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F
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buonNo.132.
 Tabl
eofCont
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F
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• BF
ARDi
rect
or,A y.AsisG.Per
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• BF
ARI
V-ARegiona
l Direc
tor
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daP azD.Manal ang.
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Noe lC.Barut.
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.1
I
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.5
I
V.Howt
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V
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• Aca nthur i
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• Be l
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• Ca esioni dae. .
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• Ca r
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• Chiroc e ntridae..
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• Clupe ida e. ..
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• Cory pha enidae....
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• Da sya da e. .
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•L eiog na thidae...
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•L ethr i
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•L utjanida e. .
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30

i
i
i
Ta
b l
eofCont
ent
s

V
. F
ishAt
las(
connuaon)
• Me ga lopi dae...
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• My ctophi dae...
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• Ne mipt e ridae..
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• Ophi cht hida e..
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.40
•P aralicht hy i
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• Platyce pha l
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dae. .
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.41
• Plotos idae. ..
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.41
• Priaca nthi dae..
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.42
•P se odi da e...
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.43
•S carida e. ...
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.43
•S combr i
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.46
•S errani da e. .
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.49
•S i
g anida e. .
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.53
•S i
llagini da e...
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•S parida e....
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.56
•S phy ra enida e..
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.56
•S ynodon da e..
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.57
•T erapon da e...
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.58
•T ric
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.59
Ref
erenc
es.
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.60
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.61

i
v
 For
ewor
d

I
ti
swi
thg
reatpl
eas
uret
hatIac
cept
edt
owr
it
eaf
orewor
dfort
hisbook
.

Ihavetwor easonsforthis
.Thefir s
treas
onisIspentmostofmypr ofes
sional
l
ifei nthe Phili
ppines,and Ik now how muc hg ui
destol ocalfishe r
ies
resourc
e sareneede d,notleastbypers
onneltask
edwi t
hidenf yingfishi n
view ofe nt
eri
ngt heircat
chinna ona lfisher
iescatchst
a sc s.Thus,this
“Atl
asofCommonF ishesofTayabasBay,Quez
onPr ovi
nce,Phil
i
ppines”wi l
l
benotonl yappre c
iatedbyt helocall
oversoffis h,butalsobyfis he r
ies
profess
ionals
.

The ot
herreason i
st ha
tt hi
s book ma
kesexhausveand ex
cel
lent
use of Fi
shBase,the globa
l onli
ne enc
ycl
opedi
a of fis
hes (
se e
h p:
//www.
fishbase.
org
).

Fi
shBase,now a ninterna ona l
lyr espec
tedg l
obalresourceforscienst
s,
res
ourcema nager
s ,s
tude ntsandt hepubli
c,wasconcei
v edinthela
te1980s
attheInterna onalCe nterf orL i
v i
ngAqua cRe sour
c esMa nagementin
Ma ka ands ubsequentlyimpl eme ntedbya nint
erna ona lt
eami nwhich
Fi
li
pinosplayedac ruci
a lrole.Thisi ssllthecasenow t hatFi
shBasehas
becomeg l
obalandha snode si
nc ountri
esrangi
ngfrom ChinatoSweden.

Ic
ongra
tulat
etheauthor
softhi
sbooka
ndl
ookf
orwa
rdt
omor
eli
kei
tfor
ot
herpart
softhePhi
l
ippi
nes
.

DANI ELPAULY
Fis
heri
esCentr
e
Uni
vers
ityofBrishColumbi
a
Vancouver
,Cana
da

v
Mes
sage

F
is
rto l
fal o
,Icnr
gau
tlt
a h
etePhi
l
ippn
ieBue
rauo i
fFshei
re n
sa dAquac
Reo
surcs(
e BFAR)Rego
inIVA,s
- pecifia
clyt
l heNao na t
lSockAse
sssment
Po
rgrm(
a NSAP)o
frco n
migup wt
ih h
tisA
“ta
lsofCommonFih
sesofTy
aabas
Ba,
yQuezonPo
rvine
c,Phi
l
ippn
ies”
.

Asde f
i rom cons r
eving and protecng t he P l
hippi
i nes’boun fulfis hery
resources,wene edtoma nage andu e
s them propery
l.We can do h
t s
iusing
enoug ha ndaccurateinformaon.T heRe publcAc
i t8 550ort heP l
hippi
i ne
Fihe
s re
isCodeof1 998ma ndt
aed BFAR o
t fom
r ulae
t polc
iy measue
rs forthe
ma nageme n,c
t onsev
r aon, and po
r e
tc on of fishery reo
surces bae
sd on
accurateinor
f maon. Secon s7,8 and9 provd
ie tht
a t
sockasses
sm entsu
td e
is
shouldbedonei nor dert oobtan
i infom
r a on foruse n
i the es ma on of
Ma ximum Susti
anableYield(MSY)and Totl
a Al
lowableC t
ach(TAC),h
tree e
ya s
r
ae t
risimpleme naona
t nde vr
eythreeyearsthereae .
r

NS sone ofs
AP i uchs udi
t eswhicha re dei
sgnedt og eneraec
t on nuous
n
i or
f maononfis heryreour
s ceswhih
c s
ivt
ialtothe management ofourfis r
hey
reour
s cs
e ti
.I sdei
sgnedto sr
tengh
ten and n
is tu onalz
iethe capabl
iieso f
theRegi l
ona Officesonr s
eourcesas
sesm
s ent,ma nae
gm ent,anddevelopment.
Theinor
f maon- s
bae dresouc
re management s
i needed towardssustan
iabe
l
deveopme
l ntandu liaonoft
z hecounty
r’smarineresoure
cs.

T sa
hi l
taswilde
l t
finiel ni
ybe a mpora
tntree
frnc
e ethatwe c ef
n us
a or
i
poly
c-mak yf
ngnotonl
i h
ortesad
iB y
a buto
froh
terae
rasaswe.
l
l

a
Agi c
n, r
ongaul
t oNS
a onst ARRe
APofBF g V-
onI
i Af hi
ort c
sac s
i
omplhme !
nt

ATTY
.ASISG.PEREZ
Dir
ect
or,BF
AR

v
i
 Mes
sage

WehavenowoneofthemostimportantachievementsoftheRegionandof
thePhi
li
ppi
neBur eauofFi
sheri
esandAquacRe sourc
es(BFAR)
.T hi
s“Atl
asof
CommonF is
he sofTaya
basBay,QuezonProvi
nce
,P hi
l
ippi
nes”repres
en t
st he
effort
sandresourcesofBFARma nag
ementa ndst
affs i
ncetheNaona lStock
Asse
ssmentProgra
m( NSAP)s
tar
tedin2003.

TheNaona lSt
ockAs sessmentP r
ograma s
sessesthefis her
yr es
ourcesof
TayabasBa ywhichwasoneoft hepriori
tyareasundertheF i
sher
iesS e
ctor
Progra
mi n1 987andint heF i
sheri
esResour
ceMa nagemen tPr
ojec
tin1 99 7.
NSAP e nabl
esust or ec
omme nd and f
ormula
tepolic
yme as
urest o be er
ma nage,conse
rvea ndpr ote
ctourfis heryres
ourcesforthebe nefitoft he
fisherf
olksandotherst
akeholders
.

Wit
hthisAtl
as,wehopet oide
nf yfishspe c
ieswhichneedtobepr ot
ect
e d,
manag
eda ndc onser
vedb yimposi
ngthec orr
ectme shsi
zesoffishi
ngg e
a rs
orbyimplemen ngac l
osedseas
ontoa ll
o wg r
owthofjuveni
lefishes.This
At
laswil
lbe ofus eto pol
icymak
ers
,r es
e ar
chers
,a nd ot
hergroupswi t
h
i
niav esonfishe r
yresour
cema na
geme nt
.

Iwoul
dli
ketoc
omme ndt
heBFARRe
gionI
V-ANSAPt
eamforthi
sac
hie
veme
n t
a
sthisi
sal
soamil
est
onef
ort
heBFARRegi
onIV-
Amanag
eme nt
!

E
SMERALDAPAZD.MANALANG,CES
OV
Reg
iona
lDi
rec
tor
,BF
ARIV-A

v
ii
Me
ssag
e

T
heNaona
lFi
she
rie
sRe
sea
rcha
ndDe
vel
opme
n tI
nst
ute(
NFRDI
)ast
he
resea
rcha r
m oft heBur eau ofF i
sheriesand AquacRe sources( BFAR)is
commie dtoi mpleme nttheNaona lS t
ockAs ses
smentPr
og r
am( NSAP )
.In
parc ul
ar,NF RDIs ha l
lge nera
tes ci
e n fic inf
ormaon,t echnol
og ie
sa nd
knowledgethatwillbeus edb yBFARasba s
isofcomingupwithma na g
e ment
str
ategi
es,polici
esa nd r
e gul
aons f ort he conser
vaon and s ustaina
ble
ma nagementofthec ountr
y’sma r
inere
s ources
.

Ast heBF ARa ndNF RDIstri

vetoa a insus
tai
nabl
efis her
iesmanage
me nt
,the
NSAPwa slaunchedin1 996andev e
n t
ual
lyi
nstu onali
zedatt
heBFARRegional
offic esinmi d-2000.T heNS APs tanda
rdi
zedtheme t
hodol
ogyofg a
theri
ng
fishe r
iesda taforthewhol ec ountry;t
hus,pr
ovidi
nga nint
egra
tedresearc
h
re
s ulttha twi l
ls upporta ma nagementpoli
cya ppl
ica
bleforthe enr e
ma ri
ner esourcesofthecountr
y.

