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Sensory and Motor System
Sensory and Motor System
December 5, 2018
Sensory and Motor System
All bilaterally symmetric organism’s posses a sensory system, and the development of
any species’ sensory system has been driven by natural selection; thus, sensory systems
differ among species according to their history of natural selection. For example, the
shark, unlike most fish predators, is electrosensitive- that is, sensitive to electrical fields
produced by other animals in its environment.
Humans and many other vertebrates have at least five special senses: olfaction (smell),
gustation (taste), equilibrium (balance and body position), vision, and hearing.
Additionally, we possess general senses, also called somatosensation, which respond to
stimuli like temperature, pain, pressure, and vibration.
Description of structure and function of the organ system (can use the textbook as main
document, and other relevant information about the organ system.
The sensory system detects signals from the outside environment and communicates it
to the body via the nervous system. The sensory system relies on specialized sensory
receptor cells that translate external stimuli into changes in membrane potentials. If the
changes in membrane potential are sufficient to induce an action potential, then these
action potentials are then communicated along neurons within the afferent division of
the PNS to the CNS for information processing. The CNS integrates and interprets the
incoming signals to effect a response to the appropriate body systems via the efferent
division of the PNS.
Sensory receptor cells can be:
A light touch activates only the mechanoreceptors near the upper layer of the skin, while
a firmer touch activates mechanoreceptors deeper in the skin, in addition to the
mechanoreceptors near the surface of the skin. A firmer touch will also activate a greater
number of receptors, and may induce more frequent action potentials in the receptors
than a lighter touch.,
Frequency is the number of waves per unit of time, which is heard as pitch.
Frequency is also related to wavelength, where high-frequency (15,000 Hz) sounds
are higher-pitched and shorter wavelength than low-frequency, long wavelength
(100 Hz) sounds. Most humans can perceive sounds with frequencies between 30
and 20,000 Hz. Dogs detect up to about 40,000 Hz; cats, 60,000 Hz; bats, 100,000
Hz; and dolphins 150,000 Hz, and some fish can hear 180,000 Hz.
Amplitude, or the dimension of a wave from peak to trough, in sound is heard
as volume. The sound waves of louder sounds have greater amplitude than those of
softer sounds. For sound, volume is measured in decibels (dB). In the figure below,
the softest sound that a human can hear is the zero point. Humans speak normally at
60 decibels.
Outer ear:
Sound waves are collected by the external, cartilaginous part of the ear
Sound waves then travel through the auditory canal and cause vibration of
the ear drum (tympanic membrane)
Middle ear:
The eardrum transmits sound to the middle ear by vibrating
the ossicles, the three small bones of the middle ear which collect force
and amplify sounds. The three ossicles are unique to mammals.
Inner ear:
The ossicles transmit the vibrations to a thin membrane called the oval
window, which is the outermost structure of the inner ear.
The vibrations of the oval window create pressure waves in the fluid inside
the cochlea. The cochlea is a whorled structure, like the shell of a snail,
and it contains receptors for transduction of the mechanical wave into an
electrical signal
Balance and Movement: The Vestibular System
The stimuli associated with the vestibular system are linear acceleration (gravity) and angular
acceleration and deceleration. Gravity, acceleration, and deceleration are detected by evaluating
the inertia on receptive cells in the vestibular system. In vertebrates, gravity is detected through
head position. Angular acceleration and deceleration are expressed through turning or tilting of
the head.
The vestibular system in vertebrates has some similarities with the auditory system. It utilizes
hair cells located within the ear in a structure called the vestibular labyrinth (located adjacent to
the cochlea), but it activates them in a different way compared to the auditory system. Hair cells
in the vesibular labyrinth detect signals in two ways:
Some hair cells lie below a gelatinous layer, with their stereocilia projecting into the gelatin.
Embedded in this gelatin are calcium carbonate crystals, like tiny rocks, that move in
response to gravity. Any time the head is tilted at an angle or is subject to acceleration or
deceleration, these crystals cause the gelatin to shift, bending the stereocilia. The bending of
the stereocilia stimulates the neurons, and they signal to the brain that the head is tilted,
allowing the maintenance of balance.
Some hair cells project into a gelatinous cap called the cupula. When the head turns, the fluid
in the canals shifts, thereby bending stereocilia and sending signals to the brain. When
movement stops, the movement of the fluid within the canals slows or stops.
Photoreceptors: Vision
As with auditory stimuli, light travels in waves. The pressure waves that create sound must travel
in a medium: a gas, a liquid, or a solid. In contrast, light is composed of electromagnetic waves
and needs no medium; light can travel in a vacuum (this is why we can see stars from space.)
The behavior of light can be discussed in terms of the behavior of waves and also in terms of the
behavior of the fundamental unit of light: a packet of electromagnetic radiation called a photon.
Humans can perceive just a small slice of the entire electromagnetic spectrum, which includes
radiation that we cannot see as light because it is below the frequency of visible red light and
above the frequency of visible violet light, which are the limits of vertebrate light detection.
