‘Appendix
Crop Growth Requirements
Douglas A. Hopper, Gary W. Stutte, Ann McCormack, Daniel J Barta,
Royal D. eins Jol E. Eran, and Theodore W.
ibbitts
‘The following table of growth requirements
isforplant species commonly grown in enclosed
environments and provides suggested condi-
tions to simulate average growth in field envi-
ronments and commercial facilities. These are
general suggestions that will provide good
growth; it should be recognized, however, that
conditions may differ for specific cultivars and
‘may differ in particular commercial production
facilities to fulfil specific quality or other re-
quirements, Detailed information on nutrient
solutions, lighting system lamp types, growing
media, ete, is contained in the previous chap-
ters of this handbook
ENVIRONMENTAL CONDI-
TIONS
MrpiA aNd Nurernion
Plants usually willbe grown ina soilless pot-
ting medium or liquid culture. Several commer-
cially available mixtures provide suitable media
for plant growth, as discussed in the previous
chapters. When plants are grown in a soilless
‘media or liquid culture, use of a complete nutri-
ent solution (approximating half strength
Hoagland) is recommended, with the nitrogen
concentrations modified as indicated in the table.
Nitrogen is emphasized because this isthe prin-
cipal nutrient that i altered for a particular spe-
cies o at particular stages of growth. If a soil-
contai
1g medium is used, itis recommended
that the plants be fertilized with a commercial
‘water-soluble fertilizer such as 20-10-20. When
nitrogen concentrations are modified with a
‘commercial fertilizer, P and K concentrations
will also be modified. Recommendations for pH
assume that a range of 5.5 to 6.5 is acceptable
for most plants, and a pH of 4.5 to 55 is accept-
able for crops requiring low pH.
Lieut
The recommended photosynthetic photon
flux (PPF) is for the top of the canopy. When light
is indicated to be 12-20 hours, there is no spe-
«cific daylength requirement for this species. The
use of the longer period will provide a greater
amount of total lighting, and thus an intensity
toward the lower end of the recommended PPE
zange can be utilized to achieve good growth. If
a short photoperiod is recommended, it is nec-
essary to provide an intensity toward the high
side of the recommended PPF range. No recom-
mendations are provided on light source because
chamber configuration, plant species, and de-
sired response can all affect lamp selection, Us-
ers are referred to specific chapters in the text
for detailed lighting design considerations in
controlled environments. Special light require-
‘ments for specific species are noted in the com-
‘ments column of the table.
TeMpeRATURE
Recommendations are for air temperature at
canopy height. The temperature for most plants
can be set2'C above orbelow the indicated tem-
peratures and still produce normal plants, al-
though growth rates will be altered. A light/dark
fluctuation is recommended, and is required
‘when shown in bold letters; most plants, how-
ever, will develop normally with a constant tem-
perature that isan average of the indicated light
and dark temperatures. The temperature period
required to break bud dormancy of woody spe-
2728
cies is cultivar dependent; thus, a range of time
is indicated,
Revarive Humiprry
Control between 60 and 70% relative humid-
ity is recommended for most crops and can be
achieved readily in most controlled environ-
ment chambers. Exceptions are noted in the
‘comments column.
Staces of Crop DEvELoP-
MENT
‘The environmental conditions required for
successful plant growth in controlled environ
ments are often not the same throughout all
stages of plant growth, For that reason, the table
has been divided into four, somewhat arbitrary,
stages of development: Propagation, Vegetative,
Flower Initiation/Development, andl Fruit/Seed
Development. These categories may not be tech-
nically correct forall the plants described, but they
should provide functional guidelines for maintaine
ing a desired growth rate fr the listed species.
Propagation is considered the period from
planting to emergence of the frst true leaf. Ea-
vironmental conditions for acclimation of tissue-
culture transplants, seedlings, and rooting of ext-
tings are also included in this section when they
are considered tobe routine propagation methods.
Vegetative is the period when reproductive
structures are not actively developing on the
plant. This typically incluces the period of rapid
stem elongation and leaf expansion that pre-
cedes flower formation. For woody species, this
period may include the period of stem and leaf
growth that occurs when flower buds are present
but dormant.
lower iitiationédewelopmentis the period from
the initiation of flowers through shedding of
pollen (anthesis), flower formation, pollination,
and fertilization. However, for many perennial
woody plants, initiation of flower buds occurs
ata time separate from flower development. The
recommended environment for breaking dor-
mancy of the initiated flower buds of these
‘woody species is indicated in the comments col-
‘uma at the right margin.
