STAPHYLINIDAEOFLATINAMERICAzt 04621 P 406

Zootaxa 4621 (1): 001–406 ISSN 1175-5326 (print edition)
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Monograph ZOOTAXA
ISSN 1175-5334 (online edition)
https://doi.org/10.11646/zootaxa.4621.1.1
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ZOOTAXA
4621
Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis,

annotated catalog, diversity and distribution
ANGÉLICO ASENJO1, ULRICH IRMLER2, JAN KLIMASZEWSKI 3, DONALD S. CHANDLER4,

HUGO EDUARDO FIERROS-LÓPEZ5 & JULIANA S. VIEIRA6
1
Museu Paraense Emílio Goeldi - MPEG, Av. Perimetral, 1901. CEP: 66077-530. Terra Firme, Belém, Brazil.
E-mail: pukara8@yahoo.com
2
Department of Applied Ecology, Institute for Ecosystem Research, Christian Albrecht University, 24098 Kiel, Germany.
E-mail: uirmler@ecology.uni-kiel.de
3
Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 du P.E.P.S., P.O.
Box 10380, Stn. Sainte-Foy, Québec, Quebec, Canada G1V 4C7. E-mail: jan.klimaszewski@canada.ca
4
Department of Biological Sciences, University of New Hampshire, Durham, NH 03824, USA. E-mail: Donald.Chandler@unh.edu
5
Centro de Estudios en Zoología, CUCBA, Universidad de Guadalajara, Apdo. Postal 234, C.P. 45100, Zapopan, Jalisco, México.
E-mail: hugofierros@yahoo.com.mx
6
Napa Valley College, California, USA. E-mail: juliana_svieira@yahoo.com.br
Magnolia Press
Auckland, New Zealand
Accepted by Z.-W. Yin: 31 Dec. 2018; published: 21 Jun. 2019

ANGÉLICO ASENJO, ULRICH IRMLER, JAN KLIMASZEWSKI, DONALD S. CHANDLER, HUGO
EDUARDO FIERROS-LÓPEZ & JULIANA S. VIEIRA
Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity
and distribution
(Zootaxa 4621)
406 pp.; 30 cm.
21 Jun. 2019
ISBN 978-1-77670-691-4 (paperback)
ISBN 978-1-77670-692-1 (Online edition)
FIRST PUBLISHED IN 2019 BY

Magnolia Press
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ISSN 1175-5326 (Print edition)

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2 · Zootaxa 4621 (1) © 2019 Magnolia Press ASENJO ET AL.

Table of Contents
Abstract . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6
Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7
Material and methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7
Results . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
Subfamily GLYPHOLOMATINAE Jeannel, 1962a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13
Subfamily MICROSILPHINAE Crowson, 1950 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14
Subfamily OMALIINAE W.S. MacLeay, 1825 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14
Tribe Anthophagini Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14
Tribe Corneolabiini Steel, 1950a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15
Tribe Omaliini W.S. MacLeay, 1825 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15
Subfamily PROTEININAE Erichson, 1839b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18
Tribe Austrorhysini Newton & Thayer, 1995 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18
Tribe Proteinini Erichson, 1839b. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18
Tribe Silphotelini Newton & Thayer, 1995 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19
Subfamily MICROPEPLINAE Leach, 1815. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19
Subfamily NEOPHONINAE Fauvel, 1905 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20
Subfamily PSELAPHINAE Latreille, 1802 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20
Supertribe Batrisitae Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20
Tribe Batrisini Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20
Subtribe Batrisina Reitter, 1882b. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20
Incertae sedis in Subtribe Batrisina Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22
Supertribe Clavigeritae Leach, 1815 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23
Tribe Clavigerini Leach, 1815 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23
Subtribe Clavigerodina L.W. Schaufuss, 1882 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23
Supertribe Euplectitae Streubel, 1839 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23
Tribe Bythinoplectini L.W. Schaufuss, 1890. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23
Subtribe Bythinoplectina L.W. Schaufuss, 1890 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23
Subtribe Pyxidicerina Raffray, 1904b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 25
Tribe Dimerini Raffray, 1908a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26
Tribe Euplectini Streubel, 1839 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26
Tribe Jubini Raffray, 1904b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28
Tribe Mayetiini Winkler, 1925 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31
Tribe Metopiasini Raffray, 1904b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32
Subtribe Metopiasina Raffray, 1904b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32
Subtribe Rhinoscepsina Bowman, 1934 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33
Tribe Trichonychini Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34
Subtribe Panaphantina Jeannel, 1950 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34
Subtribe Trichonychina Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 39
Subtribe Trimiina Bowman, 1934 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 42
Incertae sedis in Tribe Trichonychini Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 50
Tribe Trogastrini Jeannel, 194 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 50
Subtribe Phtegnomina O. Park, 1951 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 50
Subtribe Rhexiina O. Park, 1951 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 51
Subtribe Trogastrina Jeannel, 1949 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 51
Incertae sedis in Tribe Trogastrini Jeannel, 1949 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 54
Supertribe Faronitae Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 54
Supertribe Goniaceritae Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 56
Tribe Brachyglutini Raffray, 1904b. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 56
Subtribe Baradina O. Park, 1951 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 56
Subtribe Brachyglutina Raffray, 1904b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 57
Subtribe Decarthrina O. Park, 1951. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 65
Subtribe Eupseniina O. Park, 1951 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 66
Subtribe Pselaptina O. Park in O. Park et al., 1976. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 67
Tribe Goniacerini Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 68
Tribe Iniocyphini O. Park, 1951 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 70
Subtribe Iniocyphina O. Park, 1951 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 70
Tribe Proterini Jeannel, 1949. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 72
Supertribe Pselaphitae Latreille, 1802. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 73
Tribe Arhytodini Raffray, 1890b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 73
Tribe Attapseniini Bruch, 1933c . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 74
Tribe Ctenistini Blanchard, 1845 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 74
Tribe Phalepsini Jeannel, 1949 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 76
STAPHYLINIDAE OF LATIN AMERICA Zootaxa 4621 (1) © 2019 Magnolia Press · 3

Tribe Pselaphini Latreille, 1802. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 76
Tribe Tyrini Reitter, 1882b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 77
Subtribe Somatipionina Jeannel, 1949 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 77
Subtribe Tyrina Reitter, 1882b. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 80
Subfamily PHLOEOCHARINAE Erichson, 1839b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 82
Subfamily TACHYPORINAE W.S. MacLeay, 1825 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 83
Tribe Mycetoporini Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 83
Tribe Tachyporini W.S. MacLeay, 1825 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84
Tribe Vatesini Seevers, 1958 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88
Subfamily HABROCERINAE Mulsant & Rey, 1877. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88
Subfamily ALEOCHARINAE Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 89
Tribe Aleocharini Fleming, 1821. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 89
Subtribe Aleocharina Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 89
Incertae sedis in Tribe Aleocharini Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92
Antillusini Pace, 2012 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 92
Tribe Athetini Casey, 1910a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93
Subtribe Athetina Casey, 1910a. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93
Subtribe Schistogeniina Fenyes, 1918. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 104
Subtribe Thamiaraeina Fenyes, 1921a. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 104
Incertae sedis in Tribe Athetini Casey, 1910a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 104
Tribe Autaliini Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 114
Tribe Cordobanini Bernhauer, 1910 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 115
Tribe Corotocini Fenyes, 1918 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 115
Subtribe Abrotelina Seevers, 1957 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 115
Subtribe Corotocina Fenyes, 1918. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 116
Subtribe Eburniogastrina Jacobson et al., 1986 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 117
Subtribe Termitogastrina Bernhauer & Scheerpeltz, 1926 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 118
Subtribe Termitoiceina Jacobson, Kistner & Pasteels, 1986 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 120
Subtribe Termitopithina Jacobson, Kistner & Pasteels, 1986 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 121
Subtribe Timeparthenina Fenyes, 1921a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 121
Incertae sedis in Tribe Corotocini Fenyes, 1918. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 122
Tribe Crematoxenini Mann, 1921 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 123
Tribe Diestotini Mulsant & Rey 1871 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 125
Tribe Diglottini Jacobson, 1909 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 127
Tribe Ecitogastrini Fenyes, 1918 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 128
Tribe Falagriini Mulsant & Rey, 1873b. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 128
Tribe Geostibini Seevers, 1978 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132
Tribe Gymnusini Heer, 1839 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 133
Tribe Homalotini Heer, 1839 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 134
Subtribe Bolitocharina Thomson, 1859. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 134
Subtribe Dinardopsina Bernhauer & Scheerpeltz 1926. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 140
Subtribe Gyrophaenina Kraatz, 1856b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 140
Subtribe Homalotina Heer, 1839 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143
Subtribe Silusina Fenyes, 1918 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145
Incertae sedis in Tribe Homalotini Heer, 1839 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 145
Tribe Hoplandriini Casey, 1910 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 148
Subtribe Hoplandriina Casey, 1910a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 148
Subtribe Platandriina Hanley, 2002c . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 149
Incertae sedis in Tribe Hoplandriini Casey, 1910a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 150
Tribe Hygronomini Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 151
Subtribe Hygronomina Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 151
Subtribe Saphoglossina Bernhauer & Scheerpeltz, 1926 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 152
Incertae sedis in Tribe Hygronomini Thomson, 1859. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 152
Tribe Hypocyphtini Laporte, 1835 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 152
Tribe Liparocephalini Fenyes, 1918 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 154
Tribe Lomechusini Fleming, 1821. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 154
Subtribe Lomechusina Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 154
Subtribe Myrmedoniina Thomson, 1867a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 155
Subtribe Termitozyrina Seevers, 1957 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 168
Incertae sedis in Tribe Lomechusini Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 169
Tribe Mesoporini Cameron, 1959 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 170
Tribe Mimecitini Wasmann, 1917 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 170
Subtribe Labidopullina Jacobson & Kistner, 1991 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 170
Subtribe Leptanillophilina Fenyes, 1918 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 170
Subtribe Mimecitina Wasmann, 1917 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 172

Subtribe Mimonillina Bernhauer & Scheerpeltz, 1926 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 173
Tribe Myllaenini Ganglbauer, 1895. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 174
Tribe Oxypodini Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 175
Subtribe Dinardina Mulsant & Rey, 1873a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 175
Subtribe Meoticina Seevers, 1978 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 176
Subtribe Microglottina Fenyes, 1918 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 178
Subtribe Oxypodina Thomson, 1859. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 178
Sbtribu Phloeoporina Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 181
Incertae sedis in Tribe Oxypodini Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 182
Tribe Paglini Newton & Thayer, 1992 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 189
Tribe Paradoxenusini Bruch, 1937 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 190
Tribe Philotermitini Seevers, 1957 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 190
Tribe Phytosini Thomson, 1867a. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 190
Tribe Placusini Mulsant & Rey, 1871 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 191
Tribe Pronomaeini Mulsant & Rey, 1873b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192
Tribe Sceptobiini Seevers, 1978 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 193
Tribe Tachyusini Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 193
Tribe Taxicerini Lohse in Lohse & Lucht, 1989 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 196
Tribe Termitohospitini Seevers, 1941 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 196
Subtribe Termitohospitina Seevers, 1941 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 196
Tribe Termitonannini Fenyes, 1918 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 197
Subtribe Perinthina Bernhauer & Scheerpeltz, 1926. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 197
Subtribe Termitonannina Fenyes, 1918 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 199
Tribe Termitusini Fenyes, 1918 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 200
Subtribe Termitospectrina Seevers, 1957 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 200
Tribe Trichopseniini LeConte & Horn, 1883 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 200
Incertae sedis in Subfamily Aleocharinae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 201
Subfamily SCAPHIDIINAE Latreille, 1807 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 201
Tribe Cypariini Achard, 1924 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 201
Tribe Scaphidiini Latreille, 1806 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 202
Tribe Scaphisomatini Casey, 1893 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 203
Subfamily PIESTINAE Erichson, 1839a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 205
Subfamily OSORIINAE Erichson, 1839a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 206
Tribe Eleusinini Sharp, 1887a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 206
Tribe Leptochirini Sharp, 1887a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 207
Tribe Osoriini Erichson, 1839a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 208
Subtribe Osoriina Erichson, 1839a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 208
Tribe Thoracophorini Reitter, 1909 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 213
Subtribe Clavilispinina Newton & Thayer, 1992 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 213
Subtribe Glyptomina Newton & Thayer, 1992 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 214
Subtribe Lispinina Bernhauer & Schubert, 1910. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 215
Subtribe Thoracophorina Reitter, 1909 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 218
Subfamily OXYTELINAE Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 220
Tribe Blediini Ádám, 2001 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 220
Tribe Coprophilini Heer, 1839 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 221
Tribe Deleasterini Reitter, 1909 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 222
Tribe Euphaniini Reitter, 1909 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 222
Tribe Oxytelini Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 222
Tribe Thinobiini J. Sahlberg, 1876 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 226
Incertae sedis in Subfamily Oxytelinae. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 230
Subfamily OXYPORINAE Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 231
Subfamily MEGALOPSIDIINAE Leng, 1920 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 231
Subfamily SCYDMAENINAE Leach, 1815. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 232
Supertribe Cephenniitae Reitter, 1882d . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 232
Tribe Cephenniini Reitter, 1882d . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 232
Tribe Eutheiini Casey, 1897. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 233
Supertribe Mastigitae Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 233
Tribe Clidicini Casey, 1897 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 233
Supertribe Scydmaenitae Leach, 1815 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 234
Tribe Chevrolatiini Reitter, 1882d. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 234
Tribe Glandulariini L.W. Schaufuss, 1889 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 234
Tribe Scydmaenini Leach, 1815 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 244
Subfamily STENINAE W.S. MacLeay, 1825 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 246
Subfamily EUAESTHETINAE Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 247
Tribe Alzadaesthetini Scheerpeltz, 1974 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 247

Tribe Austroesthetini Cameron, 1944b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 247
Tribe Euaesthetini Thomson, 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 248
Tribe Stenaesthetini Bernhauer & Schubert, 1911 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 250
Subfamily SOLIERIINAE Newton & Thayer, 1992 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 250
Subfamily LEPTOTYPHLINAE Fauvel, 1874b. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 251
Tribe Metrotyphlini Coiffait, 1963a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 251
Tribe Neotyphlini Coiffait, 1963a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 251
Subfamily PSEUDOPSINAE Ganglbauer, 1895 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 252
Subfamily PAEDERINAE Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 253
Tribe Cylindroxystini Bierig, 1943 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 253
Subtribe Cylindroxystina Bierig, 1943 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 253
Tribe Lathrobiini Laporte, 1835 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 253
Subtribe Astenina Hatch, 1957 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 253
Subtribe Echiasterina Casey, 1905 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 254
Subtribe Lathrobiina Laporte, 1835 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 256
Subtribe Medonina Casey, 1905 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 260
Subtribe Scopaeina Mulsant & Rey, 1878c . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 267
Subtribe Stilicina Casey, 1905 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 269
Subtribe Stilicopsina Casey, 1905 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 271
Tribe Paederini Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 273
Subtribe Cryptobiina Casey, 1905 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 273
Subtribe Dicaxina Schomann & Solodovnikov, 2017 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 276
Subtribe Dolicaonina Casey, 1905. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 276
Subtribe Paederina Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 277
Incertae sedis in Tribe Paederini Fleming, 1821 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 279
Tribe Pinophilini Nordmann, 1837 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 280
Subtribe Pinophilina Nordmann, 1837 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 280
Subtribe Procirrina Bernhauer & Schubert, 1912 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 282
Subfamily STAPHYLININAE Latreille, 1802 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 283
Tribe Diochini Casey, 1906 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 283
Tribe Platyprosopini Lynch-Arribálzaga, 1884 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 284
Tribe Staphylinini Latreille, 1802 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 284
Subtribe Acylophorina Outerelo & Gamarra, 1985. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 284
Subtribe Amblyopinina Seevers, 1944 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 285
Subtribe Anisolinina Hayashi, 1993 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 288
Subtribe Cyrtoquediina Brunke & Solodovnikov in Brunke et al., 2016 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 288
Subtribe Erichsoniina Brunke & Solodovnikov in Brunke et al., 2016 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 289
Subtribe Hyptiomina Casey, 1906 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 290
Subtribe Philonthina Kirby in Richardson, 1837 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 290
Subtribe Philothalpina Chatzimanolis & Brunke in Chani-Posse et al., 2018a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 299
Subtribe Quediina Kraatz, 1857a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 299
Subtribe Staphylinina Latreille, 1802 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 301
Subtribe Tanygnathinina Reitter, 1909 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 304
Subtribe Xanthopygina Sharp, 1884 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 305
Incertae sedis in Tribe Staphylinini Latreille, 1802 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 312
Tribe Xantholinini Erichson, 1839a. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 312
Incertae sedis in family Staphylinidae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 321
Taxonomic considerations in Subfamily Pselaphinae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 341
Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 341
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 341
Abstract
The taxonomic diversity of species and genera of Staphylinidae and their distribution in Latin America are discussed and
listed/tabulated based on published literature. In total, 11,675 species in 1075 genera are reported from Latin America.
South America has the most recorded species for Latin America, 7773 including 7079 endemic species. The four subfam-
ilies with the highest total number of genera including endemic genera are Aleocharinae, Staphylininae, Pselaphinae, and
Paederinae. All recorded genera and their synonyms are listed alphabetically within the subfamilies of Staphylinidae, pro-
viding a catalog of published genera for Latin America. Two hundred forty-four color or black and white images of species
representing different genera are provided.
Key words: diversity, distribution, species, Latin America, Staphylinidae

Introduction
Biological diversity is an asset of the world, and an understanding of this asset is crucial for sustaining life on our
planet. The recognition and comprehension of biological diversity would not be possible without taxonomic
research, which is the foundation of biological science. Taxonomists combine theory and practice of discovering,
describing and classifying organisms to help in understanding the natural world.
The study of the Latin American fauna of the family Staphylinidae started with Fabricius (1787) when he
described Smilax pilosa (Fabricius) from specimens captured in French Guiana by the Prussian botanist, Julius
Philip Benjamin von Rohr. Later, Fabricius (1793) described Plochinocerus fulguns and Olivier (1795) described
Taenodema aenea from French Guiana and Suriname respectively.
Throughout time essential contributions were made in the study of the Latin American fauna of the family,
such as the Staphylinidae of the Amazon Basin (Sharp 1876a), the study of Staphylinidae of Central America
(Sharp 1883–1887a,b), the monograph of the West Indies beetles (Blackwelder 1943), and many others. Stimulated
to comprehend the coverage of these data for Latin America, it was necessary to organize all this information in an
extensive catalog initially based on the compilations of Bernhauer & Schubert (1910–1912, 1914, 1916),
Bernhauer & Scheerpeltz (1926), Csiki (1910, 1919), Scheerpeltz (1933, 1934), Raffray (1911), Blackwelder
(1944), Newton & Chandler (1989), Newton & Thayer (1992), Löbl (1997), Herman (2001b–h), Newton et al.
(2005), and all taxonomic publications in the family.
Excellent works that summarize the history of the study of the Latin American fauna of Staphylinidae can be
consulted in Navarrete-Heredia et al. (2002), Chani-Posse & Thayer (2008), Asenjo et al. (2013), and Irmler &
Asenjo (2018).
The primary purpose of this paper is to provide comprehensive information on the taxonomic diversity of the
family Staphylinidae in Latin America. This contribution is a compilation of published data on the species reported
from Latin America.
Material and methods
Taxonomic list. This section presents the arrangement of subfamilies, supertribes, tribes, and subtribes following
Bouchard et al., (2011). The most recent taxonomic changes in the subfamilies are incorporated: e.g., Aleocharinae
by Elven et al. (2010, 2012), Osswald et al. (2013); in Oxytelinae by Khachikov (2012); in Paederinae by
Schomann & Solodovnikov (2017); in Staphylinini by Chatzimanolis et al. (2010), Chatzimanolis (2104b), Brunke
et al. (2016), Chani-Posse et al. (2018a); and in Scydmaeninae by Jałoszyński (2016a).
Genera are listed in alphabetical order within their higher taxon. Valid genera are numbered continuously
through the family and printed in boldface, followed by the author, year, and page number of the original
description and the type species with author and year. All valid subgenera are listed alphabetically under their
genus, with corresponding details; synonymous genera or subgenera are listed alphabetically under their valid
names. Synonymic names are followed by “[synonymized by author, year, and page number]” that cited the first
publication as the synonym of the valid name, but for a few synonymies a secondary source of the synonymy is
given due to the difficulty of locating papers with the initial synonymy, cited as [author, year, and page number].
An asterisk in brackets “[*]” indicates a valid subgenus with species cited from Latin America. The
information on the subgenera from Latin America with “[*]” should be regarded with caution because many
species were not assigned to subgenera in the original descriptions or subsequently.
Under the heading “Characters/Description” is a chronological list of all subsequent publications (author, year
and page number) that provide a redescription and/or relevant characters for the genus or subgenera. The “Key”
section lists in chronological order papers that have keys for species found in Latin America.
Distribution. Latin America is subdivided into the West Indies, Central America and South America, and
additionally is cited its distributions in other regions of the world. The composition of the different areas of the
world follow the concepts used in the catalog of Herman (2001a–h). Species distributions in Latin America are
based on the database of the first author and follow the catalog of Blackwelder (1944), which is summarized below
(countries arranged North to South, and West to East):
West Indies: Bahamas, Cuba, Cayman Is., Turks & Caicos Isl., Jamaica, Navassa Is., Haiti, Dominican

Republic, Puerto Rico, US Virgin Isl., British Virgin Isl., Anguilla, Saint-Martin, Sint Maarten, Saint-Barthelemy,
Saint Kitts & Nevis, Antigua & Barbuda, Montserrat, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent &
Grenadines, Barbados, Grenada, Trinidad-Tobago, Aruba, Curacao, Bonaire.
Central America: Mexico, Guatemala, Belize, Honduras, El Salvador, Nicaragua, Costa Rica, Panama.
South America: Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Galapagos Isl., Peru,
Bolivia, Brazil, Paraguay, Uruguay, Chile, Argentina, Falkland Isl.
The total number of valid names (species and/or subspecies) for each genus in Latin America is cited, together
with any relevant information (record of fossil species, comments in the distribution, etc.).
The records of Latin America countries and other external regions for each genus, is based on a compilation of
records in catalogs, and in the taxonomic and ecological literature. Data in the distribution section does not include
records of taxa that lack specific species or subspecies determinations.
TABLE 1. Number of species and (genera in parentheses) by subfamily in Latin America and its subregions.
Images. The habitus images are a compilation of the species recorded in Latin America based on the literature
plus images of some species from the following insect collections: CEMT (Setor de Entomologia da Coleção
Zoológica da Universidade Federal de Mato Grosso, Departamento de Biologia e Zoologia, Cuiabá, Mato Grosso,
Brazil), CNC (Canadian National Collection of Insects, Ottawa, Canada), MEKRB (Museo de Entomologia
"Klaus Raven Büller", Universidad Agraria La Molina, Lima, Peru), MUSM (Entomological Collection of Museo

de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Lima, Peru), VMDC (Victor Manuel
Diéguez Insect Collection, Santiago, Chile). Genera that have images of species in the plates are marked in the text
with and number figure.
TABLE 2. Number of species and genera by geographical regions and country in Latin America
Results
The information provided herein on the taxonomic diversity of species, genera and subfamilies of the megadiverse
beetle family Staphylinidae in Latin America promotes the knowledge and understanding of the Latin American
fauna. This publication presents a synthesized baseline of taxonomic biodiversity data on the distribution of species
and genera, and their attributable habitats.
In Latin America, there are approximately 11,675 recorded species (including subspecies) in 1075 genera,
representing the 18% of total world biodiversity for the family. The four subfamilies with the highest total number
of species and genera are: Aleocharinae (2887 species in 407 genera), Staphylininae (1745 species in 127 genera),
Pselaphinae (1622 species in 256 genera), and Paederinae (1516 species in 96 genera). The genera with the highest
number of species are Stenus (Steninae) 620 species, Euconnus (Scydmaeninae) with 615 species and Atheta
(Aleocharinae) with 355 species. In this publication we compare species and generic diversity of the three
geographic regions of Latin America. The majority of species are in South America with 7773 species in 855
genera, followed by Central America with 3660 species in 530 genera, and the West Indies with 1070 species in
255 genera. Considering the size of the landmass and diversity of habitats of the three zoogeographic regions, these
large numbers of species and genera are thoroughly understandable (Table 1).

