Microbiological activity clearly affects the mechanical
strength of leaves. Although it cannot be denied that with
most species the loss of mechanical strength is the result of
both invertebrate feeding and microbiological breakdown,
the example of Fagus sylvatica illustrates loss without any
sign of invertebrate attack being evident. Fagus shows little
sign of invertebrate attack even after being exposed for eight
months in either a lake or stream environment, but results of
the rolling fragmentation experiment show that loss of
mechanical strength, even in this apparently resistant
species, is considerable.
Most species appear to exhibit a higher rate of degradation
in the stream environment than in the lake. This is perhaps
most clearly shown in the case of Alnus. Examination of the
type of destruction suggests that the cause for the greater
loss of material in the stream-processed leaves is a
combination of both biological and mechanical degradation.
The leaves exhibit an angular fragmentation, which is
characteristic of mechanical damage, rather than the
rounded holes typical of the attack by large particle feeders
or the skeletal vein pattern produced by microbial
degradation and small particle feeders. As the leaves
become less strong, the fluid forces acting on the stream
nylon cages cause successively greater fragmentation.
Mechanical fragmentation, like biological breakdown, is to
some extent influenced by leaf structure and form. In some
leaves with a strong midrib, the lamina breaks up, but the
pieces remain attached by means of the midrib. One type of
leaf may break cleanly, whereas another tears off and iseasily destroyed after the tissues are weakened by microbial
attack. In most species, the mechanical breakdown will take
the form of gradual attrition at the margins. If the energy of
the environment is sufficiently high, brittle species may be
broken across the midrib, something that rarely happens with
more pliable leaves. The result of attrition is that where the
areas of the whole leaves follow a normal distribution, a
bimodal distribution is produced, one peak composed mainly
of the fragmented pieces, the other of the larger remains. To
test the theory that a thin leaf has only half the chance of a
thick one for entering the fossil record, all other things being
equal, Ferguson (1971) cut discs of fresh leaves from 11
species of leaves, each with a different thickness, and
rotated them with sand and water in a revolving drum. Each
run lasted 100 hours and was repeated three times, but even
after this treatment, all species showed little sign of wear. It
therefore seems unlikely that leaf thickness alone, without
substantial microbial preconditioning, contributes much to
the probability that a leaf will enter a depositional
environment in a recognizable form. The results of
experiments with whole fresh leaves show that they are
more resistant to fragmentation than leaves exposed to
microbiological attack. Unless the leaf is exceptionally large
or small, leaf size and thickness are not likely to be as critical
in determining the preservation potential of a leaf type as the
rate of microbiological degradation.
1. The passage is primarily concerned with ...A. Why leaves disintegrate
B. An analysis of leaf structure and composition
C. Comparing lakes and streams
D. The purpose of particle feeders
E. How leaves’ mechanical strength is affected by
microbiological activity
2. The conclusion that the author reached from Ferguson's
revolving drum experiment was that:
A. Leaf thickness is only a contributing factor to leaf
fragmentation.
B. Leaves submerged in water degrade more rapidly than
leaves deposited in mud or silt.
C. Leaves with a strong midrib deteriorate less than leaves
without such a midrib.
D.Microbial attack is made worse by high temperatures.
E. Bimodal distribution reduces leaf attrition.
3. The tone of the passage is :
A. Persuasive
B. Biased
C. Objective
D. Argumentative
E. Disparaging
4. Which would be an example of “energy of the
environment” mentioned in the passage?A. Wind and rain
B. Sunlight
C. Animals that eat leaves
D. Lumberjacks
E. Fuel that may be harvested
Questions 5-7 are from the following passage:
Multinational corporations frequently encounter
impediments in their attempts to explain to politicians,
human rights groups, and (perhaps most importantly) their
consumer base why they do business with, and even seek
closer business ties to, countries whose human rights
records are considered heinous by United States standards.
The CEOs propound that in the business trenches, the issue
of human rights must effectively be detached from the wider
spectrum of free trade. Discussion of the uneasy alliance
between trade and human rights has trickled down from the
boardrooms of large multinational corporations to the
consumer on the street who, given the wide variety of
products available to him, is eager to show support for
human rights by boycotting the products of a company he
feels does not do enough to help its overseas workers.
International human rights organizations also are pressuring
the multinationals to push for more humane working
conditions in other countries and to, in effect, develop a code
of business conduct that must be adhered to if the American
company is to continue working with the overseas partner.The president, in drawing up a plan for what he calls the
“economic architecture of our times,” wants economists,
business leaders, and human rights groups to work together
to develop a set of principles that the foreign partners of
United States corporations will voluntarily embrace. Human
rights activists, incensed at the nebulous plans for
implementing such rules, charge that their agenda is being
givenlow priority by the State Department. The president
vociferously denies their charges, arguing that each situation
is approached on its merits without prejudice, and hopes that
all the groups can work together to develop principles based
on empirical research rather than political fiat, emphasizing
that the businesses with experience in the field must initiate
the process of developing such guidelines. Business leaders,
while paying lip service to the concept of these principles,
fight stealthily against their formal endorsement because
they fear such “voluntary” concepts may someday be given
the force of law. Few business leaders have forgotten the
Sullivan Principles, in which a set of voluntary rules regarding
business conduct with South Africa (giving benefits to
workers and banning apartheid in the companies that
worked with U.S. partners) became legislation.
5. Which of the following best states the central idea of the
passage?
A. Politicians are quixotic in their assessment of the
priorities of the State Department.
B. Multinational corporations have little if any influence on
the domestic policies of their overseas partners.C. Voluntary principles that are turned into law are
unconstitutional.
D. Disagreement exists between the desires of human rights
activists to improve the working conditions of overseas
workers and the pragmatic approach taken by the
corporations.
E. It is inappropriate to expect foreign corporations to adhere
to American standards.
6. According to the passage, the president wants the
voluntary principles to be initiated by businesses rather than
by politicians or human rights activists because...
A. Businesses have empirical experience in the field and thus
know what the conditions are and how they may/should be
remedied.
B. Businesses make profits from the labor of the workers
and thus have a moral obligation to improve their employees’
working conditions.
C. Workers will not accept principles drawn up by politicians
whom they distrust but may agree to principles created by
the corporations that pay them.
D. Foreign nations are distrustful of U.S. political intervention
and are more likely to accept suggestions from multinational
corporations.
E. Political activist groups have concerns that are too
dramatically different from those of the corporations for the
groups to be able to work together.7. Which of the following statements about the Sullivan
Principles can best be inferred from the passage?
A. They had a detrimental effect on the profits of those
corporations doing business with South Africa.
B. They represented an improper alliance between political
and business groups.
C. They placed the needs of the foreign workers over those
of the domestic workers whose jobs would therefore be in
jeopardy.
D. They will be used as a model to create future voluntary
business guidelines.
E. They will have a chilling effect on future adoption of
voluntary guidelines.