Fruitjseed development isthe period following
fertilization to maturity of the seed.Propagation Vegetative Foner iiaionDev. [Fused Do.
conments
aad te | ae rete | Temp Pros | Temp. Prato: | Temp prev: | Tero.
Connon tame vase | owt | coy | une | pees | coh | vane | pated | coy | uae | peed | ec)
(Cours an) fio | ub srt | vine stom | um toby | vie
stan att
Sats om wu} ov foe faves] u | we fav] o | we | coves Lett os
in | {eye
oan
Aavatmroutran [au |u| m | tem | aso) w | ao w | sao | asizo
ft
tte | Lat eg tp
de sia uw fu] om | sear | asrao | m | sean | acize| m | oe | ass] | ow | esas | Bvenseretictey,
Astensta Pesien Uy ‘etn of ie.
Aatoonert a fn] om | oe fos] uw foe we | oe | anne Focounn bea
i eyes ferpmuibe e166
sent Bis
Foamnat cfu fm | see favs | mw | 22 | wie] uw | a0 | orm) w | 2a
Ave
Mcsdoestcasian — |e | n 4 | sem | asiao] a | seen | esizo] | 200 | 25120 | emndeomansy smote msatec
sree
‘nope ne mtn fe | a fare] cf 2 fara} uf] se favre] 2 | ste | soz | amine gomintn,
Carn
cae
Forceareiceaia. — | w | n u | seam | asco] uw | seem | anise] uw | 220 | 25/0 | nawanessinnces toning
on Sm afro, 20 5p84.
Pea p mw fof oe | oe fess] ou | ote fossa} uw | so | asses Schram oer dese ater
ion
sary
eco wie mw fn fou | re fer] ow | se fas] aw | sear | ase] we | roa | zor
unto vg Phan
ee oa NS tama 5-55 Nol
i > vest (ogemm DS tag 4588 0 souls!
HS tag tunes
Bt Auial ns
{So oymoln
61zPropagation Teoeatve Ter ltaiooDev| Freed be
comers
PLANT selon roe | Tone Prat | Tenp. rates | Tong. rave | Tom
Gemmmnare J malate age | oad | cy | uate | pend | co) | uae | awed | cy | tase | aes | co)
es sey | love Aro | uve sow | ufooe| | voy | uae
con Londoner
Psat wat udu] a | seen | xize | om | sean | ezre| | r2ao | corm] ur | seco | asin | pean ants
Bary Hath fr man 0 2:00
Vesumenmoom. | 1 w]e fara] nw | ee fears] oe | seen | cov | teetimantoarnas:c eb
fob pet nares
Baer tre bial dma b 0Ha7C
Vanruhcmecn [cfu] wm | rear | ase | mn | te fasran] | seco fiasco] | an | esr | Rrtotman cm ps
pero dann ta
ante
eae uw wf ea | esrao] m | ee | esrao] w | 2 | ase | eeaeuscmn, 0 wateoneat4c.
cats
em
comet
cas Teksirg Conny rd Cnn
iemertees udu lw fos [aa] w | ow fae] mw | ae | oor Sion
Soa
aed Poutns Para ‘hore ante sug: ten
Cima temas | utw| of 2 lao] w | ow fans] wu | a fans Soest
vs Se [an
conan
Giomeneyant. [fw] ef oe [as] w | ae fas | ow | ome | wre «eis bs
ony
Pome. uf we | ea fase] nm | rear | reo | s2e0 | azo | sean demany 70 cnn wsat6c
Cingsnterun
GRismamepanstom |x fx} o | w faves] mw] w fare] a | wo | are Snags wn 250 pm8A
fanaa
anon "
MS ma ma OD aoe
i 2 fc pont tw es ‘ 2) aint
D fenatopeet t 10 onan
ti 2a nyt!
fl ‘So mountsPropagation Vegetative Flower hitation’Dev FrlSeod Dev.
comments
Prato | Ten, Prao- | Teno Tonp,
ug | pmred | 6) paicd | 6) (0)
sry | Uva Asia | uC idan
Ingostartsis stage, donot alow
w | 2 | aie we | ma oot
[Masson ox UHer) DC.
| sae | oaias 220 | 28/28 20/28 | waar evssinsces fowahg
enum sumer mw | we | asia 620 | 25/20 25/20
w]e | enias 220 | 3/28 20128
Coanes plana Cau w | we | enras 1 | asi
Most catars wt ewer unter ary
[Gosspum hrsutt. | i220 | asia 220 | 23/28 20/25 | phtopar Som cutrars require
Shot ye forfonar natn,
wu | 220 | asia 16 | 25128 2518
igh trp. ibis gemini, post
o | o |e 2 | aia anes sage; ont alow wt
ibe coved at 5 Cor nes
12 | zoi20 noices tonsa,
|
T
Lf ae jas <2 | io Sem tings wn 2500 ppm BA
Noma 65-65 D Nig
lw 45-55 v 5D 180pmol’s!
ie 160 2e0qimalme*
MS Medium 280 -4S0ymalmr's?
HE 450 -ToOjmatmes
WzPropagation Vegetative Flower Iitation/Dev FrultSeed Dev.
Comments
aoe HPs Proto- | Tone. rac- | Toro. Prot | Temp, Proto | Tomp.
Cammenname | PMATTOH une | porta | rc) | unt | ped | 0) | ut | peice | co} | uate | pated | 9)
(corso isi0y | uae is/0 | oem testy | w/oa ero | w/a
srr
Patugonin sp. w dw] m | i220 | 20] m | reo | asrz0] uw | twa | 25/20
Getor (Tana! Dain)
Serer janesont wfu] oc | oe fase] om | oe Josie] om | oe | asi ode shout aya
Baus Hook [|
ben
‘Srnngi spose wd] oc | se | ano] aw fo aio] m | se | asi20 ‘Seeds vccvced np of mode,
(Led Ham
Grae
Vins. wd] a | 220 | aso | oH | tee | asizo] ow | tee | 2520 1220 | 25720 | oreakbuddomany 7501 S000ms aC
Horbane
Myaeyamas ger c fut om fa fsa] m | 6 fases] mu | ws | zsve0
tanga Sem cuts wih 2500 pm IBA. Ba pes.
Pytargee mops = ow Jum om | ste | zsias | om | ou facies fom | eta | assis Pic ps6 BC or hs romired fr fower
(hunt) Se | ovetpen’aferntaton
Ketancon
Katretoobossisora = | HTN] a | ote [anes | wm | om | asi} mw | feo Sewd 57 om evtings
Pan.
Larbscueers
(Chenepedam abun. wef] om | oom | esian | om | om fasizo] mw | ate | 25ve0 <2 | esra0
Loe To bam atigh gt rdhigh AL
Lace sabe. ww] um | r2e0 | 2520] om | tzeo | asvzo] uw | za | 25/20 220 | 26/20 | s7 maim? day’ of PP exposes
Lani ayoaber Sear dis f coms.
Use sn. PC ee 2Cler Sw eles ong pad
Foren
‘nation "gt Photehete Proton Fu PPE)
Les Loa Spe) Tormal 55°65 D= Ow NOM
Meum 1009p low 45-55 VS Voqytov 50 1S0ymaims
Ho Hon topen Le low 160 = 250nolm's*
MS Moun 280 - 450 maim
He High S60. 700qmoims*Propagation Vegetative Flower ilatowDev. | __FriSeed Dw.
- comment
ser waa Pro | Tarp roe: | Tene. Pras | tore. pra | erp
Comannare [a] are | peed | oc) | uit | pent | cc | unit | pars |) | uae | pers | 0)
esse A su | urbe sim | uioee sity | vom sry |
aig
Tages eta mfx} wm | reeo | asieo | om | sez0 Jasco] om | 20 | asian
oats
Avra saat wf] we | rez J esieo | ow | szzo | asron] aw | sae | asin] om | re20
oe Foyt ton 72019250
Oe axczaal un w]e fase] x | ren | are] ow | ream | eave | areca ey ery
Distal tents bce
wf} me | roan | asres | ou | ean | asias} wm ] teen | anris] om | r2eo | zur | manmumionpemnrs e226
Panspesica.) user | ot | ny | sean J asizo] a | rezo | asieo] | sea | 25/20 | seactudsomany 20 ators asc
eon | Btw dre pos.
Ach page mf} o | wa pases | a | ean fasias | om | rezo | soas] ow | teen | soras | Storey cease vet nr
ew
Pe carmunt. wn H | ream | asian] w | t2z0 | asio] | r220 | 25/2 | searrossomans: rot asoomsaec
Beppe
Copsemanramey — | wn] mw | saan | asrao | w | s2ao | oso] uw | tae | asia] m | roan | 2sr20
vera |
ean
Pernraescomtioe | c|n| m | a fess) m | ve farm] w | s | zoo] mw | | core
Peta
Panna xiynseiim — | w |v] om | sea | esi} om | t2eo | asi] mw | teen | asiz0
Tow Bia) Tema 5-83 Oe Oat Nolg
Mata 0 pp) ton 45555 V2 Veytoe "sa 150 yt!