TABLE 3. Number of species and genera recorded or described from Latin America by decades.

TABLE 4. Number of species and genera reported from Latin America by different authors that described over 30
species.
Biodiversity is organized by country and geographic regions of Latin America (Table 2). For South America,
the largest number of species and genera are in Brazil (2829 species in 479 genera), Argentina (1961 species in 305
genera), and Chile (1037 species in 225 genera) (Fig. 1). In Central America, Mexico (1799 species in 367 genera),
Panama (1211 species in 292 genera), and Guatemala (1016 species in 225 genera) have the highest diversity for
the region (Fig. 1); and for the West Indies the greatest diversity is found in Cuba (312 species in 119 genera),
Jamaica (234 species in 106 genera), and Trinidad-Tobago (198 species and 108 genera) (Fig. 1).
Other interesting data are the endemicity of taxa in Latin America (Table 1 and Table 2). For practical reasons
in regards to this work, we define "endemic" as present in Latin America and not occuring elsewhere. A total of
11,229 species and 696 genera are here considered to be endemic to Latin America (Table 1). The subfamilies
Neophoninae and Solieriinae are restricted to the southern portion of South America (Chile and Argentina).
Aleocharinae (2792 species and 280 genera), Staphylininae (1623 species and 74 genera) and Pselaphinae (1593
species and 201 genera) have the highest number of endemic taxa in Latin America (Table 1). South America leads

FIGURE 1. Numbers of species and genera by regions and countries.
FIGURE 2. Number of species described by authors.

FIGURE 3. Number of species and genera described in each decade.
in the total number of endemic taxa with 7079 species and 404 genera, followed by Central America with 2846
species and 83 genera, and The West Indies has 742 species and 32 genera (Table 1). In South America Brazil leads
with 479 species and 88 genera of endemic taxa, followed for Central America by Mexico having 367 species and
25 genera, and the West Indies has 119 species and 4 genera (Table 2). Plesiomalota (Aleocharinae) and
Gnathymenus (Paederinae) are the endemic genera with the highest number of species. Plesiomalota has 99 species
found only in South America, and Gnathymenus has 79 species distributed in Central and South America.
During this compilation approximately 70 adventive species were found in Latin America, with the majority
being present in the South American countries. Philonthus (Staphylininae) and Aleochara (Aleocharinae) are the
genera with highest number of adventive species, seven and six respectively.
In Table (3) the listing of species and genera are reported chronologically (1780 to present) from Latin
America. There are three major chronological and historical peaks regarding the reported numbers of species and
genera (Fig. 3). The first extends from 1880 and 1889 (1648 species and 113 genera), the second from 1980 to 1989
(1112 species and 68 genera), and the last from 2010 to the present with 1317 species and 63 genera. These counts
were driven in the 19th century by David Sharp (1667 species) and Wilhelm Ferdinand Erichson (358 species); in
the 20th century by Max Bernhauer (1173 species) and Herbert Franz (794 species); and in the 21st century by
Roberto Pace (749 species) and Volker Puthz (583 species). This trend in description of new genera and species
continues and demonstrates that the Latin American fauna is not adequately studied, and that many taxa will be
discovered in the future, confirming the true megadiversity of Staphylinidae.
The list of species and genera reported from Latin America by different authors is presented in Table (4), when
species or genera were described by more than one author, then only the first is recorded. The most notable impact
on the knowledge of the Latin American fauna of Staphylinidae regarding the number of described taxa were the
publications of David Sharp, Roberto Pace, Max Bernhauer, Volker Puthz, and Herbert Franz (Fig. 2). Other
valuable contributors are listed in Table (4).

Staphylinidae is a megadiverse group with published information that is widely dispersed; we hope this work
will inspire researchers to study this fascinating group of beetles, with an understanding that what we know now is
only the tip of an iceberg.
Subfamily GLYPHOLOMATINAE Jeannel, 1962a
0001. Glypholoma Jeannel, 1962a: 482 (Fig. 4)
Type species: Glypholoma pustuliferum Jeannel, 1962

Lathrimaeodes Scheerpeltz, 1972d: 58 [synonymized by Newton 1975: 54]
Type species: Lathrimaeodes pustulipenne Scheerpeltz, 1972
6 species are included in this genus from South America.

Characters/Description: Newton 1975: 54 (characters); Thayer & Newton 1979: 29 (characters); Thayer 1997: 159
(characters).
Key: Thayer & Newton 1979: 45 (species of the genus); Thayer 1997: 159 (species of the genus).
Distribution.
South America: Chile, Argentina.
Other regions: Australian.
0002. Proglypholoma Thayer, 1997: 167 (Fig. 5)

Type species: Proglypholoma aenigma Thayer, 1997
Only one species is known from and is restricted to South America.

Characters/Description: Thayer 1997: 159 (characters in key).
Key: Thayer 1997: 159 (species of the genus).
Distribution.
South America: Chile.
Subfamily MICROSILPHINAE Crowson, 1950
0003. Microsilpha Broun, 1886: 889 (Fig. 6)

Type species: Microsilpha litorea Broun, 1886
Acruliodema Scheerpeltz, 1972d: 45 [synonymized by Newton & Thayer 1995: 291]
Type species: Acruliodema argentinum Scheerpeltz, 1972
Micragyrtes Champion, 1918: 46 [synonymized by Newton & Thayer 1995: 291]
Type species: Micragyrtes ocelligerus Champion, 1918

Characters/Description: Newton & Thayer 1995: 289 (characters in key as subfamily), 291 (characters as
subfamily); Jeannel 1962a: 484 (characters as Micragyrtes).
Distribution.

Subfamily OMALIINAE W.S. MacLeay, 1825
Tribe Anthophagini Thomson, 1859
0004. Amphichroum Kraatz, 1857b: 947

Type species: Lathrimaeum canaliculatum Erichson, 1840
Stachygraphis Horn, 1883: 285 [synonymized by Bernhauer & Schubert 1910: 70]
Type species: Stachygraphis maculata Horn, 1883
Only one species is known in this genus from Central America.

Characters/Description: Navarrete-Heredia et al. 2002: 34 (characters in key); Thayer 2003: 314 (characters in
key).
Distribution.
Central America: Mexico.
Other regions: Nearctic, Palaearctic, Oriental.
0005. Anthobium Leach in Samouelle, 1819: 175 (Fig. 7)

Type species: Omalium atrocephalum Gyllenhal, 1827
Eudeliphrum Champion, 1920: 244 [synonymized by Cameron 1930: 151]
Type species: Eudeliphrum gracilipalpe Champion, 1920
Lathrimaeum Erichson, 1839b: 624 [synonymized by Kloet & Hincks 1945: 161]
Subgenera:
Anthobium Leach in Samouelle, 1819: 175
Prionothorax Luze, 1905: 68
Type species: Lathrimaeum reflexum Reitter, 1891
2 species are included in this genus from Central America.

key).
Key: Thayer 2003: 344 (Mexican species).
Distribution.
Tribe Corneolabiini Steel, 1950a
0006. Metacorneolabium Steel, 1950b: 62

Type species: Metacorneolabium minor Steel, 1950
Metomalium Coiffait & Sáiz, 1968: 449 [synonymized by Scheerpeltz 1933: 1031]
Type species: Metomalium chilense Coiffait & Sáiz, 1968

Characters/Description: Coiffait & Sáiz 1968: 448 (characters in key as Metomalium), 449 (characters as
Metomalium); Thayer 1985: 116 (characters).
Key: Thayer 1985: 144 (South American species).

Distribution.
Tribe Omaliini W.S. MacLeay, 1825
0007. Acrolocha Thomson, 1858: 38

Type species: Omalium striatum Gravenhorst, 1802
Elonium Leach in Samouelle, 1819: 175 [invalid, rejected by ICZN 1993: 164 (Opinion 1722)]
Type species: Omalium striatum Gravenhorst, 1802
Only one species is known in this genus from Central America.

key).
Distribution.
Other regions: Nearctic, Palaearctic.
0008. Crymus Fauvel, 1904a: 92

Type species: Crymus antarcticus Fauvel, 1904
Arpediomimus Cameron, 1917a: 277 [synonymized by Steel 1970: 240]
Type species: Arpediopsis falklandicus Cameron, 1917
Arpediopsis Cameron, 1917b: 124 [synonymized by Steel 1970: 240]
Type species: Arpediopsis falklandicus Cameron, 1917
Only one species is known in this genus from South America and Falkland Isl.
Characters/Description: Steel 1964: 348 (characters).
Key: Steel 1964: 349 (species of genus).
Distribution.
South America: Argentina, Falkland Isl.
Other regions: Australian, Antarctic.
0009. Dropephylla Mulsant & Rey, 1880: 242 (Fig. 8)

Type species: Omalium lucidum Erichson, 1839
Only one species is known from Central America.

key).
Distribution.
Other regions: Nearctic, Palearctic, Oriental.
0010. Ischnoderus Fauvel, 1867: 46

Type species: Omalium insigne Fairmaire & Germain, 1862
Subgenera:
Ischnoderus Fauvel, 1867: 46 [*]
Type species: Omalium insigne Fairmaire & Germain, 1862

Walkerellus Bernhauer, 1939d: 203
Type species: Ischnoderus paradoxus Bernhauer, 1939

Characters/Description: Coiffait & Sáiz 1968: 448 (characters in key), 454 (characters).
Distribution.
0011. Omaliopsis Jeannel, 1940: 118

Type species: Omalium africanum Fauvel, 1907
3 species are included in this genus from Central and South America.
Characters/Description: Coiffait & Sáiz 1968: 449 (characters in key), 451 (characters); Navarrete-Heredia et al.
2002: 35 (characters in key); Thayer 2003: 315 (characters in key).
Distribution.
Other regions: Nearctic, Ethiopian, Madagascan, Oriental, Australian.
0012. Omalium Gravenhorst, 1802: 111 (Fig. 9)

Type species: Staphylinus rivularis Paykull, 1789
Homalium Ljungh, 1804: 74 [synonymized by Blackwelder 1952: 188]
Homalium Agassiz, 1846: 258 [synonymized by Blackwelder 1952: 188]
Homalium Gemminger & Harold, 1868: 665 [synonymized by Blackwelder 1952: 189]
Ochthexenus Motschulsky, 1860b: 546 [synonymized by Fauvel 1869: 493]
10 species are included in this genus from Latin America.

Characters/Description: Blackwelder 1943: 53 (characters); Navarrete-Heredia et al. 2002: 34 (characters in key);
Thayer 2003: 315 (characters in key).
Key: Blackwelder, 1943: 54 (species of the West Indies).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic.
Central America: Mexico, Guatemala, Honduras, El Salvador.
South America: Bolivia, Chile, Argentina.
Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian.
0013. Phloeonomus Heer, 1839: 184 (Fig. 10)

Type species: Omalium pusillum Gravenhorst, 1806
Subgenera:
Phloeonomodes Smetana, 1964: 163, 168
Type species: Omalium minimus Erichson, 1839
Phloeonomus Heer, 1839: 184 [*]
Type species: Omalium pusillum Gravenhorst, 1806

Characters/Description: Blackwelder 1943: 51 (characters); Navarrete-Heredia et al. 2002: 34 (characters in key);
Thayer 2003: 315 (characters in key).
Distribution.
West Indies: Cuba, Jamaica, Dominican Republic, Puerto Rico, Guadeloupe, Grenada, Trinidad-Tobago.
Central America: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama.
South America: Colombia, Venezuela, Brazil, Argentina.
Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, Oceanic.
0014. Phyllodrepa Thomson, 1859: 52 (Fig. 11)

Type species: Staphylinus floralis Paykull, 1789

Distribution.
South America: Venezuela, Peru, Argentina.
Other regions: Nearctic, Palaearctic, Oriental, Australian.
0015. Xenanthobium Coiffait & Sáiz, 1968: 452

Type species: Anthobium andicola Fauvel, 1867

Characters/Description: Coiffait & Sáiz 1968: 448 (characters in key).
Distribution.
0016. Xylodromus Heer, 1839: 174

Type species: Omalium depressum Gravenhorst, 1802
Drephopylla Fiori, 1900: 90, 93 [synonymized by Luze 1906: 493]
Type species: Omalium depressum Gravenhorst, 1802
Etheothassa Thomson, 1858: 38 [synonymized by Luze 1906: 493]
Type species: Omalium deplanatum Gyllenhal, 1810
Omalissus Broun, 1893: 1042 [synonymized by Cameron 1945b: 782]
Type species: Omalissus castaneus Broun, 1893
Characters/Description: Navarrete-Heredia et al. 2002: 34 (characters in key), 35 (characters in key as
Dropephylla); Thayer 2003: 315 (characters in key), 315 (characters in key as Dropephylla).
Distribution.
South America: Venezuela, Chile.
Other regions: Nearctic, Palaearctic, Australian.
Subfamily PROTEININAE Erichson, 1839b
Tribe Austrorhysini Newton & Thayer, 1995
0017. Megarthroides Steel, 1966: 298 (Fig. 12)

Type species: Megarthroides chilensis Steel, 1966

Characters/Description: Steel 1966: 292 (characters in key).
Distribution.
Tribe Proteinini Erichson, 1839b
0018. Megarthrus Curtis, 1829: 28 (Fig. 13)

Type species: Staphylinus depressus Paykull, 1789
Phloeobium Dejean, 1833: 69 [synonymized by Lacordaire 1854: 147]
Psyllius Gistel, 1834: 9 [synonymized by Blackwelder 1952: 331]
Characters/Description: Kistner 1961b: 2238 (characters); Navarrete-Heredia et al. 2002: 45 (characters in key);
Cuccodoro 2011: 122 (characters).
Key: Cuccodoro & Löbl 1996: 146 (species of the America north of Mexico).
Distribution.
Central America: Mexico, Guatemala, Honduras, Nicaragua, Costa Rica.
South America: Colombia, Peru, Argentina.
Other regions: Nearctic, Palaearctic, Ethiopian, Oriental, Australian, Oceanic.
0019. Proteinus Latreille, 1797: 9 (Figs. 14–15)

Type species: Dermestes brachypterus Fabricius, 1792
2 species are included in this genus from the West Indies and Central America.
Characters/Description: Coiffait & Sáiz 1968: 456 (characters); Navarrete-Heredia et al. 2002: 45 (characters in
key).
Distribution.
West Indies: Jamaica.
Central America: Costa Rica.
Other regions: Nearctic, Palaearctic, Ethiopian, Oriental.
Tribe Silphotelini Newton & Thayer, 1995
0020. Alloproteinus Steel, 1966: 300

Type species: Proteinus nigriceps Fauvel, 1868
Only one species is known from and is restricted to from South America.
Characters/Description: Steel 1966: 293 (characters in key).
Distribution.
Subfamily MICROPEPLINAE Leach, 1815
0021. Arrhenopeplus Blackwelder, 1952: 63 (Fig. 16)

Type species: Micropeplus tesserula Curtis, 1828
Arrhenopeplus C. Koch, 1937: 257 [nomen nudum, validated by Blackwelder 1952: 63]
Type species: [no type species designated]
Subgenera:
Arrhenopeplus Blackwelder, 1952: 63 [*]
Type species: Micropeplus tesserula Curtis, 1828
Sphaeropeplus Coiffait, 1982: 127
Type species: Arrhenopeplus quatordecimcarinatus Coiffait, 1982
1 species is included in this genus from Central America.

Distribution.
Central America: Mexico, Panama.
0022. Micropeplus Latreille, 1809: 377 (Fig. 17)

Type species: Staphylinus porcatus Paykull, 1789

Characters/Description: Campbell 1968: 228 (characters in key), 230 (characters); Campbell 1978: 1257
(characters in key); Navarrete-Heredia et al. 2002: 47 (characters in key).
Key: Campbell 1968: 231 (species of the genus); Campbell 1973c: 575 (species of the genus); Campbell 1978:
1257 (species of the genus).
Distribution.
0023. Peplomicrus Bernhauer, 1928c: 286 (Fig. 18)

Type species: Micropeplus uyttenboogaarti Bernhauer, 1928

Characters/Description: Campbell 1968: 228 (characters in key), 252 (characters in key); Campbell 1978: 1257
(characters in key); Campbell 1986: 62 (characters); Navarrete-Heredia et al. 2002: 47 (characters in key).
Key: Campbell 1968: 253 (species of the genus); Campbell 1978: 1257 (species of the genus); Campbell 1986: 68
(species of the New World); Campbell 1991b: 40 (species group and species of New World).
Distribution.
West Indies: Dominican Republic.
Central America: Mexico, Guatemala, El Salvador, Costa Rica, Panama.
South America: Colombia, Venezuela, Suriname, French Guiana, Ecuador, Peru.
Other regions: Palaearctic, Ethiopian.
Subfamily NEOPHONINAE Fauvel, 1905
0024. Neophonus Fauvel, 1905: 99 (Fig. 19)

Type species: Neophonus bruchi Fauvel, 1905

Characters/Description: Thayer 1987: 395 (characters); Newton & Thayer 1995: 290 (characters in key as
subfamily), 300 (characters as subfamily).
Distribution.
Subfamily PSELAPHINAE Latreille, 1802
Supertribe Batrisitae Reitter, 1882b
Tribe Batrisini Reitter, 1882b
Subtribe Batrisina Reitter, 1882b
0025. Arthmius LeConte, 1849: 91

Type species: Arthmius globicollis LeConte, 1849
Characters/Description: Raffray 1894: 231 (characters in key); Raffray 1898a: 441 (characters), 443 (characters in
key), 462 (characters); Raffray 1904a: 4 (characters in key); Raffray 1908a: 134 (characters in key), 149
(characters); O. Park 1942: 215 (characters in key), 215 (characters); O. Park 1944: 248, 249 (characters in
key); Becker & Sanderson 1953a: 404 (characters in key); Becker & Sanderson 1953b: 454 (characters);
Jeannel 1962b: 384 (characters); Chandler 2002: 52 (characters in key).
Key: Schaufuss 1887: 140 (some species of the genus as Batrisus); Reitter 1888: 246 (Brazilian species); Raffray
1898a: 463 (species of the genus); Raffray 1904a: 16 (species group); Fletcher 1930: 97 (some Neotropical
species); O. Park 1942: 217–219, 222–230, 233–236 (Neotropical group of species and species); O. Park
1943: 195 (Mexican species); O. Park 1944: 251 (Mexican species); Becker & Sanderson 1953b: 455 (species
of Honduras).
Distribution.
Central America: Mexico, Guatemala, Honduras, Panama.
South America: Colombia, Venezuela, Peru, Bolivia, Brazil, Paraguay, Chile, Argentina.
Other regions: Nearctic.
0026. Batoctenus Sharp, 1887b: 18

Type species: Batoctenus puncticollis Sharp, 1887
5 species are known in this genus and are restricted to Central and South America.
Characters/Description: Raffray 1904a: 7 (characters in key); Raffray, 1908a: 136 (characters in key), 165
(characters); O. Park 1942: 215 (characters in key), 247 (charcters), 249 (characters).
Key: O. Park 1942: 250 (species of the genus).
Distribution.
Central America: Panama.
South America: Bolivia, Brazil.
0027. Iteticus Raffray, 1904a: 19

Type species: Batrisus princeps Reitter, 1883
10 species are known from and are restricted to Central and South America.
Characters/Description: Raffray 1904a: 5 (characters in key); Raffray 1908a: 135 (characters in key), 155
(characters); O. Park 1942: 215 (characters in key), 246 (characters).

Key: Raffray 1898a: 432 (species of the genus as Batrisus); O. Park 1942: 247 (species of the genus).
Distribution.
South America: Bolivia, Brazil, Chile.
0028. Oxarthrius Reitter, 1883a: 378 (Fig. 28)

Type species: Oxarthrius bispinosus Reitter, 1883
Subgenera:
Baroxarthrius O. Park, 1942: 244 [*]
Type species: Oxarthrius escharus O. Park, 1942
Oxarthrius Reitter, 1883a: 378 [*]
Type species: Oxarthrius bispinosus Reitter, 1883
13 species are known from and are restricted Central and South America.
Characters/Description: Reitter 1888: 248 (characters); Raffray 1894: 230 (characters in key), 261 (characters);
Raffray 1898a: 436 (characters); Raffray 1904a: 4 (characters in key); Raffray 1908a: 134 (characters in key),
153 (characters); O. Park 1942: 215 (characters in key), 240 (characters); Chandler 2002: 52 (characters in
key); Chandler 2013: 324 (characters of Baroxarthrius); Asenjo et al. 2018: 414 (characters).
Key: Raffray 1898a: 437 (species); O. Park 1942: 241 (subgenera), 241 (species of subgenus Oxarthrius).
Distribution.
Central America: Mexico, Guatemala, Costa Rica, Panama.
South America: Peru, Bolivia, Brazil, Paraguay, Argentina.
0029. Syrbatus Reitter, 1882a: 134

Type species: Batrisus clypeatus Reitter, 1882
Subgenera:
Syrbatidius Jeannel, 1952c: 212
Type species: Syrbatus collarti Jeannel, 1950
Syrbatodes Jeannel, 1959: 318 [synonymized by Leleup 1973: 49]
Type species: Syrbatus leleupi Jeannel, 1950
Syrbatus Reitter, 1882a: 134 [*]
Type species: Batrisus clypeatus Reitter, 1882

Characters/Description: Raffray 1894: 231 (characters in key); Raffray 1898a: 443 (characters), 443 (characters in
key); Raffray 1904a: 4 (characters in key); Raffray 1908a: 134 (characters in key); O. Park 1942: 215
(characters in key), 233 (characters); Leleup 1981: 34 (characters).
Key: Reitter 1888: 244 (Brazilian species of the genus); Raffray 1898a: 443 (species of the genus); Raffray 1904a:
15 (species group); O. Park 1942: 233–236 (group of species and species).
Distribution.
South America: Brazil, Paraguay, Argentina.
Other regions: Ethiopian.
0030. Syrmocerus Raffray, 1898a: 459

Type species: Arthmius cervus Raffray, 1898
6 species are known from and are restricted to South America.

Characters/Description: Raffray 1898a: 443 (characters in key), 459 (characters); Raffray 1904a: 4 (characters in
key); Raffray 1908a: 134 (characters in key), 153 (characters); O. Park 1942: 215 (characters in key), 238
(characters); Borgmeier 1954: 201 (characters).
Key: Raffray 1898a: 459 (species of the genus); O. Park 1942: 238 (species of the genus).
Distribution.
South America: Brazil.
Incertae sedis in Subtribe Batrisina Reitter, 1882b
0031. Batriscydmaenus Parker & Owens, 2018: 220

Type species: Batriscydmaenus tishechkini Parker & Owens, 2018
Only one species is known from and is restricted to Central America.

