Silurian: Persculptus Biozone Strata (Together With Higher Beds) Disappear

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Silurian

M . G. B A S S E T T , B. J . B L U C K , R. C A V E , C. H . H O L L A N D & J . D. L A W S O N

The history of establishment of the Silurian System (and hence of the The end of the Ashgill coincided closely with the collapse of the late
Silurian Period) and its principal divisions has been reviewed by Cocks et al. Ashgill ice-cap, and the basinal rocks of the persculptus Biozone reflect a
(1971) and Holland (1984, 1989a). The System itself, three of the four Series sharp change in the depositional environment. The southern British Isles
into which it is divided, and the Stages within these three, are all based upon basins had a form which continued to the end of the Silurian, and thus the
localities in Britain. The names of all the divisions shown in Fig. 1 have been Ashgill Epoch is revealed more as the antecedent of the Llandovery than a
ratified by the International Union of Geological Sciences. The two major descendant of the Caradoc.
divisions, Lower Silurian and Upper Silurian, remain informal, but they are Localized tectonic activity occurred widely during, the Llandovery; in
useful when degree of precision in correlation does not allow reference at the Pretannia to the south, along the eastern margin of the Welsh Basin, in the
level of Series (Holland 1989a). The boundary stratotypes are all described closure of Iapetus leaving a remnant linear basin, and in the rising of the
in Holland & Bassett (1989). Graptolite Biozones have proved especially southern margin (Cockburnland) of the Midland Valley Basin (Bluck 1983).
useful for international correlation in the Silurian and the sequence of RC
Biozones used in Britain is shown in Fig. 1. Correlation of the Biozones with
the global Standard Series and Stages is shown approximately, but is not Sla: Southern British Isles: latest Ashgill-early Llandovery
known with exactitude in all cases.
The time represented by the persculptus and acuminatus biozones signalled
Standard Standard graptolite the start of the Llandovery marine transgression, and for this reason was
Series Stage biozones
chosen for the Atlas. As the base of the Silurian is formally defined at the
base of the acuminatus Biozone (Cocks et al. 1984), part of the evidence used
P~DOLI
graptolite zones with the shelly sequences is still not accurately accomplished,
o . 84 Ludfordian bohemlct~ the timing of events is not precise and is open to amendment. For example,
O. Zelnt~xzrdlnertsls
tumeaeens it is impossible to correlate the shelly shelf sequences of Garth and Llando-
scanlcn~s
ni~asonl
very on the southeast flank of the Towy (Twyi) Anticline with the graptolitic
Zudensis basinal sequence some 6 km away on the northwest flank. The persculptus
z nassa
Zundorent
mudstones thus cannot be mapped into the shelly sequence of sandstones
<
e~Zesae and mudstones to afford any sort of physical link. Even where the Towy
Sheinwoodian Zinnarssont
or ,_ rlgldus Anticline plunges steeply northward at Abbeycwmhir (Roberts 1929) the
r[eeurtonena[s persculptus Biozone strata (together with higher beds) disappear northward
Z3 .mrehtsont
centri~ugl~ from the sequence on the western flank of the anticline before sandstones
...J erenu~ata containing a Hirnantia fauna appear around the closure.
o Telychlan grlestonlensls
erispus At this time the deeper water depositional basins of Wales, the Lake
09 -J
turrlcu~atus District and Tornquist Sea persisted on either flank of the Midland
sedg~teklt
convo~utus Platform of England. The shallow shelf-seas were narrow, but were widening
Aeronian ~eptotheca with the transgression, so that terrestrial detritus carried from the Platform
magru~
trfangu~atus in rapid run-off rivers were trapped in the marine shelf environment behind
eyphus
Rhuddanian aclnaces
the advancing shoreline.
atavus The basins and their slopes, suddenly starved of coarse detritus and
acumtnagus
concomitant high energy turbidity currents, accumulated fine mud from low
density turbidity currents and a rain of carbonaceous pelagic debris. Occa-
Fig. 1. Silurian Global Standard Stratigraphy and graptolite biozones. sional floods of coarse detritus invaded the generally anoxic basins through
restricted basin margin submarine channels. Small base-of-slope bodies of
Geodetically the position of the British Isles in the Silurian remained gravel and sand formed, e.g. along the southeastern margin of the Welsh
astride the 25~ latitude, a position it had reached in the later Ordovician Basin (Drew & Slater 1910; Kelling & Woollands 1969, pp. 262-264 and
(Briden et al. 1973); movement was confined to rotation. During this time possibly Lockley 1983). In this sector the margin was sharply defined along
the south magnetic pole wandered, in terms of present geography, from near or just east of the site of the Towy Anticline. North of Abbeycwmhir the
to southwest Africa, to the Horn of Africa in early Silurian times, then back position of the basin margin is less well constrained, for the early Llandovery
along the equator to west Africa/Brazil at the end of the Silurian. is not exposed in crucial areas; a line from there passing east of Bala and
CHH, RC Llandudno is an approximate position. East of this line shelf-sea sediments,
often immature, poorly sorted sands, muds and gravels, were deposited
somewhat patchily. The faunas are dominated by brachiopods but do not
LLANDOVERY provide precise indications of age; it seems that on the east side of the
Berwyns the marine invasion of the shelf reached Meifod early in the
The Llandovery Epoch was characterized by marine transgressions. Thus in Rhuddanian, but did not reach the Welshpool to Berriew line (the Severn
the early Llandovery the English Midland Platform, much expanded by the Valley) until late in the Rhuddanian. So, while there are respects in which
late Ordovician regression, was flanked to east and west by narrow shelf- the Severn Valley here is the structural extrapolation of the Towy Anticline,
seas, whereas by the end of the epoch it was almost completely drowned by a it did not function palaeogeographically as it did near Garth, Llandovery
significant shallow sea. On the opposite side of the remnant Iapetus Ocean, and probably Haverfordwest where it separates shelly, shallow marine shelf
marine transgressions northward from the Mayo Midland Valley Basin facies from graptolitic basinal facies. Indeed in acuminatus Biozone times
encroached upon Grampian terrane. The transgressions were partly, maybe there was land at Welshpool and Berriew, while at Garth, Llandovery and
largely, eustatic and the agencies effecting them climatic and tectonic, with Haverfordwest there was a deepening marine condition, a condition which
both operating together in the early part of the epoch. Volcanism was minor had been continuous from at least mid Ashgill times. Rapid sedimentation
except along the northern edge of Pretannia from the Mendips to southern of transverse downwarps in the Platform edge was under way in places, e.g.
Ireland (Cope & Bassett 1987) and at the source of the bentonites of the Llandovery and Garth, so the actual acuminatus Zone shoreline was located
Southern Uplands Birkhill Shale. south of Haverfordwest, east of Llandovery, probably just west of Builth
The climatic effects towards the end of the Ordovician were those and through Welshpool.
attendant upon the growth of a polar ice cap over what is now the Sahara. Two palaeogeographical enigmas relate to the northern part of this area.
Sea level fell perhaps by 100 m (Brenchley 1984) and rose again when the ice- One is the problem of the divide between graptolitic and shelly facies at the
cap melted towards the end of the Ordovician. The resulting marine trans- north end of the Towy Anticline and in the west Berwyns (Fig. 2). Along this
gression was a worldwide event and it continued well into the Llandovery line the pre-griestoniensis Llandovery sequence is absent or largely absent.
(~3'phus times); the later stages were probably due to a tectono-eustatic rise In part of the west Berwyns the griestoniensis Biozone rests upon the per-
of sea level, a process which probably sustained the later Llandovery sculptus Biozone and in the Abbeycwmhir area of the Towy Anticline the
transgressions, too. It is possible that these repeated tectono-eustatic rises former zone rests upon Ashgill strata, Hirnantian and probably older. The
stemmed from closure or partial closure of the Iapetus Ocean, or from a sediments immediately above the break were graptolitic muds,.not the
ridge-trench collision (Campbell 1984) which ended southeastward subduc- deposits of a transgressive shoreline. The break is thus a non-sequence by
tion. non-deposition or by sliding and slumping within a submarine environment.
9 1992 The Geological Society 37
SILURIAN

shelf. To the north, turbidite deposition was continuous from the Ordovtctan
into the Wenlock. Further north, at Balbriggan (Rickards et al. 1973) and
marginal to the lapetus Suture at Tomgraney, Co. Clare (Rickards & Archer
1969), basin-plain graptolitic muds were deposited at least as early as the
acuminatus Biozone. The Tomgraney exposure may well lie north of the
lapetus Suture and the deposits therefore may have been oceanic pelagic
muds now associated with the margin of Laurentia.
In northwest England anoxic graptolitic muds were accumulating in the
western Lake District comparable with, and probably in continuity with,
those of the Welsh Basin and southern Ireland9 In the eastern Lake District
Cross Fell and the Howgill Fells (Rickards 1978; Burgess & Holliday 1979,
and at Horton in Ribblesdale, Arthurton et al. 1988) black mudstones
have at their base very calcareous mudstone, shelly in places, or a hard lime-
stone representing probably the persculptus and acuminatus biozones. These
deposits are thin, of shallow marine origin; the hard limestone with little
clastic detritus (e.g. in the Howgill Fells) may have been a hardground which
formed on the northern peninsular area of the Midland Platform at the end
of the glacioeustatic marine low, The condensed sequence rests in places on
sandy Hirnantian deposits and in the Cross Fell Inlier (Burgess & Holliday
1979) it has yielded Hirnantia sagittifera (Ingham & Wright 1972); this is
probably compatible with the supposed persculptus age and reveals that
there was little, if any, prior non-deposition in places. At Austwick, however
(Arthurton et al. 1988), a non-sequence does separate Hirnantian strata
from late Aeronian or Telychian rocks, so that attenuation or non-deposition
did occur in places.
The late Hirnantian/early Rhuddanian saw the growth of a major N-S
fault, the Brathay Fault (Rickards 1978) on the east side of the Lake District
and the limestone facies was developed only to the east of it. Furthermore
the greatest attenuation of the early Llandovery sequence occurs close to the
fault along its footwall. This may be a horst (Rickards 1978) but footwall
uplift on an extensional down-to-basin fault (cf. Jackson & McKenzie 1983)
is another possibility. If the eastern fault of the horst be discounted then the
submarine slope down towards the Howgill Fells 'basin' had to be steep
enough to promote slumping (R. B. Rickards pers. comm.). The Brathay
Fault, however, may have been only one of a set of approximately N-S
Llandovery extensional faults; for instance large gaps in the Llandovery
sequence commonly lie over or just east of approximately N-S structures
Fig. 2. Structural elements influential in Llandovery deposition in Wales and such as the Brathay Fault, the Western Berwyns Lineament, the Severn
Welsh borderland (partly after Temple 1988). Valley, the Church Stretton Fault (Presteigne) and the north end of the
Towy Anticline. Whereas the Taconic folding of the Shelve-Builth Basin
Furthermore the Llandovery sequence is more complete not only to the followed the cessation of southward directed subduction, these extensional
west, in the Welsh Basin, butalso to the east. In the east Berwyns the early elements may be related to collision and cessation of northwestward directed
Llandovery sediments are shelly; NW of Builth (e.g. Maesgwynne and subduction during the Llandovery (Murphy & Hutton 1986).
Trecoed; Baker & Hughes 1979) shelly muds and sandstones are considered RC
to extend down to the erispus Biozone; near Llanddewi Ystradenni pale
coloured shelly and graptolitic mudstones are reportedly as low 'Middle Slb, S2a: Southern British Isles: Llandovery (Aeronian)
Llandovery' (N. Kirk, MS). This non-sequence probably represents parts of
the early and mid-Llandovery submarine upper slope and distal shelf-edge For most of the Aeronian the palaeogeography changed little. Turbidites
(Fig. 3). dominated in parts of the basins, e.g. the Derwenlas Formation in Wales,
The second enigma is the depth index of the early Llandovery faunas east and 'distal mud' in the Stockdale Shales/Skelgill beds in the Lake District
of the Berwyns around Welshpool. A progressive marine invasion of and Howgill Fells.
subdued terrain suggests shallow water, but the Meifodia-Clorinda-Striek- The second transgressive pulse is attributable probably to a tectonic event
landia fauna of the Powis Castle Conglomerate is indicative of relatively and it commenced in convolutus times (Aeronian) but characterized more
deep water. The apparent contradiction points to a steep shore-face, con- particularly sedgwickii times. Marine conditions spread rapidly across the
trolled possibly by extensional faulting along the line of the Severn Valley subdued Midland Platform; its shoreline ran from south of the Malvern
(Fig. 3). Hills up their west side to fringe the Precambrian rocks of Shropshire. There
From south Wales the shoreline extended across St George's Channel the Longmynd formed a southwestward extended peninsula with sub-littoral
skirting the south coast of Eire. Llandovery turbidites, mud-sand rhyth- sands on its eastern side and an exposed shoreline of cliffs on its west
mites, commonly with base-missing Bouma sequences, occur in southeast (Bridges 1975).
Ireland (Briick et al. 1979); and in the extreme south base-of-slope con- North of Shropshire the position of the shoreline is ill-defined. Geophysi-
glomerates (Ballyhest Member) occur, comparable with those on the south- cal evidence (Bott 1967) suggests that a salient of the Midland Platform
east margin of the Welsh Basin. A precise age for these is not known (they projected into northern England, but the shoreline must have lain east or
may be mid Llandovery rather than early) but they coarsen and thicken southeast of Ribblesdale and thence through eastern England west of
southward suggestive of a steep basin-margin slope and a narrow marine boreholes near the east coast which penetrated Llandovery turbidites,

NW SE
Eastern limit of earlyLMndoverygraptoliticmud Western limit of early Llandoverymudsand& gravel
BALA W E S T E R NBERWYN HILLS MEIFOD WELSHPOOL
I I I

. ..--"....
9 "" B. . . . f ,fi~tonienais L A T E L L A N D O V E R Y

.''~-~Ga~l~ mud~ 'c...... andst0he'.~aio w ~ ~ , ~ I I I I


.'"~irnant Lst.- ......... /' I~aieAlshJIIre r!se,4n ! I I
, ASHGILL-

/ T T I ~ O ' ~ K i l o m e t r e s 1p~~ ;' T

Fig. 3. Non-speculative model of stratigraphical events in the southern Berwyn Hills. Vertical ruling represents non-sequences.
39
40 SILURIAN