I
nRe gionIV-
A,thea reainfocusi sTay
abasBaywhichwa soneoft hepr i
ori
ty
ar
ea sundertheF i
sheri
esS ectorP r
ogr
ami n1987andtheF i
sheri
esRe source
Mana gementProjectin1 997.T hr
oughNS AP
,BFARRegionIV-Aa s
s e
s s
edt he
st
atusoftheBaya ndc ameupwi threc
omme ndaonsforpoli
cyformul aont o
be erma naget her e
source.T his“At
lasofCommonF ishesofT a
y abasBa y
,
QuezonP r
ovi
nce, Phi
li
ppines
”i sthefirs
tstepi
na ai
ningthatobj
e cv e.

TheNF RDIex
tendsi
tshe
a reltcomme ndaontotheNSAPTe
amof BF
ARRegi
on
I
V -
Af orthi
sa c
hiev
emen ta
ndhope stha
tmores ci
en fici
nfor
maonc a
nbe
producedandaddedtotheNS APbodyofknowl
edge.

Ial
s ocongrat
ula
teandthankBFARIV-
ARe g
iona
lDire
ctorDr.E
sme
ral
daPa
zD.
Ma nala
ngf orhersuppor
tint heprepa
raona ndpubli
caonofthi
sAtl
as.
Cong r
atul
aonsa ndkeepupthegoodwork!

NOELC.BARUT
Na ona l
Coordi
nat
or
Na ona
lStockAsses
smentPr
ogr
am
BFARNFRDI

v
ii
i
 Pr
efa
ce

PursuanttoArcles6,7and12oftheFoodandAgricultureOrganizaon(FAO)
Codeof Conduc tforRe spons i
bleF is
hing(CCRF),prope ru liz
aon, conservaon,
protecon,de vel
opme nt,andma nageme ntoft hec ountry’
sc apturefis hery
resourcesre quirevita
l inf
or maonont hefis heri
es,includingbiologi
ca ls
tatusof
ma jorspe ci
e sb yarea.Likewi se,Arc les7,8a nd9ofRe publi
cAc t85 50ort he
Phili
ppineF i
she ri
esCodeof1 998spul atethatc on nuouss tocka sses
sme nt
shouldbec onduc teda stheba sisforthede t
erminaonof Ma ximumS usta
inable
Yiel
d( MS Y)a ndT otalAll
o wa bleCa t
ch(T AC)
. T
heout putoft hisres
e archwillbe
theba sisfort heformul aonofr egulatorypoli
ciesincludingthede terminaon
ofspe ci
e stha tneedc onse rvaone fforts,decl
araonofc l
ose dandope nfishing
area s
,theimpos ionof ba nningof catc
hof certai
ns peciesandt hereg ul
aonof
theme shsizesoffis hingg e a
rs.

TheP hili
ppineBureauofF is
he ri
esa ndAquacRe sources(BFAR)t hroug ht he
Naona lFisher
iesResear
cha ndDe ve l
opme ntInst ute(NFRDI)isma nda t
edt o
i
mpl eme nttheNaona lSt
ockAs s
e s
sme ntProgram( NSAP )
.InBFARRe gionIV -
A,
thepr oje
ctwa simplementedi nTayabasBayi nQue zonP rovi
ncea soneoft he
studya reas
. T
hes t
ocka s
sessme ntstudyinTayabasBa yg enera
tedfiv ey ear
sof
da t
aonpr oducon,s pe c
iescompos ion,a nds eas
ona li
tyofg ear
sa nds pecies
.
From thepr ojec
tdata,alis
tofc ommonl yc aughtfis hspecieswa sde ri
v eda nd
wa stheba si
sforcomingupwi ththispubli
caon. This“Atl
asofCommonF ishes
ofT ayabasBa y
,Que z
onP rovince,Phili
ppi
ne s”isoneoft heout putsofNS AP
i
na na e mp ttoprovidethes takeholder
sac ompr ehensiv
er ef
e r
e ncei nt he
i
de n fic aonoffis hspe c
iesandr egulangme shsizesofne tsforas usta i
nable
fisheryr es
ource.

Toful
lyimplemen tthepr oj
ect,
BFARRe gionIV-Ac ol
laborat
edwiththeF is
hBase
I
n f
ormaona ndRe sear
chGr oup,Inc
.(FIN),aPhili
ppineNGOe stabl
is
he din2003
thatma nagesFishBase,ag l
obalbiologic
a linf
ormaons ys
tem onfin fishes
.
Fis
hBasei seas
il
ya cc
ess i
bleont hewe ba ndi sf r
eeofc hargetot hepubli
c.
AMe mor a
ndum ofAg reeme ntont heP ubli
caonofa nAtl
as/Handbookof
CommonF is
hesof Tay
aba sBaywa sf
or gedbe t
we enBF ARRegi
onI V-
Aa ndFINin
August,2012.From BF AR’sinialfishphot ographs,F I
Nfaci
li
tatedc ont
acng
thei
rFishBasec
ollaboratorsandsoughtpe r
mi s
siontous ethei
rfishphot ogr
aphs
toaddt othi
sAtl
a s.

T
oa ddv al
ueandtoincreas
et heusef
ulne
ss,sc
opeofu l i
tyandreaders
hipof
t
hisAtl
as,foreachspeci
es,theasympto cle
ngth(L∞
)- t
he si
zea fishwould
aa i
nifitisal
lowedtog row,maxi
mum length(L
max
)-the s
iz
e ofthelonge
st
i
ndivi
dualrec
orded,andlengthatfirs
tma tur
it
y(Lm
)- t
he si
zeatw hi
ch afish
matures
,we r
ea l
sopr ov
idedb yFI
N.T hes
elengt
hi ndi
cator
swi l
lale
rtLGUs,

i
x
Pr
efa
ce

rese
archers
,thea c
a deme,andot herde v
elopmentpar
tnersaswe l
lasfishing
communie sthatthecaptur
eofj uvenil
esc a
nresul
ttoo ver
fishi
nga nd,thus,
stoc
kde ple on.Thisincr
e a
s e
da warenes
sa mongs t
akehol
dersofa ll
owing
fishtobe comes exual
lyma turebe f
oretheya r
eharvest
edc anbri
ngg reate
r
appreci
aonoft hev al
ueofgo ver
nme ntcontr
olmeasur
esanditsi
mpa ctonthe
fisheryres
ources.

MARIBETHH.RAMOS RODOLFOB.REYESJR.
E
SME RALDAM.ME NDOZ
A EMIL
YC.CAPULI
ALMAG.S ANTOS MARYANNP .BI
MBAO
BFARRe g
ionI
V-A Fi
shBa
seInf
orma ona nd
Res
ear
chGroup,I
nc.(
F I
N)

x
 Ac
knowl
edgme
nt

T
hea
uthor
sac
knowl
edg
e

 t
hesuppor
tandencour
agementg
ive
nbyfor
merRe
gionalDi
rec
torRos
aF.
Mac
asduri
ngthec
onceptua
li
zaonands
tar
tofi
mpl
eme nt
aon;

 BFARDirec
torA y.Asi
sG. P
erezandRe
g i
onIV-ADir
ect
orE
sme r
aldaP
azD.
Ma na
langforpr
ovi
dingtheinspi
raoninproduci
ngthi
sAt
las
,thefirs
tfish
gui
depubli
c aonfr
omNS AP;

 t
heNF RDIgroup,NoelC.Bar
ut,Mudji
eD. Sant
os,MelchorM.Tayame n,Fe
L
avapie-
Gonzales
,andEunic
eC.Bognotwhopae ntl
yencour
aged,provide
d
i
nputsandsuggesonstoimproveouroutput
,andtoSaludR.Ganadenwho ,
i
nspiteofhavi
ngrer e
dfromg ove
rnme ntse
rvi
ce,prov
idedval
uableinputs;

 Prov
incial
Fis
heryOfficerSoni
aO. El
l
oso;NSAPenumerat
ors
:Arl
e neE.Ag
or,
Monc hi
toM.Vi l
la
nea,J
inkyE.Padi
ll
o-Ri
coher
moso,Jes
usaP.Ba l
bi
n,and
Chri
stopherH.Ba usta;Res
e a
rch Assi
st
ants
:LouieV.Ga t
labayanand
Wil
fredoO. F
aja
rdoJr
.whopae ntl
yg a
there
d,anal
yze
d,andencodeddata;

 t
heenumera
tor
s,t
hefisher
s,fis
htr
a de
rs,a
ndr esi
dent
sofTa
yaba
sBayf
or
t
hei
rsuppor
tandcooper
aonduringthec onductoffie
ldacviesand
s
urv
eys;

 thephotogr
a phowne r
sandc ontri
butor
sforgr
an ngpe rmi
ssiontous etheir
fishphotosforthisAtlas
.Inthec as
eof mul-authore dwor k,permiss
ionwa s
grant
e dbyoneoft hea uthorslabel
ledwith“‡ ”:T homa sGloe r
felt-
Tarp,
FeL avapi
e-Gonza l
es,Gr egori
oV .He r
mosaJ r.,K athlee
nK esner-Reyes,
Keii
chiMatsuura‡,JohnE .Randall
,KwangTsaoS hao‡( TheF i
shDa tabaseof
Tai
wa n,h p://fishdb.si
nica.e
du.tw),Toshi
yukiS uzuki,Mi c
ha elN.T r
evor,
Demia nWil
lee ,
Jeffr e
yT .Will
iams,andRic
hardWi n terbo om;

 Y
uki
oI wats
ukioft
heUniv
ersi
tyofMiyaz
aki,Japa
na ndJ
ohnMcCos
kerof
t
heCa l
if
orni
aAcademyofS c
ienc
es,USAfort hei
rhel
pinva
li
dangt he
i
den ficaonofsomespe
ciespres
ente
dinthisAtl
as;

 Lecia Diz
on,f
orme
rly ofthe I
nte
rnaonalCent
erforLivi
ng Aquac
Res
ourcesMana
gement(
ICLARM,Maka),
foredingthi
sAt
las
;

