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An overview of the status and abundance of migratory waders in Sumatra,


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Stilt 50 (2006): 90–95 Migratory waders in Sumatra

AN OVERVIEW OF THE STATUS AND ABUNDANCE OF MIGRATORY WADERS IN


SUMATRA, INDONESIA

ANDREW C. CROSSLAND1, SULE A. SINAMBELA2, ANDY S. SITORUS3 & ARI W. SITORUS4


1
34 Chichester Street, Woolston, Christchurch 8006, New Zealand; Andrew.Crossland@ccc.govt.nz. 2Jalan. Mual Nauli IV,
No.16 Siopat Suhu, Pemantang Siantar, Sumatera Utara 21115, Indonesia. 3,4 Universitas Teknologi Yogyakarta, D.I.
Yogyakarta, Java, Indonesia

This paper provides a summary of the status and abundance of 37 species of migratory wader recorded in Sumatra,
western Indonesia. High numbers, totalling tens of thousands of birds occur along the eastern coastline of Sumatra
in the provinces of North Sumatra, Riau, Jambi, South Sumatra and Lampung. Smaller numbers occur in the
northern province of Aceh and along the west coast. Wader species occurring in thousands include Lesser Sand
Plover, Greater Sand Plover, Asian Dowitcher, Black-tailed Godwit, Bar-tailed Godwit, Eurasian Curlew,
Common Redshank, Terek Sandpiper and Curlew Sandpiper. Species found in hundreds include Pacific Golden
Plover, Grey Plover, Pintail Snipe, Whimbrel, Eastern Curlew, Marsh Sandpiper, Wood Sandpiper, Great Knot and
Red Knot. Sumatra is both a terminus for some migrants, as well as a passage zone for waders migrating towards
eastern Indonesia and probably Australia. The eastern coastline of Sumatra and adjacent islands combine with the
west coast of Peninsular Malaysia to create one of the most important areas for waders and other shorebirds in
South-East Asia.

INTRODUCTION most areas by mangrove forest, nipah swampland or,


following land reclamation, aquaculture ponds. North of
The Indonesian archipelago comprises 13,600 islands and Lake Toba, coastal rivers tend to develop extensive deltaic
forms a 5,000 km wide barrier between South-East Asia and river mouths with associated mangrove forests. East of Lake
Australasia. The coastline of this archipelago, as well as its Toba, rivers tend to carry a larger volume of coarser
inland wetlands and extensive areas of human-created rice sediments resulting in the prevalence of sandy beach and
fields support large populations of native and migratory sand spit development between the mudflat and mangrove
waterbirds. Although vast numbers of migratory waders are zones. Further south, the first of the major river estuary
assumed to overfly or transit through Indonesia on migration systems starts with the Asahan, Kuala, Bila and Barumun
(Lane 1987), many thousands more spend the northern Rivers, culminating with the Rokan River. The mouths of
winter on the archipelago's shores. these rivers are characterised by expansive areas of intertidal
The island of Sumatra (476,000 km2) straddles the mudflat, ranging in width from 500 m to upwards of 10 km.
equator at the western end of the Indonesian archipelago The coastline of south-eastern Sumatra (facing the
(Figure 1); it comprises both a passage zone for waders southern part of the Strait of Malacca, the South China Sea
migrating between northern Asia and Australasia, as well as and the Java Sea) has a complex mix of tidal cycles
a terminus for species that spend the non-breeding season in (including mixed, semi-diurnal and diurnal) and generally
the tropics. This overview summarises what is known about low-energy wave environments (Whitten et al. 2000). The
the status and abundance of migratory waders in Sumatra. mainland coastline comprises a sequence of large
This information is collated from a number of sources but embayments punctuated by major drainage systems entering
mainly derives from a decade of field observations by the the sea either via extensive deltas or through single channel
authors in north-east Sumatra, and from the published results (Amazon-like) river estuaries. The largest rivers such as the
of survey work carried out in south-east Sumatra during the Musi, Banyuasin, Bantang Hari, Indragiri and Kampar carry
1980s and 90s – principally by Silvius (1988), Verheught et enormous volumes of sediment to the coast, resulting in high
al. (1990, 1993) and Parrott & Andrew (1996). Little is rates of accretion, the development of expansive mudflats
known of wader concentrations along the west coast of and creation of mangrove-covered islands. Behind the
Sumatra as, to date, minimal survey work has taken place accreting coastline lies a wide alluvial plain with zones of
there. However the existence of large concentrations is mangrove, swampland and peat forest, in places extending
unlikely because the extent of intertidal mudflats is limited. up to 100 km inland. Large barrier islands line the coastlines
of southern Riau and South Sumatra Provinces, while the
Coastal Wader Habitats coastline of Lampung consists mainly of open shorelines.
The coastline of north-eastern Sumatra (facing the Andaman Compared with the east coast, the western coastline of
Sea and the northern part of the Strait of Malacca) is Sumatra offers limited wader habitat. The west coast is
characterised by a semi-diurnal tidal cycle and generally characterised by steep hill country, a narrow coastal plain
low-energy wave environments (Whitten et al. 2000). This, and a shoreline exposed to the high energy waves of the
combined with high loads of suspended sediments Indian Ocean. Much of the west coast comprises beaches and
transported by rivers, has allowed the development of cliffs, with mudflat habitat limited to a small number of
extensive soft mudflats along open shorelines, backed in embayments and river mouths.

