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Ape
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Main page This article is about the branch of primates. For other uses, see Ape (disambiguation).
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Apes (Hominoidea /hɒmɪˈnɔɪdiːə/) are a branch of Old World tailless simians native to Africa and Southeast Asia. They are the sister group of the Old World monkeys, together forming the catarrhine clade. They are distinguished from other primates by a wider degree of freedom of motion at the shoulder joint as evolved by the
Current events Hominoids or apes
Random article influence of brachiation. In traditional and non-scientific use, the term "ape" excludes humans, and can include tailless primates taxonomically considered monkeys (such as the Barbary ape and black ape), and is thus not equivalent to the scientific taxon Hominoidea. There are two extant branches of the superfamily
Temporal range: Miocene-Holocene
About Wikipedia Hominoidea: the gibbons, or lesser apes; and the hominids, or great apes.
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The family Hylobatidae, the lesser apes, include four genera and a total of sixteen species of gibbon, including the lar gibbon and the siamang, all native to Asia. They are highly arboreal and bipedal on the ground. They have lighter bodies and smaller social groups than great apes.
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The family Hominidae (hominids), the great apes, include four genera comprising three extant species of orangutans and their subspecies, two extant species of gorillas and their subspecies, two extant species of panins (bonobos and chimpanzees) and their subspecies, and one extant species of humans in a single extant
Contribute subspecies.[1][a][2][3]
Help Except for gorillas and humans, hominoids are agile climbers of trees. Apes eat a variety of plant and animal foods, with the majority of food being plant foods, which can include fruit, leaves, stalks, roots and seeds, including nuts and grass seeds. Human diets are sometimes substantially different from that of other hominoids
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due in part to the development of technology and a wide range of habitation. Humans are by far the most numerous of the hominoid species, in fact outnumbering all other primates by a factor of several thousand to one.
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Recent changes All non-human hominoids are rare and endangered.[citation needed] The chief threat to most of the endangered species is loss of tropical rainforest habitat, though some populations are further imperiled by hunting for bushmeat. The great apes of Africa are also facing threat from the Ebola virus. Currently considered to be the
Upload file greatest threat to survival of African apes, Ebola infection is responsible for the death of at least one third of all gorillas and chimpanzees since 1990.[4]

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What links here 1 Name and terminology Sumatran orangutan (Pongo abelli)
Related changes 2 Evolution
Special pages Scientific classification
3 Taxonomic classification and phylogeny
Permanent link Kingdom: Animalia
3.1 History of hominoid taxonomy
Page information
3.2 Changes in taxonomy and terminology Phylum: Chordata
Cite this page
Wikidata item 4 Characteristics Class: Mammalia
4.1 Distinction from monkeys
Order: Primates
Print/export 5 Behaviour
Suborder: Haplorhini
Download as PDF 5.1 Diet
Printable version 5.2 Cognition Infraorder: Simiiformes
6 See also Parvorder: Catarrhini
In other projects
7 Notes
Superfamily: Hominoidea
Wikimedia Commons 8 References
Gray, 1825
Wikispecies
8.1 Literature cited
Type species
Languages 9 External links
Homo sapiens
বাংলা
Linnaeus, 1758
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Name and terminology Families
िहन्दी
മലയാളം "Ape", from Old English apa, is a word of uncertain origin.[b] The term has a history of rather imprecise usage—and of comedic or punning usage in the vernacular. Its earliest meaning was generally of any non-human anthropoid primate, [c] as is still the case for its cognates in other Germanic languages.[5] Later, after the term †Proconsulidae
ꯃꯤꯇꯩ ꯂꯣꯟ "monkey" had been introduced into English, "ape" was specialized to refer to a tailless (therefore exceptionally human-like) primate.[6] Thus, the term "ape" obtained two different meanings, as shown in the 1911 Encyclopædia Britannica entry: it could be used as a synonym for "monkey" and it could denote the tailless human- †Afropithecidae
ਪੰਜਾਬੀ like primate in particular.[7]
த"# †Pliobatidae
!"# Some, or recently all, hominoids are also called "apes", but the term is used broadly and has several different senses within both popular and scientific settings. "Ape" has been used as a synonym for "monkey" or for naming any primate with a human-like appearance, particularly those without a tail.[7] Biologists have †Dendropithecidae
中⽂ traditionally used the term "ape" to mean a member of the superfamily Hominoidea other than humans,[1] but more recently to mean all members of Hominoidea. So "ape"—not to be confused with "great ape"—now becomes another word for hominoid including humans.[3][d] Hylobatidae