West of the Midland Platform the shoreline bounded the north side of Ashgill substrate there. Since there appears to be no local shore or shallow
Pretannia (Cope & Bassett 1987) through South Wales to south Pembroke- facies in the area, except the bed of coarse cross-bedded sandstone within the
shire, having migrated a little southwards beyond its early Llandovery Henfryn Conglomerate, these activities and the non-sequence must have
position. This suggests that the margin of Pretannia was steep at the time or been submarine.
was tectonically positive. There was a very active volcanic centre in St Over the western Berwyn Hills (Figs 2 & 3), mud deposition resumed
George's Channel, off Skomer Island (Ziegler et al. 1969) extruding basalt during griestoniensis time in a similar rhythmic, dysaerobic and anoxic style;
and rhyolitic domes eastward and southward (Bridges 1976; Cope & Bassett to the east however, where mud deposition had been much more continuous,
1987; Walmsley & Bassett 1976). These also shed detritus southwards the conditions were mainly oxic and the griestoniensis (and earlier) mud-
creating off-shore barrier islands and shallow marine lagoons. Close by to stones are red and green as well as grey. In crenulata time such conditions
the north, must have lain the basin, yet the products of volcanism are not spread widely over Laurentia and northern Gondwana while along northern
obvious in the contemporary deposits there. Immediately afterwards the Pretannia the shoreface transgressed southwards once more, drowning the
Pretannia shoreline regressed northwards, for deposits of the turriculatus griestoniensis terrain of earlier Llandovery sediments and voicanics.
Biozone are absent in Pembrokeshire. This event coincides with the shedding In parts of east Wales the pale coloured late Llandovery muds supported a
of the first of the northward directed lobes of coarse, commonly feldspathic shelly benthos, e.g. near Welshpool, just west of the Severn--Towy Anticline
and often high-matrix turbidite sands into the mid-Wales basin, i.e. the lineament, and near Llandrindod Wells, just east of it. Although the age of
Aberystwyth Grits (Cave & Hains 1986). It was at this time, therefore, that these faunas may differ from place to place, the lateral passage from shelly to
Pretannia took over from the eastern shelf as main supplier of coarse barren mudstones is abrupt. These circumstances along the shelf margin
detritus to the basin probably in direct consequence of tectonic and volcanic suggest that local shoaling, probably produced tectonically, reduced marine
activity along its northern edge. depths to within a sharply defined base of viability for the epifaunas.
During the Aeronian the Caban Canyon of Kelling & Woollands (1969) In northern England during griestoniensis time grey marine muds were
had continued to entrain coarse extra-basinal high-matrix gravels basinward being deposited. The Brathay Fault showed little sign of activity and banded
from the east, past Rhayader, but the sedgwickii-turriculatus transgression muds, rather like those of the western Berwyn Hills, extended from the
isolated the canyon-head from its supply of detritus, and shut it down. western Lake District across to the Howgill Fells, Cross Fell and Ribbles-
Slumped mudstones overlain by coarse and pebbly sandstones (J. A. dale where distal turbiditic mud rhythmites were accumulating.
Zalasiewicz, MS) indicate a brief resumption and eastward shift of canyon Deposits in central Ireland (Balbriggan; Rickards et al. 1973) were very
activity on this eastern margin in iurriculatus times, but the main input to the similar to those of the Lake District and it is likely that one large marine
basin from the east was of high frequency, low density, mud-dominated 'basin' extended across southern Ireland into the Lake District and Wales.
turbidites produced possibly by linear spill-over of shallow marine storm The imprecisely dated turbidites of southeast Ireland (Briick 1972; Penney
surges from the shelf, e.g. Devil's Bridge and Cwmsymlog formations and 1980) continued to form a fan, proximal in the southeast, extending distally
Rhayader Pale Shales, of which there are green and red facies. During this over the basin-plain to the northwest. The westward directed currents
time the redox surface lay within these deposits, below the benthic boundary indicated west of the Leinster Granite could have come from the Irish Sea
layer, and the basin floor was colonized by a burrowing and grazing benthos land-mass, but equally they might have been bathymetric deflectants of a
leaving traces such as Nereites and Dictyodora. general northward flow, for no good evidence of the land-mass comes from
In turriculatus times northeastern Wales saw the deposition of non-shelly the Welsh Basin at this time.
oxic muds punctuated by brief anoxic algal/graptolitic blooms, while the RC
shelly mud/sands facies advanced eastward beyond the earlier shoreline.
Basinal muds, often graptolitic and anoxic, also characterized the western WENLOCK
Lake District (Rickards 1978) and central Ireland (Balbriggan; Rickards et
al. 1973). Presumably the marine transgression seen in Wales and the The marine transgression eastwards and southwards in the late Llandovery
Borderland occurred here too, but minor influxes of sand recorded in the led to a palaeogeographical situation in the southern part of the British Isles
western Lake District and at Balbriggan suggest the appearance of a more which comprised a basin to the west and an extensive shelf to the east, across
active or proximal source area. the Midland Platform. This pattern prevailed through Wenlock times. The
The Brathay Fault was not active at this time and muds with graptolites basin was, however, partially divided by an area of shallower water and
were deposited across it at least as far as Ribblesdale but these eastern perhaps an island, traditionally referred to as the Irish Sea Landmass. At the
sequences are thin, presumably platform mud with a shelly calcareous beginning of the Wenlock (not shown on the maps), localized deposition of
component in the early Telychian. carbonates in the Welsh Borderland led to the formation of the Woolhope
RC Limestone. The more widespread and prominent development of carbonates
across the Midland Platform is seen in the Much Wenlock Limestone
S2b: Southern British Isles: (Telyehian) Formation of the late Wenlock. Farther to the east, and with reference to the
whole of the Wenlock, data from boreholes indicate the presence of deeper
The last of the Llandovery marine transgressions occurred in griestoniensis water and a connection with the open sea. There is no evidence for a Iapetus
time. In the English Midlands and the Welsh Borderland the siliciclastic, Suture in the Wenlock rocks of the British Isles, though information may
mainly mud/sand deposition which had been established everywhere west of have been lost along a subsequent fault following approximately the same
the Malvern Line then extended over the Midland Platform (Ziegler et al. line, and possibly with significant sinistral displacement.
1968). The shelf-sea sediments, derived presumably from Pretannia to the Definitive and general works on the Wenlock rocks of the British Isles,
south, were distributed by storm surges. Water was mostly shallow, though which also lead to other references, are as follows: Bassett et al. (1975) and
in places below wave-base (Benton & Gray 1981). Near Tortworth, in the Bassett (1989) (for the type and standard Wenlock), Bassett (1974) and
northern Pretannia orogenic zone, two basalts were extruded, both probably Holland (1989b) (for England and Wales); Rickards (1978) (for Northern
in griestoniensis time. The earlier one flowed directly over Tremadoc rocks England); Holland (1981) (for Ireland); and Walton (1983) (for Scotland).
and supports red mudstones on and near its surface. Here, therefore, may CHH
have been an early griestoniensis land surface poised for inundation.
Towards the western edge of the Midland Platform, along the line of the S3a: Southern British Isles: Wenlock (Sheinwoodian)
Church Stretton Fault at Old Radnor and Presteigne, no deposits of
griestoniensis age are preserved and coarse quartzose sands were deposited The early Wenlock successions of the central Irish inliers are commonly of
on Longmyndian rocks in crenulata time. Apparently this tract remained poorly fossiliferous graptolitic shales and tubidites. Spores are present in the
positive, possibly land, up to crenulata time thus maintaining a barrier which Slieve Bloom Inlier and that of Slievenamon. Richly graptolitic mudstones
trapped detritus on the shelf to the east. are seen at Balbriggan. Directional data are limited.
To the west, the Welsh Basin remained little changed. It received further In northern England, data from the Austwick Formation of graded
major influxes of coarse detritus from Pretannia to the south, producing greyish sandstone turbidites, alternating with mudstones, suggest a north-
vertically stacked lobes of turbiditic sand within a SSW-NNE depositional westerly flow and proximity to an east-southeasterly shoreline. Farther to
system some 100 km long (Smith 1987). Conditions were often anoxic, so the north-west in the Howgill Fells, the Lake District, and the Cross Fell
that interturbiditic, hemipelagic muds were commonly graptolitic and carbo- inlier, is the distinctive Brathay Flags lithology of blue-grey laminated
naceous. Dysaerobic and oxic intervals also occurred, developed as bio- mudstones with calcareous nodules and bentonites. The Brathay Flags are
turbated mudstones, especially in the thinner bedded sequences on the east richly graptolitic and yield orientated orthoconic nautiloids. Rickards (1978)
side of the basin and lateral to the coarse sand lobes. has suggested that laminations running through the calcareous nodules
The eastern margin of the basin continued to suffer tectonically affecting indicate a reduction in thickness from the original mud of over 40 per cent.
bathymetric gradients (particularly at the north end of the Towy Anticline) Since the earlier sedimentological work by Cummins (1957), using flute
and disturbing sediments and sedimentation in griestoniensis and crenulata moulds well seen in the Central and North Welsh turbidites of the Denbigh
times and even into the Wenlock. On the eastern flank of the anticline, east Grits Group as directional indicators, it is necessary to recognize two
of Abbeycwmhir, mud was deposited directly upon an Ordovician substrate. troughs, the Denbigh Trough and the Montgomery Trough, which come
In places (e.g. Henfryn) there was a rudaceous base, largely of a mass mud- together in the north. They were separated by the Derwen Ridge. There is
flow nature, containing late-Ordovician debris. On the western flank of the some evidence of slumping on the western margin of the Montgomery
anticline the underlying deposits up to the crispus Biozone show slumping, trough. A thinner, more graptolitic sequence is seen near the North Welsh
but away from the anticline on both sides griestoniensis sediments were coast. A graptolitic facies of flaggy silty mudstones and shales is also present
deposited upon earlier Llandovery deposits: It would seem, therefore, the at the other side of the basin in the Builth district, where Elles did the
griestoniensis mass-flows can have been nourished only from contemporary original work on the graptolitic biostratigraphy of the Wenlock.
sediments high on the eastern flank of the anticline and from the Caradoc/ On the shelf to the east are the muddy, more or less calcareous rocks
14 CAMBRIAN

Massif or on its margins. The Cambrian of Nuneaton accumulated to the rocks encountered in boreholes in Lincolnshire (Cowie et al. 1972, table 2)
southwest of this Charnian Massif. Coastal Hartshill Formation sandstones are varied--phyllitic slates, feldspathic sandstones and greywackes--and a
(cycle l, thick), through to condensed shallow water, shelly Hyolithes Cambrian age has been suggested, but without adequate faunal control. It is
limestones (cycle 2, thin) and deeper water Mn-rich shales (cycle 3, thick) assumed here that this area was part of a marginal basin that existed
exhibit a trend reciprocal with that of Shropshire and close to that of the throughout the early Lower Palaeozoic and extended from northern England
Avalon succession, SE Newfoundland. The nearby Lickey Quartzites are into the Brabant area of Belgium where clastic rocks of Comley and St
lithologically similar but are of uncertain age. The former presence of David's age are known (Vanguestaine 1974, 1978). It is not known whether
limestones is suggested by clasts in the Permian Nechells Breccia (Brasier & the Precambrian of the Charnwood Forest area was emergent; there is some
Hewitt 1981). geophysical evidence to the northeast of Charnwood Forest for the presence
The Withycombe Farm borehole near Banbury, Oxfordshire, gives an of a post-Charnian, pre-Triassic succession that possibly represents part of
indication of conditions to the east of the Malverns. Volcanic basement of the local Cambrian (Maguire 1987). A positive area in northern Norfolk was
presumed late Precambrian age is there overlain by the Withycombe Forma- formerly postulated on the presence of supposedly Precambrian volcanic
tion, a relatively thick sequence of sandstones bearing trace fossils (cf. cycle rocks encountered in the North Creak Borehole, but these are now thought
l) passing up into a thick succession of offshore shales with Aldanella sp., to be part of the east Midlands Caledonian Belt (Pharaoh et al. 1987).
Watsonella sp., Platysolenites antiquissimus, Orthotheca cylindrica and Lon- AWAR
tova-type acritarchs (Rushton & Molyneux 1990). This invites compari-
son with the Chapel Island Formation (cycle 1) or lower Bonavista Forma-
tion (cycle 2) of southeast Newfoundland, both of attleborensis age. 4~2a: Southern British Isles: Merioneth
MDB
In Scandinavia the Merioneth Series has been divided into some eight
E~lb: Southern British Isles: St David's biozones and thirty subzones, and this scheme can be applied in areas, such
as England and Wales, where the olenid trilobite biofacies is developed. The
The St David's Series is equivalent to the Middle Cambrian of European interval depicted on the map is the Olenus Biozone and more particularly its
usage, but not necessarily equivalent to that of other regions such as upper part (O. cataractes Subzone).
Australia, China or North America. The Series is subdivided according to Deposits formed in England and Wales during the Merioneth Epoch
the Swedish zonal scheme, with a lower, a middle and an upper part. In the reflect generally stable tectonic conditions. After a widespread non-sequence
Welsh Basin the lower part is represented mainly by thick turbiditic in the late St David's successions (possibly caused by a eustatic regression), a
sandstone successions; in Shropshire by thinner glauconitic sandstones and major new sedimentary cycle commenced, apparently before the end of the
mudstones, and in the English Midlands by mudstones. Biostratigraphical St David's Epoch, in both the Welsh Basin and the English Midlands
control is generally poor and correlation insecure (Cowie et al. 1972; (Rushton 1978; Allen et al. 1981).
Rushton 1974). The Welsh Basin, though deeper at the beginning of the Merioneth Epoch,
Faunal control is better in the middle part of the Series, where the was partly filled towards the end of Olenus Biozone times with alterations of
following biozones are used: shales and contourite(?) sandstones (upper Maentwrog Formation), and
those beds are overlain by thick shallow-water sandstones (Ffestiniog Flags
Ptychagnostus punctuosus Biozone,
Formation, >650m, Allen et al. 1981). There is some evidence in the
Hypagnostus parvifrons Biozone,
Harlech Dome, St Tudwars Peninsula, and in the St David's area, that the
Tomagnostus fissus Biozone,
sediments were deposited by currents flowing from the south, as in the St
Ptychagnostus gibbus Biozone.
David's Series. To the north of the Harlech Dome, however, the Merioneth
The gibbus Biozone appears to represent a widespread transgression, Series is comparatively attenuated and includes shallow-water quartzitic
possibly related to a eustatic sea-level rise (Rowell et al. 1982, p 176), but sandstones, with trace fossils (Cruziana facies) that appear to have been
that zone is not easily recognized in Britain. The overlyingfissus Biozone is derived from the north, though the indices of current orientation give a wide
more widely recognized and this interval, for which correlation is more range of directions (Crimes 1970).
secure, is represented on the map. Muddy sediments accumulated over the site of the Midlands Platform and
In the Shropshire area glauconitic sandstones and calcareous beds were it is assumed here that these (Outwoods Formation of the Nuneaton Inlier,
deposited in shallow water, and Cobbold (1927) demonstrated a local with known subsurface extension to the south and west) transgressed the
unconformity indicative of uplift along the Church Stretton fault zone at Precambrian Charnwood Forest massif. The fauna of the Outwoods Forma-
about this time. The absence of the St David's Series in the Malvern and tion includes olenid trilobites that tolerated oxygen-poor environments,
Lilleshall areas (Rushton et al. 1988) implies similar uplift, but with less together with a number of other trilobite genera of wide palaeogeographical
certainty. distribution that inhabited outer-shelf regions (Rushton 1983).
The Midlands Platform carries a thin sequence of variegated mudstones In the Welsh Borderland the Olenus Biozone is unrepresented, the earliest
deposited below wave-base (Abbey Shales, three biozones represented, representatives of the Merioneth Series being the Orusia Shales (Parabolina
< 40 m thick in total). The fauna is dominated by agnostid trilobites and spinulosa Biozone) resting unconformably on Comley and St David's rocks.
small brachiopods. Similar but thicker successions (c. 200 m total?) occur However, at Malvern there is a thin sequence of Cyclotron-bearing shales
in southwest Wales and also in North Wales, though there the mudstone that are thought to be of Olenus Biozone age. Uplift along the Church
sequence is underlain by turbiditic sandstones offissus age (the Gamlan and Stretton and Malvern lineaments is suggested by northeastwardly directed
Caered formations) that show evidence of derivation from a southerly slumps or currents in the Moor Wood Flags Formation of the Nuneaton
source (Crimes 1970). This implies a positive area to supply sediment to area (Taylor & Rushton 1972, p. 22). The derivation ofmicaceous detritus in
north Wales whilst by-passing the St David's area in the southwest. The the Llangynog area, south Wales, is believed to be from a metamorphic
presence of bentonite beds indicates contemporaneous volcanic activity, basement (Pretannia) to the south (Cope & Bassett 1987).
though the location of the activity is unknown. The later Merioneth Epoch, from Parabolina spinulosa Biozone to Peltura
There is no evidence for the St David's Series north of the Harlech Dome scarabaeoides Biozone times, was dominated by deposition of black mud-
and St Tudwal's Peninsula, and it is assumed that the Irish Sea Horst was a stones in disaerobic environments (Doigeilau, Bentleyford, Monks Park and
positive area; nevertheless, no contemporaneous sediments are known to White-Leaved-Oak formations). These pyritic, carbonaceous and uranifer-
have been derived thence. ous muds contain little or no coarse clastic material and are considered to
The non-agnostid trilobites most typical of this epoch, Paradoxides and represent a period of slow deposition; the proved olenid zones differ from
blind benthic forms (Hartshillia, Holocephalina) characterize outer-shelf locality to locality (Thomas et al..1984, p. 15), and the extent of possible
environments in Wales and elsewhere in Avalonia (southeast Newfound- breaks in deposition, comparable with those inferred in the contempor-
land, New Brunswick) and in Baltica and the margins of Gondwana aneous rocks in Scandinavia, is not certain. The black mudstones in the
(Conway Morris & Rushton 1988). Croft Borehole, Lilleshall (northern Shropshire), are unique in having many
After fissus Biozone times, mudstone deposition prevailed (except locally intercalations of current-bedded siltstone and sandstone, presumably of
in the Comley area of Shropshire). There is a widespread hiatus after shallow-water origin (Rushton et al. 1988). The source of the silt-grade
punctuosus Biozone times and before the Merioneth Epoch, the mudstone material is not known but is thought to originate from a local uplift along
succession in the Nuneaton district being the best constrained (Taylor & the Church Stretton Fault complex.
Rushton 1972). The sedimentary rocks of the southeast Irish marginal basin have poor
In southeast Ireland the marginal basin of Leinster has yielded some biostratigraphical control but include beds referable to the Merioneth Series
evidence of the St David's Series (Smith 1977, 1981). In Co Wexford (the (Ribband Group; Smith 1977, 1981). They are argillites and arenites that
Cahore and Ribband Groups) and Co Wicklow (the Bray Group), the appear to have been deposited in deep water. The mudstones and fine-
sediments were deposited in deep water (Crimes & Crossley 1968). These grained sandstones of the Askingarren Formation, interpreted as underlying
rocks yield trace-fossils, especially Oldhamia, in abundance (Dhonau & correlatives of the Ribband Group (Crimes & Crossley 1968, pp. 191,207),
Holland 1974), but acritarchs, in part of St David's age, are the only body- are considered as possibly late Cambrian in age. Flute casts indicate
fossils known; the best preserved are from the Booley Bay Formation derivation from a southerly quarter. There is no evidence for the continued
(Ribband Group) at Hook Head on the south coast (Smith 1981). Sediment existence, into the Merioneth Epoch, of the marginal arc thought to have
was derived from both the southeast (Irish Sea Horst) and the northwest, existed to the northwest of the southeast Irish marginal basin in St David's
where a positive area, possibly a marginal arc, is thought to have provided a times.
source area. There is no biostratigraphical evidence for the Merioneth Series in the
Less is known of the eastern Caledonide belt to the northeast of the eastern English Caledonides (see the St David's Series, above).
Midland Platform and facing the Tornquist Sea. The clastic sedimentary AWAR
42 SILURIAN