 FI
Nc ol
l
e a
guesChri
sne Ma r
ieV .Casa
l,Ra c
he lC.At anaci
o,Joann R.
Glor
ioso,and L
yraJoyc
e N.P a
gulay
anf orc ontr
ibu ng thei
rF i
shBase
expersea nd ex
peri
enceincoll
ang s peci
esi nf
ormaon,r esear
ch of
documen t
aon,phot
ov a
li
daon, a
ndhe l
pingwiththela
y outofthi
sAtl
as;

x
i
Ac
knowl
edgme
nt

 theFishBasearchit
ectsDanielP
a ulyandRainerFroe
sewhoin1990have
envis
ioneda ndwor kedi nt
of r
uionf orFi
s hBasetobec
omethemos t
compr ehe
nsive fish dat
abaseg lobal
ly,and Fis
hBasePr
ogra
m Ma na
ger
Nicol
asBa i
l
lyforthei
rtechni
caladvice
;

 ourfa
mili
esandfr
iendswhog
avethei
rsuppor
tandc
ont
ri
bu onf
ort
he
suc
cessi
nthepubl
i
caonofthi
sAt
las;
and

 mostofa
ll
,theAl
mig
htyGodforthes
treng
tha
ndg
oodhe
alt
hbe
stowe
don
usi
nthecours
eofdev
elopi
ngthi
sAtl
as.

x
ii
 I
ntr
oduc
tion

T
ay
aba
sBa
yisl
oca
tedi
nthemi
ddl
eof
thePhi
l
ippi
nes
,nor
thof
theVi
say
anS
ea.
I
tcov ers1 6muni ci
pal
ie sa ndonec it
yintheprovi
nce sofQuezon,Ma r
induque ,
andBa tang asinthesouthe rnpartofLuzon(Fi
g.1).I
nQue zonP rovi
nce, Lucena
Cit
yi sthel onec i
tyalongT ayabasBa yandits11muni ci
palie sareAg dang a
n,
Catanaua n,Ge neralL
una ,Ma cal
elon,Mulanay,PadreBurgos,Pagbil
ao,P itogo,
SanF ra
nc isc
o ,Sa
riaya
, a
ndUni san.Ma ri
nduquePro v
incehasfourmuni cipalies
compr i
singBoa c,Mog pog ,Sta.Cr uz,and Tor
rij
os.Ba ta
ng asha sonl yt he
muni ci
pa l
ityofS anJ uana l
ongT ayabasBay.One -thi
rdoft heba rang aysin
Quezona rec oas
tal,andt heresta r
elocate
di ni
n ter
iorareas.Therear emor e
i
n t
eriorba rangaysthancoa st
albarangaysex
ceptinP adr
eBurg osandPitog o.

T
ayabasBay,
whichformsaroughsqua r
ewithsidesofa bout6 0kminsouther
n
L
uzon,has49% ofitscoa
stl
ineinQue zonP r
ov i
nce,45 %i nMa ri
nduqueand
6%inBa t
angas
.Themuni c
ipal
ityofS t
a.CruzinMa r
induqueha sthelongest
c
oastl
i
nea ndthemos tnumbe rofc oast
albarangaysi ncl
udingsever
alsmall
i
sl
ands.I
ntheprovi
nceofQue z
on,thecit
yofLuc enahast helonge
stcoast
li
ne.

I
n2 010,TayabasBayhadatot
alc oast
alpopulaonof4 09,
000c
ompris
ing
82,
000 households(NS
CB,2010)
.F ort he whol
e Ba
y,25% were fis
hing
house
holds,
de fineda
shous
ehol
dswi t
hatleas
tonefami
lymembereng
agedin
afishi
nga cvity.

F
ig.
1.MapofT
ayabasBay
,Phi
l
ippi
nes(
hp:
//ma
p c
art
a.c
om/
15710990)
.

1
I
ntr
oduc
tion

TayabasBa ywa soneoft hepr i

ori
tyba y
sunde rtheF isheri
esS ectorProgram
i
n1 987a ndF i
sheri
esRe sour ceMa nageme ntProjectin1 997(F RMP,20 03;
DA,1 991).The seprojectswe reimpl eme ntedtoa ddressthema jorprobl
e ms
i
de n fiedi nt hes ect
ors tudyoft heAs i
anDe velopme ntBa nk( ADB,2 007):
resourcede ple oni nt hec oa s
talzone ,widespr
e ade n v
ironmen taldama ge,
pov e
rtyamongmuni cipalfis herme n,lowpr oducvityof aquacult
urea ndli
mi t
ed
ul iz
aonofoffs horea ndE xcl
usiveEc onomicZone( EEZ)wa ter
sb yc ommer c
ial
fisherme n. T herewe r
e1 2pr i
ori
tya rea
sc hose
nont hec rit
eriaofs ervi
ce
generaonr equiremen t
, e
n vironme nta l
deg r
adaonpr oblems,pove rtyl
eve
lsof
communie si nt he a r
ea ,a nd iniav esi ns elf
-r
e gul
aon b yt he Local
Gov er
nme ntUni ts(L
GUs )a ndfis herfolkassociaons .

I
nBF ARRegi
onIV-A,f
ourpri
ori
tyba
ysquali
fiedfort
hecri
ter
ianamel
y,Manil
a
Bay
,Cala
uagBay,Tayaba
sBay,andRag
a yGul
f.Taya
basBayisal
sooneofthe
pi
l
otba yswheretheUnifiedMunici
palFis
heryOrdi
nanc
ewa sadopte
da nd
i
mplementedbytheLGUssur
roundi
ngtheBaysi
nceMay2003.

Thewe st
e r
nc oastl
ineoft heBa yi smos tlyc overedwitha romaa ndc oconut
trees ,wit hs ome s ma l
lpa tchesofma ng r
ovet r
eesi nr ivermout hsa nd
refore s
taon a reas
.T hes horel
inei sr oc k
y ,inte
rspersed withg entl
es andy
be ache s
.T hes hal
low poronoft hes e abe diswi deront hiss i
deoft heBa y
.
The nor thern-c e
n t
ralc oastl
ine,f rom P agbil
aot oP adr
e Bur gosi smos tl
y
ma ng roves wa mp s,e s
pe ci
all
yf r
om K anlur anMul awi,inPa gbil
aotoS il
angang
Mul a wi,i
nP adreBur gos.Ho weveri na r
e a swi t
hfis hponds,thema ngrovecover
ha sbe enr educ edtot wot othreer owsoft r
e es.Onlythema ng r
over es
ervaon
oft heDe pa rtme ntofE nvir
onme n ta ndNa turalResources(DE NR)hasluxur
ious
co verofma ngr ovet r
ees.S eagrassbe dsa refoundont henor t
he as
toft he
Pa gbila
oGr a ndeI sl
and. Sma l
lpatche sofs e agrassareals
ofoundont hewe s
tern
sideoft hei sland.T hee asternc oastli
nei sc over
e dwitha romaa ndc oconut
trees .Mos toft hec oralre efareasi nt heBa yha vepoort of ai
rc or
alcover
(Re sourcesCombi nes,Inc.andM. S .
E .Uni v.Foundaon, I
nc.,19 96)
.

Thema j
orr esourcesofTaya
ba sBaya r
ede me rs
alandpelagi
cfishes,ma ngr
ov e
,
coral
s,s
e agrasses,
andma ri
nei nv
ertebrates.
Ho wever
,i
nre c
entyear
s,thereare
repor
tsofde cl
ini
ngc atc
hf r
om t hefis herme nwhichwa sa g
grava
tedb yt he
conver
sionofma ngroveareasintofishpond,a ndindic
aonsofc or
alre e
fand
habit
atde gr
adaon a rea pparentins omea reaswhe r
ei ll
egalfishingand
destr
ucv efishingme t
hodsa r
er ampa nt
. Thesit
uaoni sfurt
heraggravat
edb y
si
ltaonf rome rodedmoun tai
nsduringra i
nyseasons(
ResourcesCombines,I
nc .
andM. S.
E .Univ
. F
oundaon, Inc.,1996)
.

2
 I
ntr
oduc
tion

I
n2 00 3,unde rtheNaona lStockAs ses
sme ntP rogram( NS AP)oft heP hili
ppine
Burea uof Fi
sheriesandAquacRe sources(BF AR), astockas s
essme n tstudywa s
conduc t
edt ode t
e r
minea ndme asurethee ffe ctsofi nterven onsi ntr
oduc ed
andi mple me nt
edi nTa yabasBa ya nditss urroundi ngora dj
ace ntpr ovincesof
Que zona ndMa rinduque( Ra mose tal.
,201 1).Thi sAtlasi
soneof thepi one er
ing
effortsoft heNS APina na e mp tt odoc ume nta l
lma r
inefis hs pe ci
esc aught
withi
nT ayabasBa ya nd to pro vi
depe rne nti nf
ormaon t o he lp de v
elop
effe cv efis heri
e sresourc ema nag ements cheme sa ndpol i
cies.Itpr o vi
desa
snapshotof thefis heri
e sresourcesof t
heBa ytha tca nbeus eda sbe nchma r
kfor
fut
urewor k.Mor eover, t
hisfisha tlasofTa yaba sBa ywi l
l a
lsoc ontributet othe
Naona lP l
anof Aconf ortheCor al T
ria
ng leIniav eof theP hil
ippinesa sitwil
l
provideba seli
nei nf
ormaont oa ssi
stint heE c osystem Appr oachi nF isheri
es
Ma na ge
me ntpoli
cyproc esses(Republicoft heP hi l
ippines,2009 )
.

3
Me
thodol
ogy

F
rom 2003t
o2007,t
he BF
AR-
NSAP s
tudy i
den fie
d488 fis
hspe
cie
s
under98famil
i
es.Si
xty
-ei
ghtoft hefishesa
redemersa
l,57pelag
icand3 32
reef
-as
soc
iat
edwith31inv
e r
tebra
tes.T
hisAt
lasr
ecor
ds182speci
esbelong
ing
to93gener
aand43famil
iesasthemostcommonfishesi
nTayaba
sBa y
.

