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Biolympiads - Systematics
Biolympiads - Systematics
Biolympiads - Systematics
Agrobacterium Agrobacterium is a genus of Gram-negative bacteria that uses horizontal gene transfer to cause tumors in
plants. Agrobacterium tumefaciens is the most commonly studied species in this genus. Agrobacterium is well
known for its ability to transfer DNA between itself and plants, and for this reason it has become an important
tool for genetic engineering.
A. tumefaciens causes crown-gall disease in plants. The disease is characterised by a tumour-like growth or gall
on the infected plant, often at the junction between the root and the shoot. Tumors are incited by the conjugative
transfer of a DNA segment (T-DNA) from the bacterial tumour-inducing (Ti) plasmid.
Escherichia Escherichia is a genus of Gram-negative, nonspore forming, facultatively anaerobic, rod-shaped bacteria. In
those species which are inhabitants of the gastrointestinal tracts of warm-blooded animals, Escherichia species
provide a portion of the microbially derived vitamin K for their host. A number of the species of Escherichia are
pathogenic.
While many Escherichia are commensal gut flora, particular strains of some species are known as the most
common cause of urinary tract infections, significant sources of gastrointestinal disease, ranging from
simple diarrhea to dysentery-like conditions.
Rhizobium Rhizobium is a genus of Gram-negative soil bacteria that fix nitrogen. Rhizobium species form an
endosymbiotic nitrogen-fixing association with roots of legumes.
The bacteria colonize plant cells within root nodules, where they convert atmospheric nitrogen into ammonia and
then provide organic nitrogenous compounds such as glutamine or ureides to the plant. The plant, in turn,
provides the bacteria with organic compounds made by photosynthesis. Thus, it acts as a natural fertilizer for the
plants.
Salmonella Salmonella is a genus of rod-shaped (bacillus) Gram-negative bacteria of the Enterobacteriaceae family.
S. enterica subspecies are found worldwide in all warm-blooded animals, and in the environment. S. bongori is
restricted to cold-blooded animals, particularly reptiles. Strains of Salmonella cause illnesses such as typhoid
fever, paratyphoid fever, and food poisoning (salmonellosis).
Anabaena Anabaena is a genus of filamentous cyanobacteria that exist as plankton. They are known for nitrogen-fixing
abilities, and they form symbiotic relationships with certain plants, such as the mosquito fern. They are one of
four genera of cyanobacteria that produce neurotoxins, which are harmful to local wildlife, as well as farm
animals and pets. Production of these neurotoxins is assumed to be an input into its symbiotic relationships,
protecting the plant from grazing pressure.
They have heterocysts, which convert nitrogen into ammonia. Certain species of Anabaena have been used on
rice paddy fields, proving to be an effective natural fertilizer.
Under nitrogen-limiting conditions, vegetative cells differentiate into heterocysts at semiregular intervals along
the filaments. Heterocyst cells are terminally specialized for nitrogen fixation. The interior of these cells is micro-
oxic as a result of increased respiration, inactivation of O2-producing photosystem (PS) II, and formation of a
thickened envelope outside of the cell wall. Nitrogenase, sequestered within these cells, transforms dinitrogen
into ammonium at the expense of ATP and reductant—both generated by carbohydrate metabolism, a process
supplemented, in the light, by the activity of PS I. Carbohydrate, probably in the form of glucose, is synthesized
in vegetative cells and moves into heterocysts. In return, nitrogen fixed in heterocysts moves into the vegetative
cells, at least in part in the form of amino acids.
Streptomycetes are characterised by a complex secondary metabolism. They produce over two-thirds of the
clinically useful antibiotics of natural origin (e.g., neomycin, cypemycin, grisemycin, bottromycins and
chloramphenicol). Streptomycetes are infrequent pathogens, though infections in humans, such as mycetoma, can
be caused.
ARCHAEA
Methanobacterium Anaerobic, non‐motile, use carbon dioxide to oxidize hydrogen gas to make methane, rod shaped, found
in the GI tract of herbivores.
Halobacterium The genus Halobacterium ("salt" or "ocean bacterium") consists of several species of the Archaea with an
aerobic metabolism which requires an environment with a high concentration of salt; many of their proteins
will not function in low-salt environments. They grow on amino acids in their aerobic conditions. Their cell
walls are also quite different from those of bacteria, as ordinary lipoprotein membranes fail in high salt
concentrations.
They reproduce using binary fission (by constriction), and are motile.
Thermoplasm Thermoplasma is a genus of archaea. It belongs to the Thermoplasmata, which thrive in acidic and high-
temperature environments. Thermoplasma are facultative anaerobes and respire using sulfur and organic
carbon.
They do not contain a cell wall but instead contain a unique membrane composed mainly of a tetraether
lipoglycan containing atypical archaeal tetraether lipid attached to a glucose- and mannose-containing
oligosaccharide. This lipoglycan is presumably responsible for the acid and thermal stability of the
Thermoplasma membrane.
Sulfolobus Sulfolobus species grow in volcanic springs with optimal growth occurring at pH 2-3 and temperatures of
75-80 °C, making them acidophiles and thermophiles respectively. Sulfolobus cells are irregularly shaped
and flagellar.
EUKARYA
PROTISTA
Euglenozoa Euglena Euglena is a genus of single-celled flagellate Eukaryotes. Species of Euglena are found in fresh and
salt waters.