Distribution.
Supertribe Clavigeritae Leach, 1815
Tribe Clavigerini Leach, 1815
Subtribe Clavigerodina L.W. Schaufuss, 1882
0032. Fustiger LeConte, 1866: 109 (Fig. 22)

Type species: Articerus brasiliensis Westwood, 1856
Afrofustiger Jeannel, 1952b: 18 [synonymized by Besuchet 1977: 261]
Type species: Afrofustiger pleuralis Jeannel, 1952
Commatocerus Raffray, 1882: 1 [synonymized by Besuchet 1977: 261]
Type species: Commatocerus elegantulus Raffray, 1882
Corynotellus Jeannel, 1960a: 175 [synonymized by Coulon 1982: 50]
Type species: Corynotellus tubifer Jeannel, 1960
Pseudfustiger Reitter, 1884: 168 [synonymized by Besuchet 1986: 263]
Type species: Articerus stricticornis Reitter, 1883

Characters/Description: L.W. Schaufuss 1882b: 197 (characters); Reitter 1884: 168 (characters in key as
Pseudfustiger and Fustiger), 168 (characters in key as Commatocerus); Raffray 1905: 446 (characters in key
as Commatocerus), 447 (characters in key), 447 (characters in key as Pseudfustiger); Raffray 1908a: 418
(characters in key), 419 (characters in key as Pseudfustiger), 429 (characters), 433 (characters as
Pseudfustiger); O. Park 1942: 353 (characters in key), 353 (characters in key as Pseudfustiger); O. Park 1944:
242 (characters in key); Bierig 1945: 4 (characters), 6 (characters in key); Besuchet 1986: 263 (characters);
Chandler 2002: 49 (characters in key).
Key: L.W. Schaufuss 1882b: 198 (some South American species); O. Park 1944: 251 (Mexican species); O. Park et
al. 1976: 72 (species of the West Indies as Pseudfustiger), 73 (species of the West Indies).
Distribution.
West Indies: Cuba, Jamaica, Haiti, US Virgin Isl., St. Vincent & Grenadines.
Central America: Mexico, Honduras, Costa Rica.
South America: Peru, Bolivia, Brazil, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Oriental, Oceanic.
0033. Neofustiger Bruch, 1929: 157 (Fig. 27)

Type species: Neofustiger cochlearis Bruch, 1929

Distribution.
South America: Argentina.
Supertribe Euplectitae Streubel, 1839
Tribe Bythinoplectini L.W. Schaufuss, 1890
Subtribe Bythinoplectina L.W. Schaufuss, 1890
0034. Besucheteidos Comellini, 1985: 748

Type species: Besucheteidos guaimi Comellini, 1985
Characters/Description: Comellini 1985: 710 (characters in key); Coulon 1989: 188 (characters in key), 189
(characters); Chandler 2002: 54 (characters in key).
Distribution.
Central America: Mexico, Costa Rica, Panama.
South America: Venezuela, Ecuador.
0035. Bythinoplectoides Comellini, 1985: 739

Type species: Bythinoplectoides gigas Comellini, 1985

Distribution.
0036. Bythinoplectus Reitter, 1882b: 209

Type species: Bythinoplectus foveatus Reitter, 1883

Characters/Description: Raffray 1897: 229 (characters); Raffray 1898b: 204 (characters in key); Raffray 1904b:
501 (characters in key); Raffray 1908a: 22 (characters in key), 24 (characters); O. Park 1944: 242 (characters
in key); Becker & Sanderson 1953a: 403 (characters in key), 404 (characters); Comellini 1985: 710
(characters in key), 716 (characters); Coulon 1989: 85 (characters in key), 151 (characters); Chandler 2002:
54 (characters in key).
Key: O. Park 1944: 251 (Mexican species); O. Park 1945a: 278 (species of the genus); O. Park 1952a: 7 (species of
the genus); Becker & Sanderson 1953a: 406 (species of Honduras); O. Park et al. 1976: 5 (species of the West
Indies); Newton & Peck 2006: 133 (species of Galapagos Isl.).
Distribution.
West Indies: Bahamas, Cuba, US Virgin Isl., Grenada, Trinidad-Tobago.

Central America: Mexico, Guatemala, Honduras, Costa Rica, Panama.
South America: Suriname, French Guiana, Galapagos Isl., Peru, Brazil, Paraguay, Argentina.
0037. Dichocoryna Comellini, 1985: 742

Type species: Dichocoryna longipalpis Comellini, 1985
Characters/Description: Comellini 1985: 710 (characters in key), 742 (characters); Coulon 1989: 188 (characters in
key), 191 (characters); Chandler 2001: 75 (characters in key), 76 (characters).
Distribution.
Central America: Costa Rica, Panama.
South America: Venezuela.
Other regions: Oriental, Australian.
0038. Hendecameros Comellini, 1985: 754

Type species: Hendecameros costaricense Comellini, 1985
(characters).
Distribution.
South America: Colombia, Venezuela, French Guiana, Brazil.
0039. Octomeros Comellini, 1985: 714

Type species: Octomeros sulcatum Comellini, 1985
(characters).
Distribution.
0040. Poeciloceras Comellini, 1985: 712

Type species: Poeciloceras portentosum Comellini, 1985

(characters).
Distribution.
South America: Brazil, Paraguay.
0041. Pyxidion Comellini, 1985: 744

Type species: Pyxidion acinaces Comellini, 1985

(characters).
Distribution.
0042. Schizocoryna Comellini, 1985: 740

Type species: Schizocoryna wagneri Comellini, 1985

(characters).
Distribution.
Subtribe Pyxidicerina Raffray, 1904b
0043. Orlandia Comellini, 1985: 758

Type species: Orlandia parki Comellini, 1985

(characters); Chandler 2001: 75 (characters in key), 81 (characters); Chandler 2002: 54 (characters in key).
Distribution.
Tribe Dimerini Raffray, 1908a
0044. Barroeuplectoides O. Park, 1942: 101

Type species: Barroeuplectoides zeteki O. Park, 1942
2 species are known from and are restricted to Central America.

Characters/Description: O. Park 1942: 69 (characters in key); O. Park 1952b: 88 (characters in key); Grigarick &
Schuster 1980: 19 (characters), 54 (characters in key); Coulon 1989: 17 (characters); Chandler 2002: 54
(characters in key).
Distribution.
Central America: Mexico, Guatemala, Panama.
0045. Ocabaraja Schuster & Grigarick, 1980: 39

Type species: Ocabaraja romingeri Schuster & Grigarick, 1980
2 species are known from and are restricted to the West Indies and South America.
Characters/Description: Coulon 1989: 17 (characters).
Distribution.

0046. Tuberoplectus O. Park, 1952b: 96

Type species: Tuberoplectus plaumanni O. Park, 1952
1 species is included in this genus from South America.

Characters/Description: O. Park 1952b: 88 (characters in key); Grigarick & Schuster 1980: 19 (characters), 54
(characters in key); Chandler 2001: 72 (characters); Chandler 2002: 54 (characters in key).
Distribution.
Tribe Euplectini Streubel, 1839
0047. Euplectamecia O. Park, 1952b: 102

Type species: Euplectamecia hidalgensis O. Park, 1952
(characters in key); Chandler 2002: 57 (characters in key).
Distribution.
0048. Euplectus Leach, 1817: 80, 82

Type species: Euplectus reichenbachii Leach, 1817
Diplectellus Reitter, 1909: 207 [synonymized by Besuchet 1974: 317]
Type species: Euplectus afer Reitter, 1881
Diplectidius Jeannel, 1956: 31 [synonymized by Besuchet 1974: 317]
Type species: Euplectus theryi Guillebeau, 1893
Endoplectus Raffray, 1910: 254 [synonymized by Jeannel 1950: 100]
Type species: Euplectus doderoi Reitter, 1884
Euplectellus Reitter, 1909: 207 [synonymized by Jeannel 1950: 100]
Type species: Euplectus hummleri Reitter, 1906
Euplectinus Jeannel, 1956: 31 [synonymized by Jeannel 1959: 145]
Type species: Euplectus afer Reitter, 1881
Euplectoides Jeannel, 1954: 106 [synonymized by Besuchet 1974: 317]
Type species: Euplectoides pearcei Jeannel, 1954
Pareuplectus Jeannel, 1952a: 211 [Löbl & Besuchet 2004: 321]
Type species: Euplectus delamarei Jeannel, 1949
Phthartomerus Sawada, 1956: 102 [synonymized by Nomura 2001: 142]
Type species: Phthartomerus nanus Sawada, 1956
Stanosthetus Dejean, 1821: 25 [synonymized by Newton 2017: 7]
Type species: Pselaphus karstenii Reichenbach, 1816
14 species are included in this genus from Latin America. Euplectus antennatus and Euplectus cordicollis
described by Motschulsky (1855) from Panama are nomina nuda.
Characters/Description: Raffray 1898b: 212 (characters in key), 249 (characters); Raffray 1904b: 527 (characters
in key); Raffray 1908a: 41 (characters in key), 84 (characters); O. Park 1942: 68, 69 (characters in key), 99

(characters); O. Park 1944: 248 (characters in key); O. Park 1952b: 88 (characters in key), 89 (characters); O.
Park et al. 1976: 26 (characters); Grigarick & Schuster 1980: 22 (characters), 53 (characters in key); Chandler
1985: 110 (characters in key); Chandler 2001: 57 (characters in key), 63 (characters); Chandler 2002: 57
Key: Raffray 1904b: 539 (species groups); Raffray 1908a: 85 (subgenera and species groups); O. Park 1942: 100
(Neotropical species of the genus); O. Park 1944: 251 (Mexican species); O. Park 1952b: 89 (Neotropical
species of the genus); O. Park et al. 1976: 26 (species of the West Indies).
Distribution.
West Indies: Guadeloupe, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
Central America: Mexico, Guatemala, Costa Rica.
South America: Colombia, Venezuela, Brazil, Chile.
Other regions: Nearctic, Palaearctic, Ethiopian, Oriental, Australian.
0049. Mitrametopus Raffray, 1911: 50

Type species: Mitracephala longipennis Raffray, 1890
Mitracephala Raffray, 1890b: 107 [synonymized by Raffray 1911: 50]
Type species: Mitracephala longipennis Raffray, 1890
Characters/Description: Raffray 1904b: 587 (characters in key as Mitracephala); Raffray 1908a: 115 (characters in
key as Mitracephala), 128 (characters as Mitracephala); O. Park 1942: 67 (characters in key as
Mitracephala), 83 (characters as Mitracephala).
Distribution.
Tribe Jubini Raffray, 1904b
0050. Arctophysis Reitter, 1882b: 211

Type species: Arctophysis gigantea Reitter, 1883

Characters/Description: Raffray 1893a: 2 (characters in key), 46 (characters); Raffray 1904b: 508 (characters in
key); Raffray 1908a: 26 (characters in key), 28 (characters); O. Park 1942: 40 (characters in key); O. Park
1952a: 15 (characters in key).
Distribution.
South America: Colombia.
0051. Auxenocerus Jeannel, 1962b: 324

Type species: Auxenocerus bispina Jeannel, 1962

Characters/Description: Jeannel 1962b: 321 (characters in key).
Key: Jeannel 1962b: 324 (species of the genus).
Distribution.
0052. Balega Reitter, 1882b: 211

Type species: Balega elegans Reitter, 1883

4 species are known from and are restricted to the West Indies and Central America.
Characters/Description: Raffray 1893a: 3 (characters in key); Raffray 1893c: 187 (characters); Raffray 1904b: 509
(characters in key); Raffray 1908a: 26 (characters in key), 31 (characters); O. Park 1942: 40 (characters in
key); O. Park 1944: 243 (characters in key); O. Park 1952a: 15 (characters in key); Chandler 2002: 55
Key: O. Park 1944: 251 (Mexican species); O. Park et al. 1976: 8 (species of the West Indies).
Distribution.
West Indies: Jamaica, Dominican Republic, US Virgin Isl.
0053. Barrojuba O. Park, 1942: 59

Type species: Barrojuba albertae O. Park, 1942
14 species are known from and are restricted to Latin America.

Characters/Description: Chandler 1983b: 98 (characters); Chandler 1989: 371 (characters); O. Park 1942: 39
(characters in key); O. Park 1952a: 14 (characters in key); Chandler 2002: 55 (characters in key).
Key: Chandler 1983b: 100 (species of the genus); Chandler 1989: 373 (species of the genus).
Distribution.
West Indies: Trinidad-Tobago.
South America: Colombia, Brazil.
0054. Endytocera Sharp, 1887b: 45

Type species: Endytocera vestita Sharp, 1887

key); O. Park 1952a: 14 (characters in key); Chandler 2002: 55 (characters in key).
Distribution.
0055. Germainites Jeannel, 1962b: 330

Type species: Germainites crassus Jeannel, 1962

Characters/Description: Jeannel, 1962b: 321 (characters in key).
Distribution.
0056. Jubomorphus Raffray, 1891: 299

Type species: Jubomorphus simoni Raffray, 1891

Distribution.

0057. Jubus L.W. Schaufuss, 1872: 247
Type species: Jubus spinicollis L.W. Schaufuss, 1877
Duciola Reitter, 1882b: 211 [synonymized by Raffray 1890b: 87]
Type species: Duciola tetratoma Reitter, 1883
Gamba L.W. Schaufuss, 1872: 247 [synonymized by Raffray 1890b: 87]
Type species: Gamba brucki L.W. Schaufuss, 1875
Gasola Reitter, 1882a: 150 [synonymized by Raffray 1890b: 87]
Type species: Gasola simoni Reitter, 1882

key), 48 (characters); O. Park 1944: 243 (characters in key); O. Park 1952a: 15 (characters in key); Chandler
2002: 55 (characters in key).
Key: Raffray 1893c: 159 (some species of the genus); Raffray 1904b: 509 (species groups); O. Park 1942: 48
(group of species); O. Park 1944: 251 (Mexican species); O. Park 1952a: 22 (species of the genus); O. Park et
al. 1976: 12 (species of the West Indies).
Distribution.
West Indies: Guadeloupe, Grenada.
South America: Colombia, Venezuela, Guyana, Suriname, Bolivia, Brazil, Paraguay, Argentina.
0058. Kuscheliotes Jeannel, 1962b: 321

Type species: Kuscheliotes rugosus Jeannel, 1962
6 species is known from and is restricted to South America.

Characters/Description: Jeannel 1962b: 320 (characters in key); Jeannel 1963: 357 (characters).
Distribution.
0059. Macta Raffray, 1890b: 85, 87

Type species: Macta constricta Raffray, 1890

Distribution.
0060. Morphogenia Parker, 2014: 59

Type species: Morphogenia struhli Parker, 2014

Distribution.

0061. Phamisus Aubé, 1844: 94
Type species: Phamisus reichenbachii Aubé, 1844
Canthoderus Motschulsky, 1855: 15 [synonymized by Chandler 2000: 175]
Type species: Canthoderus villosus Motschulsky, 1855
Stratus L.W. Schaufuss, 1872: 246 [synonymized by Chandler 1997: 35]
Type species: Stratus ursinus L.W. Schaufuss, 1877

Characters/Description: Reitter 1888: 235 (characters in key); Raffray 1893a: 3 (characters in key), 3 (characters in
key as Stratus); Raffray 1893c: 188 (characters), 191 (characters as Stratus); Raffray 1904b: 509 (characters
in key as Stratus), 509 (characters in key); Raffray 1908a: 27 (characters in key), 27 (characters in key as
Stratus), 32 (characters), 32 (characters as Stratus); O. Park 1942: 40 (characters in key as Stratus), 40
(characters in key as Phamisus); O. Park 1944: 243 (characters in key as Stratus); O. Park 1952a: 16
(characters in key as Phamisus and Stratus); Chandler 2002: 55 (characters in key).
Key: O. Park 1944: 251 (Mexican species as Stratus); O. Park 1952a: 38 (species of the genus as Stratus), 41
(species of the genus); O. Park et al. 1976: 10 (species of the West Indies as Stratus).
Distribution.
South America: Colombia, Suriname, Bolivia.
0062. Pselaphomorphus Motschulsky, 1855: 15

Type species: Pselaphomorphus sculpturatus Motschulsky, 1855
1952a: 15 (characters in key); Chandler 2002: 55 (characters in key); Vásquez-Vélez 2016: 4 (characters).
Key: Raffray 1893a: 43 (some species of the genus); O. Park 1952a: 16 (species of the genus); Vásquez-Vélez
2016: 8 (species of the genus).
Distribution.
Central America: Mexico, Belize, Honduras, Costa Rica, Panama.
South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Argentina.
0063. Sebaga Raffray, 1891: 300

Type species: Sebaga centralis Raffray, 1891
1944: 243 (characters in key); O. Park 1952a: 15 (characters in key); Chandler 2002: 55 (characters in key).
Key: Raffray 1893a: 50 (some species of the genus); O. Park 1942: 47 (species of the genus); O. Park 1944: 251
(Mexican species); O. Park 1945b: 332 (some species of the genus); O. Park 1952a: 17 (species of the genus).
Distribution.
Central America: Mexico, Guatemala, Honduras, Nicaragua, Panama.
South America: Venezuela, Suriname, Brazil.

Tribe Mayetiini Winkler, 1925
0064. Mayetia Mulsant & Rey, 1875b: 87

Type species: Mayetia sphaerifera Mulsant & Rey, 1875
Subgenera:
Mayetia Mulsant & Rey, 1875b: 87
Type species: Mayetia sphaerifera Mulsant & Rey, 1875
Metamayetia Coiffait, 1962a: 249 [*]
Type species: Mayetia nevesi Jarrige, 1949
Promayetia Coiffait, 1962a: 249
Type species: Mayetia minima Coiffait, 1955

Characters/Description: Coiffait 1955: 9 (characters); Orousset 1995: 57 (characters).
Distribution.
South America: Venezuela, Peru.
Tribe Metopiasini Raffray, 1904b
Subtribe Metopiasina Raffray, 1904b
0065. Barrometopia O. Park, 1942: 210

Type species: Barrometopia quasimoda Park, 1942
Only one species is known from and is restricted to Central and South America.
Characters/Description: O. Park 1942: 205 (characters in key).
Distribution.
0066. Bibrax Fletcher, 1927: 149

Type species: Bibrax bradleyi Fletcher, 1927

Characters/Description: O. Park 1942: 284 (characters in key), 284 (characters); Comellini 1979: 689 (characters in
key).
Distribution.
0067. Chandleria Comellini, 1998: 345

Type species: Chandleria angusta Comellini, 1998
Distribution.

0068. Metopias Gory, 1832: unnumbered page
Type species: Metopias curculionoides Gory, 1832
Marnax Laporte, 1835: 137 [synonymized by Lacordaire 1854: 169]
Type species: Metopias curculionoides Gory, 1832

Characters/Description: Reitter 1885a: 334 (characters); Raffray 1908a: 187 (characters in key and characters); O.
Park 1942: 204 (characters in key); Comellini 1993: 921 (characters).
Distribution.
South America: Suriname, French Guiana.
0069. Metopiasoides Comellini, 2000: 765 (Fig. 24)

Type species: Metopiasoides werneri Comellini, 2000
Characters/Description: Comellini 2000: 765 (characters); Asenjo 2016: 587 (characters).
Distribution.
South America: Ecuador, Peru, Bolivia, Brazil.
0070. Metopiellus Raffray, 1908a: 188 (Fig. 25)

Type species: Metopias hirtus Reitter, 1885

Characters/Description: Raffray 1908a: 187 (characters in key), 188 (characters); O. Park 1942: 205 (characters in
key); Asenjo et al. 2017: 122 (characters).
Key: Asenjo et al. 2017: 122 (species of the genus).
Distribution.
South America: Brazil, Argentina.
0071. Metopiosoma Raffray, 1908a: 188

Type species: Metopias pacificus Westwood, 1856

key); Comellini 1994: 107 (characters).
Distribution.
South America: Venezuela, Brazil, Argentina.
0072. Metopioxys Reitter, 1885a: 334

Type species: Metopioxys gladiator Reitter, 1885
20 are known from and are restricted to South America.

key), 205 (characters); Comellini 1983: 438 (characters).
Key: O. Park 1942: 205 (species of the genus); Comellini 1983: 438 (species group).
Distribution.
South America: Colombia, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Uruguay, Argentina.

Subtribe Rhinoscepsina Bowman, 1934
0073. Rhinoscepsis LeConte in Schwarz, 1878: 382

Type species: Rhinoscepsis bistriatus LeConte in Schwarz, 1878
Subgenera:
Rafrhisis O. Park, 1945a: 283 [*]
Type species: Rhinoscepsis orbis O. Park, 1945
Rhinoscepsis LeConte in Schwarz, 1878: 382 [*]
Type species: Rhinoscepsis bistriatus LeConte in Schwarz, 1878
in key); Raffray 1908a: 36 (characters in key), 55 (characters); O. Park 1942: 68 (characters in key), 85
(characters); O. Park 1944: 248 (characters in key); O. Park 1945a: 282 (characters); Becker & Sanderson
1953a: 403 (characters in key), 412 (characters); Grigarick & Schuster 1980: 17 (characters), 53 (characters in
key); Chandler 2002: 54 (characters in key).
Key: Raffray 1898b: 223 (some species of the genus); O. Park 1942: 86 (species of the genus); O. Park 1944: 251
(Mexican species); O. Park 1945a: 285 (species of the genus); O. Park 1945b: 341 (some species of the
genus); O. Park 1952b: 120 (subgenera and species of Neotropical region).
Distribution.
Central America: Mexico, Honduras, Panama.
South America: Suriname, Brazil, Paraguay, Argentina.
Tribe Trichonychini Reitter, 1882b
Subtribe Panaphantina Jeannel, 1950
0074. Acotreba Reitter, 1882b: 209 (Fig. 20)

Type species: Acotreba simoni Reitter, 1883

Characters/Description: Raffray 1898b: 205 (characters in key); Raffray 1904b: 519 (characters in key); Raffray
1908a: 36 (characters in key); O. Park 1952b: 114 (characters in key).
Distribution.
0075. Allodalminiastes H. Franz, 1996: 104

Type species: Allodalminiastes abnormipenis H. Franz, 1996

Distribution.
0076. Andiotes H. Franz, 1996: 99

Type species: Andiotes malalcahuelloi H. Franz, 1996

Distribution.
0077. Araucaniotes Jeannel, 1962b: 371

Type species: Araucaniotes parallelus Jeannel, 1962

Distribution.
0078. Biblomelba O. Park, 1952b: 106

Type species: Biblomelba profunda O. Park, 1952

Distribution.
0079. Biblomimus Raffray, 1904b: 545

Type species: Biblomimus minutus Raffray, 1904
Characters/Description: Raffray 1904b: 529 (characters in key); O. Park 1942: 66 (characters in key), 93
(characters); O. Park 1952b: 104 (characters in key); Grigarick & Schuster 1980: 34 (characters), 56
Key: O. Park 1942: 93 (species of the genus); O. Park 1952b: 108 (species of the genus); O. Park et al. 1976: 24
(species of the West Indies).
Distribution.
West Indies: Guadeloupe, St. Vincent & Grenadines, Grenada.
Central America: Mexico, Honduras.
0080. Bibloplectopsis Jeannel, 1962b: 343

Type species: Bibloplectopsis chilensis Jeannel, 1962

Distribution.
0081. Bibloplectus Reitter, 1882c: 529

Type species: Pselaphus ambiguus Reichenbach, 1816
Subgenera:
Bibloplectodes Jeannel, 1949: 49
Type species: Euplectus ruficeps LeConte, 1863
Bibloplectus Reitter, 1882c: 529
Type species: Pselaphus ambiguus Reichenbach, 1816
Minitiola Karaman, 1969: 53 [synonymized by Löbl & Besuchet 2004: 291]
Type species: Minitiola eubeae Karaman, 1969

in key); Raffray 1908a: 43 (characters in key), 96 (characters); Grigarick & Schuster 1980: 37 (characters), 55
Distribution.
0082. Caligrua Jeannel, 1962b: 352

Type species: Caligrua thoracica Jeannel, 1962

Distribution.
0083. Caligruacmes Jeannel, 1963: 359

Type species: Caligruacmes convexicollis Jeannel, 1963

Distribution.
0084. Dalminiastes Jeannel, 1962b: 373

Type species: Dalminiastes semirugosus Jeannel, 1962

Distribution.
0085. Dalminiomus Jeannel, 1962b: 378

Type species: Dalminiomus araucanus Jeannel, 1962

Distribution.
0086. Etopias Jeannel, 1962b: 357

Type species: Etopias longulus Jeannel, 1962

Distribution.

