gig ol pl (2007) 86-76 :(2)38 — 48 el 3h pad Hae GALA NAS gal gall cna Gan B pbleally 49) ci SN ally Cull Cl gS ysis giant) Aaa seg i gas pal Prd AS Satan gd hd 4 . Ss day Fel PAS Eyal yf ak oe catia A SagDIG) LST ARO GAGA yell Se yea 53) oe Mae ITE y SG ae bY SSG! ce Das Ea Lt a lal Gage oS IS scala Ja GV! Rane yf 2000- 1993 gle ge Gaal aglye UIT GM aed ll ie pay UH GF yal HS Qo ay atl Aha ae juny tay 5D FAL INy ABA Gay AbD Se O5sly Dill se ON gaily ABE By cess Ug eli Opel ky Raul Reg SL i gid pas ge oa aD Dipl hey dead) SA da hs Lata 2001 SAS jal land! ga 8 Gace (General Linear Model) oS ha) 3 p91 ie BES Ga) eSyail Uil fy AB 2 Get CIs Sa sth MIVQUE ih cis 18 6 Chal (asl Gan BN sy Co aga Chill lg U8 ( BLUP ) janie yb has 555 Jad) ysid (1991) Harvey. ea Sid Lal ( Mixed model ) Hep Gijph SAH Uandy oS 53.69 Sal Hayy AAS BB SE AD ae jelly 6 AS 34.28 Ql Dhl se Gal a Baga Ob Be sins as oS YD hla oA «HOI Dall pans Ysine Seg a ba a hl se pall Lye 123.46 49 ob 438.64 0.03 5 0.06 50.1) cal se ay Rag S250 By HE5A SLIM Abi ey Hl Se GIA AN AHL CA HY ste as GS sis he 0.23 006 5 O21 50:16 5 0.16 tes ial stad i Sal 3g e ASA Ci pa Cay Ls OL 5 0.05 5 GS AF 10.83 591 45.69 5 5.175 (58593 4 iS LOT GeV Le Gi Sik DLS AB gl Fae Lee GS Qaly soe ws BAN yl Cy a le aly Seg A jg GM Janey a SSM oT we ally Thal we 5,0 ge Bia pil le Late gh hg ABN Gaal hy yy «(hd 7.65 ~ 5 pt 28.62~ 5 5S-4.50~ 5 AAS4.10 — pS 24 Y 3A DOL 5-58 0.98 ~5 pS 0.013 ~ 5 53.83 — Gal Lae gy SAO y phe 2461 y p55 1.02 y AS 00012 5 a5 3.35 036 Ba ual LS Gal AY wel 28a ALM A ABLE J oh gay Soe pes GP CLD 8 Gs le A 640 — The Iraqi Journal of Agricultural Sciences Al-Samarai & Al-Anbari ESTIMATES OF VARIANCE COMPONENTS TO MATERNAL AND. DIRECT EFFECTS FOR SOME GROWTH TRAITS OF HOLSTEIN CATTLE AbSamarai F.R* Al-Anbari N.N.** * College of Veterinary Medicine - University of Baghdad =* Department of Animal Resources ~ of Agriculture College ~ University of Baghdad Abstract This experiment included 3412 records.of bieth weight and 1378 records of weaning weight , gain , daily gain and age at weaning related to 27 sire ,through the period from 1993 to 2000, at the Eshagi dairy cattle station . The purpose was to study genetic evaluation of sires and dams according to their calves birth weight, weaning weight, gain daily gain , weaning age before adjusting to some fixed effects and to estimate heritability . The General Linear Model by using SAS (2001) program was used to study the effect of fixed factors ( season and year of birth , parity ‘and sex ) on the studied traits . Components of variance for the random effects in the employed mixed modet were estimated by the Minimum Variance Quadratic Unbjased Estimation .The Harvey program (1991) was also used to estimate BLUP values for 27 sires and 1164 dams. The effect of all fixed factors were significaat , except parity . The overall means of birth weight , weaning weight gain , daily gain and age at weaning were 34.28 kg , 88.58 kg , 53.69 kg , 438.64 g and 123.46 d , respectively. The heritability of maternal effects of che studied traits were 0.06 , 0.06 , 0.03 + 0.05 and 0.01 , respectively and for direct effects were 0.16 , 0.16, 6.21 . 