Most species of Euglena have photosynthesizing chloroplasts within the body of the cell, which
enable them to feed by autotrophy, like plants. However, they can also take nourishment
heterotrophically, like animals. Thus, they are mixotrophs.
They possess chloroplasts through secondary endosymbiosis of green algae (thus their chloroplasts
have three membranes).
Stramenopiles Paheophyta (Brown Algae) Sargasum Any number of the normally benthic
species may take on a planktonic,
often pelagic existence after being
removed from reefs during rough
weather.
Flower
• Pedicel jointed,
• Bracteate,
• Large,
• Showy,
• Pentamerous,
• Bracteolate,
• Bisexual,
• Dichlamydeous,
• Actinomorphic,
• Complete and
• Hypogynous and
• Mucilage exists in floral parts.
Euphorbiaceae Euphorbia The Euphorbiaceae, the spurge family,
is a large family of flowering plants.
Bambusa
Cyperaceae Cyperus The Cyperaceae are a family of
monocotyledonous graminoid
flowering plants known as sedges,
which superficially resemble grasses
and rushes.
In older classification systems, most amoebas were placed in the class or subphylum Sarcodina, a
grouping of single-celled organisms that possess pseudopods or move by protoplasmic flow.
Amoebae move and feed by using pseudopods, which are bulges of cytoplasm formed by the
coordinated action of actin microfilaments pushing out the plasma membrane that surrounds the cell.
Eumycetozoa Dictyostelium Slime mold or slime mould is an informal name given to several kinds of unrelated eukaryotic
(Slime molds) organisms that can live freely as single cells, but can aggregate together to form multicellular
reproductive structures. Slime molds were formerly classified as fungi but are no longer considered part
of that kingdom.
Many slime molds, mainly the "cellular" slime molds, do not spend most of their time in this state. As
long as food is abundant, these slime molds exist as single-celled organisms. When food is in short
supply, many of these single-celled organisms will congregate and start moving as a single body. In this
state they are sensitive to airborne chemicals and can detect food sources. They can readily change the
shape and function of parts and may form stalks that produce fruiting bodies, releasing countless spores,
light enough to be carried on the wind or hitch a ride on passing animals.
Fungi Zygomycota Mucor Zygomycete hyphae may be coenocytic, forming septa only where gametes are formed or to wall off
dead hyphae.
Zygomycota are defined and distinguished from all other fungi by sexual reproduction via zygospores
following gametangial fusion and asexual reproduction by uni-to-multispored sporangia within which
nonmotile, single-celled sporangiospores are produced.
Zygomycota, like all true fungi, produce cell walls containing chitin. They grow primarily as mycelia,
or filaments of long cells called hyphae. Unlike the so-called 'higher fungi' comprising the Ascomycota
and Basidiomycota which produce regularly septate mycelia, most Zygomycota form hyphae which are
generally coenocytic because they lack cross walls or septa.
The unique character (synapomorphy) of the Zygomycota is the zygospore. Zygospores are formed
within a zygosporangium after the fusion of specialized hyphae called gametangia during the sexual
cycle. A single zygospore is formed per zygosporangium. The mature zygospore is often thick-walled,
and undergoes an obligatory dormant period before germination. Most Zygomycota are thought to
have a zygotic or haplontic life cycle. Thus, the only diploid phase takes place within the zygospore.
Ascomycota Claviceps The Ascomycota, or sac fungi, is monophyletic and accounts for approximately 75% of all described
fungi. It includes most of the fungi that combine with algae to form lichens, and the majority of fungi
that lack morphological evidence of sexual reproduction. Among the Ascomycota are some famous
fungi: Saccharomyces cerevisiae, the yeast of commerce and foundation of the baking and brewing
industries (not to mention molecular developmental biology), Penicillium chrysogenum, producer of
penicillin, Morchella esculentum, the edible morel, and Neurospora crassa, the "one-gene-one-
enzyme" organism.
The shared derived character that defines the Ascomycota is the ascus. It is within the ascus that
Penicillium
nuclear fusion and meiosis take place. In the ascus, one round of mitosis typically follows meiosis to
leave eight nuclei, and eventually eight ascospores. Ascospores are formed within the ascus by an
enveloping membrane system, which packages each nucleus with its adjacent cytoplasm and provides
the site for ascospore wall formation.
Ascomycota are heterotrophs and obtain nutrients from dead or living organisms. The textbook
Ascomycota can make spores sexually (ascospores or meiospores) and asexually (condia or
mitospores). Following meiosis, the ascospores take shape inside the ascus when new cell walls
surround each nucleus.
Ascomycota are either single-celled (yeasts) or filamentous (hyphal) or both (dimorphic). Yeasts grow
Saccharomyces by budding or fission and hyphae grow apically and branch laterally. Most yeasts and filamentous
Ascomycota are haploid, but some species, Saccharomyces cerevisiae for example, can also be diploid.
Mitospores may simply reproduce the parent, or may also act as gametes to fertilize a compatible
partner. Some Ascomycota must outbreed (heterothallic), others can also self, and some can only self
(homothallic).
Basidiomycota Agaricus Symbiotic lifestyles (intimate associations with other living organisms) are well developed in the
Basidiomycota. Symbiotic Basidiomycota include important plant pathogens, such as "rusts"
(Uredinales) and "smuts" (Ustilaginales), which attack wheat and other crops. Other symbiotic
Basidiomycota cause diseases in animals, including humans.
Basidiomycota are unicellular or multicellular, sexual or asexual, and terrestrial or aquatic. The most
Puccinia diagnostic feature is the production of basidia (sing. basidium), which are the cells on which sexual
spores are produced, and from which the group takes its name. A long-lived dikaryon, in which each
cell in the thallus contains two haploid nuclei resulting from a mating event, is another characteristic
feature.
The basidium is the cell in which karyogamy (nuclear fusion) and meiosis occur, and on which
haploid basidiospores are formed (basidia are not produced by asexual Basidiomycota). Many
Basidiomycota produce basidia on multicellular fruiting bodies (e.g., mushrooms), but basidia can also
be formed directly from yeasts or other single cells. Typically, four spores are produced on each
basidium, at the tips of minute stalks called sterigmata. Each spore usually contains one or two of the
haploid meiotic products.
Basidiospores germinate to form hyphae (filaments) or yeast cells that are typically haploid and
uninucleate. The hyphae of Basidiomycota are septate. Ultrastructural features of the septa, visible
with transmission electron microscopy, have been important in developing phylogenetic hypotheses in
Basidiomycota. Mating in Basidiomycota involves fusion of haploid cells, but fusion of the nuclei is
usually delayed until the basidia are formed. Thus, the dominant phase of the life cycle in most
Basidiomycota is a dikaryon, in which the two nuclei brought together in mating exist side-by-side in
each cell.
Metazoa Porifera Spongia Sponges (phylum Porifera) are an exclusively aquatic and, with a few exceptions.
(Animalia)
It is also often mentioned that sponges lack many characteristics associated with other animals,
including a mouth, sensory organs, organized tissues and neurons and muscle cells, which are
otherwise ubiquitous in Metazoa. It is difficult to say, however, whether the lack of aforementioned
features represents a primitive condition of sponges or a secondary loss due to their sedentary and
water-filtering lifestyle.
The body plan of a sponge is simple: a Traditionally, sponges have been
single outer layer of cells (the regarded as a monophyletic group
pinacoderm) separates the inner cellular defined by several synapomorphies,
region (mesohyl) from the external including the presence of:
environment. The pinacoderm lines the
internal canals and is eventually • choanocytes
replaced by the choanoderm, a layer of • an aquiferous system with
characteristic flagellated collar cells external pores
(choanocytes) grouped in chambers. • mineral spicules
Choanocytes make up the principle • high cellular mobility and
‘pump’ and’ filter’ of the system, totipotency
driving water through the sponge, • absence of tissues
trapping and phagocytizing suspended
bacteria and other particulate food,
which is then digested and nutrients
distributed among the cells of the
mesohyl that facilitate the functions of
feeding, respiration and reproduction.
The flow of water inside a sponge is
unidirectional: the water is drawn in
through tiny pores (ostia) in the
pinacoderm and exits through one or
more larger openings (osculae). The
aquiferous system of a sponge is usually
supported by a combination of two
types of skeletal elements: mineral
spicules (either calcareous or siliceous)
and special protein fibers (spongin),
although either one or both of these
elements can be absent.
Cnidaria Anthozoa Anemonia The exclusively aquatic phylum Cnidaria is represented by polyps such as
sea anemones and corals, and by medusae such as jellyfish. A polypoid or a
medusoid cnidarian is a radially or biradially symmetrical, uncephalized
animal with a single body opening, the mouth. The mouth is surrounded by
tentacles studded with microscopic stinging capsules known as
nematocysts that are the agents of offense and defense. The possession of
intrinsic nematocysts is the defining characteristic of the phylum;
nematocysts are the most diverse and widespread of three types of cnidae
(cnidos = thread) -- hence the preferred name of the phylum.
Corallium
Scyphozoa Aurelia
Cnidarians are diploblastic -- that is, the body and tentacles consist of two
cell layers, the endoderm (sometimes referred to as the gastrodermis) and
the ectoderm (the epidermis). Between the two cell layers is the mesoglea,
which ranges from little more than a glue to bind the layers (for example, in
Hydra) to the vast bulk of the animal (for example, in jellyfish of Class
Scyphozoa). The body encompasses a single sac-like body space, the
coelenteron (koilos = cavity; enteron = intestine), which communicates
with the surrounding medium through the mouth. The less preferred name
of the phylum, Coelenterata, is based on this attribute. The coelenteron
(also termed the gastrovascular cavity) serves for gas exchange and
digestion.
All cnidarians are carnivorous, with cnidae and tentacles active in prey
capture. Because polyps are typically sessile, and only some medusae
possess sensory structures, cnidarians are generally believed to be passive
predators, feeding on prey items that blunder into their tentacles. Some
cnidarians can absorb dissolved organic matter directly from seawater, but
it is not known how widespread this ability is. Living within the tissues of
anthozoans of many species and hydrozoans and scyphozoans of a small
number of species are unicellular algae from which the animals derive
reduced carbon. Dinoflagellate symbionts, termed zooxanthellae, are by far
the most common algal symbionts; they are exclusively marine. Green
algal symbionts, termed zoochlorellae, occur in both marine and freshwater
cnidarians.
The cnida, or nematocyst, which is the sine qua non of the phylum, is
secreted by the Golgi apparatus of a cell termed a cnidoblast. A cnida
therefore is technically not an organelle, but, rather, the most complex
secretory product known. Upon receiving the appropriate physical and/or
chemical stimulus, a cnida fires, everting a tubule many times the length of
the capsule. The tubule may deliver a toxin, may stick to a prey item, or
may entangle an object, depending on the type of cnida. A cnida can fire
but once. There are three major types of cnidae: nematocysts, spirocysts,
and ptychocysts. Nematocysts occur in all classes of Cnidaria, but some of
the 30-plus varieties of nematocysts are restricted to members of certain
classes. Spirocysts are found only in Anthozoa; they are adhesive in nature.
Ptychocysts are the most taxonomically restricted in distribution, occurring
only in the anthozoan order Ceriantharia; their function is to entangle bits
of mud among their robust tubules to form the feltwork that constitutes the
tube of these burrowing animals.
Two body forms are characteristic of cnidarians -- the polyp and the
medusa. With a few exceptions, a columnar polyp is sedentary, being
attached to or burrowed into the substratum by the end opposite the mouth.
Thus, its tentacles are typically considered to point upward and outward.
Polyps of some species propagate vegetatively, forming colonies (if the
progeny remain attached to one another) or clones (if the progeny separate).
Polymorphism occurs in colonies of some species of hydrozoans and
anthozoans, the polyps being specialized for functions such as feeding,
defense, and sexual reproduction. Polyps of some taxa form a skeleton
within or external to their tissues; some skeletons are mineralic (of calcium
carbonate), others are organic (of chitin or another carbohydrate), and some
are both. The spheroidal or discoidal medusae are solitary, and those of
most species are pelagic. Although typically depicted as living with mouth
and tentacles pointing down, medusae assume all orientations in the water.
Medusae of few species possess the ability to propagate vegetatively. The
common name of medusae, jellyfish, alludes to the massive amount of
mesoglea that contributes to their buoyancy.
Ctenophora Mertensia Ctenophores (Greek for "comb-bearers") have eight "comb rows" of fused cilia arranged along the
sides of the animal, clearly visible along the red lines in these pictures. These cilia beat synchronously
and propel ctenophores through the water. Some species move with a flapping motion of their lobes or
undulations of the body. Many ctenophores have two long tentacles, but some lack tentacles
completely.
Ctenophores, variously known as comb jellies, sea gooseberries, sea walnuts, or Venus's girdles, are
voracious predators. Unlike cnidarians, with which they share several superficial similarities, they lack
stinging cells. Instead, in order to capture prey, ctenophores possess sticky cells called colloblasts. In a
few species, special cilia in the mouth are used for biting gelatinous prey.
Bilateria Platyhelminthes Trematoda Schistosoma Characteristics of Platyhelminthes:
1)Bilaterally symmetrical.
2)Body having 3 layers of tissues with organs
and organelles.
3)Body contains no internal cavity.
4)Possesses a blind gut (i.e. it has a mouth but
no anus)
5)Has Protonephridial excretory organs instead
of an anus.
Turbellaria Pseudoceros 6)Has normally a nervous system of
longitudinal fibres rather than a net.
7)Generally dorsoventrally flattened.
8)Reproduction mostly sexual as
hermaphrodites.
9)Mostly they feed on animals and other
smaller life forms.
10)Some species occur in all major habitats,
including many as parasites of other animals.
The muscles are activated by two nerves that run the length of the
nematode on both the dorsal (back) and ventral (belly) side. Unlike
other animals, where the nerves branch out to the muscle cells, a
nematode's muscle cells branch toward the nerves. The ventral nerve
has a series of nerve centers along its length, and both nerves connect
to a nerve ring and additional nerve centers located near the head.
The head of a nematode has a few tiny sense organs, and a mouth
opening into a muscular pharynx (throat) where food is pulled in and
crushed. This leads into a long simple gut cavity lacking any muscles,
and then to an anus near the tip of the body. Food digested in the gut
is not distributed by any specialized vascular system, and neither is
there a respiratory system for the uptake or distribution of oxygen.
Rather, nutrients and waste are distributed in the body cavity, whose
contents are regulated by an excretory canal along each side of the
body.
Many nematodes are able to suspend their life processes completely
when conditions become unfavorable; in these resistant states they
can survive extreme drying, heat, or cold, and then return to life when
favorable conditions return. This is known as cryptobiosis, and is a
feature nematodes share with rotifers and tardigrades.
Arthropoda See below
ARTHROPODA
Chelicerata Opiliones Phalangium The subphylum Chelicerata is one of the five subdivisions of the phylum Arthropoda, with members
characterized by the absence of antennae and mandibles (jaws) and the presence of chelicerae (a pincer-like
mouthpart as the anterior appendage, composed of a base segment and a fang portion). Extant chelicerates
include spiders, scorpions, ticks, and mites (class Arachnida), horseshoe crabs (class Xiphosura or
Merostomata), and sea spiders (class Pycnogonida).
As with all arthropods, chelicerates are characterized by the possession of a segmented body, a pair of
jointed appendages on each segment, and an exoskeleton.
