs . l, 2.
1947. Elymni,as hypermn,es.tra undularis (Drury) Talbot. Fauna of British I"dia (Bulle/flies) 2 : .386.
General Ecology : This is the ,commonest and Inost widely distributed sp,ecies of the
Palmwflies, the only one to be found in Peninsular lndia. It is also found along the warm
valleys of Himalayas. Resident of pahn forests, avoid bright sunlight but fond of chequered
shade. Flight is weak, male often settling on pahn tre _. The female is less seen than the Inale,
and when on wings is an exc,eUent mimi,c of the Plain Tiger, Danalls cJlI:.vsipPll~ or the
Common Tiger, Danaus genutia.
Diagnostic Characters : Antennae,
head, thorax and abdom,e n brown;
abdomen paler beneath . .Male : Upper
sIde: W'ings blackish-brown with purple
gloss. Forewing with a sub@marginal
s,eries of blue spots, ,curving strongly
inwards ,and becoming more elongate
near apex, forming an oblique bar to
costa. Hindwing w'ith outer margin
broadly bright chestnest, sometimes with
a paler blue spots. Under side : Pale
Upper side Male
brown withfin,e striations. Forewing with
a sub-apical, broadly triangular, pale
purplish.white markings; both wings
with a purplish-white area. Hindwing
with a small white spot ,and toothed in
both the sexes. Female : Upper side :
Reddish-brown with apex and borders
dark brown, spotted with white.
Forewing with broad white band below
apex. Under side : Markings similar to
male, the pa'ie whitish markings more
extensive; inner margin broadly with
Upper side Female
striae.
Wood-Mason (1887) records that the males emit a strong odour resembling vanilla,.
Wingspan : 72e86 mm.
Distribution : This subspe'cies is common froOm Dehra Dun to Madhya Pradesh, West
Bengal and ASSalTI . Also occurs in Myanmar. Uttarakhand : Almora, Haridwar, Bageshwar
and DehraDun districts.
KUMAR: Handbook ,on Common Butterflies of UtlarakJutnd 77
Family 6. ERYCINJDAE
'The members of falnily Erycinidae ,are small in size. They are brown on upper side with
,a tawny horizontal band oOn each wing; the under side of hindwings are usually gr'ey and
protectively marked with dark brown dots and patches. The most obvious chara'cter that
makes them unmistakable is the enormous prolong,ation .of the palpi to front of the head,
giving the appear.ance of a be,ak, thus 'commonly known as 'The Beaks, The Punches and The
Judies.
All the species are confined to hilly, wood,ed country, preferring thick forest and shaded
areas. 'They are found around streams and in ,clearings.
The flight of ,aU the m'embers is rapid, keeping in ,air for a short period at a time. The
Beaks generally settle with the wings closed together over the back and the forewing drawn
into the hindwings. The Judiesalways settle with wings half open.
The members of this family never visit flowers but can be encountered near wat'er
sourc'es. The sexes are different in appearanc,e but none .of the mal,es have ,any se,condary
characteristi,cs.
The food-plants belong to the genus Libythea Urticaceae family.
The common examples found in the State are, the Common Beak (Libythea /eptia)1 The
Club B'eak (Libythea myrrha), The Common Punch (Dodona durga) .and The Tailed Punch
(Dodona eugenes).
General Ecology : A small sized, very COlnlnon butterfly found near str'ealns in fo rests,
settling very readily on damp patches. It is
always w,ary, gets frightened and flies off
rapidly. It visits flowers ofAnaphesis species
in 'Garhwal Himalaya"
Diagnostic Charac(,ers : Antenn,ae, palpi
and body dark brown, paler below. The
,colour of cilia throughout .ochraceous brown.
sub .. apical orange yeHow spots placed obliquely, sometimes separated, sometimes joined into
a band, giving a club-like appearance, apex cut off at the tip, with a margin 'produced into
a tooth at vein 4 and falcate below it. Hindwing with an orange-yellow broad discal band,
wide ,and diffus,e dat the abdominal margin but els'ewhere well defined and not r'eaching the
costa'~ outer margin wavy. Under side : Paler, forewing with a tawny marking as above, the
discal streak wider, occupying almost the entire basal area of the cell.~ the apex (rroTated with
purplish. Hindwing with no tawny discal band, irrorat,ed throughout with purplish. Fe,male
: pa' er throughout, tbe tawny markings larger.
Wingspan : 45-55 mm.
1857.. Libythea /.eplia Moore, In : Hor field and Moore's Cat.. Lep. Mus. E.l.C.. t : 240, no. 519
G,eneral Ecology: A little butterfly, abundantly found, usually in the deep of forest
anywhere where there is sunlight. It has a variety of flights like rapid flight, skipping and
sailing. It is commonest around forest streams in the dry season and may seen settling in
large numbers on damp sand ' at their edges.
Diagnostic Characters: Male and female: Upper side: Wings dark brown. Forewing
with an or.ange-y,ellow streak in the cell, divided into two portions widening in the apex of
cell, a large square spot of the same colour
beyond it on the disc in interspace 2 and 3, a
sub-costal white spot and a pre-apicaJ double
spot, the low,er portion orange..yellow, the upper
portion white. Hindwing uniform, with a disco-
,cellular orange patch near vein 2 to vein 6.
Under side : Paler. Forewing markings as on
upper side but the pve- apical double spot white;
costal margin and apex of wing greyish
Hindwing more or less cov'ered with grey scales
and blotches of green and dark brown. Ulp per side Ma e
Distribution : The spe,cies is re,corded from Kashmir to Assam and South India and
asc'ends to about 3,000 to 9,000 Ceet in North. . West Himalayas. Uttarakhand : Debra Dun,
Haridwar, Pauri, Tehri, Pithoragarh andUttarkashi districts.
KUMAR: H,a ndbo,o k 011 Common Bulter/lies of Unarakhand 79
General Ecology : A brown coloured butterfly with tawny spots on w'ings. It inhabits
forests of North India often near water stream and flies in the sunshine. It is a quick flier
but settles frequently with its wings ,closed. The males visIt damp patches and 'flowers as
well.
I
Diagnostic Characters: Male : Upper side: Both wings dark brown with numerous dark
yellow spots. Forewing with yellow short streaks across the cell, another at its end and a
large one beyond fro'm the Icosta, yellow spots near the bases of the medi,an intersp,aces,
below the ceH, post discal and sub-marginal areas~ five linear spots from below apex to
torna} angle. Hindwing with the disco-cellular marked with ,an 9chraceous line~ an irregular
discal series of spots, a sub-marginal and marginal linear series coalesc'e above the anal lobe.
Under side with the spots large and pale yellow throughout. Forewing with the base and
inner margin fuscous, the base of costa ochraceous, the base of cell enclosing a black spot.
other m,a rkings as abov,e. Hindwing with ochraceous markings. Female: Larger than m,aJe,
wings broader, outer margin of forewing more convex" apex less produced, markings similar
throughout as in the male.
General Ecology : The TaHed Punch is an aggressive butterfly. It can be found along
'w,at,er streams and shaded areas. It sits with wings c'losed and sucks moisture, Its flight is
w,e akand short and also visits flowlers of Anpha/is.
Diagnostic Characters: Male ': Upper side: Wings dark, blackish brown. Forewing with
a narrow line acros& the middle, a curved streak near the hind angle, and about thirteen small
spots equally distributed over the apical half of the wing; all these marks are whitish. Hind
wing with the outer portion traversed by four indistinct brownish-tawny lines; at the apex
,are two black spots, edged with the light brown; on the broad square lobe at the anal angle
is quadrat,e black spot, a slender tail arising from its outer edge Under side: Tawny brown
with silky shiny strips and spots .on the hindwing and ,costa of forewing.
Female ,: Large, the wings broader, the apex of the forewing more rounded, and the outer
margin .of the wings convex; aU the markings larger and paler.
Wingspan : 35-45 mm
Family 7. NYMPHALIDAE
Brushfooted Butterflies
The members of this family are the most beautiful alnongst butterflies. Always bright and
brilliantly, coloured, most of them have tawny background with black markings which may
be banded, spotted, blotched, striped, or variegated with very different patterns. The wings
are of varied shape, the hind wings may be tailed, toothed, scalloped, sharply angled and
indented, or evenly rounded. The bodies are robust, the thorax stoutly built and abdomen
comparatively short.
The fore legs are imperfect in both the sexes and the cell open in both the wings.