90
Stilt 50 (2006): 90–95 Migratory waders in Sumatra

Figure 1. Sumatra, showing geographical regions referred to in text.

Inland Wader Habitats progressively surveyed the eastern coastline of North


Sumatra Province from Belawan in the north to the Asahan
Inland wader habitats include the banks of large rivers, lake
Rivermouth in the south.
shores, water reservoirs, aquaculture ponds, freshwater
Other information on wader numbers and distribution in
swamps and rice fields. The last is the most significant with
Sumatra (including limited survey data from the west coast
tens of thousands of hectares under cultivation across
of Sumatra) comes from the Asian Waterbird Census (Lopez
Sumatra. Although difficult to survey, rice fields are known
& Mundkur 1997; Li & Mundkur 2004), from the Sumatra
to support sizeable wader populations, especially of snipe
Bird Report periodically published in Kukila (Holmes 1996),
and some of the sandpipers and stints.
and from accounts in various regional field guides.
PREVIOUS SURVEYS
SPECIES ACCOUNTS
South-eastern Sumatra has had the best survey coverage,
“Summer” and “winter” in these accounts refer to northern
mainly as a result of fieldwork by M. Silvius and colleagues
hemisphere seasons. The summer months, June to August,
who completed wader surveys along the eastern coastlines of
refer to the northern hemisphere breeding season. The other
Riau, Jambi and South Sumatra provinces over the period
months comprise the non-breeding season; winter refers to
1984 to 1986 (Silvius 1988). Subsequently, Verheught et al.
that part of the non-breeding season in which there is no
(1990, 1993) carried out aerial and ground surveys of the
migration, usually December to February.
South Sumatra Province tidal lowlands and floodplains in
1988 and 1989. Milton (1985) and Parrott & Andrew (1996) Oriental Pratincole Glareola maldivarum
reported on wader observations made at Way Kambas
A rare passage wader in the north-east (Crossland in prep.);
National Park in Lampung Province in 1985, and in 1988
an uncommon migrant and winter visitor elsewhere,
and 1989, respectively. Crossland & Sinambela (2005)
including islands (MacKinnon & Phillipps 1993). Flocks of
reported on a survey of waders on Batam Island in the Riau
up to 60 observed in the north-east (Crossland in prep.); up
Archipelago in 2001 and 2002.
to 87 in the south-east (Silvius 1988) and 24 in the west
Wader populations in north-eastern Sumatra have
(Holmes 1996).
received much less survey coverage. Crossland (2000)
reported on wader populations observed at three coastal Pacific Golden Plover Pluvialis fulva
wetlands in the northern tip of Aceh Province in December
1995. Over the period 1994–2006 the authors have A common migrant with flocks of low hundreds in the north-
east, with highest numbers occurring in winter (Crossland in