0.06 and 0.23 , respectively. There was a ‘wide range in BLUP values estimated which reflected the importance of genetic additive variance. This could be used latter via selection methods to improve the mentioned characteristics of Holstein eattle in Iraq, 2007/3/26 Cana Sl G38 2006/1 1/8 Saal 220 gs, 6* 76gly lata Bey ay Wy SEE 8 Liye ly Gu 8 hil ae adeP Uy Gah LS Ul at yyy LMI Les oll Ws Gey a (8) Bourdon BB sll Qed gyal ala) cli Saad 98 aa LI cory les Ga ally a (im) at ot Is (bra) demas ci gt aed hey LOU Bg Jan af pai Ge had abil se bay Dual we yo gN gy gal Clee Se gals Kyaglh 4530 FIM Jonas HE! HIT, china a gua fell yal yi Sy tid Sel Babs At Jie yy chy SUB sd Ode Dae 3412 Shand! dial Spats chal 27 gh oy AY! Glial) Wp 1378) Dud we ete MGR BET Ge Spee LS 1164 y By ED Shed GU Ay Gad hed GBD py polly gent allay Cag le Lets PAS3S=3 LS pS 1 Sane cash ADU 3S pe Bp D eye Oe net UA dial Sil Use la JN fs 6 ta id GAS yb Gana 91S WE Ss JVI Ge Sod pe ahd oY one a peed CH De oy Sey iy Sl pds Gad oy J Tels haat ge 57 Vet AAT oe LN Gals SN pal Gd a yl pL) Baa LS SD HIS Jaieny Yay Ie SDE! pds d pest cds ce % 10 Iain pas Vs Salyd pun Saad ua al HN Gala asia J ‘thal oss 4B yd Led O35) Nd Id BS 7 (2007) <86-76 :(2)38 — 481 al He1j30 sled Up Aaa ete SEAS Hal yal) che as Sly YDS ay Dy cok BA a ellen cle Gy) Gf 1M 6 gabeaciv PSP A ol ath GLY! ly 0 Legg pun ob (Mo. Fler ssi act ta Hime pS yall thal Al Gaal ll del ey AE oe Li yyy gd eles AMF 5G ge hak Uelens ih Ssal 8G Cae gail ee AD CUS galt LL GT Ge sal 3) oneal ag chinad 44d ED y (imag) 5 pM ld Gy He PW 20s 10) AAV, pall BU Ga gad OF sy ala JB gb eS plc Gas ole a gh bad Wal ye ge FY co iS Chel Direct te Sis sSe end CHES lina gual wee Audi ogi Sle yb Gas al (component ) ) A che il Cal ola Ge Lamy J YI 6 43) 2 oe 5s lg asaitry ( Maternal component eM ld OI as A DS, Ge WB GH gle ABD ples al os: Toke GAN td op gaa ee Uae ML MD. (8) Hat sek tS AYE Sy 6 aly lity Legh SI ila DE Ge Sal pa OI Sale say A SE Gane lel tb Povim paas aa ties Legd De 2 5a BM AF SSD Hast (19) or.8s fash op Shad WN day sl Anal pti Gs Dall Aad dl Jilly Gancle 3 ( Excepted progeny differences Aad Sty LAS, ( Transmitting ability ) & yd Stal Ge BY Id ow On LID ey ) Mada all CADIS ust gt eA MLE ew aah SH gh Shay ah eA! toa ( Dystocia ) Eler hey Aad se gis 32) (8)( Survivability) 3) Ae 3B pt SSE, po gb FS Aeye sae ages od (II) ostghey ol pad (2007) 86-76 :(2)38 — 4a Bees ak nas A IED Lay) 8 (23) ay Lge gh ae lll OLD i Jess Gas Ua Age Gy le 6 algal Gans HYD daly 40 +S 80-75 A gal ) GEM ais Steatuly Shaan! sail! jal , SAS Jalst gain Gas ( General Linear Model Yijklm = p+ Si+Rp+ Ak-+XD+ efjklm Kk) k s¥ yb dubs cis = Ak ¢( 2000™ 1993 > : - igs STAT) 1 oslydl ots gts X16 (866-1 = 3 Dyed ae 5g) Sisky my saabd G49 “Vij GS vay Sh gS) used Us = eijklme (2 angel agg) Sa al UT lhl se 5h CnNSg Nine shay dass Winns nts Leigh lege hay FD age Bap al Aa ae sal Pees Wy Lap ty Uday k BY dss j Oud Minimum) MIVQUE 4 ij casas Ti) 1 Saad oat 8 = Sj Akad te baa 21) ( Variance Quadratic Unbiased Estimation «(BE = dsl oss) ce 1 (d= LSI ae Ea pte co ya Gat Ob Se td J~ age) Ge 3 UT T) ed eg Sd pd LES CS gel Gil fly LN Le AL Rp (ill Sass Syl) Gada ds ( 2 AN eb capt Sis ge (hha) ag) ) Apia 8 gantsiny (B= Tm) j Dh ae Yifkimno = p + Si+Rj + Ak + X1+ Fm + Dn + eijklmno, PD ASAD 3 Lak M1 5A ay Dg Gat gts ig gil Fim otal SMT cha 8 lead Syed (9) ABN) CMake Gane le ( hm ) ged W'S +FD/+ 0S + 0D + CE, oP = ota + o%m +E, 0'S=% oA, a= 4 0°S OD=% oat Fm, oD=o'S + om, om =o7D-o'S ,GE= oe-20°S we (gS 35,17) Nelasiy { giS 35.37 ) Lewy all yt cecal le = ota « (GAD) Gp glad gala = *P (3S 34.70 ) Vinny ( pS 33.52 ) Ua gd Sogh al us cee als = De NI sl Ge alah = oS 6 aad ja gal (3) gl aegs LI MALL Ral ode th HOPE 6 MN 8D ae ca eel gh MOLI Sa thee cll ital a Lad ols = OFe «ust a 3838.03 ) lett ( p85 38.16) ny cals ci ps ( 35 36.87 ) Binoy ( pif 36.50 ) Heys US, ( chat A al Sle EE Gl AE spe Se Rea ail Jans alee te Shia Spell op Lee SM) gle Ue Halal Ga il GS pe WD tie le dino JS SBM Se) cl ji PS Abell YOY A I gay ee St te! SA 3 Ae Jal }sS 34.28 Du! Soe DSN) 6S By pen GaY ean Dyk 4S 34.96) (2) all ose y Ll lia ny (1 nn Hage Gala (14) yal G39 Joes Gass (end (8S 36.3) (17) Hermiz hyst oe dy ( j1) Chal jal gine gS Milly Syl) ha fs a Bod A cstatlsed 55 ( 98S 36.72) (3) nts Hillers Ageeb citi ea GY We Ve (2455 52 Bhad A Dd deal goer boy WI (6) AEA LS UL ee cals yd ae gk BE OID BS ay Diet Day Leydig a OD aad Els Gk Lad ge OS 78GATS ght ) EDs Gb ss8) Sy SF yine ashy ain hs Aggal 13 ay ( 2S 33.64) GUI le ( as 35.74 BH oie I Se. (8 5 ioral at dag a Apadll Oe Gamal Oat J tle Go Sd Bas US 6 (7) gail $35 oh HA GD See ee og BLS Gt Lb LS 6S ded eb LV aS (18) 485g) ee (2007) 86-76 :(2)38 — 481-8 Getj 30 pslell Une 1993 fe ( pS 3748) Wed cab Oya Ap Sb LD J + 2000 ple ( 55 31.38), Basin GB OND Byes Shpall ge GBR LAY GeaVly DEY! ass Roba Ley hyd Goal, Ga ye Gyles SG PHD Labs ts of es OS Lyd (6 Sos sal A hess Li dail ge qa) Oop Radke a pd (98) Se Ae Cl ail LAR gel ee age dn eb had a yaad Sta se iat se sist dig 049 23428 Mie wae 2 0l6E 3507 a5 4 2020 + 3537 as ~ 0.3 #3470 Tin SS COS £33.52 936 Ss a Ded ae E 2031 23748 Ey 195 2 e035 = 3570 : 160 oy €020 33.71 cu 95 OS + 35.74 ‘S83 1998 0.20 + 35.87 amt 1997 ©0419 + 3529 80 198 ~ bOI 3236 9 DP 024 + 3138 35 my |e _ 3013 + 3445 ms aa aQIS + 34.63 83 yn 0.18 + 35.02 Ei 0 8021 = 3500 406 aa gl 20.28 + 34.73 235 Dany a 0.284 34.32 239 cl ade oe a 0.13 235.74 1604 Ss b ans 33.64 1808 7 Yh 5 Se 5e ylay LA Lyne GURY Goyal Jule JS hygiene Me ge JaSS (ll loa sal 79gay pa (2007) 86-76 #(2)38 ~ 43.29 Ke; asad dns CGD EL Die ae Og) gd FGA all hl es 2 Jose Stas al dase Bplay CAB ae *9537.77 3 Dy dead °° 1439.25 6 yall Ria 39.84 5. BS Juba #6 3315.22 1 alg yap 18.54 3395 ae Uns (001> i)** Si dhaaad taj GD se SHLD Sa ge gia BEANO ey Sad Sie gall Lay ae CoS 4S J iy So EL Baad USM GI 583) OF ai SAN Quen Gallia oy Gam gd jy JN) DL Gy ald ase dash J US cle Aja ell Ald Gal Be Ge (1019) S81, (359) 38a Aang G59 53) Senay pli Se Gj A Hagin Ge Regd 455) 5205) baste FS yam Oe Une (2) a LE gh pS On Sd phy 2 438.64 SM MI ce Lat gh de 500 G GB tb sie cet) gla) sill Gh Lh ued hae «bey 123.46 80 lh oe Gl oll Leesa 3 ase Ge ey gibi GB Dsl go S53) HLM 5 AS 8858 GL (4 Spe) Ded eedy Lysine |B ay «as 53.69 57.82) 4p9h Sab fy (AS 92.07) (Ua oy lel aly 8S) Gia CH pal Gd Sa Lak a. (aS OS A le AS 50.52 ) 5 (985 86.36 ) lett Jes J Sang | OSiinall IS gi Ligins Dodd Au ahs hed Heeb ASM, IOI shee Gls GI alls 8S OS Lak gates ot IS END Jabs HS oI ayy Se jp) Gb Daal gis 6 Lysine olga Goig ce ( 5S57.94 ) Aaja) Sab jy ( pS 92.27 ) Lin ee CH! Si J AS 48.70 5 5 85.30 Jyad Soalia LW! ag) yoy gD csthad otle A 33Sgy le (2007) .86-76 :(2)38 - Malad Kei 3 psa Une © fed) Haase Seta Sass (AS) (MN ae Gig A AMUN + ig fel) Slay a usta 3 dgey AO TL 5 gad ayy asia Aaja bay [ae ost AL ae $B dd 043 + 53.69 058 # 88.55 137 ee | Be eat | £0.62 + 50.52 0888636. an eat Le b 0.63 £ 53.62 bossss.37 | 292 es cos estat | coms 6735 “6 rey 2.0.67 + 57.82 1089 + 92.07 208 ae ae con a8) i)" Aang peal ctl lth y phd col yall eA nag Gl Gl gSa 7 Jose em ta [oP a oe an 1 “Trote ot | 016 [20.16 14.83 [2.13 0.06 0.16 87.56 68.20 /5.36 0.03 [021 fi6a9 38.24 3.21 00s [0.06 745.95 027.28 13.19 oor [0.23 8154 288.73 4.93 atl ROPE AM Aas odd = 07m «eee Ais! oat = ofa cea Ag eA = Wim. Ash gal = Wha (AS) gpl Gal = oP chest 8 doe geal desl Hag sae dina { [8241 | 852.56 igh Bal | Aan] AZ 3.96 ag Lj at Ba | ol 4.00 | ai ae pad 84 (0.01 > 1) **eBay lal (2007) «86-76 :(2)38 ~ AA aN e130 9 ad Hp ADU Kany a cha (BLUP) 11398 Jan La ily yh cl ast 9 dae fos re ea gi [aja Linn ye Ej pu tad as 2.41 - 51.97 Dy ae aby e410 = AS O38 3 54.50 ~ Hai3gh 8:30 = 28.62 ~ Agi jgd Bay 3t Jane ag 7.65 — pial) le pat sb Kay piel China! (BLUP) G29! 5 sat Lak ia gel cd pes 10 dys Pas aid yt Pal Jauis yer | Lyud aa 2s 3.83 - [| as335 Dual te a5al so013- | 0.012 phil oe G5 350,980 ~ a 1.02 ponds tif BSD 2 Holdall a Jos eagle 1982. Gas cued « Dye 4 Jged Guanis 106 5 48 dainal 5235 S-Abanikannda O.T.F. , ©. Olutogun and A.O.Leigh . 2001. Genetic parameter estimates for yearling traits of N’Dama cattle in humid tropics of Nigeria.J. Dairy Sci.85, (Suppl.1:1287 , 6-Ageeb ,A.G, and J.K Hillers.2005.Production and reproduction characteristics of Butana and Kenana cattle of the Sudan. swww.fao.org/agiaga/agap/ rg/Feedback/war/ut2 00bo. 7-Ataal —J.1969.Level_ of testosterone androstenedione , estrogen and estradiol in the testes of fetal sheep.Endocrinology ,85:280 — 289 85 2101 ~ 224.61 2 Haigh Fay dame | 29 6.40 ~ 29 5.40 AbiN Se yd” ileal Lo pdt any 2002 gacad gag) peas gH} Hine. Shp JG) sal yall Cline Ga ne aE BS 188 ~ 183 (2) 33. Ail pal Gel py 2004 . ae Jey aby « Bahl ~2 ia IHF oy HDL BN) By la oe any OEE Dad J Asa ALS Gd geal CGY CL wl, yell, 50 Rained pil path Sha « ad Ne Sy A Talal -3 2006. SM 25 «gas Ay cite oe gad Gb Help qghll CS Aula dla Dud ae Wy 37-28 1(3)6.HAS ol eat (2007) «86-76 :(2)38 ~ 48 px X61 439 pgteh Uae Program.tsers Guide for LSMLMW.The Olio University ,Ohio.USA,P: 62. \7-Hermiz. .H.N..K.H.Juma_,8.S.Khalaf and T.Sh Aldoor.2005.Genetic parameters of production , reproduction and growth traits of Holstein cows Dirasat ,32:157 ~ 162. 