The taxonomic term hominoid is derived from, and intended as encompassing, the hominids, the family of great apes. Both terms were introduced by Gray (1825). The term hominins is also due to Gray (1824), intended as including the human lineage (see also Hominidae#Terminology, Human taxonomy). Hominidae
75 more
The distinction between apes and monkeys is complicated by the traditional paraphyly of monkeys: Apes emerged as a sister group of Old World Monkeys in the catarrhines, which are a sister group of New World Monkeys. Therefore, cladistically, apes, catarrhines and related contemporary extinct groups such as sister: Cercopithecoidea
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Parapithecidaea are monkeys as well, for any consistent definition of "monkey". "Old World Monkey" may also legitimately be taken to be meant to include all the catarrhines, including apes and extinct species such as Aegyptopithecus,[8][9][10][11][citation needed] in which case the apes, Cercopithecoidea and Aegyptopithecus Synonyms
emerged within the Old World Monkeys.
Proconsuloidea
The primates called "apes" today became known to Europeans after the 18th century. As zoological knowledge developed, it became clear that taillessness occurred in a number of different and otherwise distantly related species. Sir Wilfrid Le Gros Clark was one of those primatologists who developed the idea that there were
trends in primate evolution and that the extant members of the order could be arranged in an ".. ascending series", leading from "monkeys" to "apes" to humans. Within this tradition "ape" came to refer to all members of the superfamily Hominoidea except humans.[1] As such, this use of "apes" represented a paraphyletic
grouping, meaning that, even though all species of apes were descended from a common ancestor, this grouping did not include all the descendant species, because humans were excluded from being among the apes.[e]

Traditionally, the English-language vernacular name "apes" does not include humans, but phylogenetically, humans (Homo) form part of the family Hominidae within Hominoidaea. Thus, there are at least three common, or traditional, uses of the term "ape": non-specialists may not distinguish between "monkeys" and "apes", that is, they may use the two terms
interchangeably; or they may use "ape" for any tailless monkey or non-human hominoid; or they may use the term "ape" to just mean the non-human hominoids.

Modern taxonomy aims for the use of monophyletic groups for taxonomic classification;[12][f] Some literature may now use the common name "ape" to mean all members of the superfamily Hominoidea, including humans. For example, in his 2005 book, Benton wrote "The apes, Hominoidea, today include the gibbons and orang-utan ... the gorilla and chimpanzee ...
and humans".[3] Modern biologists and primatologists refer to apes that are not human as "non-human" apes. Scientists broadly, other than paleoanthropologists, may use the term "hominin" to identify the human clade, replacing the term "hominid". See terminology of primate names.

See below, History of hominoid taxonomy, for a discussion of changes in scientific classification and terminology regarding hominoids.

Evolution

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Although the hominoid fossil record is still incomplete and fragmentary, there is now enough evidence to provide an outline of the evolutionary history of humans. Previously, the divergence between humans and other extant hominoids was thought to have occurred 15 to 20 million years ago, and several species of that time period, such as Ramapithecus, were once
thought to be hominins and possible ancestors of humans. But, later fossil finds indicated that Ramapithecus was more closely related to the orangutan; and new biochemical evidence indicates that the last common ancestor of humans and non-hominins (that is, the chimpanzees) occurred between 5 and 10 million years ago, and probably nearer the lower end of
that range; see Chimpanzee–human last common ancestor (CHLCA).

