Aquatic Insects

June 2005; 27(2): 113 – 123

Contribution to the knowledge of the genus Metrichia Ross

from Argentina (Trichoptera: Hydroptilidae:
Ochrotrichiini)

ELISA B. ANGRISANO & JULIETA V. SGANGA

Universidad de Buenos Aires, Facultad de Ciencias Exactas y Naturales, C1428EHA, Buenos Aires,
Argentina

Abstract
Metrichia cuniapiru, sp. n. (tribe Ochrotrichiini, family Hydroptilidae) from Misiones Province is
described. Main characters useful for recognition of females of different species of Metrichia and keys for
the males and larvae are provided. The pupa of M. neotropicalis is also described.

Keywords: Hydroptilidae, larva, pupa, female, Neotropical

Introduction
Metrichia Ross 1938, Ochrotrichia Mosely 1934, Rhyacopsyche Müller 1879 and Paratrichia
Angrisano 1995 (see Angrisano 2002) are the four Neotropical genera of Hydroptilidae that
currently constitute the tribe Ochrotrichiini, of the subfamily Hydroptilinae.
The classification criteria have changed over the years: Flint (1972) considered the species
of Metrichia and Ochrotrichia to be congeneric and grouped them together in the genus
Ochrotrichia, but maintained the subgeneric status, although this was not generally accepted
for many years. Flint and Bueno-Soria (1998) raised both taxa to generic status, which we
follow in this paper.
Metrichia is distributed from the southwestern United States, throughout Central America,
the West Indies, and North and West South America to Patagonia. In Argentina it is
represented by six species, all recorded for the west, from Salta to Santa Cruz and Córdoba
Provinces, which is the presently known range of distribution.
The males of Metrichia may be recognized by having the ninth segment open dorsally and
closed ventrally; aedeagus with two portions separated by a median constriction, usually with
hooks or spines in the distal section; claspers big, wide and long, rounded or subrectangular.
Descriptions of females are few, they are brief and scarcely comparative. Only females of
M. cafetalera Botosaneanu from Cuba, M. fontismoreaui Botosaneanu from Haiti and M. juana
(Flint) from Puerto Rico, have been described.

Correspondence: Julieta V. Sganga, Laboratorio de Entomologı́a, Departamento de Biodiversidad y Biologı́a Experimental, Facultad
de Ciencias Exactas y Naturales, Universidad de Buenos Aires, C1428EHA, Pabellón II, Ciudad Universitaria, Buenos Aires,
Argentina. E-mail: jsganga@bg.fcen.uba.ar

ISSN 0165-0424 print/ISSN 1744-4152 online ª 2005 Taylor & Francis

DOI: 10.1080/01650420500062782
114 Elisa B. Angrisano & Julieta V. Sganga

The knowledge of the larvae is very poor, as happens with so many other aquatic insects. Of
the 57 known species of Metrichia, the larvae of only two have been described: M. nigritta
(Banks) from the United States and M. juana. M. nigritta was described by Edwards and
Arnold (1961) and by Wiggins (1996), and M. juana by Flint (1964). Up to now no pupa has
been described in this genus.
Wiggins (1996) provided the first diagnosis for the larvae of Metrichia. They are very
similar to those of Ochrotrichia, differing from them by the absence of tergites on the
abdominal segments II – VIII. As in Ochrotrichia there are dorsal rings on abdominal
segments II – VIII; the setae along the anterior margin of the thoracic nota are long, curved
and directed forward. The tarsal claws are long and curved. There is a basal seta in the
tarsal claws of the forelegs that is absent in the mid- and hindlegs. The anterolateral angles
of the meso- and metanota are produced as dark lobes, as in Ochrotrichia. The knowledge of
the larvae is very important because it contributes to the solution of questions of affinities
and classification of the taxa.
The cases of Metrichia are oval and laterally compressed, built from two silken valves
sometimes covered with minerals or vegetal pieces and stuck together along the dorsal and
ventral margins, leaving the ends open. Botosaneanu and Flint (1982) described a case from
Venezuela with two dorsal chimneys, that was not associated with any species.