Wavelength (which varies inversely with frequency) is detected as hue, or color. Light at the
red end of the visible spectrum has longer wavelengths (and is lower frequency), while light
at the violet end has shorter wavelengths (and is higher frequency). The wavelength of light
is expressed in nanometers (nm); one nanometer is one billionth of a meter. Humans perceive
light that ranges between approximately 380 nm and 740 nm. Some other animals, though,
can detect wavelengths outside of the human range. For example, bees see near-ultraviolet
light in order to locate nectar guides on flowers, and some non-avian reptiles sense infrared
light (heat that prey gives off).
Wave amplitude is perceived as luminous intensity, or brightness.
Detection of light occurs through photoreceptors, cells that contain pigment-
absorbing molecules that absorb light. Photoreceptor cells are typically located in light-
collecting organs called eyes. Eyes vary in structure in different types of animals and
include:
eye cups in flatworms, which are dimple-shaped structures that detect the direction
of a light source
compound eyes of arthropods, which contain multiple lenses and detect shapes,
patterns, and movements
pinhole eyes in the nautilus, which contain no lens and forms simple, low-
resolution images
simple eyes of cephalopods and vertebrates, which contain a single lens and form
high-resolution images
Many of these types of eyes, which are present in living organisms today, may have also
represented a pathway of evolution from a simple patch of photosensitive cells to simple
lens eyes of vertebrates and cephalopods:
Detecting a taste relies on activation of specific chemical receptors in taste receptor cells
(gustatory receptors). When the specific chemical (tastant) binds the receptor, the
receptor cell becomes depolarized and releases neurotransmitter on its synapsed
afferent neuron. As in other sensory systems, specificity in taste occurs because each
taste receptor cell has only one type of protein receptor which is sensitive to either
sweet, sour, bitter, salty, or umami. The process of depolarization differs based on the
specific type of gustatory receptor:
A salty tastant (containing NaCl) provides the sodium ions (Na ) that enter the taste
+
All odors that we perceive are molecules in the air we breathe. If a substance does not
release molecules into the air from its surface, it has no smell. And if a human or other
animal does not have a receptor that recognizes a specific molecule, then that molecule
has no smell. Humans have about 350 olfactory receptor subtypes that work in various
combinations to allow us to sense about 10,000 different odors. (As you may be
expecting by now, each olfactory receptor cell has only one type of olfactory receptor
protein, meaning each receptor cell is specific to only one type of odorant.) For
comparison, mice have about 1,300 olfactory receptor types and therefore probably
sense many more odors than do humans.
Taste receptors are responsible for the sense of how salty, sweet, bitter, sour, or
savory a food is, via tastants dissolved in saliva
Olfactory receptors are responsible for the flavor of a food, via odorants detected in
the olfactory epithelium during chewing, through a process called retronasal
olfaction (the flow of air from the back of the throat up to the olfactory epithelium
via the back of the nose)
Nociceptors: Tissue Damage and Pain
Pain is the name given to nociception, which is the neural processing in response to
tissue damage. Pain is caused by both true sources of injury, such as contact with a
corrosive chemical, and also by harmless stimuli that mimic the action of damaging
stimuli, such as contact with capsaicins, the compounds that cause peppers to taste hot
(and which are used in self-defense pepper sprays and certain topical medications).
Peppers taste “hot” because the protein receptors that bind capsaicin open the same
calcium channels that are activated by heat-sensitive thermoreceptors.
There are many different types of nociceptors and we will not describe them in this
course, but the important thing to understand is that different types of nociceptors are
activated by different types of tissue damage, including extremes of hot or cold, toxic
chemicals, and extreme mechanical deformation such as stretching, cuts, and tears.
Nociception starts at the sensory receptors, but pain does not actually start until it is
communicated to the brain; pain is the interpretation of the tissue damage, not the
actual stimulus itself. There are several nociceptive pathways to and through the brain,
including through the thalamus (like most other sensory systems) and directly to the
hypothalamus , which modulates the cardiovascular and neuroendocrine functions of the
autonomic nervous system, where it can directly activate the fight-or-flight response.
sensory system:
Parkinson’s disease
-Pain, akathisia, olfactory loss, visual impairment, vestibular dysfunction, proprioceptive and
kinaesthetic dysfunction, and sensory cueing
Dystonias
-Pain, photosensitivity, alleviating manoeuvres, kinaesthetic dysfunction, abnormal temporal
and spatial discrimination
Peripherally induced dystonia, tremor, other movement disorders, and complex regional
pain syndrome
-Pain, paraesthesias
Tics and Tourette’s syndrome
-Premonitory urge phenomena, enhanced sensory perception, alleviating manoeuvres
Restless legs syndrome
-Urge phenomena, reduction of urge with bright lights
Akathisia
-Urge phenomena, reduction with passive motion (perception of movement)
Stereotypies
-Urge phenomena
Tardive Pain
-Painful mouth and vagina syndrome, and phantom dyskinesias
Leg Stereotopy Disorder
-Urge phenomena
Paroxysmal kinesigenic and non-kinesigenic dyskinesias
-Numbness, paraesthesias, crawling sensations in legs
Latest findings about the organ system (at least from two sources)
Potential Mechanisms of Sensory Augmentation Systems on
Human Balance Control
Numerous studies have demonstrated the real-time use of visual, vibrotactile, auditory, and
multimodal sensory augmentation technologies for reducing postural sway during static tasks and
improving balance during dynamic tasks.