Taxonomic classification and phylogeny

Genetic analysis combined with fossil evidence indicates that hominoids diverged from the Old World monkeys about 25 million years ago (mya), near the Oligocene-Miocene boundary.[13][14][15] The gibbons split from the rest about 18 mya, and the hominid splits happened 14 mya (Pongo),[16] 7 mya (Gorilla), and 3–5 mya (Homo & Pan).[17] In 2015, a new genus and
species were described, Pliobates cataloniae, which lived 11.6 mya, and appears to predate the split between Hominidae and Hylobatidae.[18][19][20][3] [clarification needed]
Crown Catharrhini (31) (29)   Saadanioidea (†28)
     
Cercopithecoidea (24)   Victoriapithecinae (†19)
   
 
  Crown Cercopithecoidea
Hominoidea (30)  
    Dendropithecidae (†7 Mya)
   
    Ekembo heseloni (†17 Mya)
     
      Proconsulidae (†18 Mya)
 
  Ekembo nyanzae (†17 Mya)
(29)   Equatorius (†16)
   
 
  Pliobates (†11.6 Mya)
(29) Afropithecidae (28)  
      Morotopithecus (†20)
 
  Afropithecus (†16)
Crown Hominoidea (22)   Hominidae
   
 
  Hylobatidae
Catarrhini (31.0 Mya) Hominoidea/apes (20.4 Mya) Hominidae/great apes (15.7 Mya) Homininae (8.8 Mya) Hominini (6.3 Mya)  
           

humans (genus Homo)

 
 
chimpanzees (genus Pan)

 
 

gorillas (genus Gorilla)

Traditional apes

 
 

orangutans (genus Pongo)

 
 
gibbons/lesser apes (family Hylobatidae)

 
 
Cercopithecoidea Old World monkeys

The families, and extant genera and species of hominoids are:

Superfamily Hominoidea[21]
Family Hominidae: hominids ("great apes")
Genus Pongo: orangutans
Bornean orangutan, P. pygmaeus
Sumatran orangutan, P. abelii
Tapanuli orangutan, P. tapanuliensis[22]
Genus Gorilla: gorillas
Western gorilla, G. gorilla
Eastern gorilla, G. beringei
Genus Homo: humans
Human, H. sapiens Skeletons of members of the ape superfamily, Hominoidea. There are two extant families:
Genus Pan: chimpanzees Hominidae, the "great apes"; and Hylobatidae, the gibbons, or "lesser apes".

Common chimpanzee, P. troglodytes

Bonobo, P. paniscus
Family Hylobatidae: gibbons ("lesser apes")
Genus Hylobates
Lar gibbon or white-handed gibbon, H. lar
Bornean white-bearded gibbon, H. albibarbis
Agile gibbon or black-handed gibbon, H. agilis
Western grey gibbon or Abbott's grey gibbon, H. abbotti[23]
Eastern grey gibbon or northern grey gibbon, H. funereus[23]
Müller's gibbon or southern grey gibbon, H. muelleri
Silvery gibbon, H. moloch
Pileated gibbon or capped gibbon, H. pileatus
From left: Comparison of size of gibbon, human, chimpanzee, gorilla and orangutan. Non-
Kloss's gibbon or Mentawai gibbon or bilou, H. klossii human hominoids do not stand upright as their normal posture.
Genus Hoolock
Western hoolock gibbon, H. hoolock
Eastern hoolock gibbon, H. leuconedys
Skywalker hoolock gibbon, H. tianxing
Genus Symphalangus
Siamang, S. syndactylus
Genus Nomascus
Northern buffed-cheeked gibbon, N. annamensis
Black crested gibbon, N. concolor
Eastern black crested gibbon, N. nasutus
Hainan black crested gibbon, N. hainanus
Southern white-cheeked gibbon N. siki
White-cheeked crested gibbon, N. leucogenys
Yellow-cheeked gibbon, N. gabriellae