Objectives
The aims of the present work are (1) to describe a new species of Metrichia from Misiones
Province, (2) to enlarge the known distribution of the species recorded from Argentina, (3) to
give useful characters for the recognition of the preimaginal stages of M. argentinica Schmid,
M. bidentata (Flint), M. bulbosa (Jacquemart), M. disparilis (Flint), M. neotropicalis Schmid
and M. patagonica (Flint), (4) to characterize the females of M. bulbosa, M. neotropicalis and
M. patagonica, (5) to describe a pupa of the genus for the first time, and (6) to elaborate a key
for the identification of the males and larvae of Metrichia.

Materials and methods

Males, females, pupae and last instar larvae collected in different localities in Argentina were
examined. Larvae, pupae and females were identified using the metamorphotype method.
Some females were associated because they were the only species present at the locality.
The specimens were cleared with alkali, neutralized with phenol and mounted in glycerin or
Fauré media for microscopical observations.
The types of the new species are deposited at the Museo de La Plata, Buenos Aires,
Argentina. Non-type material is deposited at the Laboratory of Entomology, Facultad de
Ciencias Exactas y Naturales, Universidad de Buenos Aires, Argentina.

Results
All the species present in Argentina are mentioned and new data reported.

Metrichia cuniapiru Angrisano sp. n. (Figures 1 – 8)

Material examined. Holotype male: Misiones, Cuñá Pirú Provincial Park, Cuñá Pirú Stream,
18/19.V.1997; allotype female, same data as the holotype; paratypes: 10 males, 5 females,
same data as the holotype.
 Genus Metrichia in Argentina (Trichoptera: Hydroptilidae) 115

Figures 1 – 8. Metrichia cuniapiru sp. n. Male: (1, 2, 3) segment IX, lateral, ventral and dorsal views; (4) aedeagus.
Female: (5, 6) dorsal and ventral views of abdominal segments VII – X showing the position of the vagina; (7) vagina;
(8) abdominal apodemes.

General coloration of the body: uniformly dark brown (specimens preserved in alcohol).
Tibial spur formula: 1-3-4.

Males. Length of forewing: 1.7 mm. Ninth abdominal segment relatively short; aedeagus
long, simple, without hooks, and with two processes after the median constriction; claspers
116 Elisa B. Angrisano & Julieta V. Sganga

short, quadrangular in lateral view, posterior margin truncate, without spurs or spines, and
with a shallow notch in the apex. The ‘‘dorsal hook’’ is long and deeply arched ventrad
(fanglike) with its end strongly pigmented, contrasting with the coloration of the other genital
structures. Cerci rounded. Tenth segment membranous, conical. Without processes in the
pregenital sternites.

Females. Length of forewing: 2.2 mm. Abdomen lightly sclerotized. With a short midventral
spine on segment VI; tergum VII with a pair of subtriangular setose sclerotized areas in the
posterior region (Figure 5) and a pair of long apodemes that reach to segment II (Figure 8);
segments VIII and IX membranous, X with a pair of apodemes that reach to segment VI when
the oviscapt is totally evaginated (Figure 8). Vagina as in Figure 7.

Larvae unknown.

Affinities. The mentioned characters clearly distinguish the new species from others recorded
from Argentina; with M. neotropicalis it shares the absence of hooks in the aedeagus, but differs
in the remaining characters, especially the shape of the claspers and the dorsal hook.

Etymology. Named for the stream where the specimens were collected.

Metrichia argentinica Schmid 1958 (Figure 19)

Metrichia argentinica Schmid 1958, p. 195. Flint and Bueno-Soria 1998. Flint, Holzenthal &
Harris 1999. Angrisano 2000.

Ochrotrichia (M.) argentinica: Flint 1974. Angrisano 1998.

Material examined. Tucumán Province, Burruyacú, Medina River, 20.VIII.93: 200 larvae.

Distribution. Tucumán, Mendoza, Catamarca and Salta Provinces.

Females unknown.

Larvae. Length: 2.3 – 2.7 mm. Pro- and mesosternum without sclerites. Subapical projection
of the foretibia with two short and thick spines; anterior tarsus with a spine, similar to that of
the tibia (Figure 19).
Without dorsal sclerites on abdominal segments I – VIII; ninth tergite similar to that of
M. bidentata.
Silk cases covered with regularly arranged pieces of filamentous algae.

Metrichia bidentata (Flint 1983) (Figures 15 – 18)

Ochrotrichia (M.) bidentata Flint 1983, p.41. Angrisano 1998.

