×Chiranthomontodendron: The Correct Nothogeneric Name for the Artificial Hybrid

Chiranthodendron ×Fremontodendron (Malvaceae: Malvoideae)

Author(s): Laurence J. Dorr
Source: Taxon, Vol. 58, No. 4 (Nov., 2009), pp. 1357-1358
Published by: International Association for Plant Taxonomy (IAPT)
Stable URL: http://www.jstor.org/stable/27757023 .
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TAXON 58 (4) November 2009: 1357-1358 Dorr Correct name for Chiranthodendron x Fremontodendron

xChiranthomontodendron: the correct nothogeneric name for the artificial

x Fremontodendron
hybrid Chiranthodendron {Malvaceae: Malvoideae)

Laurence J. Dorr

Department of Botany, MRC-166, National Museum of Natural History, Smithsonian Institution, P.O. Box
37012, Washington, D.C. 20013-7012, U.S.A. dorrl@si.edu

The nothogeneric name xChiranthomontodendron is proposed to accommodate an artificial hybrid between

species of Chiranthodendron and Fremontodendron, and the nothospecies xChiranthomontodendron lenzii

is described from cultivated material.

KEYWORDS: Chiranthodendron, xChiranthofremontia, xChiranthomontodendron, Fremontodendron,

Malvaceae, nothogenus

(1991: 239) is unsatisfactory for validation of "*Chiran

I INTRODUCTION
A close relationship between Chiranthodendron
Larreat. andFremontodendron Coville (Malvaceae: Mal
voideae) has long been recognized. The former is unispe
cific and found in southernMexico and Guatemala, the
latter comprises three species (Kelman, 1991) and is re
stricted to northernMexico (Baja California) and adjacent
California andArizona. In addition, cultivars ofFremonto
dendron, most notably 'CaliforniaGlory' (F. californicum
x F. mexicanum
(Torr.) Coville Davidson), are found in
gardens far outside the native range of both genera.
Although often placed in theSterculiaceae (see dis
cussion inKelman, 1991),morphological and molecular
data now suggest thatChiranthodendron and Fremonto
dendron may be basal in a clade includingMalvoideae
mdBombacoideae (Baum & al, 2004, as "Malvatheca").
Not surprisingly,given the close relationship between the
two genera, an artificial cross between Chiranthodendron
(male parent) and Fremontodendron (female parent) was
completed in 1981 and the hybrid is now fairlywell es
tablished in gardens in California, at least. Henrickson
(1991),who described in detail and illustrated thishybrid,
proposed to call it"xChiranthofremontiaHenrickson." He
failed, however, to followArt. H.6.2 of theICBN (McNeill
& al, 2006), which states that the condensed formula for
a bigeneric hybrid consists of a single word "using the
firstpart or thewhole [name] of one [parent], the lastpart
or thewhole of the other (but not thewhole of both) and,
optionally, a connecting vowel." For whatever reasons,
Henrickson (1991) used only "Fremontia" for the sec
ond part of his condensed formula and this is neither the
whole name nor the last part of the name of this parent.
The nomenclatural consequence of his oversight is that
"xChiranthofremontia" is not a validly published name.
Complicating matters is the fact that the combined
generic and specific description supplied by Henrickson
thofremontia lenziiHenrickson" proposed by him for the
nothospecies involved.His description is, on close inspec
tion, a description of the nothogenus only, which is not
necessary under Art. H.9 (McNeill & al., 2006). Further
more, regardless ofwhether or not thenothogeneric name
was formed correctly, it appears that the nothospecies
name was not validly published because Art. 42 (McNeill
& al., 2006) cannot apply to a nothogeneric and nothospe
cific name as the former does not have a type.
I TAXONOMIC CONCLUSIONS
In order to provide this bigeneric hybrid with a val
idlypublished name I propose the following nothogeneric
name and describe the following nothospecies:
xChiranthomontodendron Dorr, nothogen. nov. [Chi
ranthodendron Larreat. xFremontodendron Coville].
xChiranthomontodendron lenzii Dorr, nothosp. nov.
-
Type: U.S.A., California, Riverside Co., "Hybrid:
Fremontodendron 'Pacific Sunset' (female) with
Cheiranthodendron [sic], crossed with extreme dif
ficulty in June of 1981" [plant grown inChino, Cali
fornia, at the home ofAustin Griffiths], 15 Jun 1985
(fl),A. Griffiths,Jr. 17109 (holotype, RSA-583216!;
isotype, CAS).
Hybrida hortensis e Fremontodendron 'Pacific Sun
set' et Chiranthodendron pentadactylon Larreat. genita,
foliis 5-7-lobatis, (4.5-)6.5-12 x
(4-)6-10 cm, basibus
cordatis profunde, supra parce stellato-tomentosis, subtus
dense stellato-tomentosis, petiolis (2-)4-6.5(-8) cm lon
gis et aliis characteribus interparentes media.
The precise type locality is inferredfrom information
inHenrickson (1991) as the label on theholotype ispeculiar

1357

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Dorr Correct name for Chiranthodendron x Fremontodendron TAXON 58 (4) November 2009: 1357-1358

and only indirectly suggests through itsheader ("Golden

State Plants") that the specimen was collected inCalifor LITERATURE
CITED
DeWitt
nia. The label does propose a name (never published) for Baum, D.A., Smith, S., Yen, A., Alverson, W.S.,
Nyffeler, R., Whitlock, B.A. & Oldham, R.L. 2004.
the artificial hybrid, states theparentage of thehybrid and
Phylogenetic relationships ofMalvatheca (Bombacoideae
thedate the crosswas made, describes the etymology of the sensu lato) as inferred from
and Malvoideae; Malvaceae
unpublished name, states the collector, collector's number plastid DNA sequences. Amer. J. Bot. 91: 1863-1871.
and date when theholotype was collected, and provides a Henrickson, J. 1991. xChiranthofremontia, an intergeneric

general description of the hybrid plant. hybrid of Chiranthodendron pentadactylon and Fremon

To avoid additional confusion, I have not only re todendron 'Pacific Sunset'. Aliso 13: 239-248.

Kelman, W.J. 1991. A revision of Fremontodendron (Stercu

tained for the nothospecies the epithet proposed by Hen
liaceae). Syst. Bot. 16: 3-20.
rickson (1991),which commemorates Lee W. Lenz, Direc
McNeill, J., Barrie, RR., B?rdet, H.M., Demoulin, V.,
torEmeritus of theRancho Santa Ana Botanic Garden,
Hawksworth, D.L., Marhold,
K., Nicolson, D.H., Prado,
Claremont, California, but also utilized as holotype the J., Silva, P.C., Skog, J.E., Wiersema, J.H. & Turland,
same specimen as Henrickson proposed for this purpose. N.J. (eds.). 2006. International Code of Botanical Nomen
The collector's number for the latterwas miscited inHen clature (Vienna Code) Adopted by the Seventeenth Inter

rickson (1991, as "171097"). national Botanical Congress, Vienna, Austria, July 2005.
Regnum Vegetabile 146. Gantner, Ruggell.

I ACKNOWLEDGMENTS
Fred Barrie and John McNeill set me on the correct path
to giving thisnothogenus and nothospecies validly published
names. The curator of Rancho Santa Ana Botanic Garden

kindly lent the specimen that serves as the holotype.

1358

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