History of hominoid taxonomy

Further information: Human taxonomy § History

The history of hominoid taxonomy is complex and somewhat confusing. Recent evidence has changed our understanding of the relationships between the hominoids, especially regarding the human lineage; and the traditionally used terms have become somewhat confused. Competing approaches to methodology and terminology are found among current scientific
sources. Over time, authorities have changed the names and the meanings of names of groups and subgroups as new evidence — that is, new discoveries of fossils and tools and of observations in the field, plus continual comparisons of anatomy and DNA sequences — has changed the understanding of relationships between hominoids. There has been a gradual
demotion of humans from being 'special' in the taxonomy to being one branch among many. This recent turmoil (of history) illustrates the growing influence on all taxonomy of cladistics, the science of classifying living things strictly according to their lines of descent.[citation needed]

Today, there are eight extant genera of hominoids. They are the four genera in the family Hominidae, namely Homo, Pan, Gorilla, and Pongo; plus four genera in the family Hylobatidae (gibbons): Hylobates, Hoolock, Nomascus and Symphalangus.[21] (The two subspecies of hoolock gibbons were recently moved from the genus Bunopithecus to the new genus
Hoolock and re-ranked as species; a third species was described in January 2017).[24])

In 1758, Carl Linnaeus, relying on second- or third-hand accounts, placed a second species in Homo along with H. sapiens: Homo troglodytes ("cave-dwelling man"). Although the term "Orang Outang" is listed as a variety – Homo sylvestris – under this species, it is nevertheless not clear to which animal this name refers, as Linnaeus had no specimen to refer to,
hence no precise description. Linnaeus may have based Homo troglodytes on reports of mythical creatures, then-unidentified simians, or Asian natives dressed in animal skins.[25] Linnaeus named the orangutan Simia satyrus ("satyr monkey"). He placed the three genera Homo, Simia and Lemur in the order of Primates.

The troglodytes name was used for the chimpanzee by Blumenbach in 1775, but moved to the genus Simia. The orangutan was moved to the genus Pongo in 1799 by Lacépède.

Linnaeus's inclusion of humans in the primates with monkeys and apes was troubling for people who denied a close relationship between humans and the rest of the animal kingdom. Linnaeus's Lutheran archbishop had accused him of "impiety". In a letter to Johann Georg Gmelin dated 25 February 1747, Linnaeus wrote:

It is not pleasing to me that I must place humans among the primates, but man is intimately familiar with himself. Let's not quibble over words. It will be the same to me whatever name is applied. But I desperately seek from you and from the whole world a general difference between men and simians from the principles of Natural History. I certainly know
of none. If only someone might tell me one! If I called man a simian or vice versa I would bring together all the theologians against me. Perhaps I ought to, in accordance with the law of Natural History.[26]

Accordingly, Johann Friedrich Blumenbach in the first edition of his Manual of Natural History (1779), proposed that the primates be divided into the Quadrumana (four-handed, i.e. apes and monkeys) and Bimana (two-handed, i.e. humans). This distinction was taken up by other naturalists, most notably Georges Cuvier. Some elevated the distinction to the level of
order.

However, the many affinities between humans and other primates – and especially the "great apes" – made it clear that the distinction made no scientific sense. In his 1871 book The Descent of Man, and Selection in Relation to Sex, Charles Darwin wrote:

The greater number of naturalists who have taken into consideration the whole structure of man, including his mental faculties, have followed Blumenbach and Cuvier, and have placed man in a separate Order, under the title of the Bimana, and therefore on an equality with the orders of the Quadrumana, Carnivora, etc. Recently many of our best
naturalists have recurred to the view first propounded by Linnaeus, so remarkable for his sagacity, and have placed man in the same Order with the Quadrumana, under the title of the Primates. The justice of this conclusion will be admitted: for in the first place, we must bear in mind the comparative insignificance for classification of the great development
of the brain in man, and that the strongly marked differences between the skulls of man and the Quadrumana (lately insisted upon by Bischoff, Aeby, and others) apparently follow from their differently developed brains. In the second place, we must remember that nearly all the other and more important differences between man and the Quadrumana are
manifestly adaptive in their nature, and relate chiefly to the erect position of man; such as the structure of his hand, foot, and pelvis, the curvature of his spine, and the position of his head.[27]

Changes in taxonomy and terminology

See also: Hominidae and Human taxonomy

Humans the non-apes: Until about 1960, taxonomists typically divided the superfamily Hominoidea into two families. The science community treated humans and their extinct relatives as the outgroup within the superfamily; that is, humans were considered as quite distant from kinship with the
"apes". Humans were classified as the family Hominidae and were known as the "hominids". All other hominoids were known as "apes" and were referred to the family Pongidae.[28]

The "great apes" in Pongidae: The 1960s saw the methodologies of molecular biology applied to primate taxonomy. Goodman's 1964 immunological study of serum proteins led to re-classifying the hominoids into three families: the humans in Hominidae; the great apes in Pongidae; and the
"lesser apes" (gibbons) in Hylobatidae.[29] However, this arrangement had two trichotomies: Pan, Gorilla, and Pongo of the "great apes" in Pongidae, and Hominidae, Pongidae, and Hylobatidae in Hominoidea. These presented a puzzle; scientists wanted to know which genus speciated first from
the common hominoid ancestor.

Gibbons the outgroup: New studies indicated that gibbons, not humans, are the outgroup within the superfamily Hominoidea, meaning: the rest of the hominoids are more closely related to each other than (any of them) are to the gibbons. With this splitting, the gibbons (Hylobates, et al.) were
isolated after moving the great apes into the same family as humans. Now the term "hominid" encompassed a larger collective taxa within the family Hominidae. With the family trichotomy settled, scientists could now work to learn which genus is 'least' related to the others in the subfamily Ponginae.

Orangutans the outgroup: Investigations comparing humans and the three other hominid genera disclosed that the African apes (chimpanzees and gorillas) and humans are more closely related to each other than any of them are to the Asian orangutans (Pongo); that is, the orangutans, not
humans, are the outgroup within the family Hominidae. This led to reassigning the African apes to the subfamily Homininae with humans—which presented a new three-way split: Homo, Pan, and Gorilla.[30]

Hominins: In an effort to resolve the trichotomy, while preserving the nostalgic "outgroup" status of humans, the subfamily Homininae was divided into two tribes: Gorillini, comprising genus Pan and genus Gorilla; and Hominini, comprising genus Homo (the humans). Humans and close relatives
now began to be known as "hominins", that is, of the tribe Hominini. Thus, the term "hominin" succeeded to the previous use of "hominid", which meaning had changed with changes in Hominidae (see above: 3rd graphic, "Gibbons the outgroup").

Gorillas the outgroup: New DNA comparisons now provided evidence that gorillas, not humans, are the outgroup in the subfamily Homininae; this suggested that chimpanzees should be grouped with humans in the tribe Hominini, but in separate subtribes.[31] Now the name "hominin" delineated
Homo plus those earliest Homo relatives and ancestors that arose after the divergence from the chimpanzees. (Humans are no longer classified as an outgroup, but are a branch, deep in the tree of the pre-1960s ape group).

Speciation of gibbons: Later DNA comparisons disclosed previously unknown speciation of genus Hylobates (gibbons) into four genera: Hylobates, Hoolock, Nomascus, and Symphalangus.[21][24]

Characteristics

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The lesser apes are the gibbon family, Hylobatidae, of sixteen species; all are native to Asia. Their major differentiating characteristic is their long arms, which they use to brachiate through trees. Their wrists are ball and socket joints as an evolutionary adaptation to their arboreal lifestyle. Generally smaller than the African apes,
the largest gibbon, the siamang, weighs up to 14 kg (31 lb); in comparison, the smallest "great ape", the bonobo, is 34 to 60 kg (75 to 132 lb).

The superfamily Hominoidea falls within the parvorder Catarrhini, which also includes the Old World monkeys of Africa and Eurasia. Within this grouping, the two families Hylobatidae and Hominidae can be distinguished from Old World monkeys by the number of cusps on their molars; hominoids have five in the "Y-5" molar
pattern, whereas Old World monkeys have only four in a bilophodont pattern.