The family comprises more than a hundred genera from the world, of which around fifty
are found in India.
The size varies from very large Kaisers, the Great Eggfly, Oak-leaf (135 mm Wingspan)
to minute Baby Lascar (25 mm.).
They are fast fliers, difficult to catch. All the species of this family visit flowers and are
fond of sunshine and basking. SOlne species are attracted to over-riped fruits, fruits dipped
in beer, or freshly dropped dung.
The family Nymphalidae displays some interesting exalnples of Milnicry. The felnales of
the Danaid Eggfly, Hypolimnas misippus, perfectly mimic the Plain Tiger, Danaus chrysippus,
both in appearance and habits, the male being entirely different in appearance. The Oak-leaf
is beautiful and brilliantly coloured on the upper side, but very dull and completely different
coloured and marked on under side, like a dry leaf.
The sexes may be similar or different and the seasonal variation is well exhibited in sOlne
like Pansies, Oak-leafs etc.
Some common species of the family Nymphalidae found in the State are Athyma opalina
(Kollar), Neptis mahendra Moore, Neptis hylas astola Moore, Cyrestis thyodamas Boisduval,
Hypo limn as bolina (Linnaeus),) etc.
82 Zoological Survey of India
1857. Athyma opalina (Kollar) : Horsfield and Moore, Cat. Lep, Mus , E./.C. , 1 : 171 no, 351, fig. 2,
General Ecology: ,A Ylery common sp cies in the western Himalayas but rar,e towards
the East. It flies in the open areas and sunny nallahs where it can be seen in the cotnpany
of the Common and Himalayan Sail,ers. It settles on damp patches, stones leaves and 1
bushes.
Diagnostic Cha~acters ,: Male : Upper side: Wings bla'ck with creamy-white markings,.
Forewing with a narrow streak and two spots at its outer end in the ceH, a triangular spot
beyond, a sub-apical series of three spots~ a disc,at series of four spots from area three to
inner area; two sub-marginal wavy pale lines, the inner one is prominent towards the apex
and inner angle. Hindwing brownish black with creamy markings; discal white band from
the costal to inner margin, divided by the black veins; sub-marginal series of white lunules
placed from apex to inner margin.
Under side : F,orewing reddish brown; the posterior half of wings duller, the hindwing
suffused with pale lilac, the white markings as on upper side but broader, the dorsal margin
of hindwing greenish-blue. Female : Differs from the m,ale in the ground colour on both
sid,es being much paler, and all the white markings considerably large.
Distribution: The species ranges from Kashmir to Sikkim and Assam, Uttarakhand
,Almora,Chamoli, Dehra Dun, Haridwar, Pauri, Pithoragarh and Tehri districts ascending to
about 10,000 feet in Himalayas. A'Iso occurs in Myanmar.
KUMAR; Handbook on Common Butterflies of UtlarakhaJld 83
1932 Neptis hylas ,astola Moore: Evans. The ldenl{/ical,ion of Indian Butterflies. Bombay nat. Hist. Soc.,
: 166.
,Wingspan : 4S-60mm.
Distribution : The subspecies is ,common throughout tbe year in India and ranges froln
Kashmir eastwards to Myanmar, aSlcending to about 9,000 feet in Himalayas. Uttarakhand
: Almora, Dehra Dun, Haridw,ar,' Pauri, Tehri, Bageshwar, Pithoragarh and Uuarkashi
districts.
84 Zoological Surv,e y of I"dia
1969. Neptis mahendra malrendra Moore : Eliot. Bull Bri. Mus. (nal. Hisl.) en'. Suppl.. 15 : 79,
1932. Neptis yerburyi yerbury; (Buller) Evans. The Identification of Indian Butte'ilies. Bombay ll£Jt. Hi.(j/.
Soc., : 167.
General Ecology: It is found in w1eJl-wooded naHahs and ravines, fond of sunlight and
basks on rocks and leaves with it wings held wid open. In the hot, dry weather lnany of
them settle on damp patches near strealTIS.
1836. Cyl~estis lit yo dam as Boisduval, In: CUl'i,e r .$ Reg. Animal. Ins"~ 2, pI. 138, fig . 4,
General Ecology : A v'ery delicate and beautiful butterfly, aptly called Map Butterfly,
because of . ts wings having ITIap like tnarkjngs and irregular outlines. It is often seen soaring
backwafd and forward over Inountains., and streams in richly \vooded areas. It flies very
slowly with a jerky flight, keeping the wings horizontally for longer tilne than any other
Indian butterfly. On al,ert, takes a very erratic .and fast flight. In we' tern Hilnalayas., it visits
flow,ers of Budd'ieia and Ch'e stnut It also visits damp patches.
Diagnostic Characters: Male and fe.male : Upper side .: Forewing whitish in male and
yellowish in female, with map like black markings and irregular vertical dark outlines across
both wings. Wings broad and s'e mi-transparent. Tornus of forewing and apex of hindwing
cut off. Hindwing tailed at vein 4 and lobed at tomus. Tornus lobe of the hindwing yellowish
brown with a central large round black spot.
Und,er side: Paler, marked much as above, the dusky red markings at the anal angle and
lobe of the hindwing more distinct.
19 II. Hypolimnas bolina (Linna us): Tyfler. J. Bomhay 1101. His!.. Soc .. 21 (1): 54.
Gener:al Ecology A beautiful butterfly rare in the drier portions of western India,
occasionally found in gardens, abundant in the lTIoist regions, particularly in the \Varn1
valleys and sub-Inounta'in tracts. It feeds exclusively on nectar with its wings partially clos~d
whil,e sipping the nectar. It is ·a pugnacious species 'Whl,ch ails forth and cha$es O\\'Hy any
butterfly that conles close.
Diagnostic Characters Body above dark indigo-blue, h ad tnarked \vith a te\v pale
spots; palpi beneath white, abdoInen and thorax with a few whitish spots.
Male : Upper side : Wings very dark indigo blue. Forewing with a large elongated
macular white spot,lnargined with bright bluish from upper disco cellular to vein 3, with ,a
row of small white ocelli from apex to the tornus of hindwing. Hindwing with a broad
rnedial whitish fascia. Under side: Dark olivaceous brown. Forewing with the white Inarkings
as above, but with the large macular spot almostext,ending to costa, not prominently lnargined
with bluish, and followed by a small pale spot beneath the second Inedlan nerv.ure. Hind\ving
with a smaJl sub-costal pale bluish spot followed by a paler medial fascia 'iending at the inner
margin, sub .. m,arginal area pale greyish, with some dots.
Female: Upper side: Dark brown with outer diseat row of slnall, pale yellow or whitish
spots on both wings. Forewing with concave margin, having more or less obscure lnarginal
and inner terminal markings. Two blue spots in the cell in wet season form and one in dry
season form. Hindwing with broad pale yellow or white tennen, hearing a dark wavy line
or band. 'Under side : Brown with bands of white spots and patches. The female of this
species is a good mimi,c of The Common Indian Crow (Eupolea cOJ'ie).
Wingspan : 70-1l0 m'ln.
Distribution : The species is widely distributed and found ahnost throughout the plains.
It ascends nonnally to about 7,000 feet in Himalayas. Also occurs in Sri Lanka and Myanlnar.
Uttarakhand : Almora, D,ehra Dun, Haridwar, Pauri, Bageshwar, Pithor,agarh and Tehri districts.
88 Zoological Survey of India
General Ecology: It is one of the most bea uti ~u l butterfl y alnong Ny rnphal id wi th tbe :I
upper side being beautiful and brightly coloured. The leaf-like under side is lnost \\'( nderful
example of protle ctive resemblance. It is
difficult to locate the butterfly once it settles
down on a twig, riverside boulder, tre,e trunks
-or on ground with its wings closed. It is found
in the thick forests of mountainous and hilly
districts, at low or moderate elevations. This
species, a fast flier, gets attracted to ov,er-ripe
fruits and damp patches.
Diagnostic Characters : Male and female
: Upper side : Wings dark violet-blue to deep
steel blue with orange discal band and broad
black band on the forewing. Forewing ·Iarge,
sub-triangular; costa very much rounded,
slightly emigrate near the ba "e; apex less or
more acute and produced; outer margin five- Upper side Male
sixths of the length of the costa, concave
below the apex. Hindwing elongate, sub~ 1
Distribution: 'The species ranges from Kashluir eastw'ards through Kumaon to Sikkhn and
Assam, also reco~ded fr{)m 'Orissa and Maharashtra. Also occurs in Nepal and Myanlnar.