Thelis
toffis hs peci
esinthi
sAtlaswa scompil
edfrom datagat
he r
edb ythe
enume r
atorsf r
om t he t
hreela nding center
si n Dal
ahic
an,L ucena City,
Ma t
andangS abangandMa tandangK a
nlura
ninCa t
anauan,Quezon.F oreach
s
pe ci
es,theAt l
a spres
entsaphot o ,thes ci
en ficname ,Engl
ishandl ocal
c
ommonna mes,asy
mp t
o clengt
h( L∞)orthem ax
imum l
ength(L
ma)
x,and l
en g
th
atfirs
tma turi
ty(Lm)
.

L∞, L
max,andL mvaluesinF i
shBa s
e( h p: /
/www. fishba s
e .org,Froesea ndP auly,
201 3)arecompi l
edf r
ompubl i
caonsa ndr epor t
s.Itisalsopos sibl
et oesma te
theL m
ofspeciesusingL∞
orL ma
x
asinputi nt heL i
feHi s t
oryT oolinF i
shBas e.
Whe repos si
ble,theme di
a nL ∞
v
a l
u esfrom studies c
o nd uct
e d i
n thevic
init
y of
Tayaba sBayora reaswit
hint heP hi
lippineswe reselected. Forspe ci
eswhe reno
l
oc alL∞
val
ues ar
e avai
labl
e, them e dianof the val
u esco m pil
ed inFishBas
e for
thespe ci
esar epresent
ed. T
he sev aluesarema rk
edwi t
ha nasteris
k( *
).TheL ma
x
valuesa r
ethema xi
mumr e cordedf orthes pecies.TheL m
val
ue swe reesma ted
usingt heFishBaseL i
feHistoryT oolfrom L ∞
orfrom L max
(
Froesee ta l
.
,2 00 0,
Froesea ndBinohlan,2000).

Thephot osus edinthisAt lasca mefrom BF ARpubl i

caons ,phot ocol
l
e consof
BFARs taff,andF i
shBa s
ec oll
a borators
. Atotalof182s pe ci
esfrom4 3famil
iesare
i
nc ludedint hi
sAt las.Informaona boutt hesespe ciesarea lsopresente
di n
tabularforma sanAnne x.T herea r
ea ccountsofa nothers ixspec
iesint he
catchesfrom Tayaba sBa y,butt hesec ouldnotbev e ri
fieda tt hi
s me .Thes i
x
spe ci
esare:C all
echelysma rmo rata(Bleek er
,1 854);De capt
e rusakaadsiAbe ,
195 8;Nuche quul
ab lochii(Va l
encienne s
,1 835);Si
ll
a goin ge
n uu aMcKay,19 85;
Thy rs
it
oidesma r
leyiF owler,1 929;a ndUp eneusja ponicus( Hou uyn,1 782).
Althought heses peciesoc curi nc oun t
riesa dja
cen ttot heP hil
i
ppi
ne s
,t hei
r
occur r
enceinTa y
a basBa ysl lneedstobec onfirme d.

4
 Pr
ospec
ts

T
hi
sAt
lasc
ont
ainsba
sici
nfor
maona
boutt
hemos
tcommonfis
hspe
cie
s
caughtinTa yabasBa y
,Que zonP rovi
nce.Itc anbeu l izedb yBFARa ndothe r
agencie
s,communie sandindiv
idua l
sengage dinpoli
c y
,conservaon, r
e s
earch,
l
iveli
hoods,a ndot heri niav esa nchoredonfis heryr esourc
es.Itoffe r
sa
ref
e r
enceda ndv ali
da t
edfis hs peciesiden fic aong uide.Thea vai
labi
li
tyof
asymp t
o cle ngth( L∞)
,ma xi
mum l ength(Lma)
x,andl engthatfirstma t
urit
y(Lm),
forexample, wil
lena blefisheri
esma nager
st ode v
elops cie
nc e-
basedme shsi
z e
regul
aonswi t
hc on fidence.Dataf r
om thi
sAt lascana l
sobeus edtopr oducea
vari
etyofI
n formaon, Educaona ndCommuni caon( I
EC)ma ter
ial
ssuchasthe
fishruler
sa ndma turit
ypos t
ers(INCOFISHe ta l
.
,200 8).

ThisAtl
aspresen t
sapr ot
otypefiel
dg uideoffishesf oraspe ci
fica r
e a.
Itishoped
thatsimil
ariniav esforotherareasintheP hil
ippineswillde v
elopt hei
ro wn
fishiden ficaong ui
deatlas
esunde rtheNSAP.Re adersarea l
soe ncouragedto
produceothe rIECma ter
ial
susingtheda tacontainedint hisAtlas.Wes tar
t e
d
withones ma llbayinthecountry
,butwee nvis
iont hatthis“AtlasofCommon
FishesofTay
a basBa y
,Que z
onP r
ovince,Phil
i
ppine s”wil
lpro v
idethei mpe t
usto
l
a unchsimil
ariniav esforeachpro v
inceswithc oast
alfis heryresourcesfrom
Ba t
anestoTawi -t
awi.

5
Howt
oUs
e

HOW TO USETHI
SATLAS

S
cie
n ficNa
me At
ulemat
e
Aut
hora
ndY
ear (
Cuv
ier
,1833)
CommonNames Y
ell
owt
ail
sca
d
(
Engl
is
handl
ocal
) K
alapat
o
As
ympt
o cl
eng
th L
∞:30.
5cmT
L
L
eng
tha
tfir
stma
tur
it
y L
m:18.
0cmT
L
Rodol
foB.Re
yesJ
r.

Thefi shesi nt hi
sAtlasarelist
edi nalphab ec alorderb yfamil
ya ndb y
s
p eci
esn ame .Thel engt
h sp r
e s
enteda ret o t
alleng t
hs( TL)
,wh ichis
me as
u r
e df romthe po fthes nouttot he po fth ecaudalfi n.Forthe
l
o nesn gray,
the“leng
th”givenisdis
kwi dth(WD) ,t
heme asur
eme ntacr
oss
t
h e widtho ftheb ody,b et
we ent he p so fthep e c
toralfins.Wh ere
possi
ble,t heme dianL∞va l
uesf r
oms tu di
esc onductedint hev i
cini
tyof
TayabasB ayo rar
easwithi
nt hePhil
ippi
n ewe reselec
ted .F
orspecieswhere
nol ocalL ∞ val
uesa r
ea vai
labl
e,t heme d i
ano fthev al
u e
sc omp i
ledin
Fi
shBasef o rthespeci
esa represented.The s
ev aluesa r
ema r
kedwi t
ha n
ast
e r
is
k( * )
.T helengt
hsa tfirstma t
urity(Lm)we reesma tedfrome it
her
L∞ orL xu
ma s
ingtheF i
shBaseL i
feHi s
toryT ool.No t
et hatt heL∞ isnot
avai
labl
ef ora l
lspec
ies,
wh i
chind i
cat
esag apino urkno wledge.

As
ymp to cl
ength(L
∞)i
sthel
engththatfis
hesofapopul
aonwoul
dre
ach
i
ftheywereal
lowedtog
rowindefini
tel
y.

Maxi
mu mlengt
h(L
ma)
x i
sthel
onge
sti
ndi
vi
dualr
ecordedforast
ock.T
h eL
max
i
ndi
cat
edinthi
sAtl
as,isthes
iz
eofthel
onge
stindi
vi
du a
le v
errec
ordedfor
t
hespeci
es.

Lengthatfir
stma tur
it
y(Lm)isthea
veragel
e ng
tht hatfishesofap opul
ao n
matureforthefi r
st me .Thisisdet
erminedb ye xamini
ngt hegonadsof
fis
h esofdi
fferentsi
zesandn on gt
helengthatwh i
chhalfofthespec
imens
ex
ami nedcar
ryma tur
eeg g
so rmil
tanda r
eread ytos pa
wn .Lmcanalsobe
esma t
edfr
oma nemp i
ri
calrel
ao ns
hipbetweenl engt
ha tfirs
tmaturi
tyand
as
ymp to cleng
th( L
∞)o rma xi
muml ength(Lma)u
x si
ngt heF i
shBas
eL i
fe
Hi
storyTool
(Fr
o es
ea ndBinohl
an,2000)
.

6
7
FI
SH ATLAS

ACANTHURI
DAE

Ac
ant
hur
usl
ine
atus
(
Linna
eus,1758)
L
inedsur
ge onfis
h
Mangadl
it

J
ohnE.Randal
l

Ac
ant
hur
usni
gri
cauda
Duncker&Mohr,1929
Epaule es
urg
eonfish
Labahi
ta

J
effr
eyT
.Wi
l
li
ams

Nas
ouni
cor
nis
(Fors
skål
, 1775)
Bluespi
neuni cornfis
h
Suraan,Labahita,
Tidl
uan,Tur uan

K
wang-
TsaoS
hao‡

8
 FI
SH ATLAS

BELONI
DAE

Abl
enne
shi
ans
(
Val
enciennes
, 1846)
F
latneedlefish
Bat
alay,Kambabalo,
Haba,Kambilawan K
wang-
TsaoS
hao‡

Tyl
osur
usme
lanot
us
(Bl
eek
er,1850)
Ke
el-
jawedneedl
efis h
Haba
K
wang-
TsaoS
hao‡

CAESI
ONI
DAE

Cae
sioc
aer
ulaur
ea
Lacepède,1801
Blueandg ol
dfusil
ie
r
Dalagangbuk i
d(Bi
log),
Burgi
s

K
wang-
TsaoS
hao‡

Cae
sioc
uni
ng
(Bl
och,1791)
Redbell
yyel
lowtai
lfusi
l
ier
Dalagangbuki
d( L
apad),
Burgis

K
wang-
TsaoS
hao‡

9
FI
SH ATLAS

CARANGI
DAE

Al
ect
isi
ndi
ca
(
Rüppell,
1830)
I
ndia
nt hrea
dfish
T
alaki
tok,Sebo
,Mus
lo,
S
alamin

K
wang-
TsaoS
hao‡

Al
epe
sdj
edaba
(
Forss
kål
,1775)
S
hrimpscad
S
alay-
sal
ay,Sal
ayas
o

K
wang-
TsaoS
hao‡

Al
epe
svar
i
(
Cuvier
,1833)
Herr
ingscad
Sal
ay-s
alay
,Sal
ay-
sal
aybat
ang

K
eii
chi
Mat
suur
a‡

10
 FI
SH ATLAS

CARANGI
DAE

At
ropusat
ropos
(
Bloch&S chnei
der,
1801)
Cle bell
ytr
eval
ly
T
alak
itok,S
ebo,Muslo

K
wang-
TsaoS
hao‡

At
ulemat
e
(Cuvi
er,
1833)
Y
e l
l
owtail
scad
Kal
apato

Rodol
foB.Rey
esJ
r.