Further, in comparison with Old World monkeys, hominoids are noted for: more mobile shoulder joints and arms due to the dorsal position of the scapula; broader ribcages that are flatter front-to-back; and a shorter, less mobile spine, with greatly reduced caudal (tail) vertebrae—resulting in complete loss of the tail in extant
hominoid species. These are anatomical adaptations, first, to vertical hanging and swinging locomotion (brachiation) and, later, to developing balance in a bipedal pose. Note there are primates in other families that also lack tails, and at least one, the pig-tailed langur, is known to walk significant distances bipedally. The front of
the ape skull is characterised by its sinuses, fusion of the frontal bone, and by post-orbital constriction.

Distinction from monkeys

See also: Monkey § Historical and modern terminology

Cladistically, apes, catarrhines, and extinct species such as Aegyptopithecus and Parapithecidaea, are monkeys,[citation needed] so one can only specify ape features not present in other monkeys.

Unlike most monkeys, apes do not possess a tail. Monkeys are more likely to be in trees and use their tails for balance. While the great apes are considerably larger than monkeys, gibbons (lesser apes) are smaller than some monkeys. Apes are considered to be more intelligent than monkeys, which are considered to have
more primitive brains.[32]
Like those of the orangutan, the
shoulder joints of hominoids are
adapted to brachiation, or movement by Behaviour
swinging in tree branches.
Major studies of behaviour in the field were completed on the three better-known "great apes", for example by Jane Goodall, Dian Fossey and Birute Galdikas. These studies have shown that in their natural environments, the non-human hominoids show sharply varying social structure: gibbons are monogamous, territorial pair-
bonders, orangutans are solitary, gorillas live in small troops with a single adult male leader, while chimpanzees live in larger troops with bonobos exhibiting promiscuous sexual behaviour. Their diets also vary; gorillas are foliovores, while the others are all primarily frugivores, although the common chimpanzee hunts for meat.
Foraging behaviour is correspondingly variable.

Diet
Apart from humans and gorillas, apes eat a predominantly frugivorous diet, mostly fruit, but supplemented with a variety of other foods. Gorillas are predominately folivorous, eating mostly stalks, shoots, roots and leaves with some fruit and other foods. Non-human apes usually eat a small amount of raw animal foods such as insects or eggs. In the case of humans,
migration and the invention of hunting tools and cooking has led to an even wider variety of foods and diets, with many human diets including large amounts of cooked tubers (roots) or legumes.[33] Other food production and processing methods including animal husbandry and industrial refining and processing have further changed human diets.[34] Humans and other
apes occasionally eat other primates.[35] Some of these primates are now close to extinction with habitat loss being the underlying cause.[36][37]

Cognition

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All the non-human hominoids are generally thought of as highly intelligent, and scientific study has broadly confirmed that they perform very well on a wide range of cognitive tests—though there is relatively little data on gibbon cognition. The early studies by Wolfgang Köhler demonstrated exceptional problem-solving abilities in
chimpanzees, which Köhler attributed to insight. The use of tools has been repeatedly demonstrated; more recently, the manufacture of tools has been documented, both in the wild and in laboratory tests. Imitation is much more easily demonstrated in "great apes" than in other primate species. Almost all the studies in animal
language acquisition have been done with "great apes", and though there is continuing dispute as to whether they demonstrate real language abilities, there is no doubt that they involve significant feats of learning. Chimpanzees in different parts of Africa have developed tools that are used in food acquisition, demonstrating a
form of animal culture.[38]

See also
Dawn of Humanity (2015 PBS film) List of individual apes (for notable non-fictional non-human apes) List of fictional primates
Declaration on Great Apes from the Great Ape Project List of primates by population