Hap open) UE tn 0. 2opmdms!
ME Men 20 450 pts!
He ge” et Tomaswe
Vegetative Flower itation/Dev. FrulvSeed Dev.
Comments
ean) le Poe | Temp. ene Pra Prate- | Temp
conmonvizre | ma" HT ue | perc | coy ce), | batt | pr une | pros | cc)
(Gensspecns A) 5/0 | uo we iso | isrow | ida
Pri
Pra IL) Rts cfu] e | jess] om |e fasres} mw | 8 | soi
Pawoed
Amat 59. wf] om | oe | eso] om | oi | aso] om fs | asi20
Prine
Exghotiaputronne ww] oe | se | aso] om J ot fasrof om | to | asi ‘Som catinge win 2500 pp BA.
‘Wi eet
Pott, Sweet quran ar fo storage roa fomaton
omeataastjtan fou fu] om | seen | asia} oe | ct | asia] om | ore | asias] | oe | 2sv25 | Hiner lvl tne vegetative on equtos
ah
Patt, Whe gure asfor barat. Long days
Solu uberosum wf} free | ese] om | teen | esse] om | rea | esse] mM | 2a | 25/8 | wimiow PPF das ubetcaten pies
ree ‘She ay opal deg or
Oryza sate. wf] om | rez | s0720 | om | tza0 | sora] ou | teao | soya] oe | 12 | 20/20 | oneting aes wih cut
ose
Fost mut fora Thar, ww] oc | we | ees} om | we fare] om | ve | eve Sem atngs wit 2500 om A,
Preiss
(lum muon an wf} om | rea | ave] om | tea | ans] om | ae | asi
save |
Satasooraunsstower | cfu] om | 2 | ass} ow | v2 fas] mw | a | ase
Seutes -
Seophulaa
Smpfuiremaindeal. | cw} c | a jas} ow |e jam] wu | | anv
wan —__ YH
Lo La apan N= Noma 55-65 ole
= Mesum HOO ppm) Cl lw 45-55 50 150 mols!
HS High 2pm) 460 20uma ms!
250 somal’
460 70umal ms!Propagation Vegetative FloneriniationfSev. | _FruliSed ev.
Comments
on rete | Tan. Pro Prot. | Tone, Pro. | orp.
cco ial PY ye | pect | ec) | uate | pia petod | 6) | Lane | pared | 0)
(cous pecs a) eit | usta fy toy | uae stm | Doe
Shere
Sie area. uf] oo wa fess} u fe Jao] m | w | aie
Shapis
‘Sapa cfu] mw | rer | ares | mw | see feof uw | 2 | ave
Snptgin ‘ew sod ono of okt ma,
‘Annan mais we] ow foe fava} ow | oe Josie] wf sie fare Fate og yr toveog.
Soytean ‘Ser a pete dang Roeirg
(here man a mfx] | fares} mw | tezo |asran] ou | ea fae} ow | 2 | ares | vaeswnonar
Spach | old empratues
Space oboe. mtn fou foe Jaro fom |e famof w | ots fares} om | ots | s5r28 | cxcouage carer tower,
Strbary For you eta ey eppra omns
Fragaaonancssa wf] om | aa free | om | ea fas] ow | ae fas] mw | roan | zor | maemsatacunsimaai net,
Dic
Toba
Mota abcun wef] om | saa | asia | om | sea | asi] | 2a fai] om | eae | asiz0
Toro Fagus bk anda
Uropescinoscoimam | a fn] a | r220 | asia | oH | sze0 | asi] | tea | asiao} ow | roan | 25/20 | regnrurtennayinc fut et
West We beatae cod tatmat
Tcwmaostvunt. wfw] me freer | asia} om | ram | aos] mw | ee | ase] Hf rea | 2sr | omsesan tr tal née Lo
otras in eweng.
*tudten 4
Lo Law SOpp0 Toma 55-55 De Oak Noli
Mesum 1009-18) low 45-55 Vs Veylow "0" tSpnolms!
ig (20epni Le aw 150 2stqolms!
M= Ncsum 250 = Oqiet ms!
a ee
sw
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