Car
angoi
desf
erdau
(F
or s
skå
l,1775)
Bl
uetreval
ly
T
alak
itok,Sebo,Muslo

K
wang-
TsaoS
hao‡

11
FI
SH ATLAS

CARANGI
DAE

Car
ang
oide
she
dlande
nsi
s
(Whit
ley
,1934)
Bumpnosetrev
all
y
Tal
aki
tok,S
ebo

K
wang-
TsaoS
hao‡

Car
anxt
ill
e
Cuvie
r,1833
Til
l
etreval
ly
Tal
akit
ok,Sebo

J
ohnE
.Randal
l

De
capt
erusmac
ros
oma
Bleeker
,1851
Shori nscad
Galunggong,Ti
nabak
o

K
wang-
TsaoS
hao‡

12
 FI
SH ATLAS

CARANGI
DAE

De
capt
erusr
uss
ell
i
(
Rüppel
l,
1830)
I
ndi
anscad
Gal
unggong

K
wang-
TsaoS
hao‡

Me
gal
aspi
scor
dyl
a
(
Li
nna eus,
1758)
T
orpedoscad
Or
il
es,
Pak-an,Malaguno

K
wang-
TsaoS
hao‡

Sc
ombe
roi
dest
ala
(
Cuvi
er
,1832)
Barr
edqueenfish
T
alang-
tal
ang,Lapi
s

Gr
egor
ioV
. He
rmos
aJr
.

Sc
ombe
roi
dest
ol
(Cuv
ier
,1832)
Ne
edl
esc
aledqueenfish
Lapi
s

K
wang-
TsaoS
hao‡

13
FI
SH ATLAS

CARANGI
DAE

Se
larboops
(
Cuvier
,1833)
Oxe
y esc
ad
Matambaka,Bur
aw

Rodol
foB.
Rey
esJ
r.

Se
larc
rume
nopht
hal
mus
(Bl
och,
1793)
Big
eyesca
d
Matambaka,
Bur
aw

K
wang-
TsaoS
hao‡

Se
lar
oide
sle
ptol
epi
s
(
Cuvie
r,1833)
Ye
ll
owstri
pescad
Gi
nto-
ginto

J
ohnE
.Randal
l

14
 FI
SH ATLAS

CHI
ROCENTRI
DAE

Chi
roc
ent
rusdor
ab
(F
ors
skå
l,1775)
Dor
abwolf
-her
ring
Bali
la

K
wang-
TsaoS
hao‡

CLUPEI
DAE

Herkl
ots
ichthys
quadri
maculatus
(Rüppel
l
,1837)
Bl
ue s
tri
peherr
ing
T
amban,
Tapulok,
Tunsoy

J
ohnE.Randal
l

Sar
dine
llafimbr
iat
a
(Val
enci
enne
s,1847)
F
r i
nge
scal
esar
dinel
la
Tamban

K
wang-
TsaoS
hao‡

15
FI
SH ATLAS

CLUPEI
DAE

Sar
dine
llagi
bbos
a
(
Bleeker
,1849)
Goldst
ri
pesardi
nel
l
a
Tunsoy

De
mianWi
l
lee

Sar
dine
llal
emur
u
Ble
eker
,1853
Bal
isar
dinel
la
Tamban,Tuns
oy

Rodol
foB.
Rey
esJ
r.

Spr
ate
lloi
desde
lic
atul
us
(Be
nne ,1832)
Del
ica
ter
oundhe r
ri
ng
Bal
akwas
, Di
li
s

Ri
char
dWi
nte
rbo om

Spr
ate
lloi
desgr
aci
li
s
(T
emmi nc
k&S c
hle
gel
,1846)
Si
lve
r-s
tri
per
oundher
ring
Bal
akwas,Di
l
is

Ri
char
dWi
nte
rbo om

16
 FI
SH ATLAS

CORYPHAENI
DAE

Cor
yphae
nahi
ppur
us
Linnaeus
,1758
Commondol phi
nfish
Dor
ado ,
Lamarang

Ri
char
dWi
nte
rbo om

DASYATI
DAE
Ne
otr
ygonkuhl
ii
(
Müll
er&Henle
,1841)
Bl
ue-
spo e
dsng ray
Pagi

K
wang-
TsaoS
hao‡

ELOPI
DAE
El
opshawai
ens
is
Rega
n, 1909
Ha
waii
anladyfish
Bid-bi
d

J
effr
eyT
.Wi
l
li
ams

17
FI
SH ATLAS

ENGRAULI
DAE

Enc
ras
ichol
inapunc
tif
er
Fowler
,1938
Bucca
ne e
ranchov
y
Dil
i
s,Boli
naw

Gr
egor
ioV
.He
rmos
aJr
.

Stol
ephorus
commersonnii
Lace
pède,1803
Comme rs
on'sa
nchov
y
Tuwakang

J
ohnE
.Randal
l

St
ole
phor
usi
ndi
cus
(
vanHass
elt
,1823)
I
ndi
ananchovy
T
uwakang

T
homasGl
oer
fel
t-
Tar
p

EPHI
PPI
DAE

Pl
ataxboe
rsi
i
Bl
eeke
r,1853
Gol
denspadefis
h
Bay
ang

J
ohnE
.Randal
l

18
 FI
SH ATLAS

GEMPYLI
DAE

Ruve uspr
eti
osus
Cocc
o,1833
Oilfish
Mal
ai
gi
t,Ni
ng-ning

J
ohnE
.Randal
l

GERREI
DAE
Ge
rre
ser
ythr
our
us
(
Bloch,
1791)
De
ep-bodi
e dmojar
ra
Manabon,Yamas

J
ohnE
.Randal
l

Ge
rre
sfil
ame
ntos
us
Cuvi
er
,1829
Whi
pfinsil
ver
-bi
ddy
Manabon,Yamas

J
ohnE
.Randal
l

19
FI
SH ATLAS

GERREI
DAE

Ge
rre
soye
na
(F
ors
skål
,1775)
Commons i
lve
r-bi
ddy
Manabon,Yamas

K
wang-
TsaoS
hao‡

HAEMULI
DAE

Plec
torhi
nchus
chaet
odonoide
s
La
cepède,1801
Harl
equi
ns wee
tli
ps
Al
atan,L
abian

J
ohnE.Randal
l

Ple
ctor
hinchus
chr
ysotae
nia
(Ble
e k
er,
1855)
Yell
ow-st
ri
pedswe
etl
i
ps
Alatan

J
ohnE.Randal
l

20
 FI
SH ATLAS

HAEMULI
DAE

Pl
ect
orhinc
hus
diagr
ammus
(Li
nnaeus,
1758)
S
tri
peds weet
li
ps
Al
atan,Hundon

J
ohnE.Randal
l

Pl
ect
orhi
nchusgi
bbos
us
(La
c epède,1802)
Harryhotl
ips
Al
atan,Puyongdagat

J
ohnE.Randal
l

Pl
ect
orhi
nchusl
ine
atus
(
Linnaeus,
1758)
Y
ell
owbandeds weet
li
ps
Alatan,Hundon

K
wang-
TsaoS
hao‡

21
FI
SH ATLAS

HAEMULI
DAE

Pl
ect
orhi
nchuspi
ctus
(T
ortones
e,1936)
Tr
outs weet
li
ps
Alat
an

J
ohnE.Randal
l

Ple
ctor
hinchus
pol
ytaeni
a
(Bleek
er,
1853)
Ribbonedswee
tli
ps
Alatan

J
ohnE.Randal
l

HEMI
RAMPHI
DAE
He
mir
amphusf
ar
(F
ors
skål
,1775)
Bl
ack-
bar
redhalea
k
Bugi
w

K
wang-
TsaoS
hao‡

Hypor
hamphusquoyi
(
Vale
ncie
nnes,1847)
Quoy'
sgarfis
h
Si
l
iw

K
eii
chi
Mat
suur
a‡

22
 FI
SH ATLAS

HOLOCENTRI
DAE

Myr
ipr
ist
ishe
xagona
(
Lacepè
de,1802)
Doubl
etoot
hs ol
di
e r
fish
T
angi
slawi
n,Si
gangbatuhan

J
ohnE.Randal
l

I
STI
OPHORI
DAE
I
sti
ophoruspl
atypterus
(
Shaw,1792)
Indo-
Paci
ficsai
lfish
Malasugi

K
wang-
TsaoS
hao‡

LABRI
DAE
Che
ili
nusf
asc
iat
us
(Bl
och,1791)
Re
dbr
ea s
tedwrasse
Dulasan

J
ohnE.Randal
l

23
FI
SH ATLAS

LABRI
DAE

Che
ili
nust
ril
obat
us
Lacepède,1801
Tr
ipletai
lwrass
e
Dulasan,Mame ng

K
wang-
TsaoS
hao‡

Oxyc
hei
li
nusdi
gramma
(L
ac e
pè de
,1801)
Cheekli
nedwr a
sse
Dulasan,I
sdangbat
o

K
wang-
TsaoS
hao‡

LEI
OGNATHI
DAE

Aur
ige
quul
afas
ciat
a
(
Lac
epè de,
1803)
S
tri
pedponyfish
S
ap-sap,Wali
ng

Gr
egor
ioV
.He
rmos
aJr
.