0087. Fanoridius H. Franz, 1997: 168
Type species: Faronidius andinus H. Franz, 1996
Faronidius H. Franz, 1996: 106 [synonymized by H. Franz 1997: 168]
Type species: Faronidius andinus H. Franz, 1996

Distribution.
0088. Faronoma Raffray, 1893b: 259 (Fig. 21)

Type species: Rybaxis cavangula Reitter, 1889

Characters/Description: Raffray 1904b: 588 (characters in key); Raffray 1908a: 114 (characters in key), 116
(characters); O. Park 1942: 60 (characters in key), 71 (characters); O. Park 1952b: 55 (characters in key);
Jeannel 1962b: 373 (characters in key).
Distribution.
0089. Frutillariotes Jeannel, 1962b: 381

Type species: Frutillariotes kuscheli Jeannel, 1962

Distribution.
0090. Lioplectus Raffray, 1898b: 260

Type species: Lioplectus nitidus Raffray, 1898

1908a: 43 (characters in key), 99 (characters); O. Park 1942: 67 (characters in key); O. Park 1952b: 88
Key: Raffray 1908b: 64 (species of the genus); O. Park 1942: 98 (species of the genus); O. Park 1952b: 100
(species of the genus).
Distribution.
0091. Mexigaster O. Park, 1952b: 110 (Fig. 26)

Type species: Mexigaster boneti O. Park, 1952

Distribution.

0092. Mildana Jeannel, 1964: 9
Type species: Mildana castrii Jeannel, 1964

Distribution.
0093. Nothoplectus Jeannel, 1963: 360

Type species: Nothoplectus depressus Jeannel, 1963
Distribution.
0094. Platyplectus Jeannel, 1962b: 377

Type species: Platyplectus chilotus Jeannel, 1962

Distribution.
0095. Pseudocaligrua H. Franz, 1996: 96

Type species: Pseudocaligrua edentata H. Franz, 1996

Distribution.
0096. Pteroplectus Raffray, 1898b: 259

Type species: Euplectus grandicornis L.W. Schaufuss, 1880

(characters in key); Jeannel 1962b: 348 (characters in key), 356 (characters).
Distribution.
0097. Thesiastes Casey, 1893: 457

Type species: Euplectus debilis LeConte in Schwarz, 1878

(characters); O. Park 1952b: 88 (characters in key), 93 (characters); Grigarick & Schuster 1980: 36
(characters), 55 (characters in key); Chandler 2002: 57 (characters in key).

Key: Raffray 1904b: 543 (species group); O. Park 1952b: 93 (species of the genus); O. Park et al. 1976: 25 (species
of the West Indies).
Distribution.
West Indies: Jamaica, US Virgin Isl., Grenada.
South America: Guyana, Suriname, Chile.
Other regions: Nearctic, Ethiopian, Oriental, Oceanic.
0098. Thesium Casey, 1884b: 117

Type species: Euplectus cavifrons LeConte, 1863
Apothinus Sharp, 1887b: 35 [synonymized by Brendel & Wickham 1891: 48]
Type species: Apothinus impressifrons Sharp, 1887

Characters/Description: Raffray 1898b: 205 (characters in key as Apothinus), 219 (characters as Apothinus);
Raffray 1904b: 519 (characters in key as Apothinus); Raffray 1908a: 36 (characters in key as Apothinus), 53
(characters as Apothinus); O. Park 1942: 66 (characters in key), 67 (characters), 89 (characters); O. Park 1944:
243 (characters in key); O. Park 1952b: 114 (characters in key), 114 (characters); O. Park et al. 1976: 22
(characters); Grigarick & Schuster 1980: 13 (characters), 54 (characters in key); Chandler 1985: 110
Key: O. Park 1942: 90 (species of the genus); O. Park 1944: 251 (Mexican species); O. Park 1952b: 115 (species of
the genus); O. Park et al. 1976: 22 (species of the West Indies).
Distribution.
West Indies: Jamaica, Trinidad-Tobago.
0099. Trepacmes Jeannel, 1962b: 360

Type species: Trepacmes germaini Jeannel, 1962

Distribution.
0100. Tumucania H. Franz, 1997: 168

Type species: Tucumania nogalitoi H. Franz, 1996
Tucumania H. Franz, 1996: 107 [synonymized by H. Franz 1997: 168]
Type species: Tucumania nogalitoi H. Franz, 1996

Distribution.

Subtribe Trichonychina Reitter, 1882b
0101. Andiplectops Jeannel, 1962b: 358

Type species: Andiplectops temporalis Jeannel, 1962

Distribution.
0102. Chrestomera Jeannel, 1962b: 344

Type species: Chrestomera longicollis Jeannel, 1962

Distribution.
0103. Daliacmes Jeannel, 1962b: 354

Type species: Daliacmes insularis Jeannel, 1962

Key: Jeannel 1962b: 355 (species of the genus); Jeannel 1963: 362 (species of the genus).
Distribution.
0104. Mexiplectus O. Park, 1943: 174

Type species: Mexiplectus emersoni O. Park, 1943

Schuster 1980: 28 (characters), 54 (characters in key); Chandler 2002: 56 (characters in key).
Key: O. Park 1944: 251 (Mexican species).
Distribution.
0105. Oropodes Casey, 1893: 453

Type species: Oropodes orbiceps Casey, 1893

Characters/Description: Raffray 1898b: 211 (characters in key), 246 (characters); Raffray 1908a: 41 (characters in
key), 81 (characters); Grigarick & Schuster 1980: 27 (characters), 54 (characters in key); Chandler 2002: 57
(characters in key); Chandler & Caterino 2011: 428 (characters).
Key: Chandler & Caterino 2011: 432 (males), 434 (females).
Distribution.

0106. Panaramecia O. Park, 1942: 94
Type species: Panaramecia williamsi O. Park, 1942

Characters/Description: O. Park 1942: 67, 71 (characters in key); O. Park 1952b: 88 (characters in key); Grigarick
& Schuster 1980: 29 (characters), 54 (characters in key).
Distribution.
0107. Parapteracmes Jeannel, 1962b: 361

Type species: Parapteracmes kuscheli Jeannel, 1962

Distribution.
0108. Pteracmes Raffray, 1890b: 102

Type species: Pteracmes schaufussi Raffray, 1890

(characters in key); Jeannel 1962b: 348 (characters in key).
Distribution.
0109. Pteracmidius Jeannel, 1962b: 365

Type species: Pteracmidius frontalis Jeannel, 1962

Distribution.
0110. Thesiectus O. Park, 1960a: 8

Type species: Thesiectus probus O. Park, 1960
Only one species is known from and is restricted to West Indies.

Characters/Description: Grigarick & Schuster 1980: 29 (characters), 54 (characters in key).
Distribution.
0111. Tiomomus Jeannel, 1962b: 341

Type species: Tiomomus rostratus Jeannel, 1962

Distribution.
0112. Trichonyx Chaudoir, 1845: 164

Type species: Pselaphus sulcicollis Reichenbach, 1816

(characters); Grigarick & Schuster 1980: 27 (characters), 54 (characters in key).
Distribution.
South America: Suriname, Brazil.
Other regions: Nearctic [adventive], Palaearctic.
0113. Verabarolus O. Park, 1942: 104

Type species: Verabarolus subdendrus O. Park, 1942

Schuster 1980: 29 (characters), 54 (characters in key); Chandler 2002: 56 (characters in key).
Distribution.
Subtribe Trimiina Bowman, 1934
0114. Actiastes Casey, 1897: 561

Type species: Trimium foveicolle LeConte in Schwarz, 1878

Characters/Description: Jeannel 1962b: 333 (characters in key); Grigarick & Schuster 1971: 28 (characters);
Grigarick & Schuster 1980: 38 (characters), 55 (characters in key).
Key: Grigarick & Schuster 1971: 29 (species of the genus).
Distribution.
0115. Actionoma Raffray, 1898b: 235

Type species: Actionoma obesum Raffray, 1898

1908a: 39 (characters in key), 69 (characters); O. Park 1942: 69 (characters in key); O. Park 1944: 250
(characters in key); O. Park 1952b: 124: (characters in key); Chandler 2002: 58 (characters in key).
Distribution.

0116. Actium Casey, 1886b: 201
Type species: Trimium californicum LeConte in Schwarz, 1878
Basolum Casey, 1897: 571 [synonymized by Chandler 1994: 48]
Type species: Trimium impunctatum Brendel, 1865
Proplectus Raffray, 1890b: 107 [synonymized by Raffray 1898b: 232]
Type species: Proplectus decipiens Raffray, 1890
in key); Raffray 1908a: 39 (characters in key), 68 (characters); O. Park 1942: 67, 69 (characters in key), 107
(characters); O. Park 1944: 249, 250 (characters in key); O. Park 1952b: 124: (characters in key); Becker &
Sanderson 1953a: 404 (characters in key), 416 (characters); Jeannel 1962b: 333 (characters in key); Grigarick
& Schuster 1971: 3 (characterrs); Grigarick & Schuster 1980: 36 (characters), 56 (characters in key);
Key: Raffray 1904b: 534 (species group); O. Park 1942: 108 (species group of the genus); O. Park 1944: 251
(Mexican species); O. Park 1952b: 125 (species of the genus); Grigarick & Schuster 1971: 4 (species groups),
5 (species of wawonaensis group), 8 (species of barri group), 10 (species of pacificum group), 14 (species of
angustum group), 17 (species of fastigium group), 24 (species of californicum group); O. Park et al. 1976: 42
Distribution.
West Indies: Jamaica, Puerto Rico.
0117. Allobrox Fletcher, 1928: 205

Type species: Allobrox dampfi Fletcher, 1928
Cutrimia O. Park, 1945b: 336 [synonymized by Grigarick & Schuster 1980: 38]
Type species: Cupila mexicana O. Park, 1945

Characters/Description: O. Park 1942: 68 (characters in key), 71 (characters); O. Park 1944: 250 (characters in
key); O. Park 1952b: 56, 88 (characters in key); Grigarick & Schuster 1980: 38 (characters), 55 (characters in
key); Chandler 1985: 110 (characters in key); Chandler 2002: 57 (characters in key).
Distribution.
0118. Allomelba O. Park, 1954b: 16

Type species: Allomelba antennata O. Park, 1954
Only one species is known from and is restricted to the West Indies.
Distribution.
West Indies: Bahamas.

0119. Allotrimium O. Park, 1943: 187
Type species: Allotrimium michoacanensis O. Park, 1943

Characters/Description: O. Park 1944: 250 (characters in key); O. Park 1952b: 124: (characters in key); Grigarick
& Schuster 1980: 42 (characters), 56 (characters in key); Chandler 1985: 110 (characters in the key); Chandler
Key: O. Park 1944: 251 (Mexican species); Chandler 1985: 110 (some species of the genus).
Distribution.
0120. Dalmomelba O. Park, 1954b: 6

Type species: Dalmomelba cazieri O. Park, 1954
Distribution.
0121. Dalmoplectus Raffray, 1890b: 102

Type species: Dalmodes batrisoides Reitter, 1883

1908a: 37 (characters in key), 60 (characters); O. Park 1942: 70 (characters in key), 111 (characters); O. Park
1952b: 124: (characters in key); O. Park 1954b: 6 (characters in key).
Distribution.
0122. Dalmosanus O. Park, 1952b: 126

Type species: Dalmosanus inoculus O. Park, 1952
Pygmactium Schuster & Grigarick, 1968: 112 [synonymized by Chandler 2000: 169]
Type species: Pygmactium steevesi Schuster & Grigarick, 1968
Triangusella O. Park, 1952b: 132 [synonymized by Grigarick & Schuster 1980: 40]
Type species: Triangusella acuta O. Park, 1952
2 species are included in this genus from the Central America.

Characters/Description: O. Park 1952b: 124: (characters in key as Dalmosanus and Triangusella); O. Park 1954b:
6 (characters in key); Grigarick & Schuster 1980: 36 (characters as Pygmactium), 40 (characters), 56
(characters in key as Pygmactium), 56 (Characters in key); Chandler 2002: 58 (characters in key).
Distribution.
0123. Haasellia J.A. Wagner, 1984: 282

Type species: Haasiella medicina O. Park in O. Park et al., 1976

Haasiella O. Park in O. Park et al., 1976: 35 [synonymized by J.A. Wagner 1984: 282]
Type species: Haasiella medicina O. Park in O. Park et al., 1976
Characters/Description: Grigarick & Schuster 1980: 46 (characters).
Distribution.
West Indies: Puerto Rico.
0124. Hanfordia O. Park, 1960a: 10

Type species: Hanfordia absoluta O. Park, 1960
Distribution.
0125. Hispanisella O. Park in O. Park et al., 1976: 32

Type species: Hispanisella haitiana O. Park in O. Park et al., 1976
2 species are known from and are restricted to the West Indies.
Characters/Description: Grigarick & Schuster 1980: 46 (characters).
Key: O. Park et al. 1976: 32 (species of the West Indies).
Distribution.
West Indies: Haiti.
0126. Latomelba O. Park, 1955: 111

Type species: Latomelba quadrisicca O. Park, 1955
Characters/Description: Grigarick & Schuster 1980: 41 (characters), 55 (characters in key); O. Park et al. 1976: 44
(characters).
Distribution.
0127. Lemelba O. Park, 1953a: 1

Type species: Lemelba davisi O. Park, 1953
2 species are included in this genus from the West Indies.

Distribution.
West Indies: Cuba, Dominican Republic.
0128. Malleoceps O. Park, 1954a: 1

Type species: Malleoceps darlingtoni O. Park, 1954

Distribution.
West Indies: Dominican Republic, Puerto Rico.
0129. Melba Casey, 1897: 565

Type species: Trimium thoracicum Brendel, 1889
Subgenera:
Asymmelba O. Park, 1952b: 140 [*]
Type species: Melba clavata Raffray, 1909
Asymmelba O. Park, 1942: 121 [nomen nudum, validated by O. Park 1952b: 140]
Cismelba O. Park, 1960a: 16 [*]
Type species: Melba trifoveata O. Park, 1960
Frontelba O. Park, 1952b: 140 [*]
Type species: Melba frontalis Raffray, 1908
Frontelba O. Park, 1942: 120 [nomen nudum, validated by O. Park 1952b: 140]
Melba Casey, 1897: 565 [*]
Type species: Trimium thoracicum Brendel, 1889
Rameloidea O. Park, 1942: 121 [*]
Type species: Melba temporalis Raffray, 1909
Vertelba O. Park, 1942: 120 [*]
Type species: Melba caviceps Raffray, 1909

(characters); O. Park 1943: 183 (characters); O. Park 1944: 250 (characters in key); O. Park 1952b: 124:
(characters in key), 134 (characters); O. Park 1954b: 6 (characters in key); O. Park et al. 1976: 45 (characters);
Grigarick & Schuster 1980: 43 (characters), 56 (characters in key); Chandler 1985: 110 (characters in key);
Key: Raffray 1904b: 535 (species group); O. Park 1942: 117 (key to the subgenera), 117 (key to species of the
subgenus Asymmelba), 118 (key to species of the subgenus Frontelba), 119 (key to species of the subgenus
Melba); O. Park 1944: 251 (Mexican species); O. Park 1952b: 134 (subgenera), 135 (species of subgenus
Asymmelba and Frontelba), 136 (species of the subgenus Melba); O. Park et al. 1976: 45 (subgenera of the
West Indies), 46 (species of subgenus Frontelba from the West Indies), 46 (species of subgenus Melba of the
West Indies).
Distribution.
West Indies: Jamaica, US Virgin Isl., Antigua & Barbuda, Guadeloupe, Dominica, Martinique, Grenada.
Central America: Mexico, Guatemala.
South America: French Guiana, Brazil.
0130. Melbamima Raffray, 1909: 17

Type species: Melbamina clavicornis Raffray, 1909

Characters/Description: O. Park 1942: 70 (characters in key), 111 (characters); O. Park 1952b: 123: (characters in
key).
Distribution.
0131. Minibi Grigarick & Schuster, 1980: 40

Type species: Melba zonula O. Park, 1954
Characters/Description: Grigarick & Schuster 1980: 56 (characters in key).
Distribution.
0132. Neactium Jeannel, 1962b: 339

Type species: Pseudoplectus trimiiformis Reitter, 1885

Distribution.
0133. Neodalmus Raffray, 1891: 304

Type species: Neodalmus carinatus Raffray, 1891

1908a: 37 (characters in key), 61 (characters); O. Park 1942: 68 (characters in key), 111 (characters); O. Park
1952b: 123: (characters in key).
Distribution.
0134. Paractium Jeannel, 1962b: 334

Type species: Paractium kuscheli Jeannel, 1962

Distribution.
0135. Perimelba O. Park, 1952b: 141

Type species: Melba granulosa O. Park, 1943

Characters/Description: Grigarick & Schuster 1980: 44 (characters), 55 (characters in key); Chandler 2002: 57

Key: O. Park 1943: 183 (Central American species as subgenus of Melba); O. Park 1952b: 136 (key to species as
subgenus of Melba); O. Park et al. 1976: 47 (species of the West Indies as subgenus of Melba).
Distribution.
0136. Pseudotrimium Raffray, 1898b: 230

Type species: Pseudotrimium microcephalum Raffray, 1898

(characters in key); O. Park 1952b: 125: (characters in key); Chandler 2002: 56 (characters in key).
Distribution.
0137. Quadrelba O. Park, 1952b: 135, 140

Type species: Trimiopsis parmata Reitter, 1883
Quadrelba O. Park, 1942: 120 [nomen nudum, validated by O. Park 1952b: 135, 140]
Characters/Description: O. Park 1952b: 135 (characters in key); O. Park et al. 1976: 44 (characters); Grigarick &
Schuster 1980: 41 (characters), 55 (characters in key).
Key: O. Park 1942: 118 (key to species as subgenus of Melba); O. Park 1952b: 135 (key to species as subgenus of
Melba); O. Park et al. 1976: 44 (species of the West Indies).
Distribution.
West Indies: Puerto Rico, US Virgin Isl.
0138. Ramelbida O. Park, 1942: 112

Type species: Melba quadrifoveata Raffray, 1904
Characters/Description: O. Park 1942: 71 (characters in key); O. Park 1952b: 124: (characters in key).
Distribution.
West Indies: US Virgin Isl.
0139. Sandersonella O. Park in O. Park et al., 1976: 38

Type species: Sandersonella transversa O. Park in O. Park et al., 1976
Characters/Description: O. Park et al. 1976: 29 (characters in key).
Distribution.
West Indies: Haiti.

0140. Tomoplectus Raffray, 1898b: 266
Type species: Tomoplectus cordicollis Raffray, 1898

(characters in key); O. Park 1952b: 88 (characters in key); Grigarick & Schuster 1980: 35 (characters), 56
(characters in key); Chandler 1985: 111 (characters in key); Coulon 1989: 17 (characters); Chandler 2002: 57
Distribution.
0141. Trimioarcus O. Park, 1952b: 130

Type species: Trimioarcus incisurus O. Park, 1952

Characters/Description: ; O. Park 1952b: 125: (characters in key); Grigarick & Schuster 1980: 39 (characters), 56
(characters in key); Chandler 1985: 109 (characters in key); Chandler 2002: 56 (characters in key).
Distribution.
0142. Trimiodina Raffray, 1898b: 231

Type species: Trimium concolor Sharp, 1887

1908a: 38 (characters in key), 66 (characters); O. Park 1942: 70 (characters in key); O. Park 1952b: 124:
(characters in key); Grigarick & Schuster 1980: 41 (characters), 55 (characters in key).
Distribution.
Central America: Guatemala.
0143. Trimiopsis Reitter, 1882a: 149

Type species: Trimiopsis claviceps Reitter, 1882
in key); Raffray 1908a: 39 (characters in key), 70 (characters); O. Park 1942: 69, 70 (characters in key); O.
Park 1944: 250 (characters in key); O. Park 1952b: 125: (characters in key), 129 (characters); Grigarick &
Schuster 1980: 38 (characters), 55 (characters in key); Chandler 2002: 56 (characters in key);
Key: O. Park 1944: 251 (Mexican species); O. Park 1952b: 129 (species of the genus).
Distribution.
South America: Colombia, Venezuela.

0144. Trimiosella Raffray, 1898b: 236
Type species: Trimiopsis anguina Reitter, 1883
(characters); O. Park 1942: 71 (characters in key), 112 (characters); O. Park 1952b: 124: (characters in key).
Distribution.
0145. Trimiovillus O. Park, 1954b: 11

Type species: Trimiovillus bahamicus O. Park, 1954
Distribution.
West Indies: Bahamas, Jamaica.
0146. Zolium Casey, 1897: 562

Type species: Trimiopsis eggersi Reitter, 1883
Distribution.
West Indies: Puerto Rico, US Virgin Isl.
0147. Zonaira Grigarick & Schuster, 1980: 43

Type species: Zonaira trilinea Grigarick & Schuster, 1980

Characters/Description: Grigarick & Schuster 1980: 56 (characters in key); Chandler 1985: 109 (characters in key).
Key: Chandler 1985: 109 (species of the genus).
Distribution.
Incertae sedis in Tribe Trichonychini Reitter, 1882b
0148. Cautinia H. Franz, 1996: 108

Type species: Cautinia brevicornis H. Franz, 1996

Distribution.
0149. Mehuinia H. Franz, 1996: 107

Type species: Mehuinia inexpectata H. Franz, 1996

Distribution.
Tribe Trogastrini Jeannel, 1949
Subtribe Phtegnomina O. Park, 1951
0150. Phtegnomus Raffray, 1890b: 107

Type species: Phtegnomus oberthuri Raffray, 1890

Key: O. Park 1952b: 87 (species of the genus).
Distribution.
South America: Venezuela, Brazil.
Subtribe Rhexiina O. Park, 1951
0151. Rhexius LeConte, 1849: 102

Type species: Rhexius insculptus LeConte, 1849
Subgenera:
Rafrhexius O. Park, 1952b: 75 [*]
Type species: Rhexius elegans Raffray, 1909
Rhexinexus O. Park, 1952b: 85 [*]
Type species: Rhexius giganticus O. Park, 1952
Rhexius LeConte, 1849: 102 [*]
Type species: Rhexius insculptus LeConte, 1849
(characters); O. Park 1942: 67 (characters in key), 82 (characters); O. Park 1944: 248 (characters in key);
Grigarick & Schuster 1980: 21 (characters), 53 (characters in key); Chandler 2002: 55 (characters in key).
Key: O. Park 1942: 83 (species of the genus); O. Park 1945b: 340 (Neotropical species); O. Park 1952b: 75
(subgenera), 76 (species of subgenus Rhexius).
Distribution.
Subtribe Trogastrina Jeannel, 1949
0152. Aboeurhexius O. Park, 1952b: 62

Type species: Aboeurhexius crenulatus O. Park, 1952

Distribution.
0153. Adrogaster Raffray, 1890b: 100

Type species: Adrogaster longipennis Raffray, 1890
Distribution.
0154. Anarmodius Raffray, 1890b: 106

Type species: Trichonyx gibbus L.W. Schaufuss, 1874
(characters); O. Park 1942: 68, 70 (characters in key), 72 (characters); O. Park 1952b: 56 (characters in key).
Key: O. Park 1942: 72 (species of the genus); O. Park 1952b: 61 (species of the genus).
Distribution.
0155. Aporhexius Raffray, 1904b: 593

Type species: Aporhexius pubescens Raffray, 1904
(characters); O. Park 1942: 67 (characters), 68 (characters in key), 73 (characters); O. Park 1952b: 56
Distribution.
South America: Bolivia.
0156. Conoplectus Brendel, 1888: 301

Type species: Euplectus canaliculatus LeConte, 1849
Hexirhexius Grigarick & Schuster, 1980: 20 [synonymized by Carlton 1983: 57]
Type species: Euplectus canaliculatus LeConte, 1850
Prorhexius Raffray, 1890b: 107 [synonymized by Brendel 1892: 12]
Type species: Prorhexius sylvaticus Raffray, 1890

Characters/Description: Grigarick & Schuster 1980: 20 (characters as Hexirhexius), 54 (characters in key as
Hexirhexius); Carlton 1983: 57 (characters); Chandler 2002: 56 (characters in key).
Key: Carlton 1983: 59 (species of the genus).