91
Stilt 50 (2006): 90–95 Migratory waders in Sumatra

prep.). This species is scarce in the south-east (Silvius 1988) required to clarify the status and distribution of Common
but common in the Riau archipelago with highest numbers Snipe in Sumatra.
recorded during southward migration (Crossland &
Asian Dowitcher Limnodromus semipalmatus
Sinambela 2005). Pacific Golden Plovers are found mainly
on coastal sites, but also occur inland up to 1400 m a.s.l. An abundant and widespread migrant with flocks of up to
(Holmes 1996). 7000 recorded in the north-east (Crossland in prep.) and up
to 12,000 recorded in the south-east (Verheught et al. 1990,
Grey Plover Pluvialis squatarola
1993). Distribution and abundance insufficiently
Slightly less common but more widespread than preceding documented for the west. The most abundant wader species
species in the north-east. Low numbers during southern at a number of sites, including Pantai Sejara in North
migration; highest numbers during northward migration with Sumatra Province (Crossland in prep.) and Way Kambas in
flocks of up to 200 (Crossland in prep.). Much more Lampung Province (Parrott & Andrew 1996). The east coast
common than P. fulva in the south-east. Highest numbers of Sumatra is the principal wintering ground for this species.
observed during southward migration with a moderate influx The current estimated world population of 23,000 (Wetlands
during northward migration (Silvius 1988, Verheught et al. International 2006) is likely to need revision upwards once
1990). full survey coverage of Sumatran coastal wetlands is
achieved.
Little Ringed Plover Charadrius dubius
Black-tailed Godwit Limosa limosa
Relatively common in the Aceh area of the north-east
(Crossland 2000), but generally scarce elsewhere (Strange Generally uncommon during southward migration and
2001, Crossland & Sinambela 2005). winter in the north-east but a strong passage occurs during
northward migration, especially in March and April, with
Kentish Plover Charadrius alexandrinus
flocks of up to 3800 recorded (Crossland in prep). A quite
An uncommon visitor on all coasts (Parrott & Andrew 1996, different seasonal pattern has been observed in the south-east
Strange 2001). Sumatra is outside the principal wintering with Black-tailed Godwit being one of the most abundant
range for this species. species year-round. Single flocks of up to 25,000 recorded
during southward migration in South Sumatra Province
Lesser Sand Plover Charadrius mongolus
(Verheught et al. 1990). High numbers (thousands) have
An abundant and widespread migrant to mudflats, been recorded in summer (July-August) and these are
aquaculture ponds and marshland on all coasts. Flocks of presumed to be non-breeders and immatures (Silvius 1988).
thousands observed in the north-east (Crossland in prep.) and The origins of these summering birds are as yet unknown but
the south-east (Silvius 1988). may be other parts of Indonesia or possibly Australia.
Greater Sand Plover Charadrius leschenaultii Bar-tailed Godwit Limosa lapponica
An abundant and widespread migrant in the north-east where Locally abundant in the north-east. Most abundant during
it is often more numerous than preceding species, with flocks southward and northward migrations with substantial
of 2000+ recorded (Crossland in prep.). Much less numerous numbers also present through winter (Crossland in prep.).
in the south-east, with highest numbers during northward Also abundant and widespread in the south-east with highest
migration (Silvius 1988). numbers during southward migration but very few during
northward migration (Silvius 1988, Verheught et al. 1990).
Oriental Plover Charadrius veredus
Whimbrel Numenius phaeopus
Sumatra lies west of the usual wintering range of this species
and it has been recorded as a rare vagrant only (Silvius 1988, A widespread and common migrant in the north- and south-
MacKinnon & Phillipps 1993, Parrott & Andrew 1996). east (Crossland in prep., Silvius 1988) with flocks of up to
400 recorded. Highest numbers observed during southward
Pintail Snipe Gallinago stenura migration with much lower numbers during northward
A common migrant to marshland and rice fields where it can migration, especially in the south-east. Upwards of 3000
be locally abundant (MacKinnon & Phillipps 1993, counted on the coastlines of Jambi and South Sumatra
Crossland in prep.). Provinces during July-August 1985 indicating the
importance of the south-east Sumatra as an over-summering
Swinhoe's Snipe Gallinago megala area for non-breeders (Silvius 1988).
An uncommon visitor to Sumatra, recently recorded in North Eurasian Curlew Numenius arquata
Sumatra Province (Crossland in prep.) but a need for further
records to clarify its status and distribution throughout the Contrary to MacKinnon & Phillipps (1993), who state that
island (MacKinnon & Phillipps 1993). this species is a regular visitor to the Greater Sundas but
never numerous, recent surveys have found it to be an
Common Snipe Gallinago gallinago abundant and widespread migrant in Sumatra with flocks of
Reported by MacKinnon & Phillipps (1993) as a scarce up to 2600 recorded in the north-east (Crossland in prep.)
visitor to the Greater Sundas with doubtful records from and 7100 in the south-east (Verheught et al. 1990). Clearly,
Sumatra. As with the preceding species, further records are Sumatra is a major migration terminus for this species.