generally referred to collectively as the Wenlock Shales. They are sparingly were associated crinoids, brachiopods, ostracodes~ and gastropods. The
graptolitic but yield rich shelly faunas of brachiopods, trilobites, corals, etc. bioherms have an average diameter of 12 m, with maximum thickness of
Thin bentonites are commonly present, as, for example, in the Wenlock some 45 m. There are various inter-reef and off-reef facies of argillaceous
Shales of the Ludlow anticline. The andesites and tufts of the Mendips were and crinoidal limestones. The Wenlock Limestone at Dudley appears to
regarded by Ziegler et al. (1968) as of Wenlock age. The southern shoreline have been deposited earlier than elsewhere, i.e. at the time of the preceding
is shown as running north of Freshwater West and Freshwater East in Cvrtograptus tundgreni Biozone,
Pembrokeshire (Dyfed), as the basal part of the Wenlock is missing here Again a connection to open sea is shown in the east, Siveter & Turner
(Bassett 1974). (1982), described a microvertebrate assemblage from the Wenlock rocks of
What amounts to a very substantial knowlege of the pre-Permian floor the Tortworth inlier, which has Baltic and Siberian affinities. Another hint of
was charted by Wills (1978). A number of boreholes for coal or water have such wider connection from west to east is the occurrence of the Bohemian
penetrated Silurian rocks. The extent to which this southeastern marine area tabulate coral Stelliporella in the Homerian rocks of the Dingle Peninsula.
merged with the Welsh Basin and shelf remains uncertain. Exposures and CHH
boreholes in Belgium suggest a marine connection in this direction,
CHH LUDLOW

S3b: Southern British Isles: Wenlock (Homerian) After an initial rise in sea level the dominant theme in Ludlow times was one
of marine regression, leading to the restricted marine and fluviatile con-
Through the late Wenlock, the palaeogeographical framework remains ditions of the Pfidoli Epoch. The Welsh and Lake District basin became
similar, but there are some additional sources of evidence and there are some silted up and shallow. Terrestrial red beds were developed well before the
important changes in the facies pattern. end of the epoch in Pembrokeshire and southwest Ireland. In Scotland there
Conspicuous among the central Irish inliers is the relatively large Slieve is some palaeontological evidence for Ludlow rocks (Selden & White 1983).
Phelim-Devilsbit Mountain district (Doran 1974). Graptolites indicative of Radiometric dates on lavas of the Lower Old Red Sandstone (Thirlwall
the Monograptus ludensis Biozone are actually confined to a small area in the 1988) support the long-held view that marine influenced alluvial deposition
northwestern part of the inlier, but the repetitive sequence of greywackes, was taking place there in Ludlow times.
laminated siltstones, and mudstones in the remainder of the inlier are
confined to the immediately preceding Cyrtograptus lundgreni Biozone. A S4a: Southern British Isles: Ludlow (mid Gorstian)
quiet water r6gime was interrupted by westerly flowing distal turbidity
currents. Apart from graptolites and orthoconic nautiloids, other fossils are Over the Welsh Borderland there was a marked transgression in early
rare. Cope (1959) analysed those present to show that the greywackes carry Gorstian times indicated by mudstones with a diverse fauna of dominantly
derived thick-shelled brachiopods and crinoids, certain micaceous siltstones small brachiopods. The basin areas accumulated graptolitic muds at this
show a terrestrially derived association of plant fragments and a phyllo- time. The greatest depth appears to have been in mid Gorstian times when
carid, whereas possibly benthonic brachiopods, small bivalves, and crinoids graptolitic muds were also deposited over the outer shelf region around
occur in the more common laminated siltstones. In the very small inlier of Ludlow. Over most of the shelf, however, shelly muds (the Glassia obovata
Knochshigowna Hill, to the north of the Slieve Phelim-Devilsbit inlier and Association of Watkins 1979) were deposited. Bentonites are common but
to the west of Slieve Bloom, a similar facies of early Wenlock age is followed the source of the original volcanic ash is not known; a more distant source
by conglomerates with sandy pockets containg a Homerian shelly fauna in than the Mendips seems likely, perhaps in central Europe, Gorstian deposits
which brachiopods are the dominant constituent (Prendergast 1972). It are absent near Gorsley, south of the Woolhope Inlier, and are thin in the
appears to have been derived from a shallow water environment, though the May Hill Inlier but thicken southwards in the Brookend Borehole indicating
state of preservation of the fauna does not indicate prolonged transport. The that the thinning is due to a local uplift and not an approach to a shoreline.
narrow inlier of the Cratloe Hills to the north of Limerick City also yields There is little direct evidence of the source of the shelf sediments: being a
a pebbly sandstone with Homerian shelly fossils, interrupting the usual time of maximum transgression, probably little of the Midland Platform was
sequence of sparsely graptolitic laminated siltstones and sandstones emergent. The dominant source of material is presumed to be Pretannia to
(Holland et al. 1988). There is thus some evidence for a northerly directed the south (Cope & Bassett 1987). There is no borehole evidence of Gorstian
palaeoslope. Confirmatory evidence for the palaeogeography is seen in the rocks in eastern England but it is postulated that shallow seas covered this
very rich Homerian shelly faunas of the Dingle Peninsula (Holland 1988). area.
There are associated volcanics, chiefly acid in composition, including very West of the Church Stretton lineament there is a fourfold increase in
well displayed ignimbrites and pyroclastics. Finally, to the northeast at thickness and a marked change of facies in the Montgomery Trough of the
Balbriggan the largely unfossiliferous greywackes of the Skerries Formation Welsh Basin. The main stratigraphical unit is the Bailey Hill Formation
contain fragmentary graptolites of very late Wenlock and possibly younger which consists of alternations of hemipelagic carbonaceous laminae and
age. calcareous siltstone units which Bailey (1969) interpreted as turbidites. Tyler
There is evidence of the former presence of Wenlock rocks in the & Woodcock (1987) have suggested an alternative origin as distal storm
occurrence of derived corals in the Carboniferous (or possibly Devonian) deposits from the shelf, deposited below wave-base in the deeper water of an
Peel Sandstone of the Isle of Man (Crowley 1985). The situation in northern outer shelf and not a basin. One difficulty in this interpretation is the rarity
England remains similar to that in the early Wenlock. The Middle Coldwell of storm deposits in the mid Gorstian rocks of the Welsh Borderland area:
Beds are of Monograptus ludensis Biozone age. Their pale grey mudstones storm deposits are, however, very common in the succeeding Ludfordian
are unlaminated or thoroughly bioturbated. There are limestone lenses rich rocks but do not result in pseudo-turbidites in the east-central Wales area.
in trilobites and brachiopods, and also graptolitic bands of Brathay Flags Tyler & Woodcock suggest that the main axis of the Welsh Basin lay farther
lithology. to the west. Bailey, however, interpreted the northward flowing currents as
In North Wales, the latest part of the Wenlock is represented by the turbidity currents flowing along the axis of a trough; this model is supported
Lower Nantglyn Flags Group (Warren et al. 1984), the background sedimen- by the occurrence of slumped shelly mudstones around Builth which moved
tation of which is of 'ribbon banded mudstones'. In this facies mudstones northwestwards from the shelf and slumped siltstones in the Clun Forest
alternate with laminated muddy siltstones yielding graptolites, orthocones, areas which moved southeastwards towards the presumed axis. Despite
and bivalves. There are some lenticular beds of calcareous siltstone. The complications afforded by the Clun Forest area, the evidence seems to
whole represents relatively quiet sedimentation, with the mudstones having support the presence of a locally uplifted submarine area west of Clun
arrived from weak turbidity currents and the calcareous siltstones as distal Forest. It is indeed probable that the main axis of the Welsh Basin did lie
turbidites. Interrupting the ribbon banded mudstone sequence are the well to the west; perhaps, as suggested for the Wenlock, there was a western
disturbed (slumped) beds of the Brynsylldy Formation and, at a generally trough and an eastern trough.
lower level, the intensely bioturbated Upper Mottled Mudstone with its In North Wales (Warren et al. 1984) the Upper Nantglyn Flags and much
shelly fauna. Cummins (1959) gave detailed description of the Nantglyn of the Elwy Group are of this age. The Nantglyn Flags comprise ribbon-
Flags from the wider area from Central to North Wales. Their directional banded mudstones with laminae of silt (possibly distal turbidites) alternating
system of derivation was as for the earlier and coarser Denbigh Grits. with the autochthonous hemipelagic muds. In the Elwy Group greywacke
At Marloes Bay in Pembrokeshire, the shallow water Grey Sandstone siltstones and sandstones of probable turbiditic origin provide evidence of
'Series' yields Homerian fossils in its lower part. A little farther south at transport from the west, along the axis of the Denbigh Trough. Pene-
Freshwater West and Freshwater East the youngest Wenlock beds are contemporaneous slumps are common; the axes strike W-E and the evidence
already in Old Red Sandstone facies. At Cardiff, poorly fossiliferous favours sliding from the north. As yet, no reinterpretation of the Denbigh-
sandstones and siltstones are succeeded by a thin red crinoidal limestone shire sediments as being storm generated has been attempted.
which appears to be equivalent to part of the Much Wenlock Limestone. The Irish Sea Horst is presumed to have persisted to the west from
According to Bassett (1974), the southern margin of the Wenlock Lime- evidence before and after this epoch, forming the western boundary of
stone Platform lay close to Usk and Tortworth, where arenaceous horizons the Welsh Basin, but the land area is shown as small because of the trans-
appear. The Ware and Cliffe boreholes to the east show faunas and litho- gression.
logies of Wenlock Shales type. To the southwest of the Montgomery Trough, the Towy anticlinal area
In the richly fossiliferous Much Wenlock Limestone the bioherms are best accumulated detritus brought northwards from a delta forming on the
developed at the seaward fringe of the reef belt, and there is also an upward northern margin of Pretannia. Farther west the Ludlow rocks are cut out
increase in reefal material as the clay particles decrease. The frame builders by an unconformity at the base of the Old Red Sandstone, but in Pembroke-
were especially tabulate corals, together with stromatoporoids, branching shire the Gorstian is considered to be represented by the deltaic and intertidal
rugose corals, and fenestellid bryozoans. Stromatolites, laminar tabulate deposits of the Gray Sandstone Group and the fluviatile facies of the lowest
corals, stromatoporoids, algae, and bryozoans acted as reef binders. There Old Red Sandstone.
43
44 SILURIAN

The similarity of facies and faunas suggests that the Welsh Basin and the South Wales lay close to the northern margin of Pretannia and suffered
Lake District basin were connected. There, the laminated muddy siltstones greater clastic input. The Usk Inlier displays the 'normal' nodular silty
of the Upper Coldwell Beds are succeeded by the thick Coniston Grits; these limestones on the east but there is a westward decrease in lime content. A
have been interpreted as turbidites with evidence of derivation mainly from little to the south at Cardiff (Waters & Lawrence 1987), poorly calcareous
the northwest. Furness (1965) suggested that the abundance of spilite and shelly siltstones dominate. Farther west, near Llandeilo, the postulated delta
chert fragments in the Coniston Grits indicated a derivation from the of mid Gorstian times had prograded northwards to give the conglomerates
Ballantrae Volcanic Group of southern Scotland. Alternatively a marginal and quartzitic sandstones of the Trichrfig Beds (Potter & Price 1965), with
arc of Ballantrae type on the northern edge of the Southern Britain plate is red beds indicating periodic exposure. Marginal marine beds are character-
postulated here. ized by a Lingula-bivalve-gastropod association suggesting salinity vari-
In Ireland (Holland 1981), there are greywackes around Balbriggan and it ations which discouraged articulate brachiopods and other stenohaline
is assumed that the graptolitic siltstones and shales of the Wenlock rocks of forms. Northwards along the Towy Anticline the sediments fine to siltstones,
south-central Ireland persisted into the Gorstian. In the Dingle area of still poorly calcareous, and brachiopods become commoner.
southwestern Ireland graptolitic shales pass westwards into calcareous Outside the area of this map (i.e. Pembrokeshire, Lake District, Ireland)
siltstones with a shelly fauna, mainly brachiopods, similar to that of the the conditions of depositon were similar to those of mid Gorstian times.
Welsh Borderland shelf. JDL
JDL
S5a: Southern British Isles: (early Ludfordian)
S4b: Southern British Isles: Ludlow (late Gorstian)
In early Ludfordian times the carbonate shelf environment was replaced by
In late Gorstian times a gradual change took place in the Welsh Borderland clastic silt sedimentation. The basal sediments of the Lower Leintwardine
area from a muddy to a carbonate shelf: the change from 'Lower Ludlow Formation are still limestones, well developed in a higher energy carbonate
Shales' to 'Aymestrey Limestone' in the older terminology. The Lower belt near the shelf edge, e.g. Leintwardine and Aymestrey, where they are
Bringewood Formation (Atkins 1979) comprises mainly bioturbated cal- succeeded by shelly laminated shales. The limestones form the top part of
careous siltstones characterized by a fauna of large brachiopods, particularly the 'Aymestrey Limestone' of the traditional classification but it is notable
strophomenids. The Mesopholidostrophia laevigata Association (Watkins that the Kirkidium knightii-tabulate coral Association has disappeared. The
1979) is dominant but Atkins also recognizes a shelf-edge Shagamella succeeding 'Mocktree Shales' contain a less diverse benthic assemblage
ludloviensis Association and an inner shelf Atrypa reticularis-Sphaeri- dominated by Shagamella minor and Dayia navicula and show a transition to
rhynchia wilsoni Association. The map mostly covers the succeeding Upper the basin facies. At Leintwardine submarine channels descended westwards
Bringewood Formation, the most calcareous part of the shelf Ludlovian. At into the basin and slumping to the northwest is developed from Aymestrey
the shelf-edge along a N-S band from Craven Arms to Aymestrey bioclastic southwards.
limestones are well developed, showing cross-bedding in places. Shell banks Over most of the shelf the Lower Leintwardine Formation averages 50 m
crowded with disarticulated valves of Kirkidium knightii (the K. knight# in thickness. Basal nodular silty limestones are overlain by bioturbated
Association of Watkins & Aithie 1980) confirm the high-energy shallow flaggy calcareous siltstones with thin limestone bands. Watkins (1979) called
water conditions. At Aymestrey, the limestone thins and passes rapidly into this the coquinoid siltstone facies containing his Sphaerirhynchia wilsoni
calcareous siltstones and then into laminated siltstones involved in westward Association. The fossils occur as disturbed neighbourhood assemblages
slumping. often in monotypic lenses due to environmental stress (Cherns 1988) on
The main shelf region seems to have been protected by the shelf-edge a storm-influenced subtidal shelf.
ridge, although there is no evidence of any of the long-standing positive The inner shelf sees a thinning of the deposits, particularly near the
blocks being emergent and undergoing erosion. In this quieter water the M. Gorsley axis, and the common occurrence of shelly conglomeratic lime-
laevigata Association is dominant in nodular silty limestones. In the inner stones, some of which show evidence of hardground formation (Cherns
shelf area around the Malverns the influx of fine silts was increased and an 1980); these conglomerates are well seen at May Hill, South Woolhope and
Atrypa reticularis-coral Association occurs (Watkins & Aithie 1980). The in the Brookend Borehole but also occur in central England, Wenlock Edge,
near shore aspect is confirmed by the complete absence of the Upper Malvern and Usk. They evidently indicate interrupted deposition and perio-
Bringewood Formation (and other formations) in the southern Woolhope dic erosion, mainly submarine. The formation as a whole is characterized by
Inlier and the May Hill Inlier. Uplift in the Gorsley area (Lawson 1954) a Sphaerirhynchia wilsoni-Isorthis orbicularis Association over the inner
evidently resulted in non-deposition and/or erosion. South of the Bristol shelf. In the higher beds thin layers of phosphatized clasts and fish remains
Channel, however, the nodular silty limestones reappear in the Brookend occur, again indicating phases of non-deposition with the enrichment of
Borehole. The suggestion by Mohamad (1981) that much of the back-barrier organic phosphates by winnowing of fine sediment in the shallow warm seas.
shelf was intertidal seems untenable in view of the abundant articulate A low-energy environment with a low sedimentation rate is postulated and
brachiopods, corals, crinoids and bryozoa. there is little evidence of the direction of transport or the source of the
The shelf-edge was to the east of the line of the Church Stretton Fault for sediment.
most of its traceable course. There are northwesterly directed slumps around From the Welsh Borderland shelf into the Welsh Basin there is an abrupt
Knill and New Radnor indicating persistence of a basinal area, not necess- change both of facies and thickness (from 50 m to 200 m, then to 450 m;
arily of great depth. Slumps in the Clun Forest area moved towards the Holland & Lawson 1963, p. 280)just east of the Church Stretton Fault. The
southeast, again suggesting the margin of a trough along which silt was laminated siltstone facies is dominant, representing a low energy environ-
being transported northwards into an area of hemipelagic muds. The fauna ment of poorly oxic muds receiving repeated influxes of silt. The Lingula
of graptolites and orthocones with a poor benthic fauna suggests that the lata-Saetograptus leintwardinensis Association is dominant: Cherns (1979)
bottom waters were not well oxygenated, and lay well below wave-base indicated that L. lata had the infaunal mode of life of Recent lingulids but in
although not necessarily at great depth. The alternation of silt and mud gave deeper and muddier seas. At the western edge of the outcrop in Clun Forest
the laminated siltstone facies (Holland & Lawson 1963; the Striped Flags slumping to the southeast persisted, with a northerly movement of sediment
of Kirk 1951) that persists northwards through the Long Mountain along "~he presumed axial region of the trough.
into Denbighshire. For the Bringewood formations together the thickness In South Wales, along the southeastern flank of the Towy Anticline,
increases from c. 80 m on the main shelf to ~<500 m in the basin. poorly calcareous flaggy siltstones and fine sandstones occur, presumably

Fig. 4. Facies relationships in the Ludlow Series of South Wales and the Welsh borderland (from Holland & Lawson 1963). Key: I, shelly olive mudstone facies; 2, muddy
siltstone facies, 3, graptotitic shale facies; 4, laminated siltstone facies; 5, turbidite siltstone facies; 6, turnescenssiltstone facies, 7, shelly siltstone facies; 8, calcareous shelly
siltstone facies; 9, slumped shelly siltstone facies; 10, strophomenid siltstone facies; 11, massive limestone facies; 12, flaggy limestone facies; 13, deltaic facies.
45
46 SILURIAN

deposits of the delta front. The conditions were those of a moderate energy In the Lake District the late Ludfordian includes a large part of the thick
marginal sub-tidal environment. In Pembrokeshire, red sandstones are Kirby Moor Flags (Shaw 1971). This formation comprises thickly flaggy,
presumed to have been deposited partly in early Ludfordian times by rivers sorted siltstones with some slightly calcareous beds and lenses of fossils. A
flowing northwards from the Pretannia massif. shallow water Salopina-Protochonetes Association occurs but with many
In North Wales the laminated siltstone facies persisted through the S. more gastropods than usual. The sediments were derived from the north-
incipiens Biozone into the S. leintwardinensis Biozone, the higher parts of west. As with Wales, the Lake District basin had been largely infilled
which are missing because of erosion. The same facies prevails in the (although there was still tectonic subsidence) to produce a shallow subtidal
Bannisdale Slates of the Lake District and a connection with the deeper moderate-energy environment.
water basin from Wales is assumed. In Ireland, it is postulated that the central region was emergent and being
Over most of Ireland (Holland 1981) there is no evidence but the pattern eroded, with rivers providing sediment for the red fluviatile sandstones of the
of earlier maps is followed by proposing a basin tongue stretching southwest- Dingle Group in Kerry.
wards across the centre of the country. In the Dingle Peninsula, however, JDL
shallow water calcareous siltstones are dominant with a benthic fauna in
which the brachiopods suggest a connection with the Welsh Borderland S6a,b, $7: Northern British Isles: Liandovery (Rhuddanian,
shelf, although the precise route is not clear. Aeronian, Telychian)
The Upper Leintwardine Formation of the shelf is thin and though
generally of similar facies to the lower formation it has a distinctive fauna The palinspastic model used in maps of the northern British Isles, for the
including some trilobites and ostracodes which suggest a Baltic connection. Ordovician Period, was developed by J. K. Ingham and his aid in applying it
This Shaleria-neobeyrichiacean Association facilitates correlation with to the Silurian Period is gratefully acknowledged. The timing of the closure
Gotland and other areas. The basin facies has a widespread Aegiria grayi- of the Iapetus Ocean, the cessation of northwestward subduction and of
Neobeyrichia lauensis Association at the same level, and the neobeyrichia- strike-slip on elemental boundaries are controversial. The maps subscribe to
ceans also occur in the Lake District. the view that Iapetus continued closing, by northwestward subduction, into
JDL the Llandovery Epoch, although during the epoch the ocean may have
become little more than an ensialic residual trough. The type of accretionary
S5b: Southern British Isles: (late Ludfordian) prism which could have resulted is depicted schematically in Fig. 5. That the
two continents Laurentia in the north and Gondwana in the south had been
By late Ludfordian times an almost uniform shallow water facies had spread on a collision course for much of the early Palaeozoic and were still
over the Welsh Borderland, east-central Wales and the Lake District and the separated by this trough in the early Silurian are probably the two least
area of deposition became more restricted with the continuing emergence of disputed factors of the palaeogeography
the surrounding land masses in the Midlands and Irish Sea areas. The Welsh
basin was probably only slightly deeper than the shelf but remained a
SE downslope NW
tectonically negative area with a great increase in thickness of Whitcliffe __ movement
sediments from c. 40 m on the shelf to a maximum of about 400 m in the
basin area of Clun Forest.
The Upper Whitcliffe Formation of the main shelf comprises bioturbated
.r i," "xr
olive-grey flaggy calcareous siltstones with shelly coquinoid lenses: wavy and
ripple bedding are not uncommon with some thin cross-bedded units in the
higher layers (Bradfield 1987). Alternations of bioturbated siltstones and -...:-.. ;.:~/
shales with coarser laminated siltstones are interpeted as storm-influenced
deposits. The shell beds may be the traction load of the initial coarse fraction
deposit (Watkins 1979). Faunally the formation is characterized by a low- Fig. 5. Evolution of Southern Uplands accretionary wedges: accretion stage with
southeast-directed thrusting (after Needham & Knipe 1986).
diversity assemblage of brachiopods and bivalves: many of the early Ludfor-
dian (Leintwardine) brachiopods have disappeared, perhaps unable to cope One reason for supposing that a collision of the two continental forelands
with an increased rate of sedimentation which would not have affected the had occurred earlier, in late Caradoc time, is the lack of direct evidence of
infaunal bivalves. A Salopina-Protochonetes Association is dominant. The arc-type volcanicity in either margin after the Caradoc. Bluck (1983) believes
overall picture is of a proximal shelf subtidal environment with frequent there to be indirect evidence for Llandovery activity on the now concealed
storm sedimentation. Antia (1980) considered that the topmost beds were Cockburnland and distal arc-type volcanism is evinced by the numerous
tidal. There was an increased sedimentation rate, increased energy level and metabentonites in the Moffat Shale, ranging in age from the late Ordovician
increased proximity to source (Watkins 1979). Bailey & Rees (1973) postu- to the maximus Subzone (early Telychian) (Merriman & Roberts 1990).
late directions of derivation from the east but not from a nearby source. However, the inference has been drawn that subduction had ceased (Murphy
The facies is similar in the inner shelf but the Whitcliffe formations & Hutton 1986). Murphy and Hutton claim that the remnant, or 'successor',
become very thin around the Gorsley uplift (2 m at Gorsley). To the south at basin (Fig. 7) was then bridged merely by clastic infill in the Llandovery. The
May Hill, Tites Point and Brookend frequent interruptions in deposition are older view (Mitchell & McKerrow 1975) holds that the Iapetus Ocean,
indicated by the occurrence of phosphatized pebble beds, layers of fish floored by oceanic crust, continued to separate the two continents by more
denticles ('bone-beds') and layers of euhedral biotite. There is also an than 1000km until late Silurian times (McKerrow & Soper 1987); from
easterly thinning towards the Midlands. Boreholes in eastern England Ordovician to Silurian time northwestward subduction continued while an
(Stowlingtoft and Lowestoft) have yielded shallow water sediments of Ordovician-Silurian thrust-sliced sedimentary prism was incrementally
probable late Ludfordian age. accreted to the Laurentian margin. Although volcanicity in North Wales,
To the west of the Church Stretton Fault system the equivalent strata southeast Ireland and the Lake District implies that southeastward directed
increase rapidly in thickness due to more rapid deposition in a subsiding subduction of oceanic crust, beneath northern Gondwana, continued until
tectonic basin. The facies changes are, however, subtle: the siltstones are late Caradoc time there is no documented evidence for the consequential
somewhat darker and muddier but still with wavy bedding, low angle cross- accretionary sedimentary prism on the Avalonian foreland. It is supposed
bedding and lenses of shells. The brachiopods of the Salopina-Protochonetes therefore that sometime during the Silurian any such prism underthrust the
Association are slightly but noticeably smaller than on the shelf and there is Laurentian foreland (e.g. Leggett et al. 1983; Stone et al. 1987), or was
a more diverse molluscan fauna. In the highest beds of Kerry and Builth strike-slipped away.
(and also on the shelf at Usk) the Loxonema Association occurs in decalci-
fied sandstones which are considered to have been deposited in a moderate Southern Uplands / L o n g f o r d - D o w n Z o n e
energy environment in a shoreface setting. In the western part of the
outcrop, in the Kerry and Long Mountain areas, Bradfield (1987) recognizes Evidence of the palaeogeography of the Laurentian continental margin
a return to the olive-grey siltstone facies characteristic of the shelf. Some thin resides in northward-dipping southward thrust slices of sedimentary rocks
slump bands occur. These facts suggest that the Bishops Castle, Knighton, which comprise the Southern Uplands in Scotland and the Longford-Down
Radnor Forest and Builth areas were situated in somewhat deeper water tract in Ireland. Two different views have been expounded. One adopts the
than the areas to the west and east. There is still some evidence of the requisites of the accretionary prism outlined by Leggett et al. (1979) with a
northwards transport of sediment (Bailey & Rees 1973). fore-arc trench, as described by Kelling et aL (1987), being dominated by a
In the Towy anticlinal area of South Wales, the separate Whitcliffe variety of depositional systems, mainly fan-type turbidites which had
formations are not easily distinguished. The late Ludfordian strata around sources to the northwest in arc terrane, already accreted arc-apron sedi-
Cwm Dwr comprise olive-grey poorly calcareous siltstones with thin layers ments, and a continental source. Upon reaching the trench these sediments,
of sandstone and decalcified shelly limestone bands (Bradfield 1987). A in Llandovery times at least, were diverted to the southwest and formed co-
Salopina-Protochonetes Association, depleted in Salopina, is present. axial turbiditic wedges. The early Llandovery sediments of each wedge are
Further south, these beds are cut out by the unconformity at the base of the recognizably different from the next, e.g. the Kilfillan Formation (KF) and
Tilestones. Money Head Formation (MHF) of Fig. 6A, suggesting that the trench was
In Pembrokeshire fluviatile deposition continued and is presumably rep- not 'a single unconstrained basin', but divided by structural barriers. Sole
resented by red sandstones of part of the Old Red Sandstone. structures including flute-casts are common.
In North Wales, erosion has removed late Ludfordian strata but pebbles These coarse detrital sediments had an oceanward limit, possibly confined
of calcareous siltstone with Whitcliffe fossils occur in the basal conglomerate by an outer trench high, so that hemipelagic anoxic muds were accumulating
of the Carboniferous rocks and are presumed to give evidence of a similar to the southeast: the Moffat Shales. Of the fault-bound sequences in each of
facies to that in east-central Wales. the accretionary slices recognized by Leggett et al. (1979, fig. 2), that now
14 CAMBRIAN