Notes
a. ^ Although Dawkins is clear that he uses "apes" for Hominoidea, he also uses "great apes" in ways which exclude humans. Thus in Dawkins 2005: "Long before people thought in terms of evolution ... great apes were often confused with humans" (p. 114); "gibbons are faithfully monogamous, unlike the great apes which are our closer relatives"
(p. 126).
b. ^ The hypothetical Proto-Germanic form is given as *apōn (F. Kluge, Etymologisches Wörterbuch der Deutschen Sprache (2002), online version, s.v. "Affe "; V. Orel, A handbook of Germanic etymology (2003), s.v. "*apōn " or as *apa(n) (Online Etymology Dictionary (2001–2014), s.v. "ape "; M. Philippa, F. Debrabandere, A. Quak, T.
Schoonheim & N. van der Sijs, Etymologisch woordenboek van het Nederlands (2003–2009), s.v. "aap "). Perhaps ultimately derived from a non-Indo-European language, the word might be a direct borrowing from Celtic, or perhaps from Slavic, although in both cases it is also argued that the borrowing, if it took place, went in the opposite
direction.
c. ^ "Any simian known on the Mediterranean during the Middle Ages; monkey or ape"; cf. ape-ward: "a juggler who keeps a trained monkey for the amusement of the crowd." (Middle English Dictionary, s.v. "ape ").
d. ^ Dawkins 2005; for example "[a]ll apes except humans are hairy" (p. 99), "[a]mong the apes, gibbons are second only to humans" (p. 126).
e. ^ Definitions of paraphyly vary; for the one used here see e.g. Stace 2010, pp. 106
f. ^ Definitions of monophyly vary; for the one used here see e.g. Mishler 2009, pp. 114

References
1. ^ a b c Dixson 1981, pp. 13.
2. ^ Grehan, J. R. (2006). "Mona Lisa smile: the morphological
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PMID 16865704 .
3. ^ a b c d Benton, M. J. (2005). Vertebrate Palaeontology .
Wiley-Blackwell. ISBN 978-0-632-05637-8. Retrieved 10 July
2011., p. 371
4. ^ Rush, J. (23 January 2015). "Ebola virus 'has killed a third
of world's gorillas and chimpanzees' – and could pose
greatest threat to their survival, conservationists warn" . The
Independent. Archived from the original on 30 March 2015.
Retrieved 26 March 2015.
5. ^ Terry 1977, pp. 3.
6. ^ Terry 1977, pp. 3–4.
7. ^ a b Chisholm, Hugh, ed. (1911). "Ape" . Encyclopædia
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and Taxonomy I—Strepsirhini. Edinburgh Univ Pubs Science
& Maths, No 3. Edinburgh University Press. p. 53.
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9. ^ Martin, W. C. L. (1841). A General Introduction to the
Natural History of Mammiferous Animals, With a Particular
View of the Physical History of man, and the More Closely
Allied Genera of the Order Quadrumana, or Monkeys .
London: Wright and Co. printers. pp. 340, 361.
10. ^ Geoffroy Saint-Hilaire, M. É. (1812). "Tableau des
quadrumanes, ou des animaux composant le premier ordre
de la classe des Mammifères" . Annales du Muséum
d'Histoire Naturelle. Paris. 19: 85–122.
11. ^ Bugge, J. (1974). "Chapter 4". Cells Tissues Organs. 87
(Suppl. 62): 32–43. doi:10.1159/000144209 . ISSN 1422-
6405 . 17. ^ Grabowski, M.; Jungers, W. L. (2017). "Evidence of a
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External links
Data related to Hominoidea at Wikispecies
Look up ape in Wiktionary,
Hominoidea at Wikibooks the free dictionary.
Pilbeam D. (September 2000). "Hominoid systematics: The soft evidence" . Proc. Natl. Acad. Sci. U.S.A. 97 (20): 10684–6. Bibcode:2000PNAS...9710684P . doi:10.1073/pnas.210390497 . PMC 34045 . PMID 10995486 . Agreement between cladograms based on molecular and anatomical data.
Human Timeline (Interactive) – Smithsonian, National Museum of Natural History (August 2016). Ape
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V ·T ·E Apes [show]

V ·T ·E Notable non-human apes [show]

V ·T ·E Extant primate families [show]

V ·T ·E Haplorhini [show]

Taxon identifiers Wikidata: Q102470 · Wikispecies: Hominoidea · ADW: Hominoidea · EoL: 4529848 · Fossilworks: 40883 · iNaturalist: 1036675 · ITIS: 943782 · MSW: 12100751 · NCBI: 314295

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Categories: Apes Extant Chattian first appearances Taxa named by John Edward Gray

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