Uttarakhand : Dehra Dun, Haridwar, Tehri :and Uttarkashi districts up to an altitude of 8,000 te'et.
90 Zoological Survey ofb,dia
1932. Juuonia hei.rta (Fabricius) Evans. Tlte IdentificatiQIJ ofl"dian Bfluelilies. Bombay nat. His.l. Soc.,
: 176.
General Ecology : A beautiful species that prefers hot and sunny places in the forests"
parti,cularly fond of dry riverbeds, stony uncultivated fields and roads; very often fly in fr'o ot
of the collector. It flies slightly just above the ground and intermittently settling and flying
not far from the collector's sight. It Inay be s'een almost throughout Ithe year.
Diagnostic ,Characters: Male: Upper side: Black. Fore-wing with broad tnedial yellow
patch, extending from the base to beyond the disco . . ceHular, and then narrowed and bent
downward.; two short paler yellow str'eaks before the apex. Hindwing black, with large broad
sub..anal yellow patch, and a 'large distinct blue sub-basal spot. Under side : Forewing pale
y,eHow brown; medial patch paler yellow and suffused outwardly, ,cross'ed by bla,ck discoidal
lines; two black discal spots. Hindwing with basal half greyish-yellow ~ discal area pale
yellow, crossed with brown lunular 'lines. Female : Upper side : Forewing dark yellowish-
brown, with the yel10w patch paler, shortened basally, and crossed by two black lines on
'middle of the cell, and a broader disco-cellular band; black ocelli present in area 2 and 5.
Hindwing with very smaU upper and lower disc.al 'Ocelli, and two marginal lunular yellowish
lines. Under side : As in male.
Distribution : 'This subspecies is very ,common at Jow elevations and plains in India, but
less common above 4,000 feet and ascends to about 9,000 feet in Himalayas,. Also occurs
in Sri Lanka and Pakistan. Uttarakhand : Dehra Dun, Haridwar, Pauri, Nainital, Pithoragarh.,
Tehri, Uttarkashi and Udham Singh Nagar districts,.
KUMAR: Handbook on Common Bune-lilies of Uuar,a khand 91
General Ecology: One of the prettiest of Indi.an butterflies, found throughout the country
but more in dry, open plains than in ·fof,ests. The dry riverbeds and stony paths areils
favourite habitats; remains active during th,e hottest part of the day.. It visits flower of
Marigold~ Bachelor's button~ Lantana. Heliotrope etc.
.Diagno ·tic Characters: Male; Upp,er side: Forewing: Basal two-thirds blue bJack, apex
pale brown with white transverse bands; outer discal area below apex shining blue; two
orange ringed ocelli in area 2 and .5. Hindwing shining blue with two ocelli near outer
margin in ·area 2 .and 5; a black basal angular patch curving across the cell towards the anal
,angle, and merging into brown on the abdomen margin; two pale marginal lines.
Under side : Dull ochraceous,with white m,ark' ngs and wavy lines. Ocelli visible in
forewing only_
~ema./e : Larger than the m.ale, both the ocelli more profninent and of a bright red. Upper
side: Basa) half area of hindwing being entire"y black. Both the sexes are variable in colour
and markings due to seasonal variation. The dry season form being pale stone grey ,colour
on under side, compared to that in wet season form.
Distribution .: The species ranges from Pakistan ·eastwards through Him.alayas to SikkitTI.
The species is common in India during March t'0 October in North-west Hitnalayas and
ascends to about 9,000 feet. Also occurs in Sri Lanka and Myanmar. Uttarakhand : D,ehra
Dun, Chamoli, Haridwar, Pauri, Rudra Prayag, Pithoragarh, 'Tehri, Uttarkashi and Udhaln
Singh Nagar districts.
92 Zoological Survey of India
Gen~ral Ecology: A som'ewhat bold butterfly, having strong territorial and aggressive
behaviour. It tli'es quickly and strongly but with less fluttering .of wings,. (t visits various
species of flowers viz,. antana, Bachelor's Bottom,) Cosmos etc. and rests on ground. On
approaching, it darts ahead with rapid wing beating, but settles soon not v,ery far.
Di.agnostic Characters: Male and female: Upper side: Forewing dark brown or greyish
brown with the cell crossed by slender
wav,ed black Iines near base, two
waved black lines enclosed a pale
yet lowish-brown band beyond the
middle, and a similarly coloured band
at about end of cell; discal and sub-
marginal series of yellowish-brown
spots extending on the tornal angle
and two blue centered red ringed
ocelli in area 2 and 5, the ocellus in
area 2 indistinct and prominent in area
5.. Hindwing dull yellowish-brown
with a large ocellus enclosing 2-
minute white spots and lower bla,ck
ae
spot.
Under side : Yellowish-brown in male, pinkish in female with sev,e ralwavy lines and
the lower ocellus in the forewing visible. The seasonal v,ariation is quite well marked., with
th,e ocelli on wings prominent in wet season form ,and reduced in dry season form.
Wingspan : 45-60 mm,
1932. Junoni,a a/mana a/mana (Linna'e us): Va rshney, Orienla/ Insect, 28 : 176.
General Ecology A cornmon and beautiful butterfly, found in cultivat,ed areas, forest
=
clearings and paths. It basks in sunshine, in the early morning. It has a highly territorial
behavior and often chases away intruders.. It visits flowers of Marigold, Lantana., Land
C,altrops etc.
Diagnostic Characters : Male and female : Upper side : Forewing orange-yellow, the
cell and the costal area to near the apex of wing crossed by four short dark bands, a pale-
centered ocellus with the two black rings and generally two pre-apical ocelli. Hindwing
with three border lines as on fore'wing but well defined, a very large pale yellow black-
ringed .ocellus spreading over interspaces 4 to 7, the center inwardly reddish-yellow with
two white spots, outw,ardly black. In some specimens a small black-ringed ocellus in
interspace 2 is pr,esent. Under side ! Leaf-like and pale yellowish brown. In wet season
form, the ocelli :a remore prominent on hindwing andm.argins of wings smooth. In dry
season form, the apex in forewing is prominently toothed and without ocelli, hindwing is
produced into a narrow tail.
Wingspan : 60-65 mm.
Distribution : This subspecies occurs throughout India, ascending t.o about 7,500 feet in
Himalayas. Also .occurs in Sri Lanka and Myanmar. Uttarakhand : Almora, Deh~a 'Dun,
Handwar, Pauri, Bageshwar and Tehri districts.
94 Zoological Survey 0.( l.ndia
1994. J"1'Ionia atlites (Linnaeus) : Corbet & Pendlebury, Butterflies of Malaya Peninsula : 59
General Ecology : Inhabits heavy rainfall regions and does not occur in dry areas; very
fond of settling on shrubs close to wet areas. It visits flowers of various plants.
Di~gnostic Characters : Male and female: Upper side :; Pale grey. Forewing with two
wavy black lines crossing the middle of the cell, and two similar ones at the end of the cell;
a very waved and fuscous line crossing the wings a little beyond the e·en, and betwe·en this
and the outer margin are two waved fuscous lines, between which the colour is distinctly
paler,and is marked with a series of six oceHated spots, of which the first, 2 and 5 are largest
and'most brightly coloured. Hlndwing with two waved fuscous lines 'c rossing end of the ceU,
and remaining m.a rkings similar and
continuous to those on the for,ewing;
only five ocelli an:~ present, some
of the ocelli in both 'w ings are
orange-yellow on the inside. Under
side: Very pale; the cell crossed by
the fuscous lines ,as above; the ocel i
more distinct; marginal and sub
marginal lines indistinct. Female
larger in size and generally darker
in colour than the male; the
ocellated spots above larger and
mor,e distin,c t, with the area on
which they are placed much paler.
Upper side Male
Wingspan : 55 .. 65 mm.
Distribution : 'T he species is very common throughout plains of .India and occurs around
paddy fields, ascends up to 7,500 feet in Himalayas. Also occurs in Sri Lanka, Myanmar,
China and Malaysia.. Uttarakhand : Almora, Dehra Dun, Haridwar, Tehri, Chamoli, Nainita'l,
Champawat, Bageshwar,Udham Singh Nagar, Pauri and Uttarkashi districts.