Massif or on its margins. The Cambrian of Nuneaton accumulated to the rocks encountered in boreholes in Lincolnshire (Cowie et al. 1972, table 2)
southwest of this Charnian Massif. Coastal Hartshill Formation sandstones are varied--phyllitic slates, feldspathic sandstones and greywackes--and a
(cycle l, thick), through to condensed shallow water, shelly Hyolithes Cambrian age has been suggested, but without adequate faunal control. It is
limestones (cycle 2, thin) and deeper water Mn-rich shales (cycle 3, thick) assumed here that this area was part of a marginal basin that existed
exhibit a trend reciprocal with that of Shropshire and close to that of the throughout the early Lower Palaeozoic and extended from northern England
Avalon succession, SE Newfoundland. The nearby Lickey Quartzites are into the Brabant area of Belgium where clastic rocks of Comley and St
lithologically similar but are of uncertain age. The former presence of David's age are known (Vanguestaine 1974, 1978). It is not known whether
limestones is suggested by clasts in the Permian Nechells Breccia (Brasier & the Precambrian of the Charnwood Forest area was emergent; there is some
Hewitt 1981). geophysical evidence to the northeast of Charnwood Forest for the presence
The Withycombe Farm borehole near Banbury, Oxfordshire, gives an of a post-Charnian, pre-Triassic succession that possibly represents part of
indication of conditions to the east of the Malverns. Volcanic basement of the local Cambrian (Maguire 1987). A positive area in northern Norfolk was
presumed late Precambrian age is there overlain by the Withycombe Forma- formerly postulated on the presence of supposedly Precambrian volcanic
tion, a relatively thick sequence of sandstones bearing trace fossils (cf. cycle rocks encountered in the North Creak Borehole, but these are now thought
l) passing up into a thick succession of offshore shales with Aldanella sp., to be part of the east Midlands Caledonian Belt (Pharaoh et al. 1987).
Watsonella sp., Platysolenites antiquissimus, Orthotheca cylindrica and Lon- AWAR
tova-type acritarchs (Rushton & Molyneux 1990). This invites compari-
son with the Chapel Island Formation (cycle 1) or lower Bonavista Forma-
tion (cycle 2) of southeast Newfoundland, both of attleborensis age. 4~2a: Southern British Isles: Merioneth
MDB
In Scandinavia the Merioneth Series has been divided into some eight
E~lb: Southern British Isles: St David's biozones and thirty subzones, and this scheme can be applied in areas, such
as England and Wales, where the olenid trilobite biofacies is developed. The
The St David's Series is equivalent to the Middle Cambrian of European interval depicted on the map is the Olenus Biozone and more particularly its
usage, but not necessarily equivalent to that of other regions such as upper part (O. cataractes Subzone).
Australia, China or North America. The Series is subdivided according to Deposits formed in England and Wales during the Merioneth Epoch
the Swedish zonal scheme, with a lower, a middle and an upper part. In the reflect generally stable tectonic conditions. After a widespread non-sequence
Welsh Basin the lower part is represented mainly by thick turbiditic in the late St David's successions (possibly caused by a eustatic regression), a
sandstone successions; in Shropshire by thinner glauconitic sandstones and major new sedimentary cycle commenced, apparently before the end of the
mudstones, and in the English Midlands by mudstones. Biostratigraphical St David's Epoch, in both the Welsh Basin and the English Midlands
control is generally poor and correlation insecure (Cowie et al. 1972; (Rushton 1978; Allen et al. 1981).
Rushton 1974). The Welsh Basin, though deeper at the beginning of the Merioneth Epoch,
Faunal control is better in the middle part of the Series, where the was partly filled towards the end of Olenus Biozone times with alterations of
following biozones are used: shales and contourite(?) sandstones (upper Maentwrog Formation), and
those beds are overlain by thick shallow-water sandstones (Ffestiniog Flags
Ptychagnostus punctuosus Biozone,
Formation, >650m, Allen et al. 1981). There is some evidence in the
Hypagnostus parvifrons Biozone,
Harlech Dome, St Tudwars Peninsula, and in the St David's area, that the
Tomagnostus fissus Biozone,
sediments were deposited by currents flowing from the south, as in the St
Ptychagnostus gibbus Biozone.
David's Series. To the north of the Harlech Dome, however, the Merioneth
The gibbus Biozone appears to represent a widespread transgression, Series is comparatively attenuated and includes shallow-water quartzitic
possibly related to a eustatic sea-level rise (Rowell et al. 1982, p 176), but sandstones, with trace fossils (Cruziana facies) that appear to have been
that zone is not easily recognized in Britain. The overlyingfissus Biozone is derived from the north, though the indices of current orientation give a wide
more widely recognized and this interval, for which correlation is more range of directions (Crimes 1970).
secure, is represented on the map. Muddy sediments accumulated over the site of the Midlands Platform and
In the Shropshire area glauconitic sandstones and calcareous beds were it is assumed here that these (Outwoods Formation of the Nuneaton Inlier,
deposited in shallow water, and Cobbold (1927) demonstrated a local with known subsurface extension to the south and west) transgressed the
unconformity indicative of uplift along the Church Stretton fault zone at Precambrian Charnwood Forest massif. The fauna of the Outwoods Forma-
about this time. The absence of the St David's Series in the Malvern and tion includes olenid trilobites that tolerated oxygen-poor environments,
Lilleshall areas (Rushton et al. 1988) implies similar uplift, but with less together with a number of other trilobite genera of wide palaeogeographical
certainty. distribution that inhabited outer-shelf regions (Rushton 1983).
The Midlands Platform carries a thin sequence of variegated mudstones In the Welsh Borderland the Olenus Biozone is unrepresented, the earliest
deposited below wave-base (Abbey Shales, three biozones represented, representatives of the Merioneth Series being the Orusia Shales (Parabolina
< 40 m thick in total). The fauna is dominated by agnostid trilobites and spinulosa Biozone) resting unconformably on Comley and St David's rocks.
small brachiopods. Similar but thicker successions (c. 200 m total?) occur However, at Malvern there is a thin sequence of Cyclotron-bearing shales
in southwest Wales and also in North Wales, though there the mudstone that are thought to be of Olenus Biozone age. Uplift along the Church
sequence is underlain by turbiditic sandstones offissus age (the Gamlan and Stretton and Malvern lineaments is suggested by northeastwardly directed
Caered formations) that show evidence of derivation from a southerly slumps or currents in the Moor Wood Flags Formation of the Nuneaton
source (Crimes 1970). This implies a positive area to supply sediment to area (Taylor & Rushton 1972, p. 22). The derivation ofmicaceous detritus in
north Wales whilst by-passing the St David's area in the southwest. The the Llangynog area, south Wales, is believed to be from a metamorphic
presence of bentonite beds indicates contemporaneous volcanic activity, basement (Pretannia) to the south (Cope & Bassett 1987).
though the location of the activity is unknown. The later Merioneth Epoch, from Parabolina spinulosa Biozone to Peltura
There is no evidence for the St David's Series north of the Harlech Dome scarabaeoides Biozone times, was dominated by deposition of black mud-
and St Tudwal's Peninsula, and it is assumed that the Irish Sea Horst was a stones in disaerobic environments (Doigeilau, Bentleyford, Monks Park and
positive area; nevertheless, no contemporaneous sediments are known to White-Leaved-Oak formations). These pyritic, carbonaceous and uranifer-
have been derived thence. ous muds contain little or no coarse clastic material and are considered to
The non-agnostid trilobites most typical of this epoch, Paradoxides and represent a period of slow deposition; the proved olenid zones differ from
blind benthic forms (Hartshillia, Holocephalina) characterize outer-shelf locality to locality (Thomas et al..1984, p. 15), and the extent of possible
environments in Wales and elsewhere in Avalonia (southeast Newfound- breaks in deposition, comparable with those inferred in the contempor-
land, New Brunswick) and in Baltica and the margins of Gondwana aneous rocks in Scandinavia, is not certain. The black mudstones in the
(Conway Morris & Rushton 1988). Croft Borehole, Lilleshall (northern Shropshire), are unique in having many
After fissus Biozone times, mudstone deposition prevailed (except locally intercalations of current-bedded siltstone and sandstone, presumably of
in the Comley area of Shropshire). There is a widespread hiatus after shallow-water origin (Rushton et al. 1988). The source of the silt-grade
punctuosus Biozone times and before the Merioneth Epoch, the mudstone material is not known but is thought to originate from a local uplift along
succession in the Nuneaton district being the best constrained (Taylor & the Church Stretton Fault complex.
Rushton 1972). The sedimentary rocks of the southeast Irish marginal basin have poor
In southeast Ireland the marginal basin of Leinster has yielded some biostratigraphical control but include beds referable to the Merioneth Series
evidence of the St David's Series (Smith 1977, 1981). In Co Wexford (the (Ribband Group; Smith 1977, 1981). They are argillites and arenites that
Cahore and Ribband Groups) and Co Wicklow (the Bray Group), the appear to have been deposited in deep water. The mudstones and fine-
sediments were deposited in deep water (Crimes & Crossley 1968). These grained sandstones of the Askingarren Formation, interpreted as underlying
rocks yield trace-fossils, especially Oldhamia, in abundance (Dhonau & correlatives of the Ribband Group (Crimes & Crossley 1968, pp. 191,207),
Holland 1974), but acritarchs, in part of St David's age, are the only body- are considered as possibly late Cambrian in age. Flute casts indicate
fossils known; the best preserved are from the Booley Bay Formation derivation from a southerly quarter. There is no evidence for the continued
(Ribband Group) at Hook Head on the south coast (Smith 1981). Sediment existence, into the Merioneth Epoch, of the marginal arc thought to have
was derived from both the southeast (Irish Sea Horst) and the northwest, existed to the northwest of the southeast Irish marginal basin in St David's
where a positive area, possibly a marginal arc, is thought to have provided a times.
source area. There is no biostratigraphical evidence for the Merioneth Series in the
Less is known of the eastern Caledonide belt to the northeast of the eastern English Caledonides (see the St David's Series, above).
Midland Platform and facing the Tornquist Sea. The clastic sedimentary AWAR
48 SILURIAN

the oceanic area was accumulating carbonaceous graptolitic muds and


continued to do so for a further c. 10 Ma. Leggett (1987) claimed that this
supported the existence of the oceanic plate, and thus Iapetus Ocean, until
the Wenlock, but there seems no reason why such deposition could not have
carried on over the continental crust of Murphy & Hutton's 'successor
basin' (Fig. 7) in its early phase (Murphy & Hutton 1986). However,
whether because of a northwestward moving oceanic plate or by south-
eastward facies progradation (Griffith & Wilson 1982, fig. 4), greywacke
appears in the Irish sequences, in sedgwickii Biozone at Coalpit Bay, in
turriculatus Biozone at Tomgraney (Rickards & Archer 1969) and possibly
in the griestoniensis Biozone in the western end of the Slieve Bernagh Inlier
(Plegg, in Leggett et al. 1979). Greywacke and minor flows of basic lava
occur at Slieve Aughty (Emo & Smith 1978), but their ages are not known
precisely.
The alternative views of Murphy & Hutton (1986) and Stone et al. (1987)
are not represented in the accompanying maps. These can account for the
same thrust-imbricate, Ordovician and Silurian submarine fan sequences of
the Southern Uplands (and the Longford-Down Zone), with their south-
eastward overlap of pelagic mud deposits (Moffat Shales), in terms of thin-
skin thrusting of the deposits of a back-arc basin; a basin which had
existed largely undeformed through much of the Ordovician into the early
Llandovery (Stone et al. 1987, fig. 3; Fig. 8). They cite the andesite-rich
composition of some of the greywacke fans and their derivation from the
south as evidence for the existence of a now concealed Ordovician/early
Llandovery active volcanic arc to the south, with a fore-arc basin beyond.
Fig. 6. Schematic diagram depicting the palaeogeographical and geotectonic evo- Kelley & Bluck (1989) have more recently suggested that this detritus is of
lution of the SW Southern Uplands during the early Llandovery (as illustrated by Cambrian (or earlier) age.
the Rhinns of Galloway and Luce Bay sections). CM, Chippermore Member;
CFM, Cairnie Finnart Member; DPM, Daw Point Member; FBF, Float Bay
Member; GF, Garheugh Formation (with a and b Formations); GPF, Grennan
Point Formation; KF, Kilfillan Formation; MHF; Money Head Formation; SBb,
Striking Bight beds. 1, 2, a, b, etc., represent earlier accreted tectono-stratigraphic
tracts (from Kelling et al. 1987, fig. 13).

Fig. 8. Back-arc basin model for Southern Uplands (from Stone et aL 1987, fig. 3).

Oblique collision of the Laurentian and Avalonian foreland margins


during the mid-late Llandovery caused cessation of northward subduction
at an island arc and produced instead northward underthrusting with the
adjacent back arc basin. Eventually the postulated arc terrane underthrust
the back-arc basin and the Avalonian foreland underthrust the Laurentian
foreland basin. The contents of the back-arc basin rose as a thin-skin,
southward propagating, thrust stack; the Moffat Shales acted as the decolle-
ment zone and the stack thrust southeastwards, ramping over the arc terrane
during latest Telychian times (Fig. 9).