92
Stilt 50 (2006): 90–95 Migratory waders in Sumatra

Highest numbers in both the north-east and south-east suggest. All congregations of Common Greenshank
recorded during southward migration and winter with lower anywhere in Sumatra should be carefully scrutinised for any
numbers during northward migration (Silvius 1988, Nordmann’s Greenshank that might be among them.
Verheught 1990, Crossland in prep.). Upwards of 3000
Green Sandpiper Tringa ochropus
recorded in summer (July-August) along the coastlines of
Jambi and South Sumatra Provinces in the south-east Sumatra lies on the edge of this species' wintering range and
(Silvius 1988), indicating that relatively large numbers may it occurs as an uncommon visitor in small numbers
stay through summer. (MacKinnon & Phillipps 1993, Verheught et al. 1993,
Holmes 1996).
Eastern Curlew Numenius madagascariensis
Wood Sandpiper Tringa glareola
Generally an uncommon visitor (MacKinnon & Phillipps
1993, Strange 2001), but localised concentrations of 100 or Said by Strange (2001) to be probably the most numerous
more have been found in the north-east (Crossland in prep.) and widespread shorebird in Indonesia. Both MacKinnon &
and up to 2600 in the south-east (Verheught et al. 1990). Phillipps (1993) and Tilford & Compost (2000) state that
Highest numbers observed in winter in the north-east and this species is common and widespread in the Greater
during southward migration in the south-east. Sundas. Holmes & Nash (1990) list this as one of the three
most common waders seen inland in Sumatra and
Lesser Yellowlegs Tringa flavipes
Kalimantan, while Parrott & Andrew (1996) report it as
A rare vagrant to Sumatra, first recorded in 1983 (Ollington common in agricultural land surrounding Way Kambas
& Parrish 1989, MacKinnon & Phillipps 1993). National Park in Lampung Province. The Wood Sandpiper is
not typically a species of intertidal mudflats so it has
Common Redshank Tringa totanus
received poor coverage in wader surveys conducted in north-
An abundant and widespread migrant in both the north-east and south-east Sumatra to date. More surveys of rice field
(Crossland in prep.) and the south-east (Silvius 1988, and freshwater wetland habitats are required before the true
Verheught et al. 1990). Sumatra is a major wintering area for abundance of this species in Sumatra can be assessed.
this species with flocks of up to 10,000 recorded
Terek Sandpiper Xenus cinereus
(MacKinnon & Phillipps 1993). Highest numbers recorded
during winter and northern migration. An abundant and widespread migrant in the north- and
south-east with flocks of over 2000 recorded (Crossland in
Spotted Redshank Tringa erythropus
prep., Silvius 1988, Verheught et al. 1990, MacKinnon &
A scarce visitor to Sumatra, recorded in September 1988 at Phillipps 1993). In the north-east high numbers observed on
Lebak Pampangan in South Sumatra Province and southward and northward migrations with relatively low
occasionally from other localities in the south-east numbers over winter. In the south-east highest numbers on
(Verheught et al. 1993). Not yet recorded from the north-east southward migration with much lower numbers recorded on
but, given the closer proximity to the southern limit of this northward migration. These patterns suggests heavy passage
species' normal wintering range (the Thai- Malay Peninsula), of Terek Sandpipers through Sumatra, probably involving
its occurrence is likely. birds moving through to Java and eastern Indonesia or
possibly further still to Australia.
Marsh Sandpiper Tringa stagnatilis
Common Sandpiper Actitis hypoleucos
A locally common migrant to the north- and south-east with
flocks of up to 300 recorded. Highest numbers recorded in A common and widespread migrant, found in a wide range
winter in the north-east (Crossland in prep.) and during of estuarine, wetland and riverine habitats from the coast up
southward migration in the south-east (Verheught et al. to 1500 m (Holmes & Nash 1990, MacKinnon & Phillipps
1990). 1993). Flocks of up to 40 have been observed in the north-
east (Crossland in prep.).
Common Greenshank Tringa nebularia
Grey-tailed Tattler Tringa brevipes
A widespread but not numerous migrant on all coasts. Flocks
in the north-east and the south-east seldom exceed 50 birds This species has not been observed in shorebird surveys in
with highest numbers present during winter and during the north-east (Crossland 2000, Crossland in prep.) or the
northward migration (Silvius 1988, Crossland in prep.). south-east (Silvius 1988, Verheught et al. 1993, Parrott &
Andrew 1996, Crossland & Sinambela 2005). However, it is
Nordmann's Greenshank Tringa guttifer
reported by both MacKinnon & Phillipps (1993) and Strange
Not identified in Sumatra until 6 birds were found at Cemara (2001) as an uncommon to rare visitor to the Sundas,
Beach (Jambi Province) in April 1986 (Silvius 1987). including Sumatra.
Subsequently found to be a regular migrant in small numbers Ruddy Turnstone Arenaria interpres
to the north-east (MacKinnon & Phillipps 1993; Crossland in
prep.) and the south-east (Silvius 1988). Typically groups of Widespread but nowhere numerous in the north-east
6 or less have been spotted but Verheught et al. (1990, 1993) (Crossland in prep.). More numerous in the south-east where
reported a flock of 21 from South Sumatra. This species is highest numbers recorded on southward migration (Silvius
probably more numerous than the limited records to date