24
 FI
SH ATLAS

LEI
OGNATHI
DAE

Equul
ite
sle
uci
scus
(
Günther
,1860)
Whipfinponyfish
Sap-sap,
Tabil
os

ThomasGl
oer
fel
t-
Tar
p

Eubl
eeke
rias
ple
nde
ns
(
Cuvi
er
,1829)
Spl
endi
dponyfish
S
aps
ap,Laway
,Kurag

J
ohnE.Randal
l

Gazzaac
hlamys
Jor
dan&S tar
ks,1917
S
ma l
l
toothedponyfish
Sap-s
ap,Wal i
ng

J
ohnE.Randal
l

25
FI
SH ATLAS

LEI
OGNATHI
DAE

Gazzami
nut
a
(
Bloch,1795)
T
oothpony
Sap-
sap,Tambokne
neng,
Wali
ng-waling

K
wang-
TsaoS
hao‡

Le i
og nathusequul
us
(Fors
sk ål
,1775)
Commonpony fis
h
Sap-sap,Laway

K
wang-
TsaoS
hao‡

Se
cut
ori
nsi
diat
or
(Bl
och,1787)
Pugnoseponyfis
h
Sap-s
ap,Dyako

K
wang-
TsaoS
hao‡

26
 FI
SH ATLAS

LEI
OGNATHI
DAE

Se
cut
orr
uconi
us
(
Ha mi
lt
on,1822)
De
eppugnoseponyfish
Sap-
sap,Dyako

K
wang-
TsaoS
hao‡

LETHRI
NIDAE

Le
thr
inusat
kins
oni
Seale,1910
P
aci
ficy
ell
owt
ail
e mpe r
or
Kanuping

Rodol
foB.Rey
esJ
r.

Le
thr
inushar
ak
(
For
s s
kå l
,1775)
T
humbpr
inte mperor
Kanuping

K
wang-
TsaoS
hao‡

27
FI
SH ATLAS

LETHRI
NIDAE

Le
thr
inusl
ent
jan
(L
acepède
,1802)
Pi
nke a
remperor
Kanupi
ng

J
ohnE.Randal
l

Le
thr
inusmi
crodon
Val
enciennes,
1830
Small
toothemperor
Kanupi
ng, L
ugso

K
wang-
TsaoS
hao‡

Le
thr
inusmi
niat
us
(
Forste
r,1801)
T
r umpetemperor
Kanupi
ng,Lugso,
Manutsot

J
ohnE.Randal
l

28
 FI
SH ATLAS

LETHRI
NIDAE

Le
thr
inusne
bul
osus
(
For
ssk
å l
,1775)
S
pangl
ede mperor
Pu ngkanuping

K
wang-
TsaoS
hao‡

Le
thr
inuss
emi
cinc
tus
Vale
nc i
ennes
,1830
Bl
ackblot
chempe r
or
Kanuping,
Amus i
n

J
ohnE.Randal
l

Le
thr
inusvar
iegat
us
Va
lenci
ennes,1830
Slenderempe r
or
K
anuping

J
ohnE.Randal
l

29
FI
SH ATLAS

LETHRI
NIDAE

Monot
axi
sgr
andoc
uli
s
(
Forssk
ål,
1775)
Humpnosebig-
eyebr
eam
Kanupi
ng

J
ohnE.Randal
l

LUTJ
ANI
DAE

Lut
janusar
gent
imac
ulat
us
(
Fors
skål
,1775)
Mangrov
eredsnappe
r
Mangagat,
Pargo

J
ohnE.Randal
l

Lut
janusde
cus
sat
us
(Cuv
ier,1828)
Checkeredsnapper
Madrigal,Maya-maya

Rodol
foB.Rey
esJ
r.

30
 FI
SH ATLAS

LUTJ
ANI
DAE

Lut
janusf
ulvi
flamma
(
Fors
skå
l,1775)
Dorysnapper
T
ali
ngan,Pi
ngaw

K
wang-
TsaoS
hao‡

Lut
janusj
ohni
i
(
Bloch,1792)
J
ohn'ssnapper
Tal
ingan

J
ohnE.Randal
l

Lut
januskas
mir
a
(F
orss
kål,
1775)
Commonbl
uest
ri
pesnapper
Dayangdang

J
ohnE.Randal
l

31
FI
SH ATLAS

LUTJ
ANI
DAE

Lut
janusl
utj
anus
Bl
och,1790
Bi
geyesnappe
r
Maya-maya,Par
go,
Bur
ara

J
ohnE.Randal
l

Lut
janusmal
abar
icus
(Bl
och&Sc
hnei
der
,1801)
Malaba
rbl
oods
napper
Pul
ahan

J
ohnE.Randal
l

Lut
janusmonos
tigma
(
Cuvi
er,1828)
One
-spotsnapper
Maya-
may a,Tal
i
ngan

Gr
egor
ioV
. He
rmos
aJr
.

32
 FI
SH ATLAS

LUTJ
ANI
DAE

Lut
janusqui
nque
line
atus
(Bloc
h,1790)
F
ive
-li
nedsnapper
Tal
ingan

J
ohnE.Randal
l

Lut
janusr
uss
ell
ii
(
Bleeker
,1849)
Russel
l'
ssnapper
Tal
ingan,Pi
ngaw

K
wang-
TsaoS
hao‡

MEGALOPI
DAE

Me
gal
opsc
ypr
inoi
des
(
Brouss
onet,1782)
I
ndo-Pac
ifictar
pon
Buwan-buwan

K
wang-
TsaoS
hao‡

33
FI
SH ATLAS

MENI
DAE

Me
nemac
ulat
a
(Bl
och&S c
hne
ider
,1801)
Moonfish
Chabi
ta,
Hiwas
,Pater
os

J
ohnE.Randal
l

MUGI
LIDAE

Che
lonmac
rol
epi
s
(S
mith,1846)
Lar
gesc
a l
emulle
t
Banak,
Aguas,Ali
gas
in

J
ohnE.Randal
l

Cr
eni
mugi
lcr
eni
labi
s
(F
orss
kål,
1775)
Fri
ngel
i
pmul l
et
Anawan,Banak,
Aguas

J
ohnE.Randal
l

34
 FI
SH ATLAS

MUGI
LIDAE

El
loc
hel
onvai
gie
nsi
s
(
Quoy&Ga i
mard,1825)
Squaret
ail
mul l
et
Banaknagapang

J
ohnE.Randal
l

Mool
gar
das
ehe
li
(F
ors
s kå
l,1775)
Bl
uespotmul l
et
Banak
, Al
imugok

J
ohnE.Randal
l

Mugi
lce
phal
us
L
innaeus,
1758
F
lat
headg r
eymull
et
Aguas,
Banak

J
ohnE.Randal
l

Val
amugi
lbuc
hanani
(
Ble
eker,
1853)
Bl
ueta
ilmull
et
Banak
,Aguas,Al
igas
in

J
effr
eyT
.Wi
l
li
ams

35
FI
SH ATLAS

MULLI
DAE

Par
upe
neusbar
ber
inoi
des
(Bl
eeker
,1852)
Bic
olorgoa i
sh
Yanggut
an,Saramul
yet
e,
Manis

J
ohnE.Randal
l

Par
upe
neusbar
ber
inus
(
L ac
epè de,1801)
Dash-and-dotgoa i
sh
Yanggutan,Saramul
yet
e,
Manis

J
ohnE.Randal
l

Par
upe
neusc
ycl
ost
omus
(
L a
c e
pède,1801)
Gold-s
addlegoa i
sh
Saramuly
ete

J
ohnE.Randal
l

Par
upe
neusi
ndi
cus
(
Shaw,1803)
I
ndi
ang oa i
sh
S
almonete,Manis,
Y
anggutan,Sar
amul
yet
e

J
ohnE.Randal
l

36
 FI
SH ATLAS

MULLI
DAE

Par
upe
neusmul
tif
asc
iat
us
(
Quoy&Gai
mard,1825)
Many
bargoa ish
Sar
amulyete

Rodol
foB.Rey
esJ
r.