Fig. 7. Model for structure of Southern Uplands from Murphy & Hutton (1986.
fig. 3). CB = central belt; SB = southern belt; E - E = approximate level of
erosion.

forming Tract 4 (IV) reached the turbiditic greywacke environment of the


trench in early Llandovery time (e.g. the Pyroxenous Group of Talla and the
Money Head and Kilfillan formations; see Fig. 6A). Keiling et al. (1987,
p. 803) depict the trench environment at this time (Fig. 6). Sequences in the
more southeasterly tracts show that in persculptus/acuminatus times dark
grey graptotitic muds were being deposited in the oceanic regions beyond the
trench. It was not until early turriculatus times and griestoniensis times that
the trench-parallel strips of this ocean floor which now comprise Tracts 5
and 6 respectively of the prism reached the trench environment of greywacke
(Leggett et al. 1979).
Though less clearly exposed, the sequences in the Longford-Down Zone Fig. 9. Sketch section illustrating the southward migration of the Southern Uplands
of Ireland indicate a similar pattern. In the northern tracts, e.g. just thrust stack and initiation of the foreland basin during the late Llandovery Epoch
southeast of Orlock Bridge on the east coast, the oldest Silurian rocks, (from Stone et al. 1987, fig. 5).
probably of acuminatus age, are arenaceous greywackes of the Portavo
Formation (Griffith & Wilson 1982, p. 13). Further to the WSW, e.g. at
Strokestown, persculptus-acuminatus deposition was of turbidites and mass- Midland Valley of Scotland: south Mayo and north Galway
flow volcaniclastic conglomerates while in acuminatus times basalts were
extruded (Leggett et al. 1979). These might suggest a trench environment, Northeastward of the accretionary prism (or thrust stack of Stone et al.
but without the coarse clastic fans and axial wedges of the central belt of the (1987)) lay Laurentian continental crust. On this, in the Llandovery, there
Southern Uplands of Scotland. Thus it can be argued that those facies were was a marine basin forward of an Ordovician volcanic arc, possibly still
advancing southwestward along the trench slope axis but had only just active, and behind was a back-arc basin, or inter-arc basin of Bluck (1983;
reached the eastern coast of Ireland by acuminatus times. Fig. 10). According to Bluck the prism (or 'thrust stack') overrode north-
To the south, as far probably as Tomgraney (Rickards & Archer 1969), westward much of this Midland Valley terrane in post-Silurian times, thus
49
50 SILURIAN

concealing all but the back-arc basin tract lying through the Midland Valley of Llandovery age. The fluvial gravels were overwhelmed eventually by a
to south Mayo and north Galway (Fig. 10, ii). marine transgression and succeeded by shallow marine sands and gravels
Evidence of the concealed terrane has been found in xenoliths (Upton et with the remains of shells including brachiopods. The Gowlaun Conglomer-
al. 1983) and the northward-transported pebbles of Wenlock conglomerates ate, considered by Piper (1972) to represent a deep water base-of-slope
in the Midland Valley Silurian inliers (Bluck 1983). Bluck deduced that these deposit, has recently been reinterpreted as a shallow water deposit (Williams
were derived fluvially from a catchment which extended at least 70 km & O'Connor 1987). Thus without the need to accommodate a temporary
southward of the Southern Uplands Fault, a distance commensurate with deep water environ, strong comparison can be made with the Midland Valley
present trench-arc gaps. This Wenlock 'Cockburnland' (Bluck 1983, of Scotland except that in Ireland the Silurian rests partly on the displaced
pp. 126-7) probably commenced its emergence at the end of the Llandovery Dalradian block of Connemara.
as a series of islands. RC

Ii
S8a: Northern British Isles: Wenlock (Sheinwoodian)
- - - - - MIDLAND VALLEY ~ HIDDEN MIDLAND VALLEY --- ~ SOUTHERN UPLANDS . . . .
arc Evidence from the rocks themselves for early Wenlock palaeogeography in
x'7"k a rc fore-arc accretionary prism northern Britain is confined to the Southern Belt of the Southern Uplands of
interarc ~ 1 basin I trench
Scotland and its Irish equivalents in the inliers of Slieve Aughty and Slieve
Bernagh, to the Midland Valley of Scotland, and to the rather separate
western Irish area of south Mayo and north Galway. Otherwise the picture
must be fitted with what has gone before and what follows. Relationship
with southern Britain remains unknown, though, as there have been sugges-
tions that there is considerable similarity between some of the Silurian rocks
in the southern part of the Southern Uplands of Scotland and those in the
Lake District of northern England, these areas may have been in close
proximity.
. . . . . . . . MIDLAND VALLEY SOUTHERN UPLANDS The Southern Belt of the Southern Uplands contains faulted strips of
Wenlock rocks. Those of mid Sheinwoodian age are graded greywackes,
only occasionally thick and pebbly, siltstones, mudstones, and graptolitic
shales. These Riccarton Beds also yield orthoconic cephalopods. Tool marks
are conspicuous among the sole structures and there are also ripples indica-
ting bottom currents at variance with the axial flow of the turbidity currents.
! i In the group of faulted inliers of Slieve Aughty, the Derryfadda and
Killanena Formations, found respectively to the north and south, are
broadly of this age as indicated by spores and acritarchs (Emo & Smith
1978). The Derryfadda Formation comprises proximal turbidites associated
with siltstones and mudstones; they are cut by strike faults. The Killanena
Fig. 10. (i) Interarc basin, missing fore-arc basin (beneath Southern Uplands) and Formation is of thinner bedded and finer grained greywackes interbedded
Southern Uplands accretionary prism. (ii) Overthrusting of accretionary prism: 1 = with laminated beds. The situation in the complex of inliers of Slieve
'igneous conglomerate', 2 = 'quartzite conglomerate', 3 = 'greywacke conglomerate'. Bernagh, the Broadford Mountains, and the Arra Mountains is still not fully
1 and 2 have a source in rock suggested to have been overthrust by the greywackes
of the Southern Uplands (from Bluck 1983). described in modern terms. Turbidites appear here above the Monograptus
crispus Biozone (Telychian) and continue through the Wenlock.
The back-arc ('interarc') basin of Llandovery times is witnessed in the Anderson & Oliver (1986) have stressed the importance of the Orlock
rocks of the Midland Valley Silurian inliers including those near Girvan and Bridge-Kingledores Fault, which separates the Northern and Central belts
in the northern Irish inliers of Pomeroy, Lisbellaw, Charlestown and Clew of the Southern Uplands of Scotland and their equivalents in Ireland.
Bay in Mayo and Killary Harbour in north Galway. The earliest Silurian Hutton & Murphy (1987) have suggested that the northerly sedimentary
deposits are the persculptus-acuminatus graptolitic muds of Pomeroy. These source customarily referred to as Cockburnland has been lost at this time by
follow Ashgill deposits without a break and Rrobably represent the deepest sinistral strike-slip displacement along this fault.
parts of this basin. To the east at Craighead, northeast of Girvan (Freshney Several of the Silurian inliers of the Midland Valley of Scotland (Lesma-
1959) a marine transgression over Hirnantian rocks was under way in hagow, Hagshaw Hills, Carmichael, Pentland Hills) show a regressive
acuminatus time. Basal deposits of gravel appear to be derived from the sequence above marine Llandovery beds. Above the various conglomerates
Midland Valley arc to the north and were distributed as channel fills. It is which, except at Lesmahagow, mark the base of the Wenlock, there are
necessary therefore to invoke an E-W land-ridge of Midland Valley terrane arenaceous beds with some red sediments and horizons rich in fossil fish.
lying to the north (Bluck 1983) to provide this detritus. These coarser Wenlock deposits are seen as alluvial fans derived from a lost
Shelly sands predominated through the early Llandovery at Girvan, but source to the southeast.
in convolutus-early sedgwickii times graptolitic muds interrupted this The Louisberg-Clare Island succession in south Mayo shows cross-
deposition. Cocks & Toghill (1973) considered these muds to have been deep stratified pebbly sandstones, which Phillips (1974) showed to be derived
water deposits comparable with the Moffat Shales, but they probably record from the northwest. There is evidence of fluvial channels. Separated from the
merely a slight deepening and concomitant marine transgression of coastal above by faulting, and metamorphosed to lower greenschist facies, the
plains, such as was occurring on Avalonia at this time. Shelly muds and Croagh Patrick sequence is of shallow water, marine sediments. Farther
sands returned to the area even in the later part ofsedgwickii time with first a south in the country from Killary Harbour to Lough Mask, the Lettergesh
Lingula-Eocoelia fauna then Pentamerus and finally CIorinda, before thinly Formation is of Sheinwoodian age and has actually yielded Monograptus
bedded graptolitic sands and muds became established in early turriculatus riccartonensis at one level. The formation represents the maximum trans-
(maximus) time. These latter may indeed record some deepening of the sea; gression here. Poorly sorted sandstones with conglomerates, breccias, and
accommodation of sediment here was certainly rapid. Such sedimentation, silty mudstones were deposited from turbidity currents.
considered by Cocks & Toghill to be turbiditic, continued through griesto- CHH
niensis time when purple, red and green mudstones and sandstones were
deposited. Similar conditions at this time are reflected in the other Midland PRiDOLI
Valley inliers with palaeocurrent directions from the south in the Hagshaw
Hills and near Lesmahagow, but in the Pentland Hills, where the Haggis The earlier Silurian initiation of progradation of terrestrial sediments across
Grit and Conglomerate occurs, palaeocurrents came from the east. The marine areas of the British Isles led to the continually progressive and
deposition of this 'conglomerate' may have heralded the Wenlock gravels dominant establishment of non-marine depositional regimes through P~idoli
which record fluvial infill from an emergent Cockburnland to the south. times. By the end of this interval, only present-day areas of southeastern-
Overall it seems unlikely that these Midland Valley reaches of the basin were most England remained open to fully marine influences from the south and
ever 'deep' in the Llandovery. southeast, although much of the former Welsh Borderland-Midlands plat-
The deep parts of the basin continued to be around Pomeroy and form was susceptible to short-lived, periodic flooding by the marginal sea
Lisbellaw where graptolitic muds persisted through sedgwickii and griesto- until well into the Pfidoli. Alluvial processes of erosion and deposition were
niensis times. At Lisbellaw, Harper & Hartley (1938) recorded conglomer- persistent to the west and north, leading to the diachronous spread of Old
ates of mid-Llandovery age and likened them to the Caban Conglomerate of Red Sandstone magnafacies from those general directions. Although the
the same age in mid-Wales. Clearly a basin-margin slope existed to the Iapetus Ocean was probably closed well before the Pfidoli Epoch (e.g.
northwest of Lisbellaw and Pomeroy, for at Charlestown, in Co. Mayo, Picketing et al. 1988), and there is no evidence of any remaining marine
sands and calcareous deposits formed during sedgwickii to crenulata times separation between northern and southern Britain at this time, the precise
accompanied by shelly faunas including corals, brachiopods and trilobites. relative positions of the two regions remain uncertain. Sinistral transpres-
Similai" conditions prevailed in the extreme west around Killary Harbour sional convergence continued throughout the Pfidoli and later (Soper et al.
and Clew Bay where basal gravel deposits ofgriestoniensis age overstepped a 1987), so that it is unlikely that there was any direct north-south relation-
Dalradian and Ordovician substrate. In early and mid Llandovery times this ship and continuation of drainage and depositional systems between the two
area was land and the gravels were interpreted by Piper (1972) as fluvial, areas that are now adjacent on opposite sides of the sutural zone.
with westward followed by southward transport. An extrusive keratophyre Background references to stratigraphical successions of P~idoli age, and
was present in places below the gravel, recording volcanicity probably also their correlation, are given by Bassett et al. (1982), White & Lawson (1989),
14 CAMBRIAN