KUMAR : Handbook on Common Blillel:/7ies of Ut,t arakhand 95
1992. JUlfonia iphilO (Cramer) . Corbet & Pendlebury, .Butlerflies of Malaya Peninsula : J59.
General Ecology: It is found in wetter and w,eH-wooded hilly regions, it is seldom found
in the plains of upp,er India and never in the dri'er parts. It is fond of settling on damp
patches, w,et roads and shady places. It also visits flowers of various species,. It has a bold
flight, but not to any gr at distance, rests with \vin gs closed.
Diagnostic Characters : Male and female : Upper sid,e : Forewing brown w,ith indistinct
dark brown bands; the apical half of wing is somewhat paler and sub-,apical spotwhitish~
sub-ma~ginal lines wavy and dark. Hindwing with the ,apical half paler and 'with the band'
on the forew ing continued, but the inner one lnorew,aved:; four to five oceHated spoOts
between area 2 and 5. Sub-marginal lines as on forewing. Under side : Paler; both wings
crossed by two somewhat broad., dark, basal fascae, tbe first crossing the middle of cleH of
the forewing, curved inwardly on cell of hindwing, the second crossing forewing at the apex
of cell, and tenninating on hindwing a little beyond end of cell; oceUated spots obscure on
both the wings. In dry-season form, the apex of forewing is produced and angulated and the
hindwing lobed at tornal angle, with the under side paler and leaf-like.
Distribution : The species occurs throughout the Himalayas as far west as 'Kashtnir,
ascending up to 9,000 feet, and also found in central andsouthem India. AlsoO occurs in Sri
Lanka and Myanmar. Uttar,akhand: Almora, Chamoli, Dehra Dun., Haridwar, Nainital, Pauri,
Bageshwar, Champawat, Pithoragarh, Tehri, Uttarkasbi and Udham Singh Nagar districts.
96 Zoological Survey of flldia
Diagnostic Characters : ,Male and female : Upper side : Both wings ochraceous·red.
Forewing with basal ar,e,a ochraceous brown, an oblique irregular..shaped angular band from
Iniddle of cell to the tornal angle, before the apex is a short white band and a curved series
of four spots, margin dark brown. Hindwing with the basa1 area, a confluent discaI band and
the costal bOfder ochraceous brown, a transverse disca) row of five black spots with slightly
paler outer ring, a sub..:margina\ row ofl~nu\ar spots, and 'a marginal row of larger spots.
Under side : For,ewing similar as upper side but the basal area more red and the apex and
outer margin ochraceous, some additional black spots on forewing cell. Hindwing ochraceous,
richly spotted and marbled with deep shades of ochraceous ,and brown, an ov.al spot across
the middle of cell and the row of five discaI spots as above but developed into ocelli, the
2 and 5 largest, with blue c,enters and black outer rings.
Wingspan : 55-70 mm.
Distribution: Widely distributed and well known throughout India, as a migratory species,
all over the world. In ndia, it ascends to about 5,000 feet in Himalayas. Uttar.akhand :
Chamoli, Dehra Dun, Haridwar, Almora, Rudra Prayag, Bageshwar, Champawat, 'Udham
Singh Nagar, Nainital., Pauri, Pithoragarh, Tehri and Uttarkashi distri,cts.
KUMAR: Handbook on Common Bultelilies of UUcu'akluH,d 97
1794. Papilio atlanta indica Herbst, Nat. Schmett., 7 : 171, no. 64, pI. 180, figs. ),2.
19l1. Vanessa indica (Herbst) Tyller, J. Bombay nat. Hist. Soc. , 21(1) : 62,.
General Ecology: Inhabits wooded region, forests, and open country. disturbed hUlnan
habitations and gardens. It flies rapidly up and down, frequently settling on the ground with
the wingsthr,e,e ~fourths or half open. It is always wary and difficult to ,approach, best caught
when feeding on flowers. 'G,ets attracted to droppings, dung and rotting fruits.
1886. J{allessa ege,a (Cramer) .: De Niceville, The BuueJ:/Ues of India. 8111'/11(1 alld Ceylon. 2 : 237.
Gen€l'ai .Ecology : Inhabits grassy meadows and open hilltops at high elevations. There
is considerable difference between the spring and autumn forms, the spring forms being
pale, especially in the nortbern parts of its range . It flies rapidly and swi ftly. It frequently
settles on the ground or on leaves.
Distribution : The speci,es ranges from Baluchistan (Pakistan) eastwards to Nepal and
Bhutan and asc,e nds to about 11,000 feet in western Himalayas.. Uttarakhand Tehn and
Uttarkashi districts.
KUMAR: Handbook on COl1tmon Bultel:!lies of Uuarakhand 99
1983. Aglais cashmirensis (KoJlar) : Varshney, Rec. zoo/. Sun!. India, Occ, Paper No .. 47 : 8 .
'General Ecology: Cotnmonest species of the North .. West Hilnalayas, found an the year
round.. It can be seen in an kinds of terrain. It visits various kinds of flowers like TaraXllCUln
officina Ie, Aster sikkimensis, Gentiana carinata etc. It loves sunshine and can be seen
basking on paths with the wings widespread. The flight is rapid, always close to the ground
seldom sustained for long distance.,
Diagnostic Characters : Male and jelna/e : Upper side : Both wings coloured ri'ch
chestnut. Forewing with a quadrate black bar across the rniddle of the e,ell, then a pale
yeUow bar, then another much larger black bar, with anotber narrower pale yellow bar
beyond it, with a small bluish-white ,c ostal spot placed outwardly against it. Apex cut off at
the tip and produced and angulated at vein 6. Hindwingwith a basal area blackish~ inner
m,argin paler in basa.l half; outer margin bl,ack, bearing,.prominent bluish lunules, followed
by two paler lines:;ma~gin toothed at vein 4. Under side: Both wings brown at bases, paler
beyond, and thickened striated with black; a sub-marginal lunulated black line . Forewing
with a narrow oval black spot with pale center at the base of the cell, two fine black zig-
zag lines enclosing a black space across the lniddle of c'elL Hindwing with prolninent
ochraceous spot at the lower end of the cell, and two fine bla,c k lines enclosing a blackish
space across its middle. It shows seasonal variations, the wet season fonn being bright1er and
dry season fonn shows darker broods
Wingspan : 55 .. 65 min.
Distribution: The s'p e,e ies ranges from Waziristan (Pakistan) to Sikkiln, and i" cornlnon
in its range fro.m low ,elevations t.o 15,000 feet in itnalayas. Uttarakhand: Ahnora, Chalnoll,
Tehri, Pauri, Rudra Prayag., Bageshwar, Pithoragarh and Uttarkashi district
100 ZOO/OgiClI/ Srtrl'ey of 11Idia
] 983. Kalliska caJlace (Linnaeus) : Varshncy, Rec.. zool. Sun 1, Inc#a. Occ. Paper No. 47 : 8.
Gen~ral Ecology It is found in the wooded country of hilly regions and vicinity of
water. It flies fast :a nd straight, up and down over streatnbeds, roads paths etc. It trequendy
settl,es on dalnp patches. over ripe fruits~ sugar, hon y and sap. It rar,ely visit flowers . It has
a favourite re ting spot to \vhich it leturns again rather hard to catch.
Diagnostic Characters ; Male and jenlale Upper side Both wings deep indigo~blue,
crossed by a broad discal pater blue band with a fe·w white dots at the anterior end, the band
gradually increasing the width froln the costa to the anal ,angle, ,and bearing a series of slnaH
black spots along its outer border in the hind wing~ ·inner margin lighter; outer margin lobed
at vein 4. Forewing with the costa more or less broad blue bar. Under side: Most beautifully
vadegat.:ed and striated with hla,ck, green, ochraceous, pale violet and ferruginous, cross,ed
by a prominent discal dark broad band, with the edges irregular and scaHoped. Fore wing
with a smaU ochraceous spot ,at the outer lower end of the cell. Hind wing with a prominent
tail like projection in the middle. Female, usually larger than the m.ale., markings similar, but
the marginal blue lines usually obsolete, the angulations of the outer rnargins broader and
larger.
Distribution : The species ranges from Chitral (Pakistan) eastwards through Kum,aon to
Sikkitn and Assam., ascending to about 9,000 feet in Himalayas. Also occurs in Myanlnar.
Uttarakhand : Alrnora, Chamoli, Dehra Dun Nainital, Rudra Prayag, Bageshwar, Champawat,
Udham Singh Nagar, Pithoragarh, Tehri ,andUttarkashi districts.