18-Jainudeen| MR. and E.S.E-Hlatez:1980.Gestation. physiology _and reproduction in farm animals. Lea and Febiger, Philadetpia. p.247 ~383. 19-Lykins LE. , LK. Bertrand and , 1.P. Baker and T-E, Kiser .2000,Matemal birth weight breeding value as ant additional factor to predict calf birth weight in beef cattle.l.Anim.Sci.78:21 ~26 20-Maria G.A., K.G, Boldman and 1.D.Van Vieck .1993, Estimates of variance due to direct and maternal effects For growth traits of Romanoy sheep.J Anim Sci-71:845 — 849 21-Rao,C.R.1971. Minimum variance quadratic unbiased estimation of variance components ./ Multivariate Analysis..| :445-456, 22-Rohrer G.A. , JF. Taylor , 1.0. Sanders and RM. Thallman 1994, Evaluation of line and breed of cytoplasm effects on performance of purebred Brangus cattle . Anim Sci.72: 2789 — 2803 23-SAS.2001. SASISTAT Users. Guide for Personal Computer.Release 6.12,SAS Institute Ine.,Cary .N.C.,USA. 24-Saucedo JS. , L. Avendano , F.D. Alvarez C.Cecena , T.B.Renteria , J.P. Moreno , MF. Montano , G.E. Medina and M.P.Gallegos .2005 Performance of Holstein calves at weaning according its birth season in Mexicali valley Proceeding , Western Section American Society of Anim. Sci. 56:170 ~ 172 86 8-Bourdon , RLM. 1999. Understanding Animal Breeding . Prentice Hall , Upper Saddle River, NJ. USA, P: 223. 9-Cameron N.D.1997 . Selection indices and Prostuction of genetic merit in animal breeding Cat International .UK., P: 46-58 10- Dodenhoff J. , L-D. Van Vleck and KE.Gregory _.1999, Estimation. of direct, maternal anid grand maternal genetic effects for weaning weight in several breeds of beef cattle Anim. Sci.77:840 ~ 845 TL-Eler_ J.P. , L.D. Van Vleck , J.B.S. Ferraz and R.B. Lobo . 1995 .Estimation of variance due to direct and maternal effects for growth traits of Nelore cattle J.Anin.Sci.73:3253 3258 (2-Ferraz LBS. , JP.Eler and P.M.T. Ribeiro.2000 . Genetic study of Santa Gertrudies cattle in Brazil 12-219 13-Ferreira G.B. , M.D. MacNeil and L.D. Van Vleck .1999 . Variance components and breeding values for growth traits from different statistical models . J.Amin, Sci. 77: 2641 — 2650 « Livest, Res., 12 (2) 14-Goyache _ ,F.,..Fenandez,I.Alvarez,L.J.Roy and J.P.Gutierrez.2002.Gestation length in the Austuriana De Los Valles beef cattle breed and its relationship with birth weight and calving ease.Arch.Zootec.5:431-439. 15-Gutierrez J.P. , 1. Fernandez , LAlvarez . Loy and F.Goyache . 2005. Sire x. contemporary group interactions for birth weight, and preweaning growth trails in the Asturiana De Los Valles beef cattle breed www, Elsevier. Com locate/livprodsei. I6-Harvey ,W.R.1991.Mixed Model Least — Square and Maximum Likelihood Computer