Massif or on its margins. The Cambrian of Nuneaton accumulated to the rocks encountered in boreholes in Lincolnshire (Cowie et al. 1972, table 2)
southwest of this Charnian Massif. Coastal Hartshill Formation sandstones are varied--phyllitic slates, feldspathic sandstones and greywackes--and a
(cycle l, thick), through to condensed shallow water, shelly Hyolithes Cambrian age has been suggested, but without adequate faunal control. It is
limestones (cycle 2, thin) and deeper water Mn-rich shales (cycle 3, thick) assumed here that this area was part of a marginal basin that existed
exhibit a trend reciprocal with that of Shropshire and close to that of the throughout the early Lower Palaeozoic and extended from northern England
Avalon succession, SE Newfoundland. The nearby Lickey Quartzites are into the Brabant area of Belgium where clastic rocks of Comley and St
lithologically similar but are of uncertain age. The former presence of David's age are known (Vanguestaine 1974, 1978). It is not known whether
limestones is suggested by clasts in the Permian Nechells Breccia (Brasier & the Precambrian of the Charnwood Forest area was emergent; there is some
Hewitt 1981). geophysical evidence to the northeast of Charnwood Forest for the presence
The Withycombe Farm borehole near Banbury, Oxfordshire, gives an of a post-Charnian, pre-Triassic succession that possibly represents part of
indication of conditions to the east of the Malverns. Volcanic basement of the local Cambrian (Maguire 1987). A positive area in northern Norfolk was
presumed late Precambrian age is there overlain by the Withycombe Forma- formerly postulated on the presence of supposedly Precambrian volcanic
tion, a relatively thick sequence of sandstones bearing trace fossils (cf. cycle rocks encountered in the North Creak Borehole, but these are now thought
l) passing up into a thick succession of offshore shales with Aldanella sp., to be part of the east Midlands Caledonian Belt (Pharaoh et al. 1987).
Watsonella sp., Platysolenites antiquissimus, Orthotheca cylindrica and Lon- AWAR
tova-type acritarchs (Rushton & Molyneux 1990). This invites compari-
son with the Chapel Island Formation (cycle 1) or lower Bonavista Forma-
tion (cycle 2) of southeast Newfoundland, both of attleborensis age. 4~2a: Southern British Isles: Merioneth
MDB
In Scandinavia the Merioneth Series has been divided into some eight
E~lb: Southern British Isles: St David's biozones and thirty subzones, and this scheme can be applied in areas, such
as England and Wales, where the olenid trilobite biofacies is developed. The
The St David's Series is equivalent to the Middle Cambrian of European interval depicted on the map is the Olenus Biozone and more particularly its
usage, but not necessarily equivalent to that of other regions such as upper part (O. cataractes Subzone).
Australia, China or North America. The Series is subdivided according to Deposits formed in England and Wales during the Merioneth Epoch
the Swedish zonal scheme, with a lower, a middle and an upper part. In the reflect generally stable tectonic conditions. After a widespread non-sequence
Welsh Basin the lower part is represented mainly by thick turbiditic in the late St David's successions (possibly caused by a eustatic regression), a
sandstone successions; in Shropshire by thinner glauconitic sandstones and major new sedimentary cycle commenced, apparently before the end of the
mudstones, and in the English Midlands by mudstones. Biostratigraphical St David's Epoch, in both the Welsh Basin and the English Midlands
control is generally poor and correlation insecure (Cowie et al. 1972; (Rushton 1978; Allen et al. 1981).
Rushton 1974). The Welsh Basin, though deeper at the beginning of the Merioneth Epoch,
Faunal control is better in the middle part of the Series, where the was partly filled towards the end of Olenus Biozone times with alterations of
following biozones are used: shales and contourite(?) sandstones (upper Maentwrog Formation), and
those beds are overlain by thick shallow-water sandstones (Ffestiniog Flags
Ptychagnostus punctuosus Biozone,
Formation, >650m, Allen et al. 1981). There is some evidence in the
Hypagnostus parvifrons Biozone,
Harlech Dome, St Tudwars Peninsula, and in the St David's area, that the
Tomagnostus fissus Biozone,
sediments were deposited by currents flowing from the south, as in the St
Ptychagnostus gibbus Biozone.
David's Series. To the north of the Harlech Dome, however, the Merioneth
The gibbus Biozone appears to represent a widespread transgression, Series is comparatively attenuated and includes shallow-water quartzitic
possibly related to a eustatic sea-level rise (Rowell et al. 1982, p 176), but sandstones, with trace fossils (Cruziana facies) that appear to have been
that zone is not easily recognized in Britain. The overlyingfissus Biozone is derived from the north, though the indices of current orientation give a wide
more widely recognized and this interval, for which correlation is more range of directions (Crimes 1970).
secure, is represented on the map. Muddy sediments accumulated over the site of the Midlands Platform and
In the Shropshire area glauconitic sandstones and calcareous beds were it is assumed here that these (Outwoods Formation of the Nuneaton Inlier,
deposited in shallow water, and Cobbold (1927) demonstrated a local with known subsurface extension to the south and west) transgressed the
unconformity indicative of uplift along the Church Stretton fault zone at Precambrian Charnwood Forest massif. The fauna of the Outwoods Forma-
about this time. The absence of the St David's Series in the Malvern and tion includes olenid trilobites that tolerated oxygen-poor environments,
Lilleshall areas (Rushton et al. 1988) implies similar uplift, but with less together with a number of other trilobite genera of wide palaeogeographical
certainty. distribution that inhabited outer-shelf regions (Rushton 1983).
The Midlands Platform carries a thin sequence of variegated mudstones In the Welsh Borderland the Olenus Biozone is unrepresented, the earliest
deposited below wave-base (Abbey Shales, three biozones represented, representatives of the Merioneth Series being the Orusia Shales (Parabolina
< 40 m thick in total). The fauna is dominated by agnostid trilobites and spinulosa Biozone) resting unconformably on Comley and St David's rocks.
small brachiopods. Similar but thicker successions (c. 200 m total?) occur However, at Malvern there is a thin sequence of Cyclotron-bearing shales
in southwest Wales and also in North Wales, though there the mudstone that are thought to be of Olenus Biozone age. Uplift along the Church
sequence is underlain by turbiditic sandstones offissus age (the Gamlan and Stretton and Malvern lineaments is suggested by northeastwardly directed
Caered formations) that show evidence of derivation from a southerly slumps or currents in the Moor Wood Flags Formation of the Nuneaton
source (Crimes 1970). This implies a positive area to supply sediment to area (Taylor & Rushton 1972, p. 22). The derivation ofmicaceous detritus in
north Wales whilst by-passing the St David's area in the southwest. The the Llangynog area, south Wales, is believed to be from a metamorphic
presence of bentonite beds indicates contemporaneous volcanic activity, basement (Pretannia) to the south (Cope & Bassett 1987).
though the location of the activity is unknown. The later Merioneth Epoch, from Parabolina spinulosa Biozone to Peltura
There is no evidence for the St David's Series north of the Harlech Dome scarabaeoides Biozone times, was dominated by deposition of black mud-
and St Tudwal's Peninsula, and it is assumed that the Irish Sea Horst was a stones in disaerobic environments (Doigeilau, Bentleyford, Monks Park and
positive area; nevertheless, no contemporaneous sediments are known to White-Leaved-Oak formations). These pyritic, carbonaceous and uranifer-
have been derived thence. ous muds contain little or no coarse clastic material and are considered to
The non-agnostid trilobites most typical of this epoch, Paradoxides and represent a period of slow deposition; the proved olenid zones differ from
blind benthic forms (Hartshillia, Holocephalina) characterize outer-shelf locality to locality (Thomas et al..1984, p. 15), and the extent of possible
environments in Wales and elsewhere in Avalonia (southeast Newfound- breaks in deposition, comparable with those inferred in the contempor-
land, New Brunswick) and in Baltica and the margins of Gondwana aneous rocks in Scandinavia, is not certain. The black mudstones in the
(Conway Morris & Rushton 1988). Croft Borehole, Lilleshall (northern Shropshire), are unique in having many
After fissus Biozone times, mudstone deposition prevailed (except locally intercalations of current-bedded siltstone and sandstone, presumably of
in the Comley area of Shropshire). There is a widespread hiatus after shallow-water origin (Rushton et al. 1988). The source of the silt-grade
punctuosus Biozone times and before the Merioneth Epoch, the mudstone material is not known but is thought to originate from a local uplift along
succession in the Nuneaton district being the best constrained (Taylor & the Church Stretton Fault complex.
Rushton 1972). The sedimentary rocks of the southeast Irish marginal basin have poor
In southeast Ireland the marginal basin of Leinster has yielded some biostratigraphical control but include beds referable to the Merioneth Series
evidence of the St David's Series (Smith 1977, 1981). In Co Wexford (the (Ribband Group; Smith 1977, 1981). They are argillites and arenites that
Cahore and Ribband Groups) and Co Wicklow (the Bray Group), the appear to have been deposited in deep water. The mudstones and fine-
sediments were deposited in deep water (Crimes & Crossley 1968). These grained sandstones of the Askingarren Formation, interpreted as underlying
rocks yield trace-fossils, especially Oldhamia, in abundance (Dhonau & correlatives of the Ribband Group (Crimes & Crossley 1968, pp. 191,207),
Holland 1974), but acritarchs, in part of St David's age, are the only body- are considered as possibly late Cambrian in age. Flute casts indicate
fossils known; the best preserved are from the Booley Bay Formation derivation from a southerly quarter. There is no evidence for the continued
(Ribband Group) at Hook Head on the south coast (Smith 1981). Sediment existence, into the Merioneth Epoch, of the marginal arc thought to have
was derived from both the southeast (Irish Sea Horst) and the northwest, existed to the northwest of the southeast Irish marginal basin in St David's
where a positive area, possibly a marginal arc, is thought to have provided a times.
source area. There is no biostratigraphical evidence for the Merioneth Series in the
Less is known of the eastern Caledonide belt to the northeast of the eastern English Caledonides (see the St David's Series, above).
Midland Platform and facing the Tornquist Sea. The clastic sedimentary AWAR
52 SILURIAN

Cocks et al. (1971), and House et al. (1977). The first two of these Group and its correlatives, somewhat below the level of the Townsend Tuff;
publications also give details of environmental developments, especially for the latter horizon forms a useful lithostratigraphical datum across much of
the earlier P~idoli, so that only subsequent or additional sources of reference southern Wales and the Welsh Borderland (Allen & Williams 1981, 1982),
are quoted here. lying within the upper part of the Pfidoli rather than at the base of the
MGB Devonian (Bassett 1984, p. 301; cf. Allen & Williams 1981).
Periodic marine incursions continued to influence sedimentation across
S8b: Southern British Isles: earliest Pfidoli much of southern England and Wales throughout mid P[idoli times. Spora-
dic accumulations of bone bed strands that persist well into the Downton
Across the Welsh Borderland-Midlands platform and into eastern to south- Group (e.g. Temeside Bone Bed; Antia 1981) are absent in the Ledbury
central Wales the correlation of early P~idoli strata is reasonably well Formation, where sediments are dominated by thick mudstones with fine
constrained by the presence of an ostracode assemblage that includes siltstones and sandstones. Lingulacean brachiopods and bivalve molluscs
Frostiella groenvalliana and species of Londinia and Nodibeyrichia, together (e.g. Allen 1973) reflect the marginal marine episodes of flooding across
with conodonts and thelodonts that are also of limited vertical range. The extensive, featureless mudflats that were partly intertidal and partly river
accompanying presence of relict Ludlow shelly macrofaunas testifies to a influenced (Allen 1985). Drifted plant debris and articulated vertebrates
continuing marginal marine influence, whilst at the same time the increased become increasingly common upwards through the sequence. Widespread
occurrence of freshwater fishes, vascular land plants, and spores, accom- developments of pedogenic carbonates (calcretes) indicate the formation of
panied by a sharp fall in the earlier dominance of marine microplankton stable geomorphic surfaces that were starved of fluvial input for periods of
(e.g. acritarchs) is indicative of the pulsatory encroachment of closely some 104 years (Allen 1974), and they also suggest a relatively hot climate
adjacent land areas. The transition from fully marine through quasi-marine with comparatively low seasonal rainfall. Deposition across the alluvial
facies during this interval went hand in hand with the inversion of the plains was cyclical, with repeated incursions of gravels both from major
depositional basin, which had commenced similarly in earlier Silurian times stream systems derived from distant uplands and also from minor systems of
and was to reach a climax in Acadian shortening during the early-mid ephemeral drainage developed on slightly elevated interfluves between the
Devonian. principal rivers (Allen 1964; Allen & Williams 1979). These later Downton
Following latest Ludlow regressive shallowing, the Ludlow Bone Bed Group sediments thus record a considerable enlargement of quasi-marine to
Member at the base of the Downton Group in the Welsh Borderland non-marine environments established in earlier Pfidoli times (Allen 1985,
represents an initial brief and temporary interval of renewed shallow marine p. 94).
transgression, in which the bone accumulations formed lag concentrates To the north of the former Welsh Basin there is no evidence of continued
distributed widely across retreating strands. Local evidence of hummocky marine connection northwards. The Traeth Bach Beds of Anglesey, possibly
cross stratification in immediately overlying Downton beds suggests that of mid Pfidoli age, include calcretes within arenites and rudites that may
storm processes were active in shoreface environments showing little or no have been derived in part from local fault scarps to the southeast (Allen
evidence of tidal influence, in water depths of only a few metres (Smith & 1965), although the regional palaeoslope and drainage pattern was fairly
Ainsworth 1989); reduction in salinity is also inferred from the accom- certainly persistently from the northwest. Across southwest Wales the
panying restriction of faunas. Slightly higher beds in the Downton Castle Pfidoli alluvial systems developed partly in a major west-east aligned valley
Sandstone Formation across the platform are dominantly fine sandstones complex (Allen & Williams 1982; Allen 1985, fig. 3), with sediment input
with interbedded siltstones and mudstones; the facies are probably subtidal, from both the north and south. The whole of Ireland was probably by now
passing yet higher into thicker sand facies with structures indicative of the site of continuous alluvial red bed facies and erosive systems, with
vigorous wave action in intertidal and inshore zones (Allen 1985, p. 92). deposition of the Dingle Group in the southwest continuing throughout this
These complexes were deposited as early Pfidoli sand shoals that occupied a interval (Holland 1979; Doran et al. 1973; Horne 1974).
minimum area of some t2 000 km 2, and the diversity of transport directions The only evidence of more open marine conditions in the post-early
may point to more than one source area. Local easterly coarsening in the Pfidoli of Britain is across southern England, where micaceous sandstones
Midlands (Gornal Sandstone facies) suggests near shore environments with and siltstones in the subsurface at Little Missenden formed probably as
derivation from the east or southeast. Other sources on the platform suggest subtidal sand sheets with a restricted shelly fauna. Similar sediments occur at
westerly transport vectors, possibly in part from local highs in the remnant Lakenheath and Soham, although there are no faunal/floral data to confirm
Welsh trough (Allen 1985, fig. 3), but probably too from yet farther to the the age. As in the earlier P?idoli, an open marine connection to the southeast
west and northwest where direct evidence of P~idoli beds is lacking; how- is indicated. By the end of Pfidoli times the marginal marine influence across
ever, there is no reason to doubt that areas such as the Irish Sea platform the former Welsh trough and its bounding platform areas had ceased, and
were positive suppliers of sediment at this time. Although impersistent bone instead fluvial sedimentation then became ubiquitous.
beds are present locally along the eastern margins of the Welsh trough (e.g. In northern Britain, biostratigraphical data that permit precise recog-
Holland & Williams 1985) they are absent in the central axis where nition of Pfidoli ages in successions are even more sparse than to the south.
deposition was apparently continuous from the Ludlow to the Downton and However, recent geochronological studies (Thirlwall 1988) have radically
where non-marine facies did not become dominant finally until slightly later changed the stratigraphical assignment of some of the Old Red Sandstone
than on the bounding platform areas. In immediately post-groenvalliana deposits in Scotland; a good deal of the red terrestrial sediment assumed
Biozone times, the sea in the Welsh area had probably shrunk to a small previously to be of Devonian age is now known to be Silurian, and some
central remnant, with tidal sand shoals and extensive mud fiats to the east. basins active in mid P~'idoli times were initiated late in the Wenlock.
At the southern margin of the Welsh trough, early Pfidoli highly mica- Whilst most of the Grampian block has cooling ages of >425 Ma, rocks
ceous, coarse shelly arenites of the Long Quarry Formation (Tilestones) alongside the Great Glen Fault have cooling ages going down to 385 Ma,
represent high energy, open shoreface and barrier sands derived from and here as well as in the northern Highlands there are many cooling ages
alluvial plains developed on the planated metamorphic basement of between 420 and 400 Ma, suggesting that the mid Pi'idoli was a time of
Pretannia to the south (Cope & Bassett 1987). Old Red Sandstone facies substantial uplift in both these areas. In northern Scotland there was uplift
of southwest Wales (low-middle Milford Haven Group) accumulated on along the Moine Thrust in early Ludlow times, so that most of the ground to
coastal mud fiats flooded periodically by the sea, but at times scoured by the east of that structure was then undergoing elevation and must have
rivers and with local alluvial fans fed from either the south or southeast; provided a substantial source of sediment for any basins that may have been
reworking of older Silurian sources and of basement complexes were both forming contemporaneously.
involved (Allen & Williams 1978). Yet farther to the west, in Ireland, there is In addition, numerous granitoid intrusions are recorded from the
only limited evidence for P~idoli beds, but terrestrial red bed facies fairly Dalradian block to the southeast of the steep belt. These are thought to be
certainly persisted throughout the epoch (e.g. Holland 1979; Doran et al. high-level plutons that may have been associated with lavas. One plutonic
1973). The thick Dingle Group of southwest Ireland, comprising coarse complex occurs within the steep belt and near an area of uplift at Findhorn,
arenites and rudites of alluvial origins, is apparently conformable on Ludlow and far in the southwest the Ross of Mull granite is of Pi'idoli age. There are
beds (Holland 1987). a number of intrusions within the Midland Valley, including Distinkhorn,
Direct marine connections northwards from the Wales-Welsh Borderland Tinto, and diorite intrusions in the Ocl:fil Hills. Contemporaneous lavas are
area probably persisted in earliest P~idoli times to the Lake District, where also dated from the Pentland Hills and from Fife, so the Midland Valley is
the lower Scout Hill Flags bear a groenvalliana Biozone fauna in siltstones, seen as a magmatic province with lavas > 3 km thick. Although there are
mudstones and fine sandstones derived in part from the north. Similar lava flows over much of the Midland Valley, two major centres are
definitive evidence of age is lacking in areas to the east and southeast of the developed, in the OchiI-Sidlaw Hills and in the Pentlands; these lavas,
Midlands platform, although limited data from the subsurface suggests that accompanied by agglomerates and tufts, range from rhyolites, dacites and
marginal marine sedimentation persisted there as shoals and bars, with more andesites to basalts.
open marine environments beyond, across southeasternmost England. Intrusive rocks also occur along the northern edge of the Southern
MGB Uplands at Carsphairn, Cockburn Law, and Priestlaw, although it is not
known whether they were accompanied by lavas.
$9: All British Isles: mid Pfidoli The earliest sedimentary basin in the north is at the southwest end of the
Great Glen Fault near Oban. This basin originated in the late Wenlock with
Biostratigraphical data for identifying younger P~'idoli strata across the an early fill of coarse clastic sediment. By mid Pfidoli times the whole basin
British Isles are sparse, although limited evidence is available from fish was covered with lavas of the Lorne plateau, although these are mainly or
faunas and there is growing potential from the use of sporomorphs (e.g. wholly of Ludlow age.
Richardson & McGregor 1986; Richardson & Edwards 1989). Mid Pfidoli In the Midland Valley, palynological data from the Stonehaven Group
equivalents fall within sporomorph Biozone A (see Richardson & Edwards indicate that the beds are probably of late Wenlock age (Marshall 1989) and
1989, fig. 152), taken here within the Ledbury Formation of the Downton not within the Pfidoli or younger as assumed previously. There is probably a
53
54 SILURIAN