In the Chelicerata, the body is divided into two parts. The anterior part is called a prosoma (or
Scorpionida Pandinus cephalothorax) and is composed of eight segments plus a presegmental acron. The posterior part is called a
opisthosoma (or abdomen) and is composed of twelve segments plus a postsegmental telson.
The prosoma usually has eyes. The first two segments of the prosoma bear no appendages; the third bears
the chelicerae. The fourth segment bears legs or pedipalps, and all subsequent segments bear legs. The legs
on the prosoma are either uniramous or have a very reduced gill branch, and are adapted for walking or
swimming. The appendages on the opisthosoma, in contrast, are either absent or are reduced to their gill
branch.
Acari Sarcoptes
As in other arthropods, the mouth lies between the second and third segments, but whereas in other groups
there is usually a pair of antennae on the last preoral segment, here there are none.
The chelicerae, which give the group its name, are pointed appendages that grasp the food in place of the
chewing mandibles most other arthropods have. Most chelicerates are unable to ingest anything solid, so
they drink blood or spit or inject digestive enzymes into their prey.
Ixodes
Araneae Haplopelma
Nephila
Myriapoda Chilopoda Scolopendra Centipedes have long, flattened, worm-like, segmented bodies. They can have 12-100 body segments. They
(centipedes) have long, jointed antennae on their heads. The next segment of their bodies has a pair of modified legs.
These legs are not used for walking, they have sharp poisonous claws on them that the centipede uses to
capture and paralyze its prey. Each of the remaining segments of their bodies has a pair of jointed walking
legs. In many species, the walking legs end in a sharp claw.
Centipedes are nocturnal and spend the day in moist, dark places like under leaf litter, rotting logs, bark, or
Diplopoda Glomeris rocks.It is a foot in length and lives in South America. Most centipedes are carnivores and eat insects.
(millipedes)
Millipedes have long, round, worm-like segmented bodies. They have round heads with very short
antennae. Millipedes have two pairs of legs on each segment of their body. They are usually 1-2 inches in
length and brown or black in color.
Millipedes don't have poison claws like centipedes. When they are threatened, they usually coil up in a ball.
Some species emit a poisonous or smelly secretion. Most species of millipedes are scavengers or
herbivores and eat decaying vegetation and leaf litter.
Crustacea Branchiopoda Daphnia Species of Crustacea such as the shrimp, prawn, crab, or lobster are familiar. However, there are many
more with less common vernacular names such as the water fleas, beach fleas, sand hoppers, fish lice,
wood lice, sow bugs, pill bugs, barnacles, scuds, slaters, and krill or whale food. The Crustacea are one of
the most difficult animal groups to define because of their great diversity of structure, habit, habitat, and
development. No one character or generalization will apply equally well to all.
A dorsal shield or carapace of variable length arises from the dorsum of the third cephalic somite and
covers the cephalon and cephalothorax to varying extent. The carapace reaches its greatest development in
the malacostracan Decapoda (shrimps, lobsters, and crabs).
The chitinous cuticle covering the crustacean body is its external skeleton (exoskeleton). The chitin is
flexible at the joints, in foliaceous appendages, and throughout the exoskeletons of many small and soft-
bodied species, but it is often thickened and stiff in others. It becomes calcified in many species as a result
of the deposition of lime salts.
The paired appendages are typically biramous and consist of two branches: the endopod and exopod. The
endopod is definitely segmented in the higher Crustacea. The endopods are variously modified to serve a
variety of functions and needs such as sensory perception, respiration, locomotion, prehension and
comminution of food, cleansing, defense, offense, reproduction, and sex recognition and attraction. If
retained in the adult, the exopod may remain leaf- or paddlelike, or become flagellated structures,
facilitating swimming or aiding respiration.
Crustacea take up oxygen by means of gills, the general body surface, or special areas of it. Some sow or
pill bugs have special tracheal developments in their abdominal appendages for the same purpose.
Malacostraca Homarus Malacostraca is a large and diverse taxon (generally class, but sometimes subclass or order) of marine,
freshwater, and terrestrial crustaceans, including many of the most familiar crustaceans, such as crabs,
lobsters, shrimps, which are characterized by a maximum of 19 pairs of appendages, as well as trunk limbs
that are sharply differentiated into a thoracic series and an abdominal series. Other familiar members of the
Malacostraca are the stomatopods (mantis shrimp) and euphausiids (krill), as well as the amphipods, and
the only substantial group of land-based crustaceans, the isopods (woodlice and related species). This
Stenopus group represents two thirds of all crustacean species and contains all the larger forms.
The taxonomic status of the crustaceans has long been debated, with Crustacea variously assigned to the
rank of phylum, subphylum, and superclass level. As a result, the taxonomic status of Malacostraca is not
settled, generally being considered a class within the subphylum or superclass Crustacea, but sometimes
considered as an order or subclass under the class Crustacea.
• The head has 6 segments, with a pair of antennules and a pair of antennae, as well as mouthparts.
• They usually have 8 pairs of thoracic legs, of which the first pair or several pairs are often modified
into feeding appendages called maxillipeds. The first pair of legs behind the maxillipeds is often
modified into pincers.
Oniscus • There are 8 thoracic segments. The cephalothorax is covered by a carapace form via fusion of 3 of
them, letting the 5 other uncovered.