https://doaj-org.proxy.lib.fsu.edu/article/267b2f025bfc417da303deb359637b73
https://doaj-org.proxy.lib.fsu.edu/article/e1410a2f68f3426782639fdfb1c00a4e
The capsaicin precent in peppers activates the thermoreceptor for high temperatures. In
response to the perceived high temp, the nervous system triggers sweating to achieve
evaporative cooling
3. if you stimulated a sensory neuron electrically, how would that stimulation be perceived?
You would perceive the electrical stimulus as if the sensory receptors that regulate that neuron
had been activated. Ex: electrical stimulation of the sensory neuron controlled by the
thermoreceptor activated by menthol would likely be perceives as a local cooling
Otoliths detect the animals’ orientation with respect to gravity, providing information that is
essential in environments such as the tunnel habitat of the star nosed mole, where light cues
are absent
2. Suppose a series of pressure waves in your cochlea caused a vibration of the basilar
membrane that moves gradually from the apex toward the base. how would your brain
interpret this stimulus?
As a sound that changes gradually from a very low to a very high pitch
3.If the stapes became fused to the other middle ear bones or to the oval window, how would
this condition affect hearing? explain.
The stapes and other middle ear bones transmit vibrations from the tympanic membrane to the
oval window. Fusion of these bones would block this transmission and result in hearing loss
4. plants use statoliths to detect gravity (see Figure 39.22). how do plants and animals differ
with regard to the type of compartment in which statoliths are found and the physiological
mechanism for detecting their response to gravity?
In animals, the statoliths are extracellular. In contrast, the statoliths of plants are found within an
intracellular organelle. The methods for detecting their location also differ. In animals, detection
is by means of mechanoreceptors on ciliated cells. In plants, the mechanism appears to involve
calcium signaling.
Planarians have ocelli that cannot form images but can sense the intensity and direction of
light. Flies have compound eyes that form mosaic images and excel at detecting movement
2. In a condition called presbyopia, the eyes’ lenses lose much of their elasticity and maintain a
flat shape. explain how this condition affects a person’s vision.
the person can focus on distant objects but not close objects because close focusing requires
the lens to become almost spherical
3.Our brain receives more action potentials when our eyes are exposed to light even though
our photoreceptors release more neurotransmitter in the dark. propose an explanation.
4.Compare the function of retinal in the eye with that of the pigment chlorophyll in a plant
photosystem
Absorption of light by retinal converts retinal from its cis isomer to its trans isomer, initiating the
process of light detection. In contrast, a photon absorbed by chlorophyll does not bring about
isomerization, but instead boosts anelectron to a higher energy orbital, initiating the electron to
flow that generates ATP and NADPH.
1.Explain why some taste receptor cells and all olfactory receptor cells use G protein-coupled
receptors, yet only olfactory receptor cells produce action potentials.
Both taste cells and olfactory cells have receptor proteins in their plasma membrane that
bind certain substances, leading to membrane depolarization through a signal transduction
pathway involving G protein. However, olfactory cells are sensory neurons, whereas taste
cells are not
2. Pathways involving G proteins provide an opportunity for an increase in signal strength in the
course of signal transduction, a change referred to as amplification. how might this be
beneficial in olfaction?
Since animals rely on chemical signals for behaviors that include finding mates, marking
territories and avoiding dangerous substances, it is adaptive for the olfactory system to have a
robust response to a very small number of molecules of a particular odoran.
2+
1. Contrast the role of Ca in the contraction of a skeletal muscle fiber and a smooth muscle
cell.
In skeletal muscle fiver, Ca+ binds to the troponin complex, in smooth muscle cells Ca+ binds to
calmodulin.
2. Why are the muscles of an animal that has recently died likely to be stiff?
Rigor mortis, results from the complete depletion of ATP in skeletal muscle. Since ATP is
required to release myosin from actin and the pump Ca2+ out of the cytosol, muscles become
chronically contracted beginning about 3-4 hours after death.
Septa provide the divisions of coelom that allow for peristalsis, a form of locomotion requiring
independent control of different body segments.
2. Contrast swimming and flying in terms of the main problems they pose and the adaptations
that allow animals to overcome those problems.
The main problem in swimming is the drag; a fusiform body minimizes drag. The main problem
in flying is overcoming gravity; wings are shaped like airfoils to provide lift and airfield bones
reduce body mass
3. When using your arms to lower yourself into a chair, you bend your arms without using your
biceps. explain how this is possible.
When you grasp sides of chair, you are using a contraction of the triceps to keep your arms
extended against the pull of gravity on your body. As you lower yourself slowly onto chair, you
gradually decrease the number of motor units in the triceps that are contracted. Contracting
your biceps would jerk you down, since you would no longer be opposing gravity