stratigraphical break between these beds and overlying units of the Lower BAKER,J. W. & HUGHES,C. P. 1979. Summer (1973) Field Meeting in Central Wales,
Old Red Sandstone (e.g. D u n n o t t a r and Crawton groups), which may well 31 August to 7 September 1973. Proceedings of the Geologists" Association, 90,
span all or part of the Pfidoli Series, although no precise dating is yet 65-79.
BASSETT,M. G. 1974. Review of the stratigraphy of the WeDlock Series in the Welsh
available. These post-Stonehaven units contain boulder-bearing conglomer- borderland and South Wales. Palaeontology, 17, 745-777.
ates tfiat were discharged from the northeast. The conglomerates contain - - 1984. Lower Palaeozoic Wales--a review of studies in the past 25 years.
clasts of quartzite, greywacke, a variety of granites, and other intrusives-- Proceedings of the Geologists'Association, 95, 291-31 I.
but almost always present are the abundant and varied lava clasts. The - - 1989. The WeDlock Series in the WeDlock area. In: HOLLAND,C. H. & BASSETT,
quartzites and some of the porphyries are well rounded, the former being M. G. (eds) A Global Standard for the Silurian System, National Museum of
Wales, Geological Series No. 9, Cardiff, 51-73.
polycyclic. Lavas occur within the sequence, one being associated with - - , COCKS, L. R. M., HOLLAND,C. H., RtCKARDS,R. B. & WARREN,P. T. 1975.
slumping and possible contemporaneous faulting; others form useful strati- The o,pe WeDlock Series. Report of the Institute of Geological Sciences, No. 75/
graphical markers, as at Crawton. Haughton (1988) has shown that some 13.
conglomerates of this age were derived from the southeast and are typified - - , LAWSON,J. D. & WHITE,D. E. 1982. The Downton Series as the fourth Series
by the presence of a b u n d a n t greywacke clasts and granites, whose age and of the Silurian System. Lethaia, 15, 1-24.
BENTON, M. J. & GRAY, D. I. 1981. Lower Silurian distal shelf storm-induced
chemistry differs from those derived from the northwest and northeast; this
turbidites in the Welsh Borders: sediments, tool marks and trace fossils. Journal
indicates that the source was a previously unsuspected greywacke pile (of of the Geological Society, London, 138, 675-694.
Ordovician age) in the centre of the Midland Valley. BLVCK, B. J. 1983. Role of the Midland Valley of Scotland in the Caledonian
Elsewhere in the northern region of the Midland Valley there are orogeny. Transactions of the Royal Society of Edinburgh: Earth Sciences, 74,
abundant lavas and lava-bearing conglomerates. Some of these overlie the 119-136.

BOTT, M. H. P. 1967. Geophysical investigations of the northern Pennine basement


Lintrathen ignimbrite and could well be of mid Pfidoli age. Intrusions such
rocks. Proceedings of the Yorkshire Geological Society, 36, 139-168.
as those at Comrie may have been associated with lavas at higher structural BRADFIELD, K. E. S. 1987. Late Ludlovian and early Pfldoli palaeoenvironments of
levels. mid- Wales and the Welsh Borderland. PhD thesis, University of London.
Basins were also developing in the southern Midland Valley at this time. BRENCHLEY, P. 1984. Late Ordovician extinctions and their relationship to the
The distinctive Greywacke Conglomerate, with clasts dominated by grey- Gondwana glaciation. In: BRENCHLEY, P. (ed.) Fossils and Climate. Wiley,
wacke, is cut by felsites and also contains felsite clasts; both felsites resemble Chichester.
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that at Tinto, so the conglomerate is thought to be contemporaneous. The British Caledonides--Vl. Regional and Global Implications. Geophysical Jour-
greywacke clasts do not have a composition compatible with a source in the nal of the Royal Astronomical Society, 34, 107-134.
Southern Uplands, and often show a dispersal direction towards the south- BRIDGES, P. H. 1975. The transgression of a hard substrate shelf: the Llandovery
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- - , COLTHURSr,J. R. J., FELLY,M., GARDINER,P. R. R., PeNNEr, S. R., REEVES,
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Bailey, K. E. S. Bradfield, L. Cherns, L. R. M. Cocks, J. R. Davies, R. Evans, C. J. BURGESS, I. C. & HOLLIDAY,D. W. 1979. Geology of the country around Brough-
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Utrecht.
14 CAMBRIAN

Massif or on its margins. The Cambrian of Nuneaton accumulated to the rocks encountered in boreholes in Lincolnshire (Cowie et al. 1972, table 2)
southwest of this Charnian Massif. Coastal Hartshill Formation sandstones are varied--phyllitic slates, feldspathic sandstones and greywackes--and a
(cycle l, thick), through to condensed shallow water, shelly Hyolithes Cambrian age has been suggested, but without adequate faunal control. It is
limestones (cycle 2, thin) and deeper water Mn-rich shales (cycle 3, thick) assumed here that this area was part of a marginal basin that existed
exhibit a trend reciprocal with that of Shropshire and close to that of the throughout the early Lower Palaeozoic and extended from northern England
Avalon succession, SE Newfoundland. The nearby Lickey Quartzites are into the Brabant area of Belgium where clastic rocks of Comley and St
lithologically similar but are of uncertain age. The former presence of David's age are known (Vanguestaine 1974, 1978). It is not known whether
limestones is suggested by clasts in the Permian Nechells Breccia (Brasier & the Precambrian of the Charnwood Forest area was emergent; there is some
Hewitt 1981). geophysical evidence to the northeast of Charnwood Forest for the presence
The Withycombe Farm borehole near Banbury, Oxfordshire, gives an of a post-Charnian, pre-Triassic succession that possibly represents part of
indication of conditions to the east of the Malverns. Volcanic basement of the local Cambrian (Maguire 1987). A positive area in northern Norfolk was
presumed late Precambrian age is there overlain by the Withycombe Forma- formerly postulated on the presence of supposedly Precambrian volcanic
tion, a relatively thick sequence of sandstones bearing trace fossils (cf. cycle rocks encountered in the North Creak Borehole, but these are now thought
l) passing up into a thick succession of offshore shales with Aldanella sp., to be part of the east Midlands Caledonian Belt (Pharaoh et al. 1987).
Watsonella sp., Platysolenites antiquissimus, Orthotheca cylindrica and Lon- AWAR
tova-type acritarchs (Rushton & Molyneux 1990). This invites compari-
son with the Chapel Island Formation (cycle 1) or lower Bonavista Forma-
tion (cycle 2) of southeast Newfoundland, both of attleborensis age. 4~2a: Southern British Isles: Merioneth
MDB
In Scandinavia the Merioneth Series has been divided into some eight
E~lb: Southern British Isles: St David's biozones and thirty subzones, and this scheme can be applied in areas, such
as England and Wales, where the olenid trilobite biofacies is developed. The
The St David's Series is equivalent to the Middle Cambrian of European interval depicted on the map is the Olenus Biozone and more particularly its
usage, but not necessarily equivalent to that of other regions such as upper part (O. cataractes Subzone).
Australia, China or North America. The Series is subdivided according to Deposits formed in England and Wales during the Merioneth Epoch
the Swedish zonal scheme, with a lower, a middle and an upper part. In the reflect generally stable tectonic conditions. After a widespread non-sequence
Welsh Basin the lower part is represented mainly by thick turbiditic in the late St David's successions (possibly caused by a eustatic regression), a
sandstone successions; in Shropshire by thinner glauconitic sandstones and major new sedimentary cycle commenced, apparently before the end of the
mudstones, and in the English Midlands by mudstones. Biostratigraphical St David's Epoch, in both the Welsh Basin and the English Midlands
control is generally poor and correlation insecure (Cowie et al. 1972; (Rushton 1978; Allen et al. 1981).
Rushton 1974). The Welsh Basin, though deeper at the beginning of the Merioneth Epoch,
Faunal control is better in the middle part of the Series, where the was partly filled towards the end of Olenus Biozone times with alterations of
following biozones are used: shales and contourite(?) sandstones (upper Maentwrog Formation), and
those beds are overlain by thick shallow-water sandstones (Ffestiniog Flags
Ptychagnostus punctuosus Biozone,
Formation, >650m, Allen et al. 1981). There is some evidence in the
Hypagnostus parvifrons Biozone,
Harlech Dome, St Tudwars Peninsula, and in the St David's area, that the
Tomagnostus fissus Biozone,
sediments were deposited by currents flowing from the south, as in the St
Ptychagnostus gibbus Biozone.
David's Series. To the north of the Harlech Dome, however, the Merioneth
The gibbus Biozone appears to represent a widespread transgression, Series is comparatively attenuated and includes shallow-water quartzitic
possibly related to a eustatic sea-level rise (Rowell et al. 1982, p 176), but sandstones, with trace fossils (Cruziana facies) that appear to have been
that zone is not easily recognized in Britain. The overlyingfissus Biozone is derived from the north, though the indices of current orientation give a wide
more widely recognized and this interval, for which correlation is more range of directions (Crimes 1970).
secure, is represented on the map. Muddy sediments accumulated over the site of the Midlands Platform and
In the Shropshire area glauconitic sandstones and calcareous beds were it is assumed here that these (Outwoods Formation of the Nuneaton Inlier,
deposited in shallow water, and Cobbold (1927) demonstrated a local with known subsurface extension to the south and west) transgressed the
unconformity indicative of uplift along the Church Stretton fault zone at Precambrian Charnwood Forest massif. The fauna of the Outwoods Forma-
about this time. The absence of the St David's Series in the Malvern and tion includes olenid trilobites that tolerated oxygen-poor environments,
Lilleshall areas (Rushton et al. 1988) implies similar uplift, but with less together with a number of other trilobite genera of wide palaeogeographical
certainty. distribution that inhabited outer-shelf regions (Rushton 1983).
The Midlands Platform carries a thin sequence of variegated mudstones In the Welsh Borderland the Olenus Biozone is unrepresented, the earliest
deposited below wave-base (Abbey Shales, three biozones represented, representatives of the Merioneth Series being the Orusia Shales (Parabolina
< 40 m thick in total). The fauna is dominated by agnostid trilobites and spinulosa Biozone) resting unconformably on Comley and St David's rocks.
small brachiopods. Similar but thicker successions (c. 200 m total?) occur However, at Malvern there is a thin sequence of Cyclotron-bearing shales
in southwest Wales and also in North Wales, though there the mudstone that are thought to be of Olenus Biozone age. Uplift along the Church
sequence is underlain by turbiditic sandstones offissus age (the Gamlan and Stretton and Malvern lineaments is suggested by northeastwardly directed
Caered formations) that show evidence of derivation from a southerly slumps or currents in the Moor Wood Flags Formation of the Nuneaton
source (Crimes 1970). This implies a positive area to supply sediment to area (Taylor & Rushton 1972, p. 22). The derivation ofmicaceous detritus in
north Wales whilst by-passing the St David's area in the southwest. The the Llangynog area, south Wales, is believed to be from a metamorphic
presence of bentonite beds indicates contemporaneous volcanic activity, basement (Pretannia) to the south (Cope & Bassett 1987).
though the location of the activity is unknown. The later Merioneth Epoch, from Parabolina spinulosa Biozone to Peltura
There is no evidence for the St David's Series north of the Harlech Dome scarabaeoides Biozone times, was dominated by deposition of black mud-
and St Tudwal's Peninsula, and it is assumed that the Irish Sea Horst was a stones in disaerobic environments (Doigeilau, Bentleyford, Monks Park and
positive area; nevertheless, no contemporaneous sediments are known to White-Leaved-Oak formations). These pyritic, carbonaceous and uranifer-
have been derived thence. ous muds contain little or no coarse clastic material and are considered to
The non-agnostid trilobites most typical of this epoch, Paradoxides and represent a period of slow deposition; the proved olenid zones differ from
blind benthic forms (Hartshillia, Holocephalina) characterize outer-shelf locality to locality (Thomas et al..1984, p. 15), and the extent of possible
environments in Wales and elsewhere in Avalonia (southeast Newfound- breaks in deposition, comparable with those inferred in the contempor-
land, New Brunswick) and in Baltica and the margins of Gondwana aneous rocks in Scandinavia, is not certain. The black mudstones in the
(Conway Morris & Rushton 1988). Croft Borehole, Lilleshall (northern Shropshire), are unique in having many
After fissus Biozone times, mudstone deposition prevailed (except locally intercalations of current-bedded siltstone and sandstone, presumably of
in the Comley area of Shropshire). There is a widespread hiatus after shallow-water origin (Rushton et al. 1988). The source of the silt-grade
punctuosus Biozone times and before the Merioneth Epoch, the mudstone material is not known but is thought to originate from a local uplift along
succession in the Nuneaton district being the best constrained (Taylor & the Church Stretton Fault complex.
Rushton 1972). The sedimentary rocks of the southeast Irish marginal basin have poor
In southeast Ireland the marginal basin of Leinster has yielded some biostratigraphical control but include beds referable to the Merioneth Series
evidence of the St David's Series (Smith 1977, 1981). In Co Wexford (the (Ribband Group; Smith 1977, 1981). They are argillites and arenites that
Cahore and Ribband Groups) and Co Wicklow (the Bray Group), the appear to have been deposited in deep water. The mudstones and fine-
sediments were deposited in deep water (Crimes & Crossley 1968). These grained sandstones of the Askingarren Formation, interpreted as underlying
rocks yield trace-fossils, especially Oldhamia, in abundance (Dhonau & correlatives of the Ribband Group (Crimes & Crossley 1968, pp. 191,207),
Holland 1974), but acritarchs, in part of St David's age, are the only body- are considered as possibly late Cambrian in age. Flute casts indicate
fossils known; the best preserved are from the Booley Bay Formation derivation from a southerly quarter. There is no evidence for the continued
(Ribband Group) at Hook Head on the south coast (Smith 1981). Sediment existence, into the Merioneth Epoch, of the marginal arc thought to have
was derived from both the southeast (Irish Sea Horst) and the northwest, existed to the northwest of the southeast Irish marginal basin in St David's
where a positive area, possibly a marginal arc, is thought to have provided a times.
source area. There is no biostratigraphical evidence for the Merioneth Series in the
Less is known of the eastern Caledonide belt to the northeast of the eastern English Caledonides (see the St David's Series, above).
Midland Platform and facing the Tornquist Sea. The clastic sedimentary AWAR
56 SILURIAN

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