• The abdomen is behind and often used for swimming. There are 6 abdominal segments.
• They have compound stalked or sessile eyes.
• The female genital duct opens at the sixth thoracic segment; the male genital duct opens at the eighth
thoracic segment.
• They have a two-chambered stomach.
• They have a centralized nervous system.
However, this is a very diverse group. Although the term Malacostraca comes from the Greek for "soft
shell," the shell of different species may be large, small, or absent. Likewise, the abdomen may be long or
short, and the eyes may show different forms, being on movable stalks or sessile.
Hexapoda • 3 thoracic Collembola Podura Collembola – springtails
segments • Short legs, plump oval body, few abdominal segments
• 3 pairs of legs • Furcula ʻtailʼ held under tension under the body allows jumping
Metamorphosis
• Anamorphosis
• Simple
metamorphosis =
hemimetabolous
(Nymph, instar)
• True
metamorphosis =
holometabolous
Curculio
Psylliodes
Hymenoptera Urocerus • Includes bees, wasps, ants
• Complete metamorphosis
Greek hymen • Two pairs of wings act as one
ptera – • Hamuli hooks on hind wings
membranous attach to forewings
wing • Some mouthparts modified to
sucking proboscis
• Distinctive pinched abdomen
Agaon • Major modification of ovipositor
to sting
Rhyssa
Apis
Vespa
Macroglossum
Acronicta
Geometra
Eristalis
Drosophila
Holothuroidea Holothuria Echinoderms have tentacle-like structures called tube feet with suction pads situated at
their extremities. These tube feet are hydraulically controlled by a remarkable vascular
system. This system supplies water through canals of small muscular tubes to the tube feet
(= ambulacral feet). As the tube feet press against a moving object, water is withdrawn
from them, resulting in a suction effect. When water returns to the canals, suction is
released. The resulting locomotion is generally very slow.
Echinoderms are exclusively marine. They occur in various habitats from the intertidal
zone down to the bottom of the deep sea trenches and from sand to rubble to coral reefs
and in cold and tropical seas.
Some echinoderms are carnivorous (for example starfish) others are detritus foragers (for
example some sea cucumbers) or planktonic feeders (for example basket stars).
Reproduction is carried out by the release of sperm and eggs into the water. Most species
produce pelagic (= free floating) planktonic larvae which feed on plankton. These larvae
are bilaterally symmetrical, unlike their parents (illustration of a larvae of a sea star below).
When they settle to the bottom they change to the typical echinoderm features.
Chordata Urochordata Ascidia The Urochordata, sometimes known as the Tunicata, are commonly known as "sea
(Tunicata) squirts." The body of an adult tunicate is quite simple, being essentially a sack with two
siphons through which water enters and exits. Water is filtered inside the sack-shaped
body. However, many tunicates have a larva that is free-swimming and exhibits all
chordate characteristics: it has a notochord, a dorsal nerve cord, pharyngeal slits, and a
post-anal tail. This "tadpole larva" will swim for some time; in many tunicates, it
eventually attaches to a hard substrate, it loses its tail and ability to move, and its nervous
system largely disintegrates. Some tunicates are entirely pelagic; known as salps, they
typically have barrel-shaped bodies and may be extremely abundant in the open ocean.
Cephalochordata Branchiostoma With about twenty-five species inhabiting shallow tropical and temperate oceans, the
Cephalochordata are a very small branch of the animal kingdom. Known as lancelets or as
amphioxus (from the Greek for "both [ends] pointed," in reference to their shape),
cephalochordates are small, eel-like, unprepossessing animals that spend much of their
time buried in sand. However, because of their remarkable morphology, they have proved
crucial in understanding the morphology and evolution of chordates in general -- including
vertebrates.
As they do not have bone marrow, red blood cells are produced in the
spleen and special tissue around the gonads. They are also produced in
an organ called Leydig's Organ, which is only found in cartilaginous
fishes, although some lack it. Another unique organ is the epigonal
organ, which probably has a role in the immune system. The subclass
Holocephali, which is a very specialized group, lacks both of these
organs.
In the history of the class, the pectoral and pelvic girdles, which do not
contain any dermal elements, were considered to originally not be
connected. In later forms, each pair of fins became ventrally connected
in the middle when scapulocoracoid and pubioischiadic bars developed.
In rays, the pectoral fins have connected to the head and are very
flexible.
The oral teeth are usually not fused to the jaws, but are replaced
serially. It typically is assumed that their oral teeth evolved from
dermal denticles that migrated into the mouth, but it could be the other
way around as the teleost bony fish, Denticeps clupeoides, has most of
its head covered by dermal teeth (as do probably Atherion elymus,
another bony fish). This is most probably a secondary evolved
characteristic, which means there is not necessarily a connection
between the teeth and the original dermal scales. The old placoderms
did not have teeth at all, but had sharp bony plates in their mouth.
Barbus
Anguilla
Salmo
TETRAPODA
Amphibi Caudata Salamandra To identify an animal as an amphibian, it should have each of these characteristics:
a
• Amphibians have a backbone. They are vertebrates.
• Amphibians are cold-blooded. They cannot regulate their own body temperature.
• Amphibians spend at least part of their lives in water and on land.
• Amphibians do not have scales and their skin is permeable (molecules and gases can pass through).