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Stilt 50 (2006): 90–95 Migratory waders in Sumatra

1988, Verheught et al. 1990). Scarce on west coast with first DISCUSSION
record in May 1992 (Holmes 1996).
Wintering migrants, passage migrants and vagrants
Great Knot Calidris tenuirostris
Wader surveys completed to date in Sumatra have very
A locally common migrant in the north-east with flocks of much been preliminary work with only about half of the east
up to 400 recorded. Much less common in the south-east coast and just a fraction of the west coast covered. Much
where mainly seen on southward migration (Silvius 1988). greater survey effort is required before a definitive
Red Knot Calidris canutus understanding of abundance and distribution of waders in
Sumatra can emerge.
Described by MacKinnon & Phillipps (1993) as a rare This paper has provided a summary for 37 migratory
passage migrant, this species is generally very scarce in wader species recorded in Sumatra up until 2006.
Sumatra except for a brief passage during northward Approximately a fifth of these use Sumatra (and presumably
migration (Silvius 1988, Crossland in prep.). Flocks of up to neighbouring parts of the Indonesian archipelago) as a
400, usually comprising birds in full breeding plumage, have migration terminus and don't migrate beyond the region in
been observed mainly in late March to early April and any great numbers. Such species include Pintail Snipe,
quickly pass through. With an absence of concentrations of Swinhoe’s Snipe, Little Ringed Plover, Asian Dowitcher,
Red Knot wintering in western Indonesia, it is likely that Eurasian Curlew, Common Redshank and Nordmann's
these birds have an Australian origin. Greenshank.
Another group (comprising over half the recorded
Sanderling Calidris alba
species) overwinter in Sumatra as well as in neighbouring
A rare visitor to the north-east (Holmes 1996) and the south- Indonesian islands and Australia. For most of these species,
east (Silvius 1987, 1988; Verheught et al. 1993). Likely to influxes evident during one or both migration periods
be more common on the sandy coastlines of the west, but suggest passage through Sumatra – presumably to Java and
shorebird surveys have yet to be undertaken in areas of neighbouring islands, and possibly onwards to Australia.
suitable habitat to confirm this. These species include: Pacific Golden Plover, Grey Plover,
Greater Sand Plover, Lesser Sand Plover, Black-tailed
Red-necked Stint Calidris ruficollis
Godwit, Bar-tailed Godwit, Whimbrel, Eastern Curlew,
A widespread but generally uncommon migrant to the north- Marsh Sandpiper, Common Greenshank, Wood Sandpiper,
and the south-east with flocks exceeding 170 birds recorded Terek Sandpiper, Common Sandpiper, Ruddy Turnstone,
in Aceh Province (Crossland 2000), but seldom exceeding Great Knot, Curlew Sandpiper, Sanderling, Red-necked Stint
50 birds elsewhere (Silvius 1988, Crossland in prep.). and Broad-billed Sandpiper.
A third group comprises species which are seldom
Long-toed Stint Calidris subminuta
recorded wintering in Sumatra, but which occur as passage
A widespread but scarce visitor to Indonesia, including migrants, probably heading to (or from) eastern Indonesia or
Sumatra (Tilford & Compost 2000, Strange 2001). This Australia. This group includes Oriental Pratincole, Oriental
species does not usually frequent coastal mudflats so has not Plover, Red Knot and Grey-tailed Tattler.
been picked up on shorebird surveys in the north- and south- A fourth group consists of species for which Sumatra lies
east. on the edge of their the south-east Asian range and they
consequently occur as stragglers in very low numbers. This
Broad-billed Sandpiper Limicola falcinellus group includes: Kentish Plover, Common Snipe, Spotted
A locally common migrant in the north-east with flocks of Redshank, Green Sandpiper, Long-toed Stint and Ruff.
up to 100 observed (Crossland in prep.). A rare visitor in the Potential additions to this group which should be looked for
south-east, recorded mainly during northward migration on Sumatran wetlands include Ringed Plover Charadrius
(Silvius 1988, Verheught et al. 1993). hiaticula, Long-billed Plover Charadrius placidus,
Temminck's Stint Calidris temminckii, Spoon-billed
Curlew Sandpiper Calidris ferruginea Sandpiper Eurynorhynchus pygmaeus, Dunlin Calidris
An abundant and widespread migrant with flocks of over alpina and Red-necked Phalarope Phalaropus lobatus.
1000 recorded in the north-east (Crossland in prep.). Less
abundant but still widespread and common in the south-east, An estimate of total migratory wader numbers
particularly during winter and northward migration (Silvius The total shoreline length of Sumatra’s east coast and
1988, Verheught et al. 1990). adjacent islands exceeds 3500 km, much of which comprises
Ruff Philomachus pugnax intertidal mudflats, mangroves and swampland. In addition,
there are thousands of hectares of aquaculture ponds and tens
An uncommon migrant to Sumatra, where it occurs on of thousands of hectares of rice fields throughout the coastal
coastal and inland habitats (MacKinnon & Phillipps 1993). lowlands. Verheught et al. (1990) estimated that 500,000
Not recorded on coastal surveys in the north-east (Crossland migratory waders were dependent on the coastal mudflats of
in prep.) or the south-east (Silvius 1988, Verheught et al. South Sumatra Province. Extrapolations from this to cover
1993). the entire east coast and adjacent islands, as well as pockets
of suitable wader habitat along the west coast, suggest that