Metrichia bidentata (Flint): Flint & Bueno-Soria 1998. Flint, Holzenthal & Harris 1999.
Angrisano 2000.

Material examined. Córdoba Province, Copina, 3.XII.1982: 1 larva; Copina, 1.I.1984: 17

larvae; Copina, 15.VII.1985: 11 larvae; Copina, 16.I.1992: 2 males, 7 larvae, 3 pharate males,
 Genus Metrichia in Argentina (Trichoptera: Hydroptilidae) 117

1 pharate female; Ensenada Stream, 22.I.1989: 3 larvae, 1 pharate female. Chubut Province,
Parque Nacional Los Alerces, Irigoyen Waterfall 5.III.1995: 3 cases.

Distribution. Neuquén, Chubut and Córdoba Provinces (new record).

Females unknown.

Larvae. Length: 3.5 – 3.8 mm. Pro- and mesosternum with a lateroposterior sclerite on either
side (Figure 16). Tibia of foreleg with two short and thick spines in the subapical projection;
anterior tarsus with a long spine and a small one (Figure 18).
First abdominal segment with a dorsal sclerite that bears two setae, with the anterior margin
strongly convex and posterior margin straight, with two small lateral notches, a pair of
laterodorsal circular sclerites and a pair of lateral sclerites, each one bearing a seta (Figure 15).
Second to eighth segments with a pair of laterodorsal oval sclerites that bear a seta, and a pair
of lateral setae without sclerotization. Ninth tergite with the anterior margin straight and the
posterior strongly convex (Figure 17).
Silk cases, covered with pieces of moss epidermis of flabellate shape, of uniform size.

Metrichia bulbosa (Jacquemart 1963) (Figures 9, 12, 20)

Argentitrichia bulbosa Jacquemart 1963, p. 339.

Ochrotrichia bulbosa (Jacquemart): Mangeaud 1996 [Córdoba]. Angrisano 1998.

Figures 9 – 14. Metrichia spp., females. Segment VII, dorsal view: (9) M. bulbosa; (10) M. neotropicalis: (11) M.
patagonica. Vagina: (12) M. bulbosa; (13) M. neotropicalis; (14) M. patagonica.
118 Elisa B. Angrisano & Julieta V. Sganga

Figures 15 – 25. Larvae. M. bidentata: (15) abdominal segment I, dorsal view; (16) pro- and mesothorax, ventral
view, showing the posterolateral sclerites; (17) tergite IX; (18) anterior tarsus. M. argentinica: (19) anterior tarsus.
M. bulbosa: (20) anterior tarsus. M. disparilis: (21) anterior tibia and tarsus. M. neotropicalis: (22, 23) foreleg, external
and internal views; (24) anterior tarsus. M. patagonica: (25) anterior tibia and tarsus.

Metrichia bulbosa (Jacquemart). Flint & Bueno-Soria 1998. Flint, Holzenthal & Harris 1999.
Angrisano 2000.

Material examined. Córdoba Province, Copina, 3.XII.1982: 5 larvae; Copina, 1.I.1984: 9

larvae; Copina, 16.I.1992: 2 larvae; Huerta Grande, 28.IV.1991: 2 pharate males, 89 males
and 80 females. Tucumán Province, San Javier, 24.V.1980: 1 larva.
 Genus Metrichia in Argentina (Trichoptera: Hydroptilidae) 119

Distribution. San Juan and Córdoba Provinces.

Females. Length of forewing: 2.1 – 2.8 mm. Posterior region of seventh abdominal tergum
mesally with an oval lightly sclerotized area, and two setose lateral lobes (Figure 9). Vagina as
in Figure 12.

Larvae. Length: 2.2 – 3.1 mm. Prosternum with a lateroposterior sclerite on either side similar
to that of M. bidentata. Without mesosternal sclerites. Subapical projection of the foretibia
with two thick and short spines, tarsus with a scale-like spine, with a shallow notch in the apex,
and two small spines (Figure 20).
First abdominal segment with a pair of dorsal rectangular sclerites, and laterodorsal and
lateral circular pairs, each one bearing a seta. Second to eighth segments with a pair of dorsal
oval sclerites; the sclerites of the first to seventh segments bear a seta; those of the eighth two.
Ninth tergite similar to that of M. bidentata.
Silk cases covered with regularly arranged diatoms.