• Amphibians have gills for at least part of their lives. Some species have gills only as larvae, while
others can have gills throughout their lives.
Anura Rana • Most amphibians go through metamorphosis.
• Amphibians are frogs, toads, salamanders, caecilians and newts.
Amniota Testudine Testudo Turtles (including tortoises and terrapins) are characterized by a shell that completely encloses both of the limb
s girdles. The shell is composed of a dorsal carapace of dermal bone that incorporates endochondral
contributions from the vertebrae and ribs and a ventral plastron of clavicles and interclavicles anteriorly and
abdominal ribs posteriorly. No turtles have teeth on their jaws, and all have the external ear supported by a
large, semicircular quadrate.
Diapsida Squamata Gekko The Squamata, or the scaled reptiles, are the largest recent order of reptiles,
comprising all lizards and snakes.
Two characteristics that unite the squamates. The first is that they shed their
skin periodically. Some squamates, such as snakes, shed their skin in one piece.
Other squamates, such as many lizards, shed their skin in patches. In contrast,
non-squamate reptiles regenerate their scales by other means—for example
crocodiles shed a single scale at a time while turtles do not shed the scales that
cover their carapace and instead add new layers from beneath.
Phyton
The second characteristic shared by squamates is their uniquely jointed skulls
and jaws, which are both strong and flexible. The extraordinary jaw mobility of
squamates enables them to open their mouths very wide and in doing so,
consume large prey. Additionally, the strength of their skull and jaws provides
squamates with a powerful bite grip.
Neoaves Columba
Passer
Apus
Dendrocopos
Mammalia Monotremata Ornithorhynchus Endemic to Australasia - an important point to note as it means that they
represent a whole subclass of extant mammalian life in a single geographic
region. While the platypus is semi-aquatic, the echidnas are all terrestrial, and
their respective distributions and habitats are as follows:
Features:
• Males have a spur on their ankles, which bears poison in the platypus.
• Toothless - platypuses have a leathery electrosensory bill, with
crushing horny plates to break through the tough exoskeleton of
arthropods; echidnas have an elongate horny rostrum with a long
sticky tongue for collecting insects.
• A range of mammalian characters:
o Produce milk (lactate) from mammary glands. However, while
therians have nipples, monotremes do not, and consequently the
young suck milk from patches of mammary hairs - specialised
areas of fur positioned around the ventral openings of the
mother's mammary glands.
o Epipubic bones - two thin rod-like bones
extending anteriorly from the pubic bones of the pelvic girdle.
o Lower jaw (mandible) made up of a single bone, the tooth-
bearing dentary.
o A middle ear formed of three bones: the incus, malleus,
and stapes. While the stapes is present in the middle ear of all
living tetrapods, the incus and malleus are modified bones from
the typical amniote jaw joint. The jaws of non-
mammalian amniotesarticulate via the quadrate of the upper jaw,
and the articular of the lower jaw; in mammals, the quadrate
migrated to form the incus, while the articular became the
malleus, leaving a jaw joint formed of the dentary articulating
with the squamosal (the angular bone of the non-
mammalian amniote lower jaw is used as a bony support for the
eardrum in mammals). The following diagram illustrates these
differences:
Marsupialia Macropus Marsupials are the group of mammals commonly thought of as pouched
mammals (like the wallaby and kangaroo at left). They give live birth, but they
do not have long gestation times like placental mammals. Instead, they give
birth very early and the young animal, essentially a helpless embryo, climbs
from the mother's birth canal to the nipples. There it grabs on with its mouth
and continues to develop, often for weeks or months depending on the species.
The short gestation time is due to having a yolk-type placenta in the mother
marsupial. Placental mammals nourish the developing embryo using the
mother's blood supply, allowing longer gestation times.
Like other mammals, the marsupials are covered with hair. Mothers nurse their
young — a young kangaroo may nurse even when it has grown almost to the
mother's size.
Equus
In all species, digit III is the most
prominent on all feet, and, as the
plane of symmetry of the foot passes
through this digit, perissodactyls are
said to have a mesaxonic foot. The
first digit (equivalent the thumb or
big toe of humans) is lost in all
species. The Equidae have a single
functional toe on each foot (the third
digit), while the Rhinocerotidae have
three toes per foot. The Tapiridae -
the closest family to the ancestral
perissodactyl condition - possess
Rhinoceros four toes on the forefeet (digit V is
used on wet or marshy ground) and
three on the hind. Perissodactyls are
truly unguligrade, with the heel, sole,
and digits of the foot never touching
the ground. The ulna and fibula
(bones in the forearm and lower leg)
are reduced, simplifying the
wrist/ankle joint considerably. The
clavicle (collar bone) is absent,
allowing for efficient running - the
main driving force behind
unguligrade evolution.
Proboscidea Elephas Later proboscideans are
distinguished by tusks and long,
muscular trunks; these features are
less developed or absent in early
proboscideans.
Respiratory General Characteristics
Subphylum Class Examples Symmetry Tissues Coelome Circulatory System Nervous System Digestive System
System and Key Words
Have all of the
developmental
characteristics of other
chordates, but lack
Tunicates, vertebrae. Have been
Urochordata and
lancelets Triploblasts, Alimentary canal, extensively studied to exam
Cephalochordata n/a Bilateral Coelomate heart Primitive ?