KUMAR : Handbook on Common Bulter/lies of UUarakhaud lOt
1932. Argyreus hyperhius hyperhius (Johanssen) : Evans, Identification Qf Indian Butter/lies, : 182.
General Ecology: It is one of the beautiful species of the family Nymphalidae. It occurs
in the hilly regions and seen in open areas, ,cultivated fields and gardens in the plains. 'It
visits flow'ers of Zinnia" Aztec, Marigold, Buckwheat ,etc. It rests on the ground with wings
closed and sometimes basks with its wings three-fourths open. The male is a fairly fast flier
compared to the female.
Diagnostic Characters: Male: Upp,er s'ide : Wings bright orange yeHow with yel10wish
green base. Forewing with black sinuous cellular lines, quadrate, triangle, dentate., and round
spots; marginal lines. Hindwing pale; 7 discal spots .and Inarginal border black and dentate
and followed by a row of slender brown or bilue lunules.
1767. Papilio lalllol1;a Linnaeus, Syst. Nat. (l2 1'h ed.), 1(2) : 7,86. no, 213.
1990. Issoria lalhonia (Linnaeus) : Va rshney, Revised ,Nomenclature for Buttel:11y (,Qxa .' Jour. Bom. nat.
Hist. Soc .. 87 : 60.
Diagnostic Characters : Male = Upper side: tawny, with bla,ck spots. Forewing with the
base" costa and inner margin tawny, hindwing with basal and abdominal area widely, mottled
with darker coloured scales. Forewing with a reniform spots enclosing a portion of the
gr.ound colour, a transv,erse spot beyond and .one ,closing the end of the ,cell. H indwing with
a spot closing the cell, three angled series of black spots in continu.ation of those on the
forewing, the outer margin as on forewing.
Under side :. Paler., Forewing with the subm,arginal series of black spots pupilled with
silver, a prominent sliver spots within the upper most spot of this series, and a five apical
marginal sHver spots increasing to the third which is the 'largest. Hindwing yellowish brown
with a large silver spot.
Wingspan : 50-60mm.
Distribution: The species is one of the commonest Himalayan species of Ar-gyrelts and
ascendIng to about 5,000 to 8,000 feet in Himalayas. Also occurs in Nepal and Myanmar.
Uttarakhand : Almora, Cham.oli, Dehra Dun, P,auri, 'Tehri, Pithoragarh and Uttarkashi districts.
KUMAR : Ha11dbook 011 COmmOll Butteljlies of Uuarakltalld 103
General Ecology ': It is the finest of all Indian FritiHaries and attains largest size. It is a
North-West Himalayan species, It flies very rapidly and can be f70und along grass slop'es,
roads, in glades and in open country. It frequently s,eules on the ground and flowers' rather
an UnCOITI'm On spe,cies.
Diagnostic Characters: Male: Upper side: Forewing dark or.ange with two narrow
sinuous lines ,across the middle of the cell. Pre,-apical area pale orange; black spots in lniddle
and end of cel , the fonner not touching the medial nervule ,and followed by two sub-
marginal series of rounded spots. Hindwing as the forewing; with a streak at the end of the
cell, a discal-angular series of six spots beyond the cell, an inner sub-marginal series of five
round spots, another series of eight spots beyond, becoming sOlnewhat lunular towards the
anal angle; margin dentate. Under side : Forewing red; pre-apical area ochraceous and the
apex greenish, crossed by two silvery lines; other markings as above but more prominent.
Hindwing rich shining greenish ochraceous" crossed by 'm,any silvery lines and bands, edged
black on one or both sides; outer margin dentate. Head and body thickly c:tothed with
fulvous hairs
Female: Differs from the male only in the ground colour ,as under side being much dull.
Distribution: The species ranges from 'Chitral (Pakistan) to Kumaon and Assam. ascending
to about 7,000 to 10,000 fe,et in Himalayas. Al 0 occurs in Myanmar. 'Uttarakhand : Ahnora,
Chamoli, Nainital, Tehri, Rudra Prayag, Bageshwar, Udham Singh Nagar, and Uttarkashi
districts.
104 ZQologlc,a l Survey o/India
1990. Fabrici,QIlQ kamala (Moor'e) : Varshney, Revised Nomenclature for Bunerfly taxa, Jour. Bom. nat.
Hijt. Soc ... 87 : 59.
General Ecology It is tnuch slnal1er than Childl':ena childreni, found along roads, in
glades, at the edges of forest and in open country, frequently pitching on the ground or on
shrubs, thistles, bralubles and visits low growing flowers. The flight is fast and rapid.
Under side: Forewing with the markings as on upper sidle, apex dusted with dark gr,een
scales bearing two Hnes of silvery spots. Hindwing beautiful metalHc green with five silvery
sub-tnarginal and inner bands and a seri,es of m,arks about the basal half silvery white;
between the suh-marginal and inner band some sm,all spalces of yellow, and an indistinct r'ow
of reddish spots.
Wingspan: 65 ~ 75 mm.
1773. Papilio pJralantha Drury, II/us. l1at. Hist. 1 : pI. 21, figs. I, 2.
1997. Phatanta pllalantha pllo/allllta (Drury) : Varshney et al. . Rec. zool. Surv. India, Oec. Paper No .. 31
: 18-19.
General Ecology : A very 'Common species, occurring in the gardens and at the edg1es
of jungles. It avoids shady places. It visits flowers of Lantana, hisHes, Pigeon berry, Tridax
species and damp patches, and wet roads. Sometimes dozens of individuals assemble atlnud
puddling sit'e sin the hot season. It is fond of basking in sunshine. lts flight is fast with jerky
movements. It has a very strong territorial behaviour and chases any butterfly, which ,comes
in its range.
Distribution : The subspecies occurs throughout the year in the plains of India and
ascends to about 8,000 feet in HimaJayas. Also occurs in Sri Lanka, Pakistan and Myanmar.
Uttarakhand : Almora, Dehra Dun, Haridwar., Rudra Prayag, Chamoli, Uttarkashi, Nainitak
Pauri, Udham Singh N,agar, Pithoragarh and Tehri districts .
]06 Zoological Survey of India
1932. Ariadne merione (Cramer) : Nandi & Varshney, Rec. zool. Sun. India, 85(1) : 127.
Gen~ral Ecology: Also known as Castor Butterfly, abundant :in the hills and forested
areas. It oc'curs in castor fields and around edges and bushes. It 'Occurs throughout the year.
It is a w,eak flier and prefers to -fly close to the ground and in the neighborhood of its host
plant, i.e., Castor.
Distribution : The species is quite common (roln Kashtnir to Kumaon and ascends
to about 7,000 feet in North West Hitnalayas, recorded also from Orissa and Madhya
Pradesh. A1so occurs in Sri Lanka and Myanmar. Uttarakhand :Almora, Dehra Dun,
Haridwar, Nainital, Bageshwar, Champawat, Pauri, Pithorag,arh, Tehri and Udham Singh
Nagar districts.
KUMAR: Ha11dbook ,on Common Butte 1ilies of Ullarakhand 107
}770. Papilio ,erymanthis Drury, Exot. Ent.• J ; pI. 15, Figs. J-4.
General Ecology' Inhabitant of thickly forested areas. It flies close to the ground with
a w,e ak flight. When disturbed, it dives into the undergrowth. Both the sexes Vlsit flo\\'ers
and males occasionally to damp patches.
.Distribution ; The species occurs from Mussoorie (western Himalaya) to ASSalTI and
Sikkim., ascends to .about 5,000 feet in 'w estern Himalayas, also recorded from South India.
Also oc'c ursin Sri Lanka and Myanmar. Uttarakhand : Dehra Dun, 'Tehri, Ahnora, Bag,eshwa.r,
Champawat, Udham Singh Nagar and Nainital districts.
108 Zoo!o,g ;cal Survey of bldia
1932, Sephisa ,dlchroa (KoUar) Evans. The IdentifIcation of Il1dia1.l Butterflies. Bombay nat, Hist. Soc.
147.,
General Ecology: It is fairly cornrnon butterfly, visits damp patches, dung, flowers, over. .
rip'e fruit etc, and generally found in shaded nallahs bordering forested regions and along
strealTIS. It flights rapid and g,eneraJly returning to the same spot
Wingspan : 60 . . 75 mm.