Upe
neusmol
ucc
ens
is
(Bl
eeker
,1855)
Gol
dbandgoa ish
S
agi
ng-
sagi
ng,Sar
amulyete

J
ohnE.Randal
l

Upe
neuss
ulphur
eus
Cuvi
er
,1829
Sul
phurgoa i
sh
S
almonete
,Manis,
Sar
amuly
ete

J
ohnE.Randal
l

Upe
neusvi at
us
(
For
s s
kål,1775)
Y
ell
ows
tri
pedg oa ish
Y
apot
,Manis,Saramulyete

K
wang-
TsaoS
hao‡

37
FI
SH ATLAS

MYCTOPHI
DAE

Myc
tophum ni
tidul
um
Garman,1899
Pear
lyl
anternfis
h
Se
rom-serom

K
eii
chi
Mat
suur
a‡

NEMI
PTERI
DAE

Ne
mipt
erusbat
hybi
us
Snyder
,1911
Yel
lowbel
lyt
hre
adfinbr
eam
Bis
ugo

K
wang-
TsaoS
hao‡

Ne
mipt
erushe
xodon
(Quoy&Gaimar
d,1824)
Ornat
ethr
eadfinbream
Bis
ugo

Kat
hle
enKes
ner
-Rey
es

38
 FI
SH ATLAS

NEMI
PTERI
DAE

Ne
mipt
erusne
mur
us
(
Ble
e ke
r,1857)
Re
dspi
net
hreadfinbre a
m
Bisugo

J
ohnE.Randal
l

Sc
olops
isaffini
s
Pete
rs,1877
Pet
ers'
monoc l
ebre a
m
Gut
o-guto
,Ipot,Bi
sugo

J
ohnE.Randal
l

Sc
olops
ismar
gar
iti
fer
a
(Cuvier
,1830)
P
ear
lymonoc l
ebream
Gut
o-gut
o,I
pot
,Bisugo,Ti
ngin

J
ohnE.Randal
l

Sc
olops
ist
aeni
opt
era
(Cuv
ier
,1830)
Lac emonocl
ebre a
m
Gut
o-gut
o,I
pot,
Bisugo

J
ohnE.Randal
l

39
FI
SH ATLAS

OPHI
CHTHI
DAE

Myr
icht
hysc
olubr
inus
(
Bodda e
rt,
1781)
Har
lequinsna
keeel
I
gat,
P al
os,Ubod

J
ohnE.Randal
l

Sc
ole
cenc
hel
ysgymnot
a
(
Blee
ke r
,1857)
I
ndo-
Pa c
ificsl
ende
rwor
m-e
el
I
gat,Pal
os,Ubod

J
effr
eyT
.Wi
l
li
ams

PARALI
CHTHYI
DAE
Ps
eudor
hombusar
sius
(
Hamil
ton,1822)
L
arget
oothflounder
P
alad,T
ampal,Dapa

K
wang-
TsaoS
hao‡

40
 FI
SH ATLAS

PARALI
CHTHYI
DAE

Ps
eudorhombus
ci
nnamoneus
(
Temmi
nck&Schle
gel
,1846)
Ci
nnamonflounder
Pal
ad,
Tampal,Dapa

K
wang-
TsaoS
hao‡

PLATYCEPHALI
DAE

Coc
iel
lac
roc
odi
lus
(Cuvi
er
,1829)
Cr
ocodil
eflathead
L
ubalob

J
ohnE.Randal
l

Sunagoc
iaar
eni
col
a
(
Schul
tz
,1966)
Broa
dheadflathead
L
ubal
ob,Is
dangtuko

K
wang-
TsaoS
hao‡

PLOTOSI
DAE
Pl
otos
usl
ine
atus
(Thunbe
rg,1787)
St
ripedeel
cai sh
L
it
o,I
to-
i
to,
Patuna,Su
mb i
lang

K
wang-
TsaoS
hao‡

41
FI
SH ATLAS

PRI
ACANTHI
DAE

Hete
ropri
acant
hus
cr
uentat
us
(
Lace
pè de
,1801)
Gl
ass
e y
e
Si
ga,Di
lat

J
ohnE.Randal
l

Pr
iac
ant
hushamr
ur
(
Fors
skål,
1775)
Moonta
ilbul
l
sey
e
Si
ga,Di
lat

J
ohnE.Randal
l

Pr
iacant
hus
macr
acanthus
Cuvi
er,
1829
Redbig
eye
Si
ga,Dil
at

Gr
egor
ioV
.He
rmos
aJr
.

42
 FI
SH ATLAS

PRI
ACANTHI
DAE

Pr
iac
ant
hust
aye
nus
Ric
hardson,
1846
Pur
ple
-spo edbige
y e
S
iga,Di
lat

J
ohnE.Randal
l

PSETTODI
DAE
Ps
e ode
ser
ume
i
(Bl
och&S chnei
de r
,1801)
India
nha l
ibut
P
alad,Tampal,I
sdangdapa

K
wang-
TsaoS
hao‡

SCARI
DAE
Ce
tos
car
usbi
col
or
(Rüppel
l,1829)
Bi
colourpa
rroi sh
Bu
n-a
k,L
amon-l
a mon,
Mu lmol

J
ohnE.Randal
l

43
FI
SH ATLAS

SCARI
DAE

Chl
orur
usbowe
rsi
(S
nyder
,1909)
Bower'
sparr
oi sh
Bun-
ak,L
amo n-
lamon,
Mul
mol

J
ohnE.Randal
l

Chl
orur
usmi
cror
hinos
(Bl
eeker
,1854)
Ste
epheadparr
ots
Bun-ak

J
ohnE.Randal
l

Chl
orur
uss
ordi
dus
(F
orssk
ål,
1775)
Dais
yparroish
Bun-ak

J
ohnE.Randal
l

Sc
arusf
est
ivus
Val
enci
ennes
,1840
Fesveparr
oi s
h
Bun-
ak

K
wang-
TsaoS
hao‡

44
 FI
SH ATLAS

SCARI
DAE

Sc
arusf
renat
us
Lac
epède,1802
Br
idl
edpar
roi sh
Bun-ak

K
wang-
TsaoS
hao‡

Sc
arusghobban
For
ssk
ål,1775
Bl
ue-bar
redparr
oi sh
Isdangbat
o,Bun-ak

K
wang-
TsaoS
hao‡

Sc
arushyps
elopt
erus
Ble
e k
er,1853
Y
ell
ow-
tai
lparr
oi sh
Bun-ak

J
ohnE.Randal
l

Sc
arusni
ger
F
orsskå
l,1775
Duskyparr
oi sh
Is
dangbato

J
ohnE.Randal
l

45
FI
SH ATLAS

SCARI
DAE

Sc
arusps
i ac
us
Fors
s k
ål,
1775
Commonpa rr
ois
h
Bun-ak

K
wang-
TsaoS
hao‡

Sc
arusr
ivul
atus
Val
enci
ennes,
1840
Ri
vul
ate
dpa r
roish
I
sdangbato
,Bun-ak

K
wang-
TsaoS
hao‡

SCOMBRI
DAE

Auxi
sroc
hei
(Ri
sso,1810)
Bul
lettuna
Tul
ingan

K
wang-
TsaoS
hao‡

Auxi
sthazar
d
(
Lacepè
de ,1800)
F
rig
atetuna
T
uli
ngan,Tuli
ngangas
o

K
wang-
TsaoS
hao‡

46
 FI
SH ATLAS

SCOMBRI
DAE

Gymnos
ardauni
col
or
(Rüppell,
1836)
Dog toothtuna
T
ambak
ol,
T una,Tal
iyas
in

K
wang-
TsaoS
hao‡

Kat
suwonuspe
lami
s
(
Li
nnaeus,1758)
Sk
ipja
ckt una
Gulyasan

K
wang-
TsaoS
hao‡

Ras
tre
lli
gerbr
achys
oma
(
Ble
eker
,1851)
S
hortmacker
el
Has
a-hasa

J
effr
eyT
.Wi
l
li
ams

Ras
tre
lli
gerf
aughni
Mat
sui
,1967
I
sl
andma c
kerel
Al
umahan,
Hil
ats
ay(j
uveni
l
e)

Rodol
foB.
Rey
esJ
r.

47
FI
SH ATLAS

SCOMBRI
DAE

Ras
tre
lli
gerkanagur
ta
(Cuvi
er
,1816)
I
ndianma c
kerel
Alumaha
n,Hil
at
say(
juv
eni
l
e),
Hasa-
hasa

K
wang-
TsaoS
hao‡

Sar
daor
ient
ali
s
(
Temmi nc
k&S chl
egel
,1844)
S
tri
pedbonito
S
enori
ta,Tul
i
nganaso

K
wang-
TsaoS
hao‡

Sc
ombe
romor
usc
omme
rson
(
L ac
epède,1800)
Narro
w-ba
rredSpani
shma
cke
rel
Tangi
gue

K
wang-
TsaoS
hao‡

Sc
ombe
romor
usgu at
us
(
Bloc
h&S c
hnei
der
,1801)
I
ndo-Pac
ificki
ngmacker
el
T
angigue

J
ohnE.Randal
l

Thunnust
onggol
(
Ble
eker,
1851)
L
ongta
ilt
una
T
ambakol,T
una

K
wang-
TsaoS
hao‡

48
 FI
SH ATLAS

SERRANI
DAE

Ae
thal
ope
rcar
ogaa
(
For
sskå
l,1775)
Redmouthgrouper
L
apu-l
apu,Sigapo

J
ohnE.Randal
l

Anype
rodon
l
euc
ogr
ammicus
(
Val
enciennes,
1828)
Slendergr
ouper
L
apu-
lapu,
Sigapo,Banahan

Mi
chae
lN.
Tre
vor

Ce
phal
ophol
isar
gus
Sc
hneider
,1801
Peacoc
khind
L
apu-l
apu,Si
gapo

K
wang-
TsaoS
hao‡

Ce
phal
ophol
isboe
nak
(Bl
och,1790)
Chocol
atehind
L
apu-l
apu,
Sigapo

K
wang-
TsaoS
hao‡

49
FI
SH ATLAS

SERRANI
DAE

Cephal
opholi
s
se
xmaculat
a
(
Rüppell
,1830)
S
ixbl
otchhind
L
apu-l
apu,Bana
han
,
S
igap
ongp ul
a

J
ohnE.Randal
l

Ce
phal
ophol
iss
onne
rat
i
(
Valenci
ennes,1828)
T
oma tohind
S
igapo,Lapu-
lapu

J
ohnE.Randal
l

Epi
nephe
lusar
eol
atus
(For
sskål
,1775)
Areol
ategrouper
Lapu-
lapu,Si
gapoluba

K
wang-
TsaoS
hao‡

Epi
nephe
lusmal
abar
icus
(
Bloc
h&S chnei
der,
1801)
Mala
bargr
oupe r
L
apu-l
apu,
S i
gapo

J
ohnE.Randal
l

50
 FI
SH ATLAS

SERRANI
DAE

Epi
nephe
lusme
lanos
tigma
Schul
tz,1953
One
-bl
otchgrouper
Si
gapo,Kig ng

J
ohnE.Randal
l

Epi
nephe
lusme
rra
Bl
och,1793
Hone
ycombg r
ouper
Lapu-l
apu

J
ohnE.Randal
l

Epi
nephe
lusongus
(Bl
och,1790)
White-
str
eakedgr
oupe r
L
apu-
lapu,Si
gapo,Se
norita