Distribution.
0157. Eurhexius Sharp, 1887b: 41

Type species: Eurhexius vestitus Sharp, 1887

(characters); O. Park 1942: 67, 68 (characters in key), 74 (characters); O. Park 1952b: 56 (characters in key);
Grigarick & Schuster 1980: 22 (characters), 54 (characters in key).
Key: Raffray 1904b: 588 (species group); O. Park 1942: 75 (group of species), 76–78, 80 (species of the genus); O.
Park 1952b: 66 (species of the genus).
Distribution.
Central America: Guatemala, Panama.
South America: Colombia, Venezuela, Suriname, French Guiana, Bolivia, Brazil, Paraguay, Uruguay, Argentina.
0158. Fletcherexius O. Park, 1942: 72

Type species: Eurhexius macrodactylus Fletcher, 1928

Characters/Description: O. Park 1942: 67 (characters), 70 (characters in key); O. Park 1944: 248 (characters in
key); O. Park 1952b: 56 (characters in key); Grigarick & Schuster 1980: 20 (characters), 54 (characters in
key); Chandler 2002: 56 (characters in key).
Distribution.
0159. Rhexidius Casey, 1887: 478

Type species: Rhexidius granulosus Casey, 1887

(characters); O. Park 1942: 67 (characters); O. Park 1944: 248 (characters in key); O. Park 1952b: 56
(characters in key); Grigarick & Schuster 1980: 20 (characters), 54 (characters in key); Chandler 2002: 56
Key: O. Park 1944: 251 (Mexican species); O. Park 1952b: 63 (species of the genus).
Distribution.
0160. Rhexinia Raffray, 1890b: 106

Type species: Rhexinia angulata Raffray, 1890

(characters); O. Park 1942: 67 (characters), 68 (characters in key), 73 (characters); O. Park 1952b: 56

(characters in key); Grigarick & Schuster 1980: 22 (characters), 53 (characters in key); Chandler 2002: 56
Key: O. Park 1952b: 57 (species of the genus); Grigarick & Schuster 1975: 315 (species of the genus).
Distribution.
West Indies: Guadeloupe.
South America: Colombia, Argentina.
0161. Rhexiola O. Park, 1952b: 57

Type species: Rhexinia mexicana O. Park, 1952

Characters/Description: Grigarick & Schuster 1980: 22 (characters), 53 (characters in key); Chandler 2002: 56
Key: O. Park 1952b: 57 (species of the genus as subgenus of Rhexinia)
Distribution.
0162. Xerhius Raffray, 1891: 305

Type species: Xerhius cordicollis Raffray, 1891
Characters/Description: O. Park 1942: 70 (characters in key); O. Park 1952b: 56 (characters in key).
Distribution.
Incertae sedis in Tribe Trogastrini Jeannel, 1949
0163. Jubogaster Parker & Maruyama, 2013: 371

Type species: Jubogaster towai Parker & Maruyama, 2013

Distribution.
South America: Peru.
Supertribe Faronitae Reitter, 1882b
0164. Golasa Raffray, 1904b: 494

Type species: Sagola microcephala Reitter, 1883

(characters); O. Park 1942: 35 (characters in key); Jeannel 1962b: 308 (characters in key), 308 (characters).
Distribution.

0165. Golasidius Jeannel, 1962b: 311
Type species: Golasidius microps Jeannel, 1962

Distribution.
0166. Golasina Jeannel, 1962b: 311

Type species: Golasina robusta Jeannel, 1962

Key: Jeannel 1962b: 311 (some species of the genus).
Distribution.
0167. Golasites Jeannel, 1962b: 310

Type species: Golasites longicornis Jeannel, 1962

Distribution.
0168. Megarafonus Casey, 1897: 550

Type species: Megarafonus ventralis Casey, 1897
Subgenera:
Megarafonus Casey, 1897: 550
Type species: Megarafonus ventralis Casey, 1897
Nafonus Schuster & Marsh, 1958: 194 [*]
Type species: Megarafonus fundus O. Park, 1943
Nanorafonus Schuster & Marsh, 1958: 192
Type species: Megarafonus parvus Schuster & Marsh, 1958

(characters); O. Park 1944: 242 (characters in key); Chandler 2002: 50 (characters in key).
Distribution.
0169. Prosagola Raffray, 1904b: 494

Type species: Sagola elfridae Reitter, 1885
Chequenia H. Franz, 1996: 124 [synonymized by Hlaváč & Chandler 2005: 86]
Type species: Chequenia cekalovici H. Franz, 1996

(characters); O. Park 1942: 35 (characters in key); Jeannel 1962b: 308 (characters in key), 313 (characters);
Hlaváč & Chandler 2005: 86 (Characters).
Distribution.
0170. Salagosa Raffray, 1904b: 494

Type species: Sagola brevipennis Reitter, 1885

(characters); O. Park 1942: 35 (characters in key); Jeannel 1962b: 308 (characters in key), 315 (characters).
Distribution.
0171. Salagosita H. Franz, 1996: 86

Type species: Salagosita chomei H. Franz, 1996

Distribution.
Supertribe Goniaceritae Reitter, 1882b
Tribe Brachyglutini Raffray, 1904b
Subtribe Baradina O. Park, 1951
0172. Euphalepsus Reitter, 1883a: 378

Type species: Euphalepsus globipennis Reitter, 1883
Barada Raffray, 1891: 314 [synonymized by Chandler 2000: 171]
Type species: Barada mucronata Raffray, 1891
Tetrasemus Jeannel, 1962b: 386 [synonymized by Chandler 2000: 171]
Type species: Tetrasemus delamarei Jeannel, 1962
Characters/Description: Raffray 1904a: 13 (characters in key), 117 (characters in key as Barada); Raffray 1908a:
141 (characters in key), 183 (characters), 198 (characters in key as Barada), 255 (characters as Barada); O.
Park 1942: 215 (characters in key), 123 (characters in key as Barada), 253 (characters); O. Park 1944: 248
Key: Reitter 1883a: 380 (some species of Central and South America); Reitter 1888: 240 (some species of the
genus); Raffray 1904a: 61 (species group); O. Park 1933: 566 (species of the genus); O. Park 1942: 253–256
(groups of species and species); O. Park 1944: 251 (Mexican species).
Distribution.
South America: Colombia, Venezuela, Brazil, Paraguay, Argentina.

0173. Phalespoides Raffray, 1890b: 116
Type species: Euphalepsus punctatissimus Reitter, 1888
Phalepsoides Raffray, 1904a: 104 [synonymized by Newton & Chandler 1989: 39]
Type species: Euphalepsus punctatissimus Reitter, 1888

Characters/Description: Raffray 1904a: 13 (characters in key as Phalepsoides); Raffray 1908a: 140 (characters in
key as Phalepsoides), 184 (characters as Phalepsoides); O. Park 1942: 215 (characters in key as
Phalepsoides), 259 (characters as Phalepsoides).
Distribution.
Subtribe Brachyglutina Raffray, 1904b
0174. Achilia Reitter, 1890: 212

Type species: Bryaxis valdiviensis Reitter, 1883
Byraxis Raffray, 1890b: 123 [synonymized by Reitter 1890: 212]
Type species: Bryaxis valdiviensis Reitter, 1883
Clermontodes Jeannel, 1950: 317 [synonymized by Besuchet 1986: 259]
Type species: Clermontodes insularis Jeannel, 1950
68 species are known from and are restricted to South America. Achilia insularis (Jeannel) described from France
was mislabeled.
Characters/Description: Raffray 1904a: 110 (characters in key as Byraxis), 113 (characters in key); Raffray 1908a:
193 (characters in key as Byraxis), 195 (characters in key), 220 (characters); O. Park 1942: 126 (characters in
key), 154 (characters); O. Park 1944: 241 (characters in key); Jeannel 1962b: 391 (characters in key), 396
(characters); H. Franz 1996: 114 (characters); Kurbatov & Sabella 2015: 304 (characters); Sabella et al. 2017:
120 (charactes of Achilia crassicornis species group), 123 (characters of Achilia tumidifrons group), 127
(characters of Achilia bifossifrons group), 132 (characters of Achilia lobifera group); Kurbatov et al. 2018:
166 (characters of Achilia frontalis species group).
Key: Reitter 1885b: 324 (species of the genus as Bryaxis); Raffray 1904a: 134 (species groups); O. Park 1942: 155
(species groups); Jeannel 1962b: 397 (species groups), 399, 402, 405, 407, 409, 421, 424, 425, 431, 435, 438
(species in the groups).
Distribution.
0175. Achillidia Jeannel, 1962b: 391

Type species: Bryaxis bituberculata Reitter, 1885

Distribution.
0176. Achilliotes Jeannel, 1962b: 445

Type species: Achilia brevicornis Raffray, 1904

Distribution.
0177. Anchylarthron Brendel, 1887: 208

Type species: Decarthron cornutum Brendel, 1865
Verticinotus Brendel, 1888: 300 [synonymized by Newton & Chandler 1989: 45]
Type species: Decarthron cornutum Brendel, 1865

(characters); Becker & Sanderson 1953a: 403 (characters in key); Chandler 2002: 53 (characters in key).
Distribution.
0178. Atacamia H. Franz, 1996: 111

Type species: Atacamia paludosa H. Franz, 1996

Distribution.
0179. Berdura Reitter, 1882b: 206

Type species: Berdura excisula Reitter, 1883
Key: O. Park 1942: 164 (species of the genus); O. Park et al. 1976: 49 (species of the West Indies).
Distribution.
West Indies: Cuba, US Virgin Isl.
0180. Brachygluta Thomson, 1859: 54

Type species: Pselaphus fossulatus Reichenbach, 1816
Subgenera:
Brachygluta Thomson, 1859: 54 [*]
Type species: Pselaphus fossulatus Reichenbach, 1816
Brachyglutina Jeannel, 1950: 286 [synonymized by Karaman 1961: 128]
Type species: Bryaxis globulicollis Mulsant & Rey, 1861
Brachyglutodes Jeannel, 1956: 97 [synonymized by Karaman 1961: 128]
Type species: Bryaxis leprieuri Saulcy, 1876
Dicrobiella Jeannel, 1950: 315 [synonymized by Jeannel 1956: 82]
Type species: Bryaxis dentiventris Saulcy, 1876
Dicrobionoma Jeannel, 1956: 97 [Newton & Chandler 1989: 42]
Type species: Bryaxis caligata Saulcy, 1876

Neobrachygluta Kaszab & Székessy, 1953: 475 [synonymized by Besuchet 1963: 27]
Type species: Brachygluta dudichi Kaszab & Székessy, 1953
Nisa Casey, 1886b: 182
Type species: Bryaxis cavicornis Brendel, 1865

(characters); Chandler 2002: 53 (characters in key); Chandler et al. 2015: 7 (characters).
Key: Raffray 1904a: 150 (species group); Chandler et al. 2015: 8 (species of the genus).
Distribution.
Other regions: Nearctic, Palaearctic, Ethiopian.
0181. Braxyda Raffray, 1904a: 145

Type species: Braxyda hamata Raffray, 1904

(characters); O. Park 1942: 126 (characters in key), 159 (characters); Jeannel 1962b: 388 (characters in key),
388 (characters).
Distribution.
0182. Briaraxis Brendel, 1894: 158

Type species: Briaraxis depressa Brendel, 1894
1 species is included in this genus from Latin America.

(characters); Chandler 1992b: 16 (characters); Chandler 2002: 54 (characters in key).
Distribution.
West Indies: Cuba, Jamaica, US Virgin Isl., Barbados, Grenada, Trinidad-Tobago.
0183. Bryaxina Raffray, 1904a: 140

Type species: Bryaxis fraudatrix L.W. Schaufuss, 1880

Characters/Description: Raffray 1904a: 113, 115 (characters in key); Raffray 1908a: 195 (characters in key), 221
388 (characters).
Key: Raffray 1904a: 141 (species group); O. Park 1942: 158 (species of the genus).
Distribution.
0184. Bryaxinella Jeannel, 1962b: 394

Type species: Bryaxinella nodicornis Jeannel, 1962

Distribution.
0185. Bunoderus Raffray, 1904a: 151

Type species: Bunoderus carinicollis Raffray, 1904
Chandler 1983a: 204 (characters); Chandler 2002: 54 (characters in key).
Key: O. Park 1942: 154 (species of the genus); O. Park 1944: 251 (Mexican species); Chandler 1983a: 205 (species
of the genus, males).
Distribution.
Central America: Mexico, Nicaragua, Panama.
0186. Byraxorites Jeannel, 1962b: 389

Type species: Bryaxorites alticola Jeannel, 1962

Distribution.
0187. Bythinogaster L.W. Schaufuss, 1887: 111

Type species: Bythinogaster simplex L.W. Schaufuss, 1887
(characters); O. Park 1942: 124 (characters in key), 203 (characters); O. Park et al. 1976: 53 (characters).
Distribution.
West Indies: Cuba, Jamaica, "Hispaniola".
0188. Cryptorhinula L.W. Schaufuss, 1887: 150, 163

Type species: Cryphorhinula nodifera L.W. Schaufuss, 1887
Characters/Description: Raffray 1890b: 124 (characters); Raffray 1897: 257 (characters in key); Raffray 1904a:
112 (characters in key); Raffray 1908a: 194 (characters in key), 215 (characters); O. Park 1942: 125
(characters in key), 179 (characters).
Distribution.
0189. Drasinus Raffray, 1904a: 148

Type species: Drasinus binodulus Raffray, 1904

Subgenera:
Drasinus Raffray, 1904a: 148 [*]
Type species: Drasinus binodulus Raffray, 1904
Paradrasinus Newton & Chandler, 1989: 43 [*]
Type species: Drasinus hirsutus Fletcher, 1928
Paradrasinus O. Park, 1942: 182 [nomen nudum, validated by Newton & Chandler 1989: 43]
Becker & Sanderson 1953a: 403 (characters in key); Becker & Sanderson 1953b: 437 (characters); Chandler
Key: O. Park 1942: 181 (species of the genus); O. Park 1944: 251 (Mexican species).
Distribution.
Central America: Mexico, Belize, Honduras, Nicaragua, Costa Rica, Panama.
South America: Suriname, Brazil.
0190. Ectopocerus Raffray, 1904a: 149

Type species: Decarthron verticicornis Reitter, 1885

443 (characters).
Distribution.
0191. Eremomus Raffray, 1904a: 177

Type species: Eremomus obesus Raffray, 1904

Distribution.
0192. Globa Raffray, 1887: 37 (Fig. 23)

Type species: Globa longipes Raffray, 1887
Characters/Description: Raffray 1897: 256 (characters in key), 258 (characters); Raffray 1904a: 110 (characters in
key); Raffray 1908a: 193 (characters in key), 204 (characters); O. Park 1942: 124 (characters in key), 202
Distribution.
South America: Colombia, Venezuela, Bolivia.

0193. Leptachillia Jeannel, 1962b: 446
Type species: Leptachillia laevissima Jeannel, 1962

Distribution.
0194. Mallecoa H. Franz, 1996: 110

Type species: Mallecoa abnormis H. Franz, 1996

Distribution.
0195. Mitona Raffray, 1904a: 129

Type species: Mitona quadraticeps Raffray, 1904

(characters); O. Park 1942: 125 (characters in key), 175 (characters); O. Park 1945b: 394 (characters).
Key: O. Park 1942: 176 (species of the genus); O. Park 1945b: 395 (species of the genus).
Distribution.
South America: Colombia, Venezuela, Bolivia.
0196. Nisaxis Casey, 1886b: 183

Type species: Bryaxis tomentosa Aubé, 1833
Dranisaxa O. Park, 1945b: 391 [synonymized by Chandler 2000: 174]
Type species: Dranisaxa gloydi O. Park, 1945
Distribution.
West Indies: Cayman Is.
0197. Nondulia Newton & Chandler, 1989: 43

Type species: Bryaxis convexa L.W. Schaufuss, 1880
Nodulina Raffray, 1904a: 115 [synonymized by Newton & Chandler 1989: 43]
Type species: Bryaxis convexa L.W. Schaufuss, 1880

Characters/Description: Raffray 1904a: 115 (characters in key as Nondulia); Raffray 1908a: 197 (characters in key
as Nondulia), 232 (characters as Nondulia); O. Park 1942: 125 (characters in key), 153 (characters).
Distribution.
South America: Guyana, Suriname, Bolivia, Brazil.

0198. Panabachia O. Park, 1942: 153
Type species: Bryaxis vulnerata Sharp, 1887
Characters/Description: O. Park 1942: 125 (characters in key); O. Park 1945b: 389 (characters); Chandler 2002: 53
Distribution.
0199. Parachillia H. Franz, 1996: 112

Type species: Parachillia longicornis H. Franz, 1996

Distribution.
0200. Pseudachillia Jeannel, 1963: 367

Type species: Pseudachillia bicolor Jeannel, 1963

Distribution.
0201. Pseudachillidia H. Franz, 1996: 112

Type species: Pseudachillidia andina H. Franz, 1996

Distribution.
0202. Raxybis Raffray, 1908a: 222

Type species: Raxybis nodosa Raffray, 1908

Characters/Description: Raffray 1908a: 195 (characters in key); O. Park 1942: 126 (characters in key), 157
(characters); Jeannel 1962b: 388 (characters in key), 389 (characters).
Distribution.
0203. Reichenbachia Leach, 1826: 451

Type species: Bryaxis juncorum Leach, 1817
Dierobia Thomson, 1859: 54 [synonymized by Reitter 1885c: 68]
Reichenbachius Casey, 1906: 359 [synonymized by Newton & Chandler 1989: 44]

(characters); O. Park 1942: 124 (characters in key), 135 (characters); O. Park 1944: 241, 249, 250 (characters
in key); Becker & Sanderson 1953a: 404 (characters in key), 419 (characters); Chandler 2001: 295 (characters
in key), 349 (characters); Chandler 2002: 54 (characters in key).
Key: Raffray 1904a: 125 (species group); O. Park 1942: 136–149 (group of species and species); O. Park 1944:
251 (Mexican species); O. Park 1945b: 381 (species of Mexico and Guatemala); O. Park 1946a: 501 (species
of the genus); Becker & Sanderson 1953a: 420 (species of Honduras); O. Park et al. 1976: 57 (species of the
West Indies).
Distribution.
West Indies: Cuba, Guadeloupe, St. Vincent & Grenadines, Grenada.
South America: Colombia, Venezuela, Suriname, Peru, Bolivia, Brazil, Paraguay, Argentina.
0204. Rybaxidia Jeannel, 1962b: 392

Type species: Rybaxidia kuscheli Jeannel, 1962

Key: Jeannel, 1962b: 393 (species of the genus).
Distribution.
0205. Scalenarthrus LeConte, 1880: 185

Type species: Scalenarthrus hornii LeConte, 1880
Abryxis Raffray, 1890b: 124 [synonymized by Raffray 1904a: 208]
Type species: Bryaxis denticornis L.W. Schaufuss, 1880
Cylindrembolus L.W. Schaufuss, 1887: 103 [synonymized by Raffray 1904a: 208]
Type species: Cylindrembolus marginalis L.W. Schaufuss, 1887

Characters/Description: Reitter 1882a: 141 (characters in key); Raffray 1897: 257 (characters in key as
Cylindrembolus), 260 (characters as Cylindrembolus); Raffray 1904a: 110 (characters in key); Raffray 1908a:
193 (characters in key), 209 (characters); O. Park 1942: 125 (characters in key), 165 (characters); O. Park
1944: 249 (characters in key); Becker & Sanderson 1953a: 403, 404 (characters in key), 427 (characters);
Key: Raffray 1904a: 125 (species group); O. Park 1943: 191 (some species of the genus); O. Park 1942: 167–170
(species group and species); O. Park 1944: 251 (Mexican species); Becker & Sanderson 1953a: 427 (species
of Honduras); O. Park et al. 1976: 51 (species of the West Indies).
Distribution.
West Indies: Jamaica, Guadeloupe, St. Vincent & Grenadines, Grenada.
Central America: Mexico, Guatemala, Belize, Honduras, Panama.
South America: Bolivia, Brazil, Paraguay.
0206. Silillicus Lucas, 1920: 504

Type species: Phoberus armatus Raffray, 1904
Phoberus Raffray, 1904a: 176 [synonymized by Lucas 1920: 504]
Type species: Phoberus armatus Raffray, 1904

Characters/Description: Raffray 1904a: 115 (characters in key as Phoberus), 176 (characters as Phoberus); Raffray
1908a: 197 (characters in key as Phoberus), 241 (characters as Phoberus); O. Park 1942: 124 (characters in
key as Phoberus), 134 (characters as Phoberus).
Distribution.
0207. Strombopsis Raffray, 1904a: 133

Type species: Strombopsis breviventris Raffray, 1904

(characters); O. Park 1942: 126 (characters in key); O. Park 1944: 241 (characters in key).
Distribution.
0208. Xybarida Raffray, 1897: 260

Type species: Xybarida clavata Raffray, 1897
Characters/Description: Raffray 1897: 257 (characters in key); Raffray 1904a: 112 (characters in key); Raffray
1908a: 195 (characters in key), 217 (characters); O. Park 1942: 126 (characters in key), 160 (characters); O.
Park 1944: 249 (characters in key); Chandler 2002: 53 (characters in key).
Key: O. Park 1942: 160 (species of the genus); O. Park 1944: 251 (Mexican species).
Distribution.
0209. Xybaris Reitter, 1882a: 140

Type species: Xybaris spiniceps Reitter, 1882

Characters/Description: Reitter 1882a: 139 (characters in key); Raffray 1897: 257 (characters in key); Raffray
1904a: 111 (characters in key); Raffray 1908a: 194 (characters in key), 214 (characters); O. Park 1942: 125
Distribution.
Subtribe Decarthrina O. Park, 1951
0210. Decarthron Brendel, 1865: 30

Type species: Bryaxis formiceti LeConte, 1849
Subgenera:
Decarfuss Newton & Chandler, 1989: 5, 44 [*]
Type species: Bryaxis monoceros L.W. Schaufuss, 1882
Decarfuss O. Park, 1942: 194 [nomen nudum, validated by Newton & Chandler 1989: 5, 44]

Decarthron Brendel, 1865: 30 [*]
Type species: Bryaxis formiceti LeConte, 1849

Characters/Description: Reitter 1882a: 139 (characters in key); Raffray 1904a: 116 (characters in key); Raffray
1908a: 198 (characters in key), 249 (characters); O. Park 1942: 124 (characters in key), 185 (characters); O.
Park 1944: 242 (characters in key); Becker & Sanderson 1953a: 403 (characters in key); Becker & Sanderson
1953b: 439 (characters); O. Park 1954b: 19 (characters in key); O. Park et al. 1976: 58 (characters); Chandler
Key: Raffray 1904a: 185 (species group); O. Park 1942: 186 (group of species), 189–194 (species of the genus),
200 (species of Decarthron and Drasinus from Barro Colorado island); O. Park 1944: 230 (Brazilian species),
251 (Mexican species); Becker & Sanderson 1953b: 440 (species of Honduras); O. Park 1958: 69
(cavernicolous species from Eastern of the United States); O. Park et al. 1976: 59 (species of the West Indies).
Distribution.
West Indies: Bahamas, Cuba, Grenada.
South America: Colombia, Venezuela, Guyana, Suriname, Peru, Bolivia, Brazil, Paraguay, Chile, Argentina.
0211. Euteleia Raffray, 1904a: 183

Type species: Bryaxis recens L.W. Schaufuss, 1880
(characters); O. Park 1942: 123 (characters in key), 201 (characters); Chandler 2002: 53 (characters in key).
Distribution.
0212. Raffrayolus Jakobson, 1914: 528

Type species: Itamus laticeps Raffray, 1904
Itamus Raffray, 1904a: 196 [synonymized by Jakobson 1914: 528]
Type species: Itamus laticeps Raffray, 1904

Characters/Description: Raffray 1904a: 117 (characters in key as Itamus); Raffray 1908a: 198 (characters in key as
Itamus), 251 (characters as Itamus); O. Park 1942: 124 (characters in key as Itamus), 201 (characters as
Itamus);
Distribution.
Subtribe Eupseniina O. Park, 1951
0213. Eupsenina Raffray, 1909: 35

Type species: Eupsenina fracticornis Raffray, 1909

Characters/Description: O. Park 1942: 69 (characters in key), 123 (characters in key), 129 (characters); Chandler
Key: O. Park, 1942: 130 (species of the genus).
Distribution.
0214. Eupsenius LeConte, 1849: 90

Type species: Eupsenius glaber LeConte, 1849

Characters/Description: Reitter 1882a: 141 (characters in key); Raffray 1904a: 117 (characters in key); Raffray
1908a: 198 (characters in key), 254 (characters); O. Park 1942: 69, 71 (characters in key), 123 (character in
key), 127 (characters); O. Park 1944: 249 (characters in key); Becker & Sanderson 1953a: 404 (characters in
key), 417 (characters); O. Park 1954b: 19 (characters in key); O. Park et al. 1976: 62 (characters); Chandler
Key: O. Park 1942: 128 (species of the genus); O. Park 1944: 251 (Mexican species); Becker & Sanderson 1953a:
417 (species of Honduras); O. Park et al. 1976: 63 (species of the West Indies).
Distribution.
West Indies: Bahamas, Jamaica, Dominican Republic, US Virgin Isl., Guadeloupe, Grenada.
Subtribe Pselaptina O. Park in O. Park et al., 1976
0215. Caligocara O. Park, 1945b: 397

Type species: Caligocara carinifera O. Park, 1945

Characters/Description: Chandler 2002: 53 (characters in key).
Key: O. Park 1945b: 398 (species of the genus).
Distribution.
0216. Eutrichites LeConte, 1880: 184

Type species: Eutrichites zimmermanni LeConte, 1880
(characters); Carlton 1989: 106 (characters); Carlton & Leschen 1996: 161 (characters); Chandler 2002: 53
Key: Carlton 1989: 106 (species of the genus).
Distribution.
Central America: Mexico, Guatemala, Belize, Honduras, El Salvador, Costa Rica, Panama.
South America: Venezuela, Peru, Bolivia.