(amphioxus), sea eumatazoa one way the origin of vertebrates.
(amphioxus)
squirts Tunicates live in benthic
habitats. Lancelets keep
notochord through
adulthood, tunicates only
have it as larvae.
Notochord found in larvae
Triploblasts,
Vertebrata Fish (Jawless) agnatha, lamprey, Bilateral Coelomate Two chambered heart Complete, brain Gills Alimentary canal and adult, cartilaginous
eumatazoa
hagfish skeleton
Jaws and teeth, reduced
notochord with cartilaginous
Fish Triploblasts, Alimentary canal, vertebrae (from here on:
Vertebrata Shark Bilateral Coelomate Two chambered heart Complete, brain Gills
(Cartilaginous) eumatazoa one way notochord is present only in
embryonic stage, replaced by
vertebrae in adulthood)
Cnidaria Hydra, Radial Diploblasts, n/a None Nerve net, no None Gastrovascular None - Aquatic habitats, some have stinging cells
jellyfish, true tissues (diffusion) brain (diffusion) cavity, two (diffusion) (nemotocysts), some have life cycle that switches
sea (eumetazoa) way digestion between polyp and medusa forms, sexual or asexual
anenome, reproduction, gastrovascular cavity acts as
coral hydrostatic skeleton to aid in movement
Platyhelminths Flat worms, Bilateral w/ Triploblasts, Acoelomate None Two nerve None Gastrovascular Protonephridia - Can reproduce sexually (hermaphrodites) or
trematoda, cephalization eumatazoa (diffusion) cords, anterior (diffusion) cavity, two and flame asexually (regeneration), mainly aquatic habitats,
flukes, centralized way digestion cells parasitic lifestyles, most primitive of triploblastic
tapeworm, ganglia (brain), animals, has organs. Tapeworms specifically don't
planarian some have a true digestive tract, they just absorb food
planarians around them.
have eye spots
Nematoda Round Bilateral Triploblasts, Pseudocoloemate None Nerve chord None Alimentary None - Some have cuticle to prevent degradation by host
worm, hook eumatazoa (diffusion) and ring (diffusion) canal, one way (diffusion) digestive system, longitudinal muscles, no circular
worm, muscles, parasitic, not segmented
trichina, C.
elegans,
ascarcis
Rotifera Rotifers Bilateral Triploblasts, Pseudocoloemate None Cerebral None Alimentary Protonephridia - Not trule segmented, can reproduce sexually or
eumatazoa (diffusion) ganglia (brain) (diffusion) canal, mouth and flame parthenogenetically, mostly freshwater
w/ some and anus cells environments. Draw food and water into mouth by
nerves beating cilia.
extening
through body
Annelida Earthworm, Bilateral Triploblasts, Coelomate Closed Vental nerve None Alimentary Most have Protostome Segmented bodies, coelom is divided by septa,
leech eumatazoa circulatory chord, anterior (diffusion) canal, mouth metanephridia sexual (hermaphrodites) and asexual (regeneration)
system, ganglia (brain) and anus reproduction, longitudinal and circular muscles
multiple
pairs of
aortic
arches,
distinct
arteries and
veins
Molluska Clam, snail, Bilateral Triploblasts, Coelomate Mainly Ventral nerve Gills Complete, Nephridia Protostome Visceral mass, calcium carbonate mantle, radula
slug, squid, eumatazoa open chords and mouth and (tongue), aquatic or terrestial habitats, have
octopus, brain anus, radula hemocoel
cephalopod,
gastropod
Arthropoda Ant, Bilateral Triploblasts, Coelomate Open Fused ganglia, Spiracles One-way Malpighian Protostome Exoskeleton, jointed appendages, coelomates,three
(Insecta) dragonfly eumatazoa circulatory ventral nerve and tracheal digestion, tubules pairs of legs, chitonous exoskeleton, more species
system, chord tubes some have than any other phylum combined, metamorphosis
hemolymph salivary
glands
Arthropoda Spider, Bilateral Triploblasts, Coelomate Open Fused ganglia, Trachea or One-way Malpighian Protostome Exoskeleton, jointed appendages, coelomates, four
(Arachnida) scorpion eumatazoa circulatory ventral nerve book lungs digestion, tubules and/or pairs of legs, land habitats
system, chord some have coxal glands
hemolymph salivary
glands
Arthropoda Lobster, Bilateral Triploblasts, Coelomate Open Fused ganglia, Some have One-way Terrestrial:use Protostome Exoskeleton, jointed appendages, coelomates,
(Crustacea) crayfish, eumatazoa circulatory ventral nerve gills digestion, malpighian; aquatic habitats
crab system, chord some have Aquatic use
hemolymph salivary green glands
glands
Echinodermata Starfish, Bilateral Triploblasts, Coelomate Open, no Nerve ring and None Complete, None Deuterostome Spiny, central disk, water vascular system, tube feet,
sea urchin, (larvae), eumatazoa heart radial nerves (diffusion) mouth and (diffusion) sexual or asexual reproduction, closest related major
sea fivefold anus phyla to chordates
cucumber radial (adult)
Chordata Vertebrates Bilateral Triploblasts, Coelomate See the table above Deuterostome See 'Phylum Chordata' Sheet
eumatazoa
LICHENS
Parmelia Cladonia