8. Family LYCAENIDAE
Blues
Members of the family Lycaenidae are commonly known as "The Blues" They are small
to medium-sized butterflies, with predominantly blue wings but major colour being violet to
various shades of blue. Brown, chestnut, or blackish tinge on the upper side of wings are
not uncommon. Upper side of wings tnay be unmarked, or may be spotted or banded, with
a variety of colours. The under side is typically spotted, banded and streaked in intricate
patterns and hind wings with the outer margin smoothly curved, or the tennen produced into
a lobe or a tail, which may be long or thread like.
Both the sexes have fully developed forelegs and the pre-costal vein is absent in the hind
wing. They are very delicate, very slow fliers and found almost everywhere (meadows, open
grasslands, shrubs and forest clearing). The males are fond of sunshine and visit flowers and
dalnp patches but the females seldom venture out of their habitats.
Many species show in their early stages association with ants. The ants carry caterpillars
to their nests and nurse them for their sweet-scanted juice and in return protect the caterpillars
and chrysalis from their enemies.
Seasonal variations occur in a number of species, like change of colour and markings on
the wings sometimes being very pronounced. Some species appear so Inuch alike that often
it becomes difficult to separate them with certainty except by examination of their genitilia.
The Lycaenids are among the smallest of the butterflies. The "Grass Jewel" is the slnallest
of all the Indian butterflies, measuring about 15 mm.
SOine common species of this family occurring In the State are CastaUlIs rosilnon
(Fabricius), Lampides boeticus (Linnaeus), etc.
no ZooJogic,a! Survey of India
1907. Caslalius rosimol1 (Fabricius) : Bingham "rile f',auna of Britid, India" (Bunel/lics) , 2 : 424.
General ,Ecology : A COlnlnon speci1es of the open d,eciduous forests" scrub" grasslands
and near hUlnan habitatiQns.Aweak flier ,a nd always flies close to the ground~ avoids
thickets or shaded places, prefering edges and roads in the forests; fond of sunshine and
basking,. It visits flowers and dalnp patches.
Diagnostic Characters : Head, thorax and abdomen black above and white beneath, the
latter banded at the sides.
Male: Upper side: Both wings white, with a grey-blue base, ov'er which a brilliant
silvery irroration, varying according to the light; anterior margin ,and border blackish-brown .
Forewing with a very irregular series of square, broad, dark spots on disco-cellular ,and post-
discal area. Hindwing with spots as on forewing but narrow and not irregularly placed.
Under side: Both wings white with deep spots and streaks.
Female : Larger than male, both wings with posterior border broader, the black colour
more prominent; both wings as in the tnale. The species shows seasonal variation. Th,e
specimens on the wings during the rains are much darker and heavily marked than those
occurring in the dry s'eason.
Distribution : The species is comtTIQn throughout India including Andaman and Nicobar
Islands, ascending to about 7,000 fe,et in Himalayas. Also occurs in Sri Lanka and Myanmar.
Uttarakhand : Ahnora, Dehra Dun, Haridwar ~ehri, Bageshwar and Pauri districts .
KUMAR: Handbook on Common Butt(n1he.') (~r Uuar.akhalld
1962. Lampides boeticus (Linna'e us) : Cantlie The Lycaenida1e portion of Brig. Evans "rite Identification
of Indian Butterflies, 1932 : 6'9.
Diagnostic Characters: ,M ale: Upper side: Both wings violet~ blue with two tornal
spots on hindwing. Forewing with tbe costal.nargin narrowly and the outer Inargin more
broadly pale fuscous. Hindwing with the costal and post'erior margins p,al'e fuscous" two
large bla'c k spots at the anal angle,. Underside : Both wings p,a ie brown or whitish with
narrow brown bands. Hindwing with two orange crowned tornal black spots. In between
the brown bands a broad wh itish discal band.
Female ': Upper side : Both wings dark brownish with blue scaling at the bases;
generaHy with obscure white marginal crescent. Forewing with a disca1 bluish patch.
Hindwing has a slightly long, white tipped tail; two outer greyish sub-marginal bands .
the inner broadest; black anal angutar spots as in the tnale, distinctly m,argined with
greenish,. Under side : Both wings ,as in the male.
Wingspan : 24. . 36 mm.
Distribution : The species is common throughout the country and ascends to about
J0.,000 feet in Himalayas. Uttarakhand : Chamoli" Dehra Dun and Haridwar districts.
112 Zoologic,a l Survey of India
1932 . Lycaena pavalla (Kollar) Evans, rhe Identification of .Indian Butte,:/lies : 244 .
•
G,eneral Ecology: A fairly cornmon species that .occurs during SUlTIlner months on the
outer ranges of the west,e rn Hilnalayas. It loves sunshine and open places" ' specially waslte
lands. The flight is lively but not very fast, close to the ground and seldoln sustains. It visits
low growing flow,_rs.
Diagnostic Characters : Male and fe/nale : Upper side : Forewing rich copper red, the
,c osta narrowly, tbe out'er margin broadly black; three black spots in the discoidal cell; :a
discal seri,es of seven spots placed in the ,e chelon. Hindwing dark brown with an obscure
central copper patch; a linear black spot at the end of the 'c ell, two similar spots beyond it;
a strong marginal hand of copp'e r cresc1ents. Under side:: Forewing bright orang'e, the outer
margin cinereous, bearing an outer series of black dots and an inner series of prominent
black spots. Hindwing brownish-grey, with prolninent black spots surrounded by a narrow
whitish ring., three towards the base, thr,e,e on the dis,c, two at the end of theceH. Female:
Upper side : Forew'i ng differs froln thelnale in the outer black border being broader.
Hindwingwith.out central copper patch. Under side : Both wings like the male.
['957. Heliophortls sena (KoUar) : Wynter-Blyth, The BUllelilies of Indian Regiol1 : 303 .
Distribution : Th,e speci'es is very COITIlnOn fro III Chitral (Pakistan) to KU111aOn and
ascends toO about 9,000 feet in Hitn,alayas. Uttarakhand Ahnora, Chalno'li, D'ehra Dun~
Tehri, Pauri, Nainital, Rudra Prayag., Bageshwar, Uttarkashi and Pithoragarh districts.
114 Zoological SurvC'y (~l '"dia
1957. Heliopltorus (llldrocles contscan' (Moore) ,JVYlltcr-b~vth. rh<, Buttel:llh's of /lldi<1Il R,'gion .: 304.
Genel;al Ecology: It is one of the most beautiful of Indian butterflies, found along forest
paths, open fields and \vastelands and hUlnan habitations. A fast flier and sun loving butterfly.
It flies close Ito the ground for short distances. Both sexes are seen basking with their wings
fully open early in the morning hours . Rests in the thicket and bushes with wings closed.
It visits flower , datnp patches and droppings.
Diagnostic Characters Male Upper sidle : For,ewing brilliant rnetall ic green or blue
with the costa narrowly and increasingly, the apex very widely, and the outer margin decreasing
bJack. indwing with tbe disc and base green, the costa and outer margin broadly black, two
orange colour lunul'es at the anal angle. Under side : Both wings are yellow. with a disc.al
dark somewhat narrowlunular line; forewing has ·a disco-cellular hne, a sub-rnarginal dark
line, beyond which the wing is sometimes marked slightly with vermilion, the usual
conspicuous oval black spot inwards defined by a narrow white Jine at the anal angle. In the
hindwing the usual slnall indistinct dark dots scattered over the d·isc and base.
Female : Upper side : Dull smoky .. black. Forewing with small orange discal lunules.
Hindwingwith an orange marginal band. Under side:Both wings as in male.
1907. Pseudozizeel'ia maha (Kollar) : Bingham, The Fauna ,o f British India (Buuerflie,v) 2 : 355,
General Ecology: It is one of the COlnmonest butterflies of hHly regions. It can be seen
in open grassy patches along riv r beds, disturbed human habitations and borders of cultivated
fi,elds. It is a w ak fli ,er, does not fly very far and tlies ,close to the ground. It vis it low
growing flowers and settles on J ave with wing clos,ed basks in the Jnorning \vith 'A,jngs
1
partly open, but wh'en it feeds .or rests, it ke'eps its wings clos' d 'O ver its back~ .
Diagnostic Characters : H,ead, thorax and abdomen bluish above; antennae with while
rings.
Male: Upper side: Both wings s:ilvery bluish, w'ith the entir,e border narrow and fuscous.
Under side: For,ewing with a series of large black spots in the clell ,as well as a bar end 'c,eU,
Hindwing with a series of slnall black dots" that in area 6 lnidway between 5 and 7 and in
line. AU the dots circl,e dwith whitish.
Felnate : Upper side: Both wings dark brown to fuscous with basal blue scaling. Under
side: With darker markings.
The species shows weU-tnarked seasonal variation, the specimens" wh:ich occur during
rains are much darker than those which occur during the dry season.
Distribution : The species occurs throughout India, asclending to about 9'1000 feet in
Himalayas. Also occurs in Pakistan, Bangladesh, Nepal and Myanrnar. Uttarakhand : Dehra
Dun, Haridwar, Pauri, Tehri, Bageshwar, Uttarkashi and Udhaln Singh Nagar. districts.
1\6 Zoological SUrl'ey of India
1957. Rapala varrma (Hewitson) ,: Wynter~ Blyth, Tlte ButleJilies of India" Region : 361.
Genetal Ecology It is a beautiful and robust little butterfly, somewhat reselnbling the
Skippers in its Inanner of flight, which i. straight and very fast. It lives in thick and hilly
areas but som,etimes found on the plains. It sits with its wings closed and loves visiting
flowers. Male also visits dalnp patches,
Diagnostic Characters: Male: Upper side: Both wings dun shining dark indigo-blue or
brilliant rich deep ultra marine-blue. Forewing with costa narrowly, the apex widely and
decreas' ngly to the anal angle, where it ends in a point, black. Hindwing with costa and
outer margin narrowly black, the abdominal margin pale fuscous, the anal lobe bla,ck, inw,ardly
with a patch of dull ochraceous ferruginous scales. Under side : Both wings dull slaty-
brown, with deep black markings; forewing discal band white edged on both sides and
usually broad. Hindwing discal band generally curved and parallel to termen; bar end cell
usuaUy close to discal band.
Female : Upper side : Both wings as in male, but the blue not so brilliant. Under side
: Both wings pale ochraceous-brown, all the markings dark brown. Hindwing with the lower
portion of discal band fined in with ferruginous; a fine anticiliary white line, obsolete
anteriorly.
Wingspan : 28 .. 32 mm.
Distribution: The species is widely distributed in India, both at low el,evalions in the hills
and in the plains and ascends to about 7,000 feet in Himalayas, Also occurs in Sri Lanka
and Myanlnar. Uttarakhand : Dehra Dun and Nainital districts.
KUMAR: Handbook on Common Butterflies of Uuarakhand 117
General Ecology ': It is an extremely ,c ommon butterfly. It can be found in the gardens,
shrubs, hedges etc. and more nearer streams. The species is noticeably not fond of s'etd ing
on damp patches. The food plant of this species is Pril1ceia utilis of falnily Rosaceae.
9. Family HESPERIIDAE
S .ippers
Members of this falnily are popularly known as "Skippers" They are rnostly slnall in
size and very active in habits. They are always dark brown to black in colour and spotted
or banded with white or yellow, and the rnarkings often being se'l nj ... transparent' very few are
brightly coloured.
The bodi,es of these buttertl ies are very robust and strong. In both the sexes, the legs are
prefe,ct and functional for walking. The legs ar,e covered with vlery dense., long and thin
scales; antenna'e are placed wide apart, tapering to a point at the tip or with a curved hook.
The head is wider than the thorax and eyes are Jargle and prominent.
The flight is usually v,ery rapid, They are fond of basking in sunshine and visit tlow,ers,
damp patches; some feed on bird"s droppings. The eggs are dome shaped, smooth or ridged
longitudinally. The smooth and cylindrical caterpillars spend their life in cells made of
leaves. They feed on dicotyledon, grasses, palms, bamboos and ginger etc.
The family Hesperiidae is the third largest family of the bUUerflies in the world. In India,
about 320 species are known t'O occur, of whi,ch only 63 are re,corded from Western Himalaya
and 42 from Uttarakhand. The present account deals with only four species.
1897. CelaenorrhiJlus leucocera (Ko]Jar) : Elwes & Edward, Trans.R. ent. Lond.. 14~ pI. 4 : 116.
General Ecology: As the butterflies of this group keep their wings spread flat, they are
caned HFI,a ts"The Common Spotted Flat a is butterfly of shaded areas, ev,ergr,een and
mixed deciduous for,ests, especiaHy in low land country. It rests with wings widely open on
the under sides of the leaves, It onen visits Howlers and occ,asionaHy wet patches. Its flight
is rapid but seldom sustained for long distance.
Diagno 'tic Characters : Male and
female : Upper side : Wings dark brown.
Forewing with a white discal band formed
of 5 semi-transparent spots and 5 apical
spots" which are joined to each other. The
wings margin ,chequered with bla,ck and
white. Hindwing with a few obscure non
transparent orange o,c hra,c eous spots.
pper side Male
KUMAR: Handbook on Common Buuelllies of Ullar(lkhal1d 119
Female sirnHar to male but two large discal bands disconnected in th forew'ing. The tnale
has shaft plain white antennae where as the female has brown antennae with white cluh,
Under side : Markings as above but pale with an additional yeHowish brown small round
spot in the celL
Wingspan : 45 .. 55 mm.
Distribution : Widely distribut,ed species in the plains and asc,ends to about 8.,000 feet in
Himalayas. Uttarakhand : Ahnora, Nainital and Bageshwar districts.
1868. Pterygaspidea ransonneul Felder, Verh. Zool. Bot. Ges. Wein.. : 284 .
1987. Caprolla ralJsonnelt (Felder): lwes & Edward, Trans. R. ell!. Lond., t 4, pl. 4 : 150-151.
General Ecology : Acomlnon butterfly with angular wings, seen in the lowland forests.
It is a fast flier, flight always ,clos,e to the ground and does not sustain for long distance. It
visits flowers and dalnp pat,ches and often settles on the top of leaves . It can be 'caught while
basking in tbe sun.
Diagnostic Characters: Male and female Upper side : Wings dark brown. Forewing
with central and 'marginal golden .. brown areas having seJni~transparent, large and small
white spots in the disc and a group of small elongate spots in the apex. Hindwing dark
brown with central yellow-brown area divided by dark veins in the wet season fonn,. Th,e dry
season forms are much paler with extensive yellow-brown tnarkings.
Under side : Hindwing white with basal, discal and marginal rows of dark spots,.
DIstribution: The species ranges from peninsular Ind'a to Madhya Pradesh and Hilnachal
Pradesh to Assam. Also occurs in Bangladesh, Myanmar, and Sri Lanka. Uttarakhand :
Dehra Dun, Haridwar and Pauri distri,cts.
120 Zoologic,cll Survey of Indi,a
1877. Padl'ona gola Moore, Proc.. zool. Soc. Lond." : pI, 58, fig. 9,
1949. ,Oriens gala (Moore) : Evans, A Cata/~gue of Hesperiidae from Europe. Asia and Auslralia in
Brilish Museum (Nat.. His!.) : 372.
"
General Ecology His generally seen in the forest areas of heavy rainfall; fond of
visiting flowers. lttl ies closer to ground at low ,e levations r,epidly, but never to a great
d istance.
Diagnostic Characters : .Male and female : Upper side : Wings dark brown with tawny
yellow markings. Forewing 'with low,er part of straight tawny yellow discal band c,entral
(spot in I b is in the middle of I b, spot end cell co-joined to spot in 2) ~ con ..joined ,apical
spots 6 to 8; spot ,end cell, Inay be one above it. Hindwing with central discal band; no spot
in cell. The discal bands on the upper side are not divided by black veins.
Under side : Hindwing usually orange with prominent black spot.
Distribution : The species is common in South India, Kumaon to Sikkim ,and Assam
and ascends to about 6,500 feet in Himalayas. Also occurs in Sri Lanka and Myanmar.
Uttarakhand Dehra Dun, Haridwar, Pithorag.arh, Champawat, Nainital and Udham Singh
Nag.ar districts .
KUMAR: Handbook on Common ButleJ/lies of UUlIJ'akhand 12\
J949, Udaspes la/us (Cr.amer), Evans: A Catalogue 0/ Hesperiidae .li'om Europe. Asia and AUJlralia in
BrUisl, Museum (Nat. Hist,J : 292.
General Ecology: It is quite cOlnmon in open areas .of forests as w'ell as on tbe plains
more abundant in neighbourhood of streams where it s ttles on stones. Its flight is quick,
short and close to the ground. It can be seen basking e,a dy in th,e morning with hind wings
funy expanded and forewings partly opened. Like most of the Skippers, it is a nectar-lover.