0217. Pselaptus LeConte, 1880: 184
Type species: Pselaptus belfragei LeConte, 1880

Characters/Description: Reitter 1882a: 141 (characters in key); Raffray 1897: 257 (characters in key), 259
(characters); Raffray 1904a: 111 (characters in key); Raffray 1908a: 194 (characters in key), 210 (characters);
O. Park 1942: 125 (characters in key), 173 (characters); O. Park 1944: 248 (characters in key); Chandler 2002:
Key: O. Park 1942: 174 (species of the genus); O. Park 1944: 251 (Mexican species); O. Park et al. 1976: 51
Distribution.
West Indies: Cuba, Grenada.
Tribe Goniacerini Reitter, 1882b
0218. Adrocerus Raffray, 1890b: 132

Type species: Adrocerus cavicornis Raffray, 1890

(characters); O. Park 1942: 284 (characters in key); Comellini 1979: 689 (characters in key).
Distribution.
0219. Enneameron Comellini, 1990b: 345

Type species: Enneameron orousseti Comellini, 1990
Distribution.
South America: French Guiana.
0220. Goniaceroides Raffray, 1918: 483

Type species: Goniaceroides umbilicatus Raffray, 1918

Characters/Description: O. Park 1942: 284 (characters in key); Comellini 1979: 689 (characters in key).
Distribution.
South America: Paraguay.
0221. Goniacerus Motschulsky, 1855: 17

Type species: Goniacerus gibbus Motschulsky, 1855
Metopioides L.W. Schaufuss, 1872: 245 [synonymized by L.W. Schaufuss 1877a: 451]
Type species: Metopioides setifer L.W. Schaufuss, 1877

(characters); O. Park 1942: 284 (characters in key), 285 (characters); Comellini 1979: 689 (characters in key);
Comellini 1990a: 224 (characters).
Key: Comellini 1990a: 224 (species of the genus).
Distribution.
South America: Venezuela, French Guiana, Ecuador, Peru, Bolivia, Brazil.
0222. Goniastes Westwood, 1870: 125

Type species: Goniastes sulcifrons Westwood, 1870

(characters); O. Park 1942: 284 (characters in key); Comellini 1979: 682 (characters), 689 (characters in key);
Comellini 1981a: 249 (characters).
Distribution.
South America: French Guiana, Bolivia, Brazil, Paraguay.
0223. Heptameron Comellini, 1979: 687

Type species: Heptameron reticulatum Comellini, 1979

Characters/Description: Comellini 1979: 689 (characters in key).
Distribution.
0224. Listriophorus L.W. Schaufuss, 1872: 245

Type species: Listriophorus felix L.W. Schaufuss, 1874
Comellini 1979: 689 (characters in key); Comellini 1981b: 345 (characters); Chandler 2002: 52 (characters in
key).
Key: O. Park 1944: 251 (Mexican species); Comellini 1981b: 347 (species groups).
Distribution.
0225. Paragoniastes Comellini, 1979: 681 (Fig. 29)

Type species: Paragoniastes raffrayi Comellini, 1979

Characters/Description: Comellini 1979: 682 (characters), 689 (characters in key); Cuccodoro et al. 2012: 2
(characters).
Key: Cuccodoro et al. 2012: 2 (species of the genus).
Distribution.

Tribe Iniocyphini O. Park, 1951
Subtribe Iniocyphina O. Park, 1951
0226. Anoplobraxis O. Park, 1942: 277

Type species: Anoplobraxis guianensis Park, 1942

Distribution.
South America: Guyana.
0227. Batriphysis O. Park, 1942: 270

Type species: Batriphysis epicranis O. Park, 1942

Distribution.
0228. Batrisobryaxis L.W. Schaufuss, 1887: 148

Type species: Batrisobryaxis labialis L.W. Schaufuss, 1887
Insulomodes O. Park in O. Park et al., 1976: 13 [synonymized by Chandler 2002: 96]
Type species: Insulomodes tobagoensis O. Park in O. Park et al., 1976
Characters/Description: Chandler 2002: 52 (characters in key).
Key: O. Park et al. 1976: 14 (species of the West Indies of Insulomodes).
Distribution.
West Indies: Grenada, Trinidad-Tobago.
0229. Bythinophysis Raffray, 1908a: 266

Type species: Bythinophysis punctipennis Raffray, 1908

(characters); O. Park 1944: 249 (characters in key); Chandler 2002: 52 (characters in key).
Key: O. Park 1942: 263 (species of the genus); O. Park 1943: 205 (species of the genus); O. Park 1944: 253
(Mexican species).
Distribution.
0230. Dalmoburis O. Park, 1942: 266

Type species: Dalmoburis petrunkevitchii O. Park, 1942

Characters/Description: O. Park 1942: 261 (characters in key); Chandler 2002: 52 (characters in key).
Distribution.
0231. Dalmodes Reitter, 1882b: 210

Type species: Dalmodes rybaxides Reitter, 1883
Buris Fletcher, 1928: 227 [synonymized by Chandler 2002: 96]
Type species: Buris brunneus Fletcher, 1928

(characters); O. Park 1942: 261 (characters in key), 262 (characters), 261 (characters in key as Buris), 266
(characters as Buris); O. Park 1944: 249 (characters in key); Chandler 2002: 52 (characters in key).
Key: O. Park 1942: 262 (species of the genus), 269 (species of Buris); O. Park 1943: 200 (species of Buris); O.
Park 1944: 251 (Mexican species of Dalmodes and Buris).
Distribution.
West Indies: Antigua & Barbuda, Trinidad-Tobago.
0232. Dalmomima Raffray, 1908a: 270

Type species: Batrybraxis caviceps Raffray, 1896

(characters).
Distribution.
0233. Dalmonexus O. Park, 1942: 272

Type species: Dalmonexus seeversi O. Park, 1942
Distribution.
0234. Dalmophysis Raffray, 1897: 267

Type species: Dalmophysis cylindrica Raffray, 1897

Distribution.

0235. Iniocyphus Raffray, 1912: 435
Type species: Iniocyphus iheringi Raffray, 1912

Characters/Description: O. Park 1942: 262 (characters in key), 277 (characters).
Distribution.
0236. Phybytharsis O. Park, 1942: 274

Type species: Phybytharsis gambosis O. Park, 1942

Distribution.
0237. Trimicerus Motschulsky, 1855: 16

Type species: Trimicerus pacificus Motschulsky, 1855
Batrybraxis Reitter, 1882a: 141 [synonymized by Chandler 2000: 178]
Type species: Batrybraxis curtula Reitter, 1882
Characters/Description: Reitter 1882a: 141 (characters in key as Batrybraxis); Raffray 1904a: 257 (characters in
key as Batrybraxis); Raffray 1908a: 259 (characters in key as Batrybraxis), 270 (characters as Batrybraxis);
O. Park 1942: 262 (characters in key as Batrybraxis), 279 (characters as Batrybraxis); O. Park 1944: 249
(characters in key as Batrybraxis); Chandler 2002: 52 (characters in key).
Key: O. Park 1944: 251 (Mexican species of Batrybraxis).
Distribution.
Tribe Proterini Jeannel, 1949
0238. Euharmophola O. Park, 1960a: 23

Type species: Euharmophola carinata O. Park, 1960
Distribution.
0239. Harmomima Raffray, 1904a: 269

Type species: Harmomima grandiceps Raffray, 1904


Distribution.
0240. Harmophola Raffray, 1896b: 130

Type species: Harmophola clavata Raffray, 1896
Distribution.
0241. Rhamophorus Newton & Chandler, 1989: 50

Type species: Harmophorus manticoroides L.W. Schaufuss, 1886
Harmophorus L.W. Schaufuss, 1886: 264 [synonymized by Newton & Chandler 1989: 50]
Type species: Harmophorus manticoroides L.W. Schaufuss, 1886

Characters/Description: Raffray 1904a: 257 (characters in key as Harmophorus); Raffray 1908a: 259 (characters in
key as Harmophorus), 269 (characters as Harmophorus); O. Park 1942: 261 (characters in key as
Harmophorus), 264 (characters as Harmophorus).
Distribution.
Supertribe Pselaphitae Latreille, 1802
Tribe Arhytodini Raffray, 1890b
0242. Caccoplectinus Chandler & Wolda, 1986: 521

Type species: Caccoplectinus afoveatus Chandler & Wolda, 1886

Key: Chandler & Wolda 1986: 480 (species of the genus).
Distribution.
0243. Caccoplectus Sharp, 1887b: 22

Type species: Caccoplectus celatus Sharp, 1887
34 species (1 in Mexican amber) are included in this genus from Latin America.
Characters/Description: Raffray 1904a: 316 (characters in key); Raffray 1908a: 316 (characters); O. Park 1944:
242 (characters in key); Chandler & Wolda 1986: 480 (characters); Chandler 2002: 50 (characters in key).
Key: O. Park 1944: 251 (Mexican species); Chandler 1976a: 304 (species of the genus); Chandler 1976b: 154
(species of the genus); Chandler & Wolda 1986: 480 (species of the genus).
Distribution.
West Indies: Cuba, Jamaica, Trinidad-Tobago.

South America: French Guiana, Ecuador, Brazil.
0244. Pachacuti Besuchet, 1987: 231

Type species: Pachacuti huggerti Besuchet, 1987

Distribution.
South America: Ecuador, Bolivia.
0245. Rhytus Westwood, 1870: 126

Type species: Rhytus vestitus Westwood, 1870
Arhytodes Reitter, 1882b: 209 [synonymized by Newton & Chandler 1989: 7, 54]
Type species: Rhytus vestitus Westwood, 1870
Characters/Description: Raffray 1908a: 359 (characters in key), 412 (characters as Arhytodes); Chandler 1992a:
339 (characters); Chandler 2002: 50 (characters in key).
Key: Raffray 1905: 411 (species group); Raffray 1909: 47 (species of the genus as Arhytodes); O. Park 1942: 344
(species of the genus as Arhytodes); Chandler 1992a: 340 (species of Panama).
Distribution.
South America: Venezuela, Peru, Bolivia, Brazil, Argentina.
0246. Woldenka Chandler, 1992a: 342

Type species: Woldenka barroensis Chandler, 1992

Distribution.
Tribe Attapseniini Bruch, 1933c
0247. Attapsenius Bruch, 1933c: 27

Type species: Attapsenius chernosvitovi Bruch, 1933

Characters/Description: O. Park 1942: 348 (characters).
Distribution.
Tribe Ctenistini Blanchard, 1845
0248. Ctenisis Raffray, 1890b: 143

Type species: Desimia dispar Sharp, 1887
Characters/Description: Raffray 1897: 271 (characters); Raffray 1904a: 337 (characters in key); Raffray 1908a:
330 (characters in key), 347 (characters); O. Park 1942: 291 (characters in key), 292 (characters); O. Park
1944: 243 (characters in key); Becker & Sanderson 1953a: 403 (characters in key); Becker & Sanderson
1953b: 457 (characters); Chandler 2002: 51 (characters in key).
Key: Raffray 1897: 273 (some species of the genus); O. Park 1942: 292 (species of the genus); O. Park 1944: 251
(Mexican species).
Distribution.
Central America: Mexico, Guatemala, Honduras, Nicaragua, Panama.
South America: Colombia, Brazil, Argentina.
0249. Ctenisodes Raffray, 1897: 274

Type species: Ctenisodes laticeps Raffray, 1897
Pilopius Casey, 1897: 617 [synonymized by Chandler 2000: 168]
Type species: Pilopius lacustris Casey, 1897
4 species are included in this genus from Latin America. The record of Ctenisodes zimmermani (LeConte) for
Colombia and Brazil are doubtful.
Characters/Description: Raffray 1904a: 337 (characters in key); Raffray 1908a: 329 (characters in key as Pilopius),
336 (characters as Pilopius), 346 (characters); O. Park 1942: 291 (characters in key as Pilopius), 292
(characters in key), 293 (characters as Pilopius), 294 (characters); O. Park 1944: 243 (characters in key as
Ctenisodes and Pilopius); Chandler 2002: 51 (characters in key).
Key: O. Park 1944: 251 (Mexican species of Ctenisodes and Pilopius).
Distribution.
West Indies: Cuba.
0250. Ctenistes Reichenbach, 1816: 75

Type species: Ctenistes palpalis Reichenbach, 1816
Subgenera:
Ctenistes Reichenbach, 1816: 75
Type species: Ctenistes palpalis Reichenbach, 1816
Dionyx LePeletier & Serville in Latreille et al., 1825: 221 [synonymized by Aubé 1833: 17]
Type species: Dionyx dejeanii LePeletier & Serville, 1825
Leptoctenistes Jeannel, 1956: 171 [synonymized by Besuchet in Löbl and Smetana 2004: 29]
Type species: Ctenistes wittmeri Jeannel, 1956
Tecnesis Peyerimhoff, 1925: 59
Type species: Ctenistes vaulogeri Pic, 1904

(characters).
Key: Raffray 1904a: 339 (species group).

Distribution.
Other regions: Palaearctic, Ethiopian, Oriental.
Tribe Phalepsini Jeannel, 1949
0251. Phalepsus Westwood, 1870: 131

Type species: Phalepsus subglobosus Westwood, 1870
Characters/Description: Reitter 1888: 242 (characters); Raffray 1904a: 372 (characters in key); Raffray 1908a: 360
(characters in key), 407 (characters); O. Park 1942: 296 (characters in key), 297 (characters); Chandler 2002:
Distribution.
Tribe Pselaphini Latreille, 1802
0252. Neopselaphus Jeannel, 1951: 8

Type species: Pselaphus bizonatus L.W. Schaufuss, 1886

Characters/Description: Besuchet 1982: 798 (characters); Chandler 2002: 50 (characters in key).
Key: Besuchet 1982: 798 (species of the genus).
Distribution.
South America: Colombia, Suriname, French Guiana, Brazil, Bolivia.
0253. Pselaphellus Raffray, 1908a: 307

Type species: Pselaphus opacus L.W. Schaufuss, 1886
key), 287 (characters); Chandler 2002: 51 (characters in key).
Distribution.
0254. Pselaphus Herbst, 1792: 106

Type species: Pselaphus heisei Herbst, 1792
Faradayus Reitter, 1909: 217 [synonymized by Jeannel 1951: 6]
Type species: Pselaphus banghaasi Reitter, 1893
Pselaphinus Jeannel, 1951: 10 [synonymized by Newton & Chandler 1989: 63]

Type species: Pselaphus pilipalpis Reitter, 1883
Pselaphoptrus Reitter, 1891a: 139 [synonymized by Besuchet 1999: 58]
Type species: Pselaphoptrus kubischteki Reitter, 1891
Pselaphoxys Raffray, 1890b: 139 [synonymized by Besuchet 1999: 58]
Type species: Pselaphus delicatulus Raffray, 1882
1 species is included in this genus from the West Indies.

Characters/Description: Raffray 1904a: 317 (characters in key); Raffray 1908a: 304 (characters in key and
characters); O. Park 1942: 286 (characters in key), 286 (characters); Chandler 2002: 51 (characters in key).
Key: Raffray 1904a: 318 (species group).
Distribution.
Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental.
Tribe Tyrini Reitter, 1882b
Subtribe Somatipionina Jeannel, 1949
0255. Aploderina Raffray, 1904a: 386

Type species: Aploderina sulcicornis Raffray, 1904

(characters); O. Park 1942: 296 (characters in key), 332 (characters); Hlaváč & Chandler 2005: 91 (characters
in key).
Distribution.
0256. Cercoceroides Raffray, 1896a: 137

Type species: Cercocerus germaini Raffray, 1890
(characters); O. Park 1942: 297 (characters in key), 334 (characters); Becker & Sanderson 1953a: 403
(characters in key); Becker & Sanderson 1953b: 460 (characters); Hlaváč & Chandler 2005: 90 (characters in
key).
Key: Raffray 1905: 409 (species of the genus); O. Park 1942: 334 (species of the genus).
Distribution.
Central America: Honduras.
South America: Venezuela, Brazil, Paraguay.
0257. Cercoceropsis Raffray, 1904a: 370

Type species: Cercoceropsis longipes Raffray, 1904

in key).

Distribution.
0258. Cercocerulus Raffray, 1905: 411

Type species: Phalepsus hirsutus L.W. Schaufuss, 1886

in key).
Distribution.
0259. Circocerus Motschulsky, 1855: 16

Type species: Circocerus perplexus Motschulsky, 1855
Apharus Reitter, 1882a: 129 [synonymized by Chandler 2013: 324]
Type species: Apharus muelleri Reitter, 1882

Characters/Description: Raffray 1904a: 370 (characters in key as Apharus); Raffray 1908a: 358 (characters in key
as Apharus), 395 (characters as Apharus); O. Park 1942: 297 (characters in key as Apharus), 305 (characters
as Apharus); Becker & Sanderson 1953a: 403 (characters in key as Apharus); Becker & Sanderson 1953b:
459 (characters as Apharus); Borgmeier 1954: 208 (characters as Apharus); Chandler 2002: 51 (characters in
key), 51 (chacters in key as Apharus); Hlaváč & Chandler 2005: 91 (characters in key); Chandler 2013: 324
(characters).
Key: O. Park 1942: 306 (species of Apharus).
Distribution.
Central America: Honduras, Costa Rica, Panama.
South America: Colombia, Venezuela, Brazil.
0260. Ephimia Reitter, 1882b: 203

Type species: Ephimia simoni Reitter, 1883
Juxtahamotopsis O. Park, 1942: 303 [synonymized by O. Park et al. 1976: 66]
Type species: Juxtahamotopsis bardeni Park, 1942
(characters); O. Park 1942: 289 (characters), 296 (characters in key as Juxtahamotopsis); O. Park et al. 1976:
66 (characters); Chandler 2002: 51 (characters in key); Hlaváč & Chandler 2005: 90 (characters in key).
Key: O. Park 1942: 290 (species of the genus); O. Park et al. 1976: 66 (species of the West Indies).
Distribution.
West Indies: Cuba, US Virgin Isl., Grenada.
Central America: Nicaragua, Panama.

0261. Hamotidius Jeannel, 1962b: 476
Type species: Hamotidius hirtellus Jeannel, 1962

Characters/Description: Jeannel 1962b: 473 (characters in key); Hlaváč & Chandler 2005: 90 (characters in key).
Distribution.
0262. Hamotocellus Raffray, 1912: 444

Type species: Hamotocellus hirsutus Raffray, 1912

Characters/Description: O. Park 1942: 296 (characters in key); O. Park 1946b: 448 (characters); Borgmeier 1954:
204 (characters); Hlaváč & Chandler 2005: 91 (characters in key).
Distribution.
0263. Hamotus Aubé, 1844: 91

Type species: Hamotus latericius Aubé, 1844
Subgenera:
Hamotoides L.W. Schaufuss, 1888: 298 [*]
Type species: Hamotus tritomus Reitter, 1882
Hamotus Aubé, 1844: 91 [*]
Type species: Hamotus latericius Aubé, 1844
Mothaus Jeannel, 1962b: 477 [synonymized by Hlaváč & Chandler 2005: 86]
Type species: Mothaus delamarei Jeannel, 1962

Characters/Description: Reitter 1888: 225 (characters), 235 (characters in key); Raffray 1904a: 370 (characters in
key); Raffray 1904a: 388 (characters); Raffray 1905: 405 (characters of the subgenus Hamotoides); Raffray
1908a: 359 (characters in key), 359 (characters in key of the subgenus Hamotoides), 397 (characters), 398
(characters of the subgenus Hamotoides); O. Park 1942: 297 (characters in key as subgenera Hamotoides and
Hamotus), 308 (characters); O. Park 1944: 243 (characters in key); Jeannel 1962b: 473 (characters in key as
Hamotus and Mothaus); O. Park et al. 1976: 71 (characters); Chandler 2002: 51 (characters in key); Hlaváč &
Chandler 2005: 86 (characters), 90 (characters in key for subgenus Hamotus), 91 (characters in key for
subgenus Hamotoides).
Key: Reitter 1883a: 374 (some species of Central and South America); Reitter 1888: 230 (some species of the
genus); Raffray 1904a: 388 (species of the genus); Raffray 1905: 405 (species of the subgenus Hamotoides);
O. Park 1942: 311 (subgenera), 311 (subgeneric groups of Hamotus), 312 (subgeneric groups of Hamotoides),
312–319 (species of the genus); O. Park 1944: 251 (Mexican species).
Distribution.
West Indies: Montserrat, Guadeloupe, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
Central America: Mexico, Guatemala, Belize, Nicaragua, Costa Rica, Panama.
South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Chile,
Argentina.