Metrichia disparilis (Flint 1983) (Figure 21)

Ochrotrichia (M.) disparilis Flint 1983, p. 41. Angrisano 1998.
Metrichia disparilis (Flint): Flint & Bueno-Soria 1998. Flint, Holzenthal & Harris 1999.
Angrisano 2000.

Material examined. Tucumán Province, Burruyacú, Medina River, 20.VIII.1993: 1 pharate

male.

Distribution. Tucumán Province.

Females unkown.
Larvae. Subapical projection of the foretibia with two scale-like spines, with the apex rounded;
anterior tarsus with a spine similar to that of the tibia (Figure 21).
Larval cases entirely made of silk, with two dorsal chimneys.

Metrichia neotropicalis Schmid 1958 (Figures 10, 13, 22 – 24)

M. argentinica Schmid: Flint 1967 [misidentification].
Metrichia neotropicalis Schmid: Schmid 1958, p.195 [Tucumán, Peru]. Angrisano 2000. Flint
1967 [as M. argentinica]. Flint & Bueno-Soria 1998. Flint, Holzenthal & Harris 1999.
Ochrotrichia (M.) neotropicalis (Schmid 1958). Flint 1980 [Peru], 1990, p. 118 [El Loa, from
Cuzco to Mendoza and Talca]. Mangeaud 1996 [Córdoba]. Angrisano 1998. Medina et al.
1997. Vallania et al. 1998.

Material examined. Buenos Aires Province, Bahı́a Blanca, X.1980: a large number of males
and females; Catamarca Province, Antofalita, XII.1985: 4 pharate males. Córdoba Province,
Copina, Primero Stream, La Suela River, VII.1992: 21 larvae, 1 female and 1 pharate male;
Pinturas River, IV.1996: 1 female; Capilla del Monte, Los Mogotes, 12.VII.1971: 3 larvae;
Huerta Grande, 28.IV.1991: 7 males. Mendoza Province, Tres Saltos Stream, 8.X.2001:
10 pupae, 20 larvae; Chiquero Stream 5.II.2002: 12 larvae; Las Cuevas, 4.V.2001: 1 larva;
120 Elisa B. Angrisano & Julieta V. Sganga

Evarsa 9.VI.2000: 7 larvae; Evarsa 1.XI.2001: 5 larvae; Potrerillos, 9.VI.2000: 2 larvae;

Uspallata, X.1989: 3 females and 16 males; 8.VI.2000: 1 larva. Salta Province, La Calderilla
River, 28.X.2001: 7 larvae, 3 pharate male. San Luis Province, Dique Chico, XII.2001: 1
male metamorphotype, 1 pharate female and 1 larva. Tucumán Province, Va. Padre Monti,
20.XI.1999: 8 males.

Distribution. Tucumán, Córdoba, San Luis and Buenos Aires Provinces (new record).

Females. Length of forewing: 2.6 – 3.2 mm. Posterior region of the seventh abdominal tergum
mesally with a pentagonal area, lightly sclerotized, and two setose lateral lobes (Figure 10).
Vagina as in Figure 13.

Larvae. Length: 3.2 – 5 mm. Prosternum with a small lateroposterior sclerite on either side,
similar to that of M. bidentata. Without mesosternal sclerites. Subapical projection of the
foretibia with two thick and short spines; tarsus with a scale-like spine with the apex straight,
and two small spines (Figures 22, 23, 24).
First abdominal segment with a pair of dorsal rectangular sclerites, and lateral and
laterodorsal circular pairs, each one bearing a seta; segments II – VIII with a pair of dorsal oval
sclerites and a laterodorsal circular pair, dorsal sclerites of segment VIII bearing two setae, the
rest only one. Ninth tergite similar to that of M. bidentata.
Silk cases with a few filaments of algae or sand grains.

Pupae. Mandibles short, with a pointed end, and the inner margin serrate (Figure 28).
Abdominal terga of segments III – VII with anterior dorsal plates, each one with
approximately 15 hooks, and segments III – V with posterior dorsal plates that bear retrorse
hooks and a small anterior oval sclerite (Figures 26 and 27).

Figures 26 – 28. Metrichia neotropicalis, pupa: (26) abdomen of pharate male, dorsal view; (27) dorsal plates; (28) left
mandible.
 Genus Metrichia in Argentina (Trichoptera: Hydroptilidae) 121

Metrichia patagonica (Flint 1983) (Figures 11, 14, 25)

Ochrotrichia (M.) patagonica Flint 1983, p. 41. Mangeaud 1996 [Córdoba]. Angrisano 1998.

Metrichia patagonica (Flint): Flint, Holzenthal & Harris 1999. Angrisano 2000.

Material examined. Rı́o Negro Province, Gutierrez Lake, XII.1956: 1 female, 7 cases with larval
sclerites. Chubut Province, Nahuel Pan Stream, 11.II.1993: 12 females and 24 males.

Distribution. Córdoba, Neuquén, Rı́o Negro and Chubut Provinces.

Females. Length of forewing: 2.2 – 2.6 mm. Posterior region of the seventh abdominal tergum
mesally with a trapezoidal area lightly sclerotized, and two lateral setose lobes (Figure 11).
Vagina simple, as in Figure 14.

Larvae. Subapical projection of the foretibia with two long spines, tarsus with a spine similar
to that of the tibia (Figure 25).
Silk cases covered with very small sand grains.

Keys to Argentinean species of Metrichia

Males
1. Aedeagus without hooks .......................................................................................... 2
1’. Aedeagus with hooks ............................................................................................... 3
2. Claspers short; apex truncate, lightly notched ........................Metrichia cuniapiru sp. n.
2’. Claspers elongate, with ventrodistal margin notched, apex with a spine.M. neotropicalis
2@. End of the claspers rounded ................................................................. M. patagonica
3. Claspers without spines; end of the claspers with a deep notch ................. M. disparilis
3’. Claspers with spines ................................................................................................ 4
4. End of the claspers with two spines, aedeagus with short subapical spines.M. bidentata
4’. Claspers with a spine, aedeagus with two hooks........................................................ 6
5. Claspers expanded dorsally and notched ventroapicaly in lateral view..... M. argentinica
5’. Claspers quadrangular in lateral view ......................................................... M. bulbosa

Females
A key cannot yet be provided. All species with very long apodemes in the genital segments
that reach to the first abdominal segment. The females of M. bulbosa, M. cuniapiru sp. n.,
M. neotropicalis, and M. patagonica may be recognized by the coloration of the seventh
abdominal tergite and the shape of the vaginal sclerites.

Last instar larvae

1. Larval cases without dorsal chimneys ........................................................................ 2
1’. Larval cases made of silk, with two dorsal chimneys; subapical projection of the foretibia
with two scale-like spines, the apex rounded; anterior tarsus with a spine similar to that of
the tibia (Figure 21) ................................................................................. M. disparilis
2. Subapical projection of the foretibia with two short and thick spines .......................... 3
122 Elisa B. Angrisano & Julieta V. Sganga

2’. Subapical projection of the foretibia with two long spines; tarsus with a long spine; cases
covered with very small sand grains (Figure 25)..................................... M. patagonica
3. Pro-, mesosternum and abdominal terga I – VIII without sclerites; anterior tarsus with a
short and thick spine; silk cases covered with regularly arranged pieces of filamentous
algae (Figure 19) ................................................................................... M. argentinica
3’. Prosternum and abdominal terga I – VIII with sclerites.............................................. 4
4. Mesosternum with a lateroposterior sclerite on either side; anterior tarsus with a long
spine and a small one; first abdominal tergum mesally with a sclerite as in Figure 15;
silk cases, covered with pieces of moss epidermis of flabellate shape, of uniform size
(Figures 16 – 18).......................................................................................M. bidentata
4’. Mesosternum without sclerites; first abdominal tergum mesally with two rectangular
sclerites.................................................................................................................... 5
5. Abdominal terga II – VIII mesally with a pair of oval sclerites and a laterodorsal circular
pair; anterior tarsus with a scale-like spine with straight apex and two small spines; silk
cases with a few filaments of algae or sand grains (Figures 22 – 24) ........M. neotropicalis
5’. Abdominal terga II – VIII mesally with a pair of oval sclerites; anterior tarsus with a scale-
like spine, with a shallow notch in the apex, and two small spines; silk cases covered with
regularly arranged diatoms (Figure 20) ....................................................... M. bulbosa

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