It visits flowers of Vincoroses and Lantana, its favQuri ,te nectar sources. It also visits cow
dung rarely_
REFERENCES
Arora, O.S. & MandaI, D.K., 1977. On Scientific results of Swiss Zoological Expedition to
India. Part I. Butterflies of Oarhwal District. New/. zoo/. Surv. India, 3(4) : 229-234.
Baindur, O. 1993. The butterflies of Nanda Devi in Scientific and Ecological Expedition
Nanda Devi, 2 nd May to 22 nd July, 1993 : 35-43. Army H.Q., New Delhi.
Carter, D. J. 1992. Handbook- Butterflies and Moths: 304. Darling Kindersley Ltd. London
Chaturvedi, N. 1981. Some entomological notes from a visit to the Valley of Flowers. J.
Bombay nat. Hist. Soc., 78(2) : 402-405.
D' Abrera, B. 1982-1985. Butterflies of Oriental Region, 3 vols. (Hill House, Melbourne).
Doherty, W.1886. A list of butterflies in Kumaon. J. Asiat. Soc. Beng., 55(2) : No.3: 103-140.
Evans, W.H. 1932. The Identification of Indian Butterflies, 2 nd Revised edition, 454 pp., 32
pIs. Bombay nat. Hist. Soc., Bombay.
Hannyngton, F. 1910-11. The butterflies of Kumaon. J. BOlnbay nat. Hist. Soc., 20( 1) : 130-
142; 20(2) : 361-372; 20(3) : 871-872.
Haribal, M. 1988. Butterflies of Sikkim: Himalyas and their Natural HistolY. J. BOlnbay nat.
Hist. Soc., 85 : 271-280.
Haribal, M. 1992. The Butterflies of Sikkim : Himalaya and their Natural His/oIY, Sikkiln
Nature Conservation Faundation,Gangtok, 217 pp+60 pIs.
Kollar, V. 1844. Lepidoptera in C.F. Von Hugel "Kaschlnir und das Reich der Siek, Stuttgart,
4(2).
Kumar, Parmod & Gupta, S.K. 2004. Lepidoptera : Rhopalocera. In : Some selected fauna
ofGovind Pasu Vihar, Conservation Areas Series, 18 : 9-20. Zoo/. Surv. India, Kolkata.
Larsen Torben, B. 1987-88. The Butterflies of the Nilgiri Mountains of southern India
(Lepidoptera: Rhopalocera). J. Bombay nat. Hist. Soc., 84(1) : 26-54; 84(2) : 291-
316; 84(3) : 560-584; 85(1) : 26-43.
Mandai, O.K. 1984. Account of the Indian Tons Valley Expedition 1972. With an annotated
list of species and Redescription of Colias electo Menetries (Order : Lepidoptera)
from the Indian Palaearctic Region. Rec. zoo I. Surv. India, Occ. Pap. No., 57 : 1-31.
Mani, M.S. and Singh, S. 1962. Entomological- Survey of Himalayas. xxvi. J. Bombay nat.
Hist. Soc. 59( 1) : 77-99.
Mani, M.S. 1982. Butterflies of Himalayas. Oxford and IBH, New Delhi.
Marshall, G.F.L.and Niceville, L.de. 1883. The Butterflies of India, Burma and Ceylon, 1 :
1-321, XVII pIs; 1886, 2: IV + 332 pp., pIs. Xviii-xxiv; 3: 1-503, pIs. Xxv-Xxix
(Taylor & Francis London).
Moore F.( 1890-1907). Lepidoptera Indica. Vols. I-VII Lovell and Reeve London.
Peile H.D. 1937. A guide to collection of Butterflies in India. Staples Press London.
Shields, O. 1984. Oriental Lycaenidae, Riodinidae and Hesperidae from the Central Nepal
Himalaya. J. Bombay nat. Hist. Soc., 81(3) : 546-550.
Singh, Arun Pratap & Bhandari, R.S. 2003. Butterflies diversity in tropical moist deciduous
Sal forests of Dehra Dun Valley. Indian Forester, October: 1257-1268.
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1951 Au Garhwal Bull Mens. Soc. Linn., Lyon, 21 (3) : 181-184.
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124 Zoological Survey of India
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XV+506 pp., 2 pIs. 1 map.(Published By Taylor and Francis, London).
Varshney, R.K. 1977. Index Rhopalocera Indica: An idea of local list of butterflies from
India and neighbouring Countries, Rec. zool. Surv. India, 73 : 159-178.
Varshney, R.K. 1979-90. Revised nomenclature for taxa in Wynter-Blyth book on the
Bu"tterflies of the Indian Region. J. Bombay nat. Hist. Soc., 76( 1) : 33-40; 82(2) : 309-
321 & 87(1) : 53-61.
Varshney, R.K. 1993, Index Rhopalocera indica Pt. III. Genera of Butterflies from India and
neighbouring countries. Oriental Ins., 27 : 347-372.
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neighbouring countries. Oriental Ins., 28 : 151- 198.
Varshney, R.K. 1997. Index Rhopalocera Indica Pt. III. Genera of Butterflies from India and
neighbouring Countries. Oriental Ins., 31 : 83- 138.
Wiliams C.B.1930 Migration of Butterflies. : 296-301. Oliver and Boyd, Edinburgh, U.K.
Bombay nat. Hist. Soc., Bombay; xx + 523 pp., 72 pIs.
Wyanter-Blyth, M.A. 1957. Butterflies of Indian Region, The Bombay nat. Hist. Soc. Bombay,
XX +523pp., 72 pis.
Table 1. Distribution of Butterfly species dealt, in the State of Uttarakhand.
...,
.-.c .-
ell
~ .c ~
.c
~ ell
= """
~ """
~ ~ =
-e .--;...,
~
.;;
= """~
~ """
Q..
1:1}
~ ~
ell
~
~
.c
~
.-"""
"0 Q..
~ ~ 0
e e """
·c
.-=
~ "0 1:1}
0t:
.a """ ...,
...,""" e"""
0
...,"""
.c
0 ~ ~
.c ell
~
-<
~ ~
~ = .c "0
=
~
.c ~
ell
~ .c "0 ~
=
~
Q :t: ~ ;J u z Ci: u ~z
3 4 "5 6 7 8 9 10 11 12 13
No.
Family PAPILIONIDAE
1. Pachliopta aristolochiae + + + + +
(Fabricius)
-
~.
Q
23. Pieris brassicae nepalensis + + + + + + + + +
Doubleday
Family DANAIDAE
36. Danaus chrysippus Linnaeus + + + + + + + + + + + +
37. D. genutia genutia (Cramer) + + + + + + + + + + + + +
38. Parantica sUa (Kollar) + + + + +
39. Trrumala limniace leopardus + + + + +
(Butler)
No.
41. Parantica aglea melanoides + + + + + - .. + +
Moore
Family ERYCINIDAE
68. Libythea myrrha Godart + + + + + +
69. L. leptia Moore + + + + + + ...
81. N arne of the species 1 2 3 4 5 6 7 8 9 10 11 12 13
No.
c
70. Dodona durga (Kollar) + + + + + + +
71. D. eugenes Bates - +
Family NYMPHALIDAE
72. Athyma opalina (Kollar) + + + + + + +
73. Neptis hylas astola Moore + + + + + + + +
74. N. mahendra Moore + + + + + + + +
75. N. yerburyi yerburyi Butler + + + + + + +
76. Cyrestis thyodamas Boisduval + + + + + + + + + +
77. Hypolimnas bolina (Linnaeus) + + + + + + +
78. H. misippus (Linnaeus) + + + +
79. Kallima inachus Boisduval + + + +
80. Junonia hierta hierta (Fabricius) + + + + + + + +
81. Junonia orithya (Linnaeus) + + + + + + + + +
82. Junonia lemonias (Linnaeus) + + + + + + + + + + +
Family LYCAENIDAE
99. Castalius rosimon (Fabricius) 4- + + + + +
Family HESPERIIDAE
107. Celaenorrhinus leucocera - + + +
(Kollar) -
108. Caprona ransonnetti (Felder) + + +
109. Oriens gola (Moore) + + + + + +
110. Udaspes folus (Cramer) - + +
Note: Distribution of the species in Uttarakhand is based on the collection actually available with the author
or reported by previous workers.
KUMAR: Handbook on Common Butterflies of Uttaraklland 133
INDEX