J
ohnE.Randal
l

Epi
nephe
lusquoyanus
(
Val
enci
ennes
,1830)
Long
fingrouper
Lapu-
la
pu,Si
gapo

J
ohnE.Randal
l

51
FI
SH ATLAS

SERRANI
DAE

Epi
nephe
luss
exf
asc
iat
us
(
Vale
nciennes,1828)
S
ixbargrouper
L
apu-la
pu, Si
gapo

J
ohnE.Randal
l

Epi
nephe
lust
auvi
na
(
Forss
kål,
1775)
Greas
ygrouper
Si
gapongpu k

J
ohnE.Randal
l

Pl
ect
ropomusl
eopar
dus
(
Lac
epède,1802)
L
eopardcora
lgroupe
r
L
apu-l
apu,Bi
loan

J
ohnE.Randal
l

Var
iol
aal
bimar
ginat
a
Ba
iss
ac,1953
Whit
e-e
dg edl
yre
tai
l
Bi
l
oan

J
ohnE.Randal
l

52
 FI
SH ATLAS

SI
GANI
DAE

Si
ganusar
gent
eus
(
Quoy&Ga i
ma r
d,1825)
Stre
amli
nedspi
nefoot
Bal
iwi
s,T
il
is
,Samaral

J
ohnE.Randal
l

Si
ganusc
anal
icul
atus
(
Park,1797)
Whit
e-spo e
ds pi
nefoot
Bal
iwis
,Ti
li
s,Samaral

K
wang-
TsaoS
hao‡

Si
ganusc
oral
li
nus
(Va
lenci
ennes,1835)
Bl
ue-
spo e ds
pinefoot
T
alagbago,
Bal i
wis

F
eLav
api
e-Gonz
ale
s

Si
ganusf
usc
esc
ens
(Hou uyn,1782)
Mo l
edspinefoot
Til
i
s,Bali
wis

K
wang-
TsaoS
hao‡

53
FI
SH ATLAS

SI
GANI
DAE

Si
ganusgu at
us
(
Bloch,1787)
Goldl
inedspi
nef
oot
Samaral

K
wang-
TsaoS
hao‡

Si
ganusj
avus
(
Li
nnaeus
,1766)
S
tre
akedspi
nefoot
S
amaral

F
eLav
api
e-Gonz
ale
s

Si
ganusl
ine
atus
(
Valenci
ennes
,1835)
Golden-
li
nedspi
nefoot
Samaral

J
ohnE.Randal
l

Si
ganusve
rmi
cul
atus
(Va
lenci
ennes,1835)
Vermic
ulate
ds pi
nefoot
Samaralbato

F
eLav
api
e-Gonz
ale
s

54
 FI
SH ATLAS

SI
GANI
DAE

Si
ganusvi
rgat
us
(
Va l
enci
ennes ,1835)
Barhe
ads pinefoot
S
amar
al,
Baliwi
s,Talagbago

J
ohnE.Randal
l

Si
ganusvul
pinus
(
Schl
ege
l&Mül
ler,1845)
Foxface
T
alagbago

F
eLav
api
e-Gonz
ale
s

SI
LLAGI
NIDAE
Si
ll
agos
ihama
(
For
sskål
,1775)
Si
lv
ersil
lago
Asohos

J
ohnE.Randal
l

55
FI
SH ATLAS

SPARI
DAE

Ac
ant
hopagr
uspac
ific
us
I
watsuki
,Kume&Y os
hino,
2010
Pa
cificseabre
am
Bi
koko,
Bak ok
o,Bi
gok

T
oshi
yuk
iSuz
uki

SPHYRAENI
DAE

Sphyr
aenabar
rac
uda
(E
dwa rds
,1771)
Greatbarr
acuda
Barr
acuda,Rompe

J
ohnE.Randal
l

Sphyr
aenaj
ell
o
Cuvi
er,1829
Pic
khandlebar
racuda
Tors
il
yos,Bar
akuda,Rompe

K
wang-
TsaoS
hao‡

Sphyr
aenaobt
usat
a
Cuvi
er,1829
Obtusebarr
acuda
Tor
sil
yos

J
ohnE.Randal
l

56
 FI
SH ATLAS

SPHYRAENI
DAE

Sphyr
aenaput
namae
Jor
da n&S e
ale,1905
Sa
wt oot
hba r
racuda
T
ors
il
yos,Barak
uda,Rompe

J
ohnE.Randal
l

SYNODONTI
DAE

Saur
idagr
aci
li
s
(
Quoy&Gaimar
d,1824)
Gr
aci
lel
iz
ardfish
Kal
aso

K
wang-
TsaoS
hao‡

Saur
idat
umbi
l
(Bl
och,1795)
Gr
eate
rli
zardfish
K
alas
o,U nbundok

K
wang-
TsaoS
hao‡

Saur
idaundos
quami
s
(Ric
hardson,1848)
Brus
htoothli
zardfish
K
alaso,U nbundok

Gr
egor
ioV
.He
rmos
aJr
.

Saur
idawani
eso
S
hindo&Yamada,1972
Wani
esol
iz
ardfish
Kal
aso

Rodol
foB.
Rey
esJ
r.

57
FI
SH ATLAS

SYNODONTI
DAE

Synodusmyops
(
Fors
ter
,1801)
S
nakefish
K
alaso

Gr
egor
ioV
.He
rmos
aJr
.

Synodusvar
iegat
us
(L
acepède
,1803)
Vari
ega
tedli
zar
dfis
h
Kal
aso,
U nbundok

K
wang-
TsaoS
hao‡

TERAPONTI
DAE

Pe
lat
esquadr
ili
neat
us
(Bl
och,1790)
Fourl
inedter
apon
Bakul
e

J
ohnE.Randal
l

Te
raponj
arbua
(F
orssk
ål,
1775)
Ja
rbuaterapon
Bagaong,Gung-
gong

K
wang-
TsaoS
hao‡

58
 FI
SH ATLAS

TERAPONTI
DAE

Te
raponput
a
Cuvier
,1829
S
mal
l
-sc
a l
edterapon
Bagaong

J
ohnE.Randal
l

Te
rapont
her
aps
Cuvier
,1829
L
arg
escal
edterapon
Bugaong,Bagaong

K
wang-
TsaoS
hao‡

TRI
CHI
URI
DAE

Tr
ichi
urusl
ept
urus
L
innaeus
,1758
L
argehe
adha i
rt
ail
E
spada
K
wang-
TsaoS
hao‡

59
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60
61
62

Notes
:
Asympto clengt
hs(L
∞) mar
kedwit
ha nas
ter
isk(
*)aretheme di
anofa
llval
uesf
orthespec
ies
,wh i
l
et her
estaremedi
anofv
aluesf
rom s
tudi
esc
onduc
tedi
nthePhi
li
ppi
nes
.
TheLengt
h satfirs
tma t
uri
ty(L
m)wereesmatedfrom ei
thert
heL∞orL xu
ma si
ngtheFi
shBas
eLifeHi
storyT
ool.
 Notes
:
Asympto clengt
hs(L
∞) mar
kedwit
ha nas
ter
isk(
*)aretheme di
anofa
llval
uesf
orthespec
ies
,wh i
l
et her
estaremedi
anofv
aluesf
rom s
tudi
esc
onduc
tedi
nthePhi
li
ppi
nes
.
6

TheLengt
h satfirs
tma t
uri
ty(L
m)wereesmatedfrom ei
thert
heL∞orL xusi
ngtheFi
shBas
eLifeHi
storyT
ool.
3

ma
64

Notes
:
Asympto clengt
hs(L
∞) mar
kedwit
ha nas
ter
isk(
*)aretheme di
anofa
llval
uesf
orthespec
ies
,wh i
l
et her
estaremedi
anofv
aluesf
rom s
tudi
esc
onduc
tedi
nthePhi
li
ppi
nes
.
TheLengt
h satfirs
tma t
uri
ty(L
m)wereesmatedfrom ei
thert
heL∞orL xu
ma si
ngtheFi
shBas
eLifeHi
storyT
ool.
 Notes
:
Asympto clengt
hs(L
∞) mar
kedwit
ha nas
ter
isk(
*)aretheme di
anofa
llval
uesf
orthespec
ies
,wh i
l
et her
estaremedi
anofv
aluesf
rom s
tudi
esc
onduc
tedi
nthePhi
li
ppi
nes
.
6

TheLengt
h satfirs
tma t
uri
ty(L
m)wereesmatedfrom ei
thert
heL∞orL xusi
ngtheFi
shBas
eLifeHi
storyT
ool.
5

ma
66

Notes
:
Asympto clengt
hs(L
∞) mar
kedwit
ha nas
ter
isk(
*)aretheme di
anofa
llval
uesf
orthespec
ies
,wh i
l
et her
estaremedi
anofv
aluesf
rom s
tudi
esc
onduc
tedi
nthePhi
li
ppi
nes
.
TheLengt
h satfirs
tma t
uri
ty(L
m)wereesmatedfrom ei
thert
heL∞orL xu
ma si
ngtheFi
shBas
eLifeHi
storyT
ool.
 Notes
:
Asympto clengt
hs(L
∞) mar
kedwit
ha nas
ter
isk(
*)aretheme di
anofa
llval
uesf
orthespec
ies
,wh i
l
et her
estaremedi
anofv
aluesf
rom s
tudi
esc
onduc
tedi
nthePhi
li
ppi
nes
.
TheLengt
h satfirs
tma t
uri
ty(L
m)wereesmatedfrom ei
thert
heL∞orL xu
ma si
ngtheFi
shBas
eLifeHi
storyT
ool.
67
NAT
IONALS
TOCKASSESSME
NTPROGRAM
2014
I
SBN978-
971-
95899-
1-4