94
Stilt 50 (2006): 90–95 Migratory waders in Sumatra

upwards of one million migratory waders could overwinter Aceh, North Sumatra, West Sumatra, Bengkulu and
in or migrate through Sumatra during the course of a year. Lampung Provinces.
However, the results of more recent survey work undertaken
by Wetlands International, Wildlife Conservation Society ACKNOWLEDGEMENTS
and others require publication before a more satisfactory
population estimate can be made. Many thanks to the small groups of dedicated researchers
Eastern Sumatra supports wader populations comparable and the growing band of Indonesian and visiting
or greater than those found on the west coast of Peninsular birdwatchers who have braved the heat, soft mud, mangrove
Malaysia (Li et al. 2006). Together these coastlines on roots, mosquitoes, snakes and the occasional crocodile to
opposite sides of the Strait of Malacca comprise one of the track down the hitherto unknown haunts of Sumatra’s
most important areas for waders and other shorebirds in waders and begin the process of appreciating and protecting
Asia, comparable to areas like the Yellow Sea, the Gulf of these magnificent migratory birds.
Thailand and the coast of Bangladesh.
REFERENCES
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at the north-western tip of Sumatra, Indonesia. Stilt 36: 4–6.
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upwards of 1000 migratory waders. Surveyed sites known to the migratory wader community found on Batam Island, Riau
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has been carried out but shoreline and estuary surveys on waterbirds with notes on the Asian Dowitcher Limnodromus
nearby islands, such Bintan and Kundur are yet to take place. semipalmatus. Kukila 3: 117–137.
Strange, M. 2001. A Photographic Guide to the Birds of Indonesia.
Additional areas that require survey effort include the Periplus, Hong Kong.
northern and eastern shorelines of Rupat, Bengkalis and Tilford, T. & A. Compost. 2000. A Photographic Guide to Birds
Rangsang islands bordering the Strait of Malacca; the of Java, Sumatra and Bali. New Holland Publishers (UK) Ltd,
Kampar river mouth area; the islands in the Lingga London.
Archipelago, especially Lingga and Singkep; and the Verheught, W.J.M., H. Skov & F. Danielseon. 1993. Notes on
southern coastlines of Lampung Province. the birds of the tidal lowlands and floodplains of South Sumatra
Along the west coast, localised wader surveys have taken Province, Indonesia. Kukila 6: 53–84.
place as part of the Asian Waterbird Census in West Sumatra Wetlands International. 2006. Waterbird Population Estimates -
Province, but a comprehensive survey is needed covering Fourth Edition. Wetlands International, Wageningen, The
Netherlands.
suitable wader habitats throughout the western coastlines of Whitten, T., S.J. Damanik, & N. Hisyam. 2000. The Ecology of
Sumatra. Periplus, Hong Kong.

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