0264. Motschtyrus Chandler, 2000: 173
Type species: Tychus pilosus Motschulsky, 1855

Characters/Description: Hlaváč & Chandler 2005: 90 (characters in key).
Distribution.
0265. Phamisulus Reitter, 1888: 234

Type species: Phamisulus hetschkoi Reitter, 1888
Neophamisus Reitter, 1888: 234 [synonymized by Raffray 1890b: 158]
Type species: Phamisus horroris L.W. Schaufuss, 1886

Characters/Description: Reitter 1888: 235 (characters), 235 (characters in key, Neophamisus); Raffray 1904a: 370
key), 332 (characters); Hlaváč & Chandler 2005: 91 (characters in key).
Distribution.
0266. Pselaphocompsus Raffray, 1908a: 402

Type species: Pselaphocompsus punctatus Raffray, 1908

Characters/Description: Raffray 1908a: 359 (characters in key); Raffray 1912: 447 (characters); O. Park 1942: 297
(characters in key), 337 (characters); Hlaváč & Chandler 2005: 90 (characters in key).
Distribution.
0267. Pseudohamotus Raffray, 1890b: 154

Type species: Hamotus conjunctus Reitter, 1883
in key).
Distribution.
Central America: Guatemala, Nicaragua, Panama.
Subtribe Tyrina Reitter, 1882b
0268. Lethenomus Raffray, 1895a: 396

Type species: Aplodea villosa L.W. Schaufuss, 1886
Hualpenia H. Franz, 1996: 124 [synonymized by Hlaváč & Chandler 2005: 86]
Type species: Hualpenia relicta H. Franz, 1996

470 (characters); Hlaváč & Chandler 2005: 86 (characters), 93 (characters in key).
Distribution.
0269. Neotyropsis H. Franz, 1996: 131

Type species: Neotyropsis chomei H. Franz, 1996

Characters/Description: Hlaváč & Chandler 2005: 93 (characters in key).
Distribution.
0270. Neotyrus Raffray, 1895a: 396

Type species: Aplodea gibbicollis L.W. Schaufuss, 1886

(characters); O. Park 1942: 296 (characters in key), 299 (characters); Borgmeier 1954: 212 (characters);
Jeannel 1962b: 449 (characters in key); Hlaváč & Chandler 2005: 93 (characters in key).
Distribution.
South America: Venezuela, Brazil, Argentina.
0271. Pseudotyropsis H. Franz, 1996: 130

Type species: Pseudotyropsis pseudopalpalis H. Franz, 1996

Distribution.
0272. Termitotyrus Borgmeier, 1954: 209

Type species: Termitotyrus palpalis Borgmeier, 1954
Palodea Jeannel, 1962b: 450 [synonymized by Hlaváč & Chandler 2005: 86]
Type species: Aplodea hirta Reitter, 1888

Characters/Description: Jeannel 1962b: 450 (characters in key as Palodea); Hlaváč & Chandler 2005: 86
(characters), 94 (characters in key).
Key: Jeannel 1962b: 451 (species of the genus as Palodea).
Distribution.

0273. Tyrogatunus O. Park, 1942: 338
Type species: Tyrogatunus zeteki O. Park, 1942

Characters/Description: O. Park 1942: 296 (characters in key); Hlaváč & Chandler 2005: 93 (characters in key).
Distribution.
0274. Tyropsidius H. Franz, 1996: 132

Type species: Tyropsidius mochae H. Franz, 1996

Distribution.
0275. Tyropsis Saulcy, 1874: 80

Type species: Tyropsis chevrolati Saulcy, 1874
Aplodea Reitter, 1882b: 203 [synonymized by Raffray 1890a: 265]
Type species: Pselaphus castaneus Blanchard, 1851
Mochanus H. Franz, 1996: 125 [synonymized by Hlaváč & Chandler 2005: 87]
Type species: Mochanus insulanus H. Franz, 1996
Paratyropsis H. Franz, 1996: 128 [synonymized by Hlaváč & Chandler 2005: 87]
Type species: Paratyropsis spinosissima H. Franz, 1996
452 (characters); Hlaváč & Chandler 2005: 86 (characters), 94 (characters in key).
Key: Reitter 1885b: 321 (Chilean species as Aplodea); Reitter 1887b: 323 (some species as Aplodea); Raffray
1904a: 385 (species groups); O. Park 1942: 299 (species groups); Jeannel 1962b: 453 (species groups), 453,
455, 459, 462, 468 (species in the groups).
Distribution.
Subfamily PHLOEOCHARINAE Erichson, 1839b
0276. Charhyphus Sharp, 1887a: 709 (Fig. 30)

Type species: Charhyphus brevicollis Sharp, 1887
Chapmania Bernhauer, 1933a: 121 [synonymized by Blackwelder 1952: 351]
Type species: Chapmania paradoxa Bernhauer, 1933
Pseudeleusis Bernhauer, 1923b: 63 [synonymized by Herman 1972b: 4]
Type species: Hypotelus picipennis LeConte, 1863
Siberia Blackwelder, 1952: 351 [synonymized by Herman 1972b: 4]
Type species: Chapmania paradoxa Bernhauer, 1933
Triga Fauvel, 1878b: 182 [synonymized by Scheerpeltz 1933: 996]
Trigites Handlirsch, 1907: 731 [synonymized by Herman 1972b: 4]

Characters/Description: Herman, 1972b: 4 (characters).
Key: Herman, 1972b: 4 (species of the genus).
Distribution.
0277. Ecbletus Sharp, 1887a: 708 (Fig. 31)

Type species: Ecbletus simplex Sharp, 1887

Characters/Description: Moore 1965: 44 (characters).
Distribution.
Subfamily TACHYPORINAE W.S. MacLeay, 1825
Tribe Mycetoporini Thomson, 1859
0278. Bryoporus Kraatz, 1857a: 452

Type species: Tachinus cernuus Gravenhorst, 1806
Characters/Description: Blackwelder 1943: 509 (characters in key), 526 (characters); Campbell 1993a: 6
(characters in key), 8 (characters); Navarrete-Heredia et al. 2002: 106 (characters in key); Newton & Peck
Key: Blackwelder 1943: 527 (species of the West Indies); Campbell 1993a: 6 (some species of the genus).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl. (St. Thomas), Grenada,
Trinidad-Tobago.
South America: Colombia, Venezuela, Galapagos Isl., Brazil, Paraguay, Argentina.
0279. Carphacis Gozis, 1886: 14

Type species: Staphylinus striatus Olivier, 1795

Characters/Description: Campbell 1980: 936 (characters); Navarrete-Heredia et al. 2002: 106 (characters in key).
Distribution.
0280. Ischnosoma Stephens, 1829a: 22

Type species: Tachinus splendidus Gravenhorst, 1806
Leichotes Gistel, 1834: 9 [synonymized by Campbell 1991a: 74]

Myteroxis Gozis, 1886: 14 [synonymized by Ganglbauer 1895: 367]
Ischnosomata Strand, 1935: 293 [synonymized by Campbell 1991a: 75]

Characters/Description: Campbell 1991a: 74 (characters), 9 (characters in key); Navarrete-Heredia et al. 2002: 106
Key: Campbell 1991a: 78 (species group and species of North and Central America).
Distribution.
Other regions: Nearctic, Palearctic, Ethiopian, Oriental.
0281. Lordithon Thomson, 1859: 47 (Figs. 34–35)

Type species: Oxyporus pygmaeus Fabricius, 1777
Bobitobus Tottenham, 1939a: 225 [synonymized by Schülke 2009: 832]
Type species: Staphylinus lunulatus Linnaeus, 1760
Characters/Description: Campbell 1982: 10 (characters); Navarrete-Heredia et al. 2002: 107 (characters in key);
Schülke 2009: 832 (characters).
Key: Campbell 1982: 13 (subgenera and species of the North and Central America); Coiffait & Sáiz 1968: 403
(Chilean species as Bolitobius).
Distribution.
0282. Mycetoporus Mannerheim, 1830: 62

Type species: Tachinus punctus Gravenhorst, 1806
Schinomosa Tottenham, 1939a: 226 [synonymized by Campbell 1991a: 9]
Type species: Tachinus punctus Gravenhorst, 1806

Characters/Description: Campbell 1991a: 9 (characters), 9 (characters in key); Navarrete-Heredia et al. 2002: 106
Key: Campbell 1991a: 13 (species group and species of Mexico and North America).
Distribution.
Tribe Tachyporini W.S. MacLeay, 1825
0283. Cilea Jacquelin du Val, 1856: 25 (Fig. 32)

Type species: Staphylinus silphoides Linnaeus, 1767
Astictus Thomson, 1858: 36 [synonymized by Thomson 1861: 153]

Leucoparyphus Kraatz, 1857a: 393 [synonymized by Thomson 1861: 153]
1 species is included in this genus from the West Indies and South America.
Characters/Description: Blackwelder 1943: 509 (characters in key as Leucoparyphus), 510 (characters as
Leucoparyphus); Campbell 1975a: 199 (characters); Navarrete-Heredia et al. 2002: 106 (characters in key).
Key: Campbell 1975a: 177 (some species of America north of Mexico); Campbell 1994: 140 (some species of the
genus).
Distribution.
West Indies: Puerto Rico, US Virgin Isl., Guadeloupe, St. Vincent & Grenadines, Barbados, Grenada, Trinidad-
Tobago.
0284. Cileoporus Campbell, 1994: 126

Type species: Tachinus politus Bernhauer, 1923
Characters/Description: Campbell 1994: 141 (characters in key); Navarrete-Heredia et al. 2002: 106 (characters in
key).
Key: Campbell 1994: 143 (species of the genus).
Distribution.
Central America: Guatemala, Costa Rica, Panama.
South America: Colombia, Ecuador, Brazil, Argentina.
0285. Coproporus Kraatz, 1857a: 399 (Fig. 33)

Type species: Coproporus colchicus Kraatz, 1858
Erchomus Motschulsky, 1858b: 218 [synonymized by Ganglbauer 1895: 336]
Type species: Erchomus sanguinolentus Motschulsky, 1858
80 species (1 in Dominican amber) are included in this genus from Latin America.
Characters/Description: Lynch-Arribálzaga 1884: 108 (characters as Erchomus); Blackwelder 1943: 509
(characters in key), 512 (characters); Campbell 1975a: 177 (characters); Campbell 1994: 140 (characters in
key); Navarrete-Heredia et al. 2002: 106 (characters in key); Newton & Peck 2006: 134 (characters in key).
Key: Blackwelder 1943: 512 (species of the West Indies); Campbell 1975a: 177 (some species of America north of
Mexico).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Antigua &
Barbuda, Montserrat, Guadeloupe, Dominica, St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-
Tobago.
South America: Colombia, Venezuela, Guyana, French Guiana, Galapagos Isl., Peru, Bolivia, Brazil, Paraguay,
Uruguay, Chile?, Argentina.
0286. Leucotachinus Coiffait & Sáiz, 1968: 411

Type species: Tachinus luteonitens Fairmaire & Germain, 1862

Characters/Description: Coiffait & Sáiz 1968: 411 (characters); Campbell 1994: 141 (characters in key).
Distribution.
0287. Sepedophilus Gistel, 1856: 267, 386 (Fig. 36)

Type species: Staphylinus pubescens Paykull, 1790

Characters/Description: Campbell 1994: 140 (characters in key); Campbell 1976b: 5 (characters); H.J. Kim & Ahn
2000a: 251 (characters); Navarrete-Heredia et al. 2002: 105 (characters in key).
Key: Campbell 1976b: 9 (species group and species of the America north of Mexico); Coiffait & Sáiz 1968: 407
(Chilean species as Conosomus).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, Montserrat, Guadeloupe, St. Vincent &
Grenadines, Trinidad-Tobago.
Central America: Mexico, Guatemala, Nicaragua, Costa Rica, Panama.
South America: Colombia, Venezuela, Peru, Bolivia, Brazil, Chile, Argentina.
0288. Tachinomorphus Kraatz, 1859a: 54

Type species: Tachinus fulvipes Erichson, 1840
Physetoporus Horn, 1877: 106 [synonymized by Bernhauer & Schubert 1916: 487]
Type species: Coproporus grossulus LeConte, 1863
Characters/Description: Campbell 1973a: 1016 (characters); Campbell 1994: 140 (characters in key); Navarrete-
Heredia et al. 2002: 106 (characters in key).
Key: Campbell 1973a: 1018 (species of the genus).
Distribution.
South America: Ecuador, Peru.
0289. Tachinus Gravenhorst, 1802: 134

Type species: Staphylinus rufipes Linnaeus, 1758
Subgenera:
Latotachinus Ullrich, 1975: 282
Type species: Tachinus punctiventris Sharp, 1888
Tachinoderus Motschulsky, 1858b: 217 [*]
Type species: Tachinoderus longicornis Motschulsky, 1858
Paratachinus Blackwelder, 1952: 293 [synonymized by Ullrich 1975: 291]
Type species: Paratachinus laticollis Cameron, 1932
Paracoproporus Bernhauer, 1917b: 42 [synonymized by Campbell 1993b: 41]
Type species: Coproporus grandicollis Bernhauer, 1917
Tachinus Gravenhorst, 1802: 134 [*]
Type species: Staphylinus rufipes Linnaeus, 1758

Conosoma Kraatz, 1857a: 431 [synonymized by Ganglbauer 1895: 354]
Type species: Oxyporus bipustulatus Fabricius, 1792
Conosomus Motschulsky, 1858c: 54 [synonymized by Bernhauer & Schubert 1916: 465]
Conurus Stephens, 1829a: 22 [synonymized by Kraatz 1857a: 431]
Drymoporus Thomson, 1859: 46 [synonymized by Gemminger & Harold 1868: 554]
Type species: Tachinus elongatus Gyllenhal, 1810
Elliptoma Motschulsky, 1845: 41 [synonymized by Gemminger & Harold 1868: 554]
Type species: Staphylinus marginellus Fabricius, 1781
Hamotraho Gozis, 1886: 13 [synonymized by Bernhauer & Schubert 1916: 479]
Type species: Staphylinus subterraneus Linnaeus, 1758
Japanotachinus Ullrich, 1975: 288 [synonymized by Schülke 2015: 7]
Type species: Tachinus nakanei Ullrich, 1975
Neocharidius Marié, 1928b: 86 [synonymized by Campbell 1973b: 13]
Type species: Tachinus manueli Sharp, 1874
Neocharis Marié, 1928a: 13 [synonymized by Scheerpeltz 1934: 1508]
Type species: Tachinus manueli Sharp, 1874
Porodrymus Rey, 1882: 303 [synonymized by Duvivier 1883: 129]
Type species: Tachinus discoideus Erichson, 1839
Pterygotachinus Scheerpeltz, 1976b: 160 [synonymized by Schülke 1997: 46]
Type species: Pterygotachinus lacinipennis Scheerpeltz, 1976
Characters/Description: Lynch-Arribálzaga 1884: 105 (characters of Conosoma); Coiffait & Sáiz 1968: 406
(characters of Conosomus), 413 (characters); Blackwelder 1943: 509 (characters in key of Conosomus), 524
(characters of Conosomus); Campbell 1973b: 13 (characters); Campbell 1994: 141 (characters in key); H.J.
Kim & Ahn 2000b: 125 (characters); Navarrete-Heredia et al. 2002: 106 (characters in key).
Key: Coiffait & Sáiz 1968: 407 (Chilean species); Campbell 1973b: 15, 19 (species group and species of the North
and Central America); Campbell 1975b: 87, 88 (some species of the genus); Campbell 1976a: 196 (species of
the Mexico and Central America); Li 1995: 53 (subgenera).
Distribution.
0290. Tachyporus Gravenhorst, 1802: 124 (Fig. 37)

Type species: Staphylinus chrysomelinus Linnaeus, 1758
Subgenera:
Palporus Campbell, 1979: 11 [*]
Type species: Staphylinus nitidulus Fabricius, 1781
Tachyporus Gravenhorst, 1802: 124 [*]
Type species: Staphylinus chrysomelinus Linnaeus, 1758
Trachyopus Rey, 1882: 237 [synonymized by Bernhauer & Schubert 1916: 472]
Type species: Tachyporus tersus Erichson, 1839

Characters/Description: Campbell 1979: 5 (characters); Campbell 1994: 140 (characters in key); Navarrete-
Key: Campbell 1979: 8 (subgenera and species of the New World).
Distribution.
Other regions: Nearctic, Palearctic, Ethiopian, Oriental, Australian.
0291. Termitoplus Silvestri, 1946b: 547

Type species: Termitoplus grandis Silvestri, 1946

Distribution.
Tribe Vatesini Seevers, 1958
0292. Vatesus Sharp, 1876c: 201 (Fig. 38)

Type species: Vatesus latitans Sharp, 1876
Wasmannotherium Bernhauer, 1921a: 77 [synonymized by Wasmann 1929: 81]
Type species: Xenocephalus clypeatus Wasmann, 1887
Xenocephalus Wasmann, 1887: 411 [synonymized by Scheerpeltz 1933: 1472]
Type species: Xenocephalus clypeatus Wasmann, 1887
Characters/Description: Wasmann 1900b: 241 (characters as Xenocephalus); Bernhauer 1921a: 77 (characters);
Seevers 1958: 183 (characters); Borgmeier 1961: 189 (characters); Seevers 1965: 332 (characters); Navarrete-
Key: Seevers 1958: 186 (species of the genus); Borgmeier 1961: 192 (species of the genus); Seevers 1965: 335
Distribution.
Central America: Mexico, Belize, Costa Rica, Panama.
South America: Colombia, Peru, Bolivia, Brazil, Paraguay, Argentina.
Subfamily HABROCERINAE Mulsant & Rey, 1877
0293. Habrocerus Erichson, 1839b: 400 (Fig. 39)

Type species: Tachyporus capillaricornis Gravenhorst, 1806
Characters/Description: Erichson 1839a: 242 (characters); Coiffait & Sáiz 1965b: 222 (characters in key); Ashe &
Newton 1993: 267–286 (characters); Assing & Wunderle 1995: 309 (characters), 353 (characters in key).
Key: Assing & Wunderle 1995: 353 (species group and species of the genus).
Distribution.

0294. Nomimocerus Coiffait & Sáiz, 1965b: 217 (Fig. 40)
Type species: Tachyporus marginicollis Solier, 1849

Characters/Description: Coiffait & Sáiz 1965b: 222 (characters in key); Ashe & Newton 1993: 267–286
(characters); Assing & Wunderle 1995: 343 (characters), 353 (characters in key).
Key: Assing & Wunderle 1995: 356 (species of the genus).
Distribution.
Subfamily ALEOCHARINAE Fleming, 1821
Tribe Aleocharini Fleming, 1821
Subtribe Aleocharina Fleming, 1821
0295. Aleochara Gravenhorst, 1802: 67 (Figs. 41–42)

Type species: Staphylinus curtulus Goeze, 1777
Correa Fauvel, 1878c: 592 [synonymized by Pace 2003b: 124]
Type species: Correa oxytelina Fauvel, 1878
Fauvelia Tate, 1880: xlvi [synonymized by Blackwelder 1952: 165]
Fauvelia Tate in Holder, 1882: 78
Subgenera:
Aidochara Casey, 1906: 145
Type species: Aidochara planiventris Casey, 1906
Aleochara Gravenhorst, 1802: 67 [*]
Copiata Gozis, 1886: 12 [synonymized by Bernhauer & Scheerpeltz 1926: 775]
Baryodma Thomson, 1858: 31
Type species: Staphylinus bipunctatus Olivier, 1795
Arybodma Blackwelder, 1952: 63 [Schülke & Smetana 2015: 492]
Type species: Aleochara intricata Mannerheim, 1830
Calochara Casey, 1906: 149 [*]
Type species: Calochara rubripennis Casey, 1906
Oreochara Casey, 1906: 148 [synonymized by Klimaszewski 1984: 59]
Type species: Oreochara laramiensis Casey, 1906
Ceranota Stephens, 1839: 351
Type species: Aleochara daltoni Stephens, 1832
Ceronota Agassiz, 1846: 72 [Newton & Thayer 2003]
Type species: Aleochara daltoni Stephens, 1832
Hoplonotus Schmidt-Göbel, 1846: 245 [synonymized by Märkel 1846: 300]
Type species: Hoplonotus laminatus Schmidt-Göbel, 1846
Hylozyras Iablokoff-Khnzorian, 1960: 1882 [synonymized by Assing 2015a: 16]
Type species: Zyras stenocephalus Iablokoff-Khnzorian, 1960
Sorecocephala Bernhauer, 1902: 238 [synonymized by Smetana in Löbl & Smetana 2004: 33]

Type species: Ocalea reitteri Bernhauer, 1900
Coprochara Mulsant & Rey, 1874b: 430 [*]
Type species: Aleochara bilineata Gyllenhal, 1810
Eucharina Casey, 1906: 165 [synonymized by Blackwelder 1952: 166]
Type species: Aleochara sulcicollis Mannerheim, 1843
Funda Blackwelder, 1952: 166 [synonymized by Klimaszewski 1984: 15]
Type species: Aleochara sulcicollis Mannerheim, 1843
Mecorhopalus Solier, 1849: 347 [synonymized by Bernhauer & Scheerpeltz 1926: 775]
Type species: Mecorhopalus elongatus Solier, 1849
Skenochara Bernhauer & Scheerpeltz, 1926: 795 [synonymized by Maus 1998: 83]
Type species: Aleochara squalithorax Sharp, 1888
Echochara Casey, 1906: 176
Type species: Rheochara lucifuga Casey, 1893
Rheocharella Casey, 1906: 181 [synonymized by Klimaszewski 1984: 94]
Type species: Aleochara fenyesi Bernhauer, 1905
Emplenota Casey, 1884a: 17 [*]
Type species: Emplenota maritima Casey, 1884
Polistoma Casey, 1893: 289 [synonymized by Bernhauer & Scheerpeltz 1926: 795]
Type species: Aleochara obscurella Gravenhorst, 1806
Polycharina Reitter, 1909: 28 [synonymized by Fenyes 1921b: 415]
Type species: Aleochara grisea Kraatz, 1856
Polystoma Stephens, 1833b: 91 [synonymized by Casey 1893: 289]
Type species: Aleochara obscurella Gravenhorst, 1806
Polystomaria Reitter, 1909: 28 [synonymized by Scheerpeltz 1925: 447]
Type species: Aleochara obscurella Gravenhorst, 1806Heterochara Mulsant & Rey, 1874b: 299
Type species: Aleochara clavicornis Redtenbacher, 1849
Aphaenochara Maruyama & Hlaváč, 2003: 225 [synonymized by Tronquet 2009: 96]
Type species: Aphaenochara lackneri Maruyama & Hlaváč, 2003
Ctenochara Casey, 1906: 134 [Newton & Thayer 2003]
Type species: Aleochara clavicornis Redtenbacher, 1848
Homoeochara Mulsant & Rey, 1874b: 414
Type species: Aleochara sparsa Heer, 1839
Madagaschara Klimaszewski et al., 2000: 250
Type species: Aleochara subsulcata Pace, 1999
Maseochara Sharp, 1883: 154 [*]
Type species: Maseochara opacella Sharp, 1883
Pinalochara Casey, 1906: 177 [synonymized by Klimaszewski 1984: 103]
Type species: Pinalochara wickhami Casey, 1906
Tithanis Casey, 1884a: 16 [synonymized by Bernhauer & Scheerpeltz 1926: 797]
Type species: Aleochara valida LeConte, 1858
Mesochara Blackwelder, 1952: 239
Type species: Aleochara almorensis Cameron, 1939
Polystomota Casey, 1906: 136
Type species: Aleochara grisea Kraatz, 1856
Tinotus Sharp, 1883: 170 [*]
Type species: Tinotus cavicollis Sharp, 1883
Acrimea Casey, 1911: 14 [synonymized by Gusarov 2003e: 10]
Type species: Acrimea resecta Casey, 1911

Arpatheta Blackwelder, 1952: 62 [synonymized by Sawada 1987: 148]
Type species: Paratheta carnivora Cameron, 1920
Exaleochara Keys, 1907: 102 [synonymized by Bernhauer & Scheerpeltz 1926: 713]
Type species: Aleochara morion Gravenhorst, 1802
Paratheta Cameron, 1920a: 269 [synonymized by Blackwelder 1952: 62]
Type species: Paratheta carnivora Cameron, 1920
Triochara Bernhauer, 1901c: 373
Type species: Aleochara trisulcata Weise, 1877
Xenochara Mulsant & Rey, 1874b: 344 [*]
Type species: Aleochara decorata Aubé, 1850
Dyschara Mulsant & Rey, 1874b: 425 [synonymized by Assing 2009b: 36]
Type species: Aleochara inconspicua Aubé, 1850
Euryodma Reitter, 1909: 24 [synonymized by Assing, 2009b: 36]
Type species: Aleochara brevipennis Gravenhorst, 1806
Isochara Bernhauer, 1901a: 440 [Klimaszewski 1984: 35]
Type species: Aleochara tristis Gravenhorst, 1806
Metalea Mulsant & Rey, 1875e: 299 [synonymized by Bernhauer & Scheerpeltz 1926: 789]
Type species: Ocalea procera Erichson, 1839
Notiochara Casey, 1906: 134 [synonymized by Klimaszewski & Jansen 1993: 68]
Type species: Notiochara subaspera Casey, 1906
Ophiochara Bernhauer, 1901a: 439 [synonymized by Assing 2009b: 36]
Type species: Aleochara breiti Ganglbauer, 1897
Polychara Mulsant & Rey, 1874b: 348 [synonymized by Klimaszewski 1984: 34]
Type species: Aleochara discipennis Mulsant & Rey, 1853
Rheochara Mulsant & Rey, 1874b: 294 [synonymized by Assing 2016a: 270]
Type species: Ocalea spadicea Erichson, 1837

Characters/Description: Lynch-Arribálzaga 1884: 67, 74 (characters of subgenus Baryodma); Fenyes 1920: 389
(characters as Fauvelia), 396 (characters); Fenyes 1921b: 417 (characters of subgenus Maseochara);
Klimaszewski 1984: 7, 14 (characters of subgenus Coprochara), 24 (characters of subgenus Xenochara), 59
(characters of subgenus Calochara), 70 (characters of subgenus Aleochara), 89 (characters of subgenus
Echochara), 95 (characters of subgenus Emplenota), 102 (characters of subgenus Maseochara); Lohse et al.
1990: 125 (characters in key), 137 (characters); Klimaszewski & Maus 1999: 209 (characters of Puberula
species group), 211 (characters of Weiseri species group), 212 (characters of Richteri species group), 215
(characters of Lacustris species group), 216 (characters of Glabra species group), 217 (characters of subgenus
Aleochara), 217 (characters of Parvicollis species group), 218 (characters of Bruchi species group), 219
(characters of subgenus Maseochara), 219 (characters of Wickhami species group); Hanley 2002b: 456
(characters as Tinotus); Ashe 2002b: 124 (characters in key), 125 (characters in key of subgenus Tinotus);
Klimaszewski et al. 2002: 284 (characters of subgenus Tinotus); Gusarov 2003e: 10 (characters of subgenus
Tinotus); Pace 2008a: 364 (characters in key of subgenus Tinotus); Aballay et al. 2014: 103 (characters in
key); Yamamoto & Maruyama 2016: 83 (characters of subgenus Tinotus).
Key: Lynch-Arribálzaga 1884: 67 (species of Buenos Aires province, Argentina); Fenyes 1920: 397 (some
subgenera); Klimaszewski 1984: 14 (subgenera), 16 (species of the subgenus Coprochara), 36 (species of the
subgenus Xenochara), 60 (species of the subgenus Calochara), 71 (species of the subgenus Aleochara), 90
(species of the subgenus Echochara), 96 (species of the subgenus Emplenota), 103 (species of the subgenus
Maseochara); Klimaszewski & Ashe 1992: 400 (some species of Costa Rica); Gusarov 2003e: 11 (species of
the subgenus Tinotus).

Distribution.
West Indies: Cuba, Jamaica, Haiti, Puerto Rico, US Virgin Isl., Guadeloupe, Dominica, St. Lucia, St. Vincent &
Grenadines, Grenada, Trinidad-Tobago.
Central America: Mexico, Guatemala, Honduras, El Salvador, Nicaragua, Costa Rica, Panama.
South America: Colombia, Venezuela, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay,
Uruguay, Chile, Argentina.
0296. Ocyota Sharp, 1883: 163

Type species: Ocyota dubia Sharp, 1883
2 species are known from and are restricted to Central America. Assing & Wunderle (1997: 122), removed Ocyota
bruchi Bernhauer (described of Argentina) from the genus but did not place it in any other genus.
Characters/Description: Fenyes 1920: 362 (characters); Assing & Wunderle 1997: 119 (characters); Ashe 2002b:
Distribution.
Incertae sedis in Tribe Aleocharini Fleming, 1821
0297. Amarochara Thomson, 1858: 32

Type species: Calodera umbrosa Erichson, 1837
Amarocharella Bernhauer, 1921b: 182 [synonymized by Blackwelder 1952: 48]
Type species: Amarochara rambouseki Bernhauer, 1921
Lasiochara Ganglbauer, 1895: 100 [synonymized by Blackwelder 1952: 209]
Type species: Calodera bonnairii Fauvel, 1865
Mniobates Mulsant & Rey, 1875e: 326 [synonymized by Blackwelder 1952: 249]
Type species: Bolitochara forticornis Lacordaire, 1835
Mniobates Mulsant & Rey, 1875d: 488 [synonymized by Blackwelder 1952: 249]
Pachycerota Casey, 1906: 114 [synonymized by Klimaszewski et al., 2018b: 114]
Type species: Pachycerota duryi Casey, 1906

Characters/Description: Fenyes 1920: 325 (characters), 326 (characters as Lasiochara and Mniobates).
Distribution.
0298. Eydelusa Pace, 1997b: 45 (Fig. 57)

Type species: Eydelusa bartolozzii Pace, 1997

Distribution.
South America: Ecuador.
Antillusini Pace, 2012

0299. Antillusa Pace, 2012: 59 (Fig. 46)
Type species: Antillusa dominicana Pace, 2012
Distribution.
Tribe Athetini Casey, 1910a
Subtribe Athetina Casey, 1910a
0300. Acrotona Thomson, 1859: 38

Type species: Aleochara aterrima Gravenhorst, 1802
Eustrigota Casey, 1911: 165 [synonymized by Gusarov 2003e: 102]
Type species: Strigota seclusa Casey, 1911
Pancota Casey, 1906: 345 [Newton & Thayer 2003]
Type species: Pancota collaris Casey, 1906
Subgenera:
Acrotona Thomson, 1859: 38
Type species: Aleochara aterrima Gravenhorst, 1802
Colpodota Mulsant & Rey, 1873d: 153 [synonymized by Schülke & Smetana 2015: 506]
Type species: Homalota parens Mulsant & Rey, 1852
Colpodota Mulsant & Rey, 1873b: 175 [Newton & Thayer 2003]
Colpodota Mulsant & Rey, 1874c: 207 [Newton & Thayer 2003]
Colpodota Mulsant & Rey, 1874a: 7 [Newton & Thayer 2003]
Aremia Casey, 1910a: 145
Type species: Dolosota reclusa Casey, 1910
Arisota Casey, 1910a: 133
Type species: Arisota tetricula Casey, 1910
Dolosota Casey, 1910a: 136
Type species: Eurypronota scopula Casey, 1893
Eurypronota Casey, 1893: 334
Type species: Eurypronota discreta Casey, 1893
Neada Casey, 1910a: 152
Type species: Acrotona lubricans Casey, 1910
Phanerosphena Pace, 1991a: 138
Type species: Atheta rougemonti Pace, 1991
Reania Casey, 1910a: 146
Type species: Dolosota fontinalis Casey, 1910

Characters/Description: Lynch-Arribálzaga 1884: 58 (characters of Colpodota); Fenyes 1920: 202 (characters as
Pancota), 223 (characters); Lohse et al. 1990: 126 (characters in key), 198 (characters).
Key: Newton & Peck 2006: 139 (species of the Galapagos Isl.).
Distribution.
West Indies: Jamaica, Haiti, Guadeloupe, Saint Vincent & the Grenadines, Grenada.

Central America: Guatemala, Costa Rica.
South America: Colombia, French Guiana, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil, Paraguay, Chile,
Argentina.
0301. Adota Casey, 1910a: 67

Type species: Atheta massettensis Casey, 1910
Halostiba Yosii & Sawada, 1976: 86 [synonymized by Gusarov 2003b: 4]
Type species: Ischnopoda ushio Sawada, 1971
Panalota Casey, 1910a: 71 [Lohse & Smetana 1985: 282]
Type species: Atheta setositarsis Casey, 1910
Phyconoma Easton, 1971: 24 [synonymized by Lohse 1988: 46]
Type species: Atheta immigrans Easton, 1971

Characters/Description: Fenyes 1920: 175 (characters), 243 (characters as Panalota); Gusarov 2003b: 3
(characters).
Key: Gusarov 2003b: 8 (Nearctic species).
Distribution.
0302. Amidobia Thomson, 1858: 33

Type species: Homalota talpa Heer, 1841

Characters/Description: Fenyes 1920: 178 (characters).
Distribution.
0303. Amischa Thomson, 1858: 33

Type species: Aleochara analis Gravenhorst, 1802
Arthropycna Bernhauer, 1921c: 162 [synonymized by Blackwelder 1944: 159]
Type species: Atheta myrmecovagans Bernhauer, 1921
Colposura Casey, 1893: 336 [synonymized by Bernhauer & Scheerpeltz 1926: 592]
Type species: Colposura praelonga Casey, 1893
Characters/Description: Fenyes 1920: 178 (characters); Pace 2008a: 280 (characters in key).
Distribution.
South America: Colombia, Ecuador, Brazil, Chile, Argentina.
0304. Atheta Thomson, 1858: 36 (Figs. 48–49)

Type species: Aleochara graminicola Gravenhorst, 1806
Iotota Casey, 1910a: 95 [Newton & Thayer 2003]

Type species: Iotota tepida Casey, 1910
Megonychiella Marcuzzi, 1977: 17 [nomen nudum, Newton & Thayer 2003]
Type species: Atheta oxypodoides Marcuzzi, 1977
Paraceritaxa Marcuzzi, 1977: 16 [nomen nudum, Newton & Thayer 2003]
Type species: Atheta sexspinulosa Marcuzzi, 1977
Pseudometaxia Wüsthoff, 1940: 683 [Blackwelder 1952: 328]
Rhodeota Casey, 1911: 147 [Newton & Thayer 2003]
Type species: Ousipalia tartarea Casey, 1910
Synaptina Casey, 1910a: 131 [Newton & Thayer 2003]
Type species: Synaptina merica Casey, 1910
Subgenera:
Acromyrmecoxene Scheerpeltz, 1976a: 138 [*]
Type species: Atheta acromyrmecicola Scheerpeltz, 1976
Acrotonida Pace, 1991a: 141
Type species: Atheta invicta Fauvel, 1907
Alaobia Thomson, 1858: 36
Type species: Homalota ochracea Erichson, 1837
Anopleta Mulsant & Rey, 1873b: 4 [*]
Type species: Homalota lepida Kraatz, 1856
Ouralia Mulsant & Rey, 1873d: 174 [Smetana 2004: 372]
Type species: Ouralia picicornis Mulsant & Rey, 1873
Arctostiba Bernhauer, 1928b: 16
Type species: Atheta freyi Bernhauer, 1928
Atheta Thomson, 1858: 36 [*]
Callicerodes Iablokoff-Khnzorian, 1960: 1883 [synonymized by Assing 2005: 63]
Type species: Callicerus velox Iablokoff-Khnzorian, 1960
Elytrusa Casey, 1906: 336 [Smetana 2004: 372]
Type species: Homalota granulata Mannerheim, 1846
Hypatheta Fenyes, 1918: 23 [Benick & Lohse 1974: 191]
Type species: Bolitochara castanoptera Mannerheim, 1830
Megista Mulsant & Rey, 1873b: 591 [Benick & Lohse 1974: 191]
Attatheta Scheerpeltz, 1936: 507 [*]
Type species: Atheta anisophthalma Scheerpeltz, 1936
Badura Mulsant & Rey, 1873d: 159
Type species: Badura nudicornis Mulsant & Rey, 1873
Bessobia Thomson, 1858: 35 [*]
Type species: Homalota monticola Thomson, 1852
Thrichiota Mulsant & Rey, 1873d: 180 [synonimized by Bernhauer & Scheerpeltz 1926: 625]
Type species: Bessobia gibbera Mulsant & Rey, 1873
Ceritaxa Mulsant & Rey, 1873d: 164
Type species: Ceritaxa spissata Mulsant & Rey, 1873
Hilara Mulsant & Rey, 1873: 160 [synonimized by Lohse 1971: 77]
Type species: Microdota fulva Mulsant & Rey, 1873
Neohilara Lohse, 1971: 77 [synonimized by Assing 2015a: 16]
Type species: Microdota fulva Mulsant & Rey, 1873
Chaetida Mulsant & Rey, 1873b: 272 [*]

Type species: Aleochara longicornis Gravenhorst, 1802
Coprothassa Thomson, 1859: 38 [*]
Type species: Homalota testudinea Erichson, 1839
Hemitropia Mulsant & Rey, 1873b: 179 [synonimized by Bernhauer & Scheerpeltz 1926: 670]
Type species: Oxypoda melanaria Mannerheim, 1830
Dabura Cameron, 1948a: 241
Type species: Atheta anommatophila Cameron, 1948
Datomicra Mulsant & Rey, 1873b: 355 [*]
Type species: Homalota celata Erichson, 1837
Datostiba Yosii & Sawada, 1976: 18 [Silfverberg 1992: 29]
Type species: Atheta lewisiana Cameron, 1933
Hilarina Casey, 1910a: 128 [synonimized by Bernhauer & Scheerpeltz 1926: 666]
Type species: Datomicra particula Casey, 1910
Micromota Casey, 1910a: 127 [synonimized by Bernhauer & Scheerpeltz 1926: 666]
Type species: Datomicra filiformis Casey, 1910
Monadia Casey, 1910a: 130 [Synonimized by Bernhauer & Scheerpeltz 1926: 666]
Type species: Datomicra lucana Casey, 1910
Oligomia Casey, 1910a: 129 [synonimized by Bernhauer & Scheerpeltz 1926: 666]
Type species: Datomicra scintilla Casey, 1910
Taxicerella Casey, 1910a: 113 [Smetana 2004: 379]
Type species: Sableta remissa Casey, 1910
Delphota Casey, 1910a: 17
Type species: Atheta cephalina Casey, 1910
Diaprepota Pace, 2004c: 360
Type species: Atheta ruiliensis Pace, 1998
Diaprepota Pace, 1998: 926 [synonimized by Pace 2004c: 360]
Dimetrota Mulsant & Rey, 1873d: 165 [*]
Type species: Homalota tristicula Mulsant & Rey, 1873
Cadaverota Yosii & Sawada, 1976: 55 [Smetana 2004: 380]
Type species: Homalota cadaverina Brisout de Barneville, 1860
Coproceramius Gistel, 1857: 9 [Smetana 2004: 380]
Type species: Bolitochara impressifrons Mannerheim, 1830
Engamota Casey, 1910a: 151 [Smetana 2004: 380]
Type species: Acrotona absona Casey, 1910
Dimetrotina Casey, 1911: 143
Type species: Dimetrota vaniuscula Casey, 1911
Mycetota Ádám, 1987b: 13 [synonymized by Gusarov 2016: 395]
Type species: Aleochara laticollis Stephens, 1832
Donesia Casey, 1910a: 48
Type species: Atheta temporalis Casey, 1910
Dysanomota Pace, 2008d: 431 [*]
Type species: Homalota cinctella Erichson, 1839
Ekkliatheta Pace, 2004b: 507
Type species: Atheta aniiensis Pace, 2004
Epimella Peyerimhoff, 1914: 250
Type species: Atheta cinctuta Peyerimhoff, 1914
Eubadura Sawada, 1990: 543

Type species: Atheta akiensis Sawada, 1990
Eugluta Sawada, 1985: 93
Type species: Atheta picticollis Bernhauer, 1915
Eumorphotheta Ádám, 2008: 161
Type species: Atheta siculorum Ádám, 2008
Evaniodera Pace, 1986a: 430 [*]
Type species: Atheta apicilutea Pace, 1986
Geostibops Israelson, 1985: 13
Type species: Atheta aptera Israelson, 1985
Gyroatheta Pace, 2014c: 100 [*]
Type species: Atheta guyanicola Pace, 2014
Hummleriella Bernhauer, 1929c: 191
Type species: Atheta ponferradae Bernhauer, 1929
Hydrosmectomorpha Klimaszewski & Webster in Klimaszewski et al., 2018a: 2
Type species: Atheta meduxnekeagensis Webster & Klimaszewski in Klimaszewski et al., 2018
Hypsostiba Bernhauer, 1929a: 200 [*]
Type species: Atheta dampfi Bernhauer, 1929
Ischnoderoecia Scheerpeltz, 1974: 249
Type species: Atheta lunaris Scheerpeltz, 1974
Kamptodota Pace, 2009: 274
Type species: Atheta papuaimitatrix Pace, 2009
Kladobrachydotina Pace, 2008a: 302 [*]
Type species: Atheta inflatheca Pace, 2008
Lamiota Casey, 1910a: 17
Type species: Atheta keeni Casey, 1910
Lampratheta Scheerpeltz, 1965b: 334
Type species: Atheta malaisei Scheerpeltz, 1965
Libanostiba Scheerpeltz, 1929: 227
Type species: Atheta ebneri Scheerpeltz, 1929
Lobodota Pace, 2009: 274
Type species: Atheta nabirensis Pace, 2009
Metadimetrota Klimaszewski & Winchester, 2002: 31
Type species: Atheta cheersae Klimaszewski, 2002
Microdota Mulsant & Rey, 1873d: 160 [*]
Type species: Microdota sericea Mulsant & Rey, 1873
Anatelloleptusa Scheerpeltz, 1976c: 50 [Smetana 2004: 384]
Type species: Leptusa himalayica Scheerpeltz, 1976
Dicolyota Pace, 2004: 359 [nomen nudum, synonymized by Schülke & Smetana 2015: 542]
Type species: Atheta praelata Pace, 1986
Dicolyota Pace, 1998: 924 [synonymized by Pace 2004: 359]
Heteronoma Mulsant & Rey, 1873b: 4 [synonymized by Bernhauer & Scheerpeltz 1926: 630]
Type species: Homalota luctuosa Mulsant & Rey, 1853
Heterophaena Lynch-Arribálzaga, 1884: 45 [synonymized by Bernhauer & Scheerpeltz 1926: 630]
Type species: Heterophaena palliditarsis Lynch-Arribálzaga, 1884
Metamischa Peyerimhoff, 1938: 65 [synonymized by Pace 1990c: 915]
Type species: Homalota scrobicollis Kraatz, 1859
Meoticaemorpha Scheerpeltz, 1976c: 66 [Smetana 2004: 384]

Type species: Meoticaemorpha janetscheki Scheerpeltz, 1976
Physadota Pace, 1990c: 907 [synonymized by Pace 2004c: 359]
Type species: Atheta subscabra Cameron, 1939
Sonomota Casey, 1911: 158 [synonymized by Lohse & Smetana 1988: 270]
Type species: Sipalia lippa Casey, 1911
Microdotina Scheerpeltz, 1974: 254
Type species: Atheta xanthopyga Scheerpeltz, 1974
Nemota Casey, 1910a: 56
Type species: Atheta paganella Casey, 1910
Niphetodroma Scheerpeltz, 1947: 288
Type species: Niphetodroma obsolescens Scheerpeltz, 1947
Oligatheta Bernhauer & Scheerpeltz, 1926: 605 [*]
Type species: Atheta cordillerana Bernhauer, 1921
Micratheta Bernhauer, 1921b: 179 [synonymized by Bernhauer & Scheerpeltz 1926: 605]
Type species: Atheta cordillerana Bernhauer, 1921
Oreostiba Ganglbauer, 1895: 219
Type species: Homalota tibialis Heer, 1839
Agaphygra Tottenham, 1949c: 78 [Smetana 2004: 31]
Type species: Homalota subglabra Sharp, 1869
Orientatheta Sawada, 1985: 99
Type species: Atheta chinkiangensis Bernhauer, 1939
Oxypodera Bernhauer, 1915d: 185
Type species: Atheta kilimandjarensis Bernhauer, 1915
Pachyatheta Münster, 1925: 11
Type species: Homalota cribrate Kraatz, 1856
Paradota Sawada, 1989: 274
Type species: Atheta liberta Sawada, 1989
Parameotica Ganglbauer, 1895: 228 [*]
Type species: Homalota laticeps Thomson, 1856
Parametaxya Jeannel & Paulian, 1945: 106
Type species: Atheta jeanneliana Bernhauer, 1944
Paramidobia Bernhauer, 1908: 356 [*]
Type species: Amidobia longiceps Bernhauer, 1908
Parataxicera Brundin, 1943: 27
Type species: Atheta moczarskii Bernhauer, 1915
Peliolurga Tottenham, 1939b: 228
Type species: Bolitochara luridipennis Mannerheim, 1830
Pelurga Mulsant & Rey, 1873b: 577 [synonymized by Tottenham 1939b: 228]
Type species: Bolitochara luridipennis Mannerheim, 1830
Pelmatheta Scheerpeltz, 1965b: 335
Type species: Atheta crassitarsis Scheerpeltz, 1965
Philacrotona Pace, 1991c: 140
Type species: Atheta newar Pace, 1991
Polydota Pace, 1983d: 310 [*]
Type species: Atheta uhligi Pace, 1983
Poromicrodota Pace, 1990c: 907
Type species: Atheta subamicula Cameron, 1939
Pseudhygroecia Jeannel & Paulian, 1945: 104

Type species: Atheta elgonensis Bernhauer, 1944
Pseudobessobia Bernhauer, 1921b: 177 [*]
Type species: Atheta weiseri Bernhauer, 1921
Pseudohygroecia Bernhauer, 1929c: 189
Type species: Atheta crisiana Bernhauer, 1929
Pseudoleptonia Bernhauer, 1934c: 507 [*]
Type species: Atheta polyporina Bernhauer, 1934
Pseudophilygra Bernhauer, 1929c: 190
Type species: Atheta holtzi Bernhauer, 1929
Pseudota Casey, 1910a: 114
Type species: Pseudota dissensa Casey, 1910
Pseudothinoecia Bernhauer, 1899: 20
Type species: Atheta puellaris Bernhauer, 1899
Rhagocneme Münster, 1923: 206
Type species: Atheta fractipes Münster, 1923
Sepedomicra Sawada, 1985: 97
Type species: Atheta mycetophaga Cameron, 1920
Sipalatheta Pace, 1993: 92
Type species: Atheta difformis Pace, 1993
Spelaeolla Rambousek, 1915b: 31
Type species: Atheta absoloni Rambousek, 1915
Stictatheta Cameron, 1939a: 5
Type species: Atheta quadripunctula Cameron, 1939
Stilbatheta Scheerpeltz, 1962a: 596
Type species: Atheta speculiventris Scheerpeltz, 1962
Strobilocera Ganglbauer, 1895: 191
Type species: Homalota capitulata Eppelsheim, 1880
Suensonia Bernhauer, 1936d: 314
Type species: Atheta obsoletepunctata Bernhauer, 1936
Synaloconota Scheerpeltz, 1974: 247
Type species: Atheta cascadensis Scheerpeltz, 1974
Tachynota Bernhauer, 1901b: 113
Type species: Atheta thinodromoides Bernhauer, 1901
Tetropla Mulsant & Rey, 1873b: 492
Type species: Aleochara nigritula Gravenhorst, 1802
Thinobaena Thomson, 1859: 39
Type species: Aleochara quisquiliarum Gyllenhal, 1810
Parabessobia Scheerpeltz, 1958: 32 [synonymized by G. Benick 1973: 187]
Type species: Atheta serricauda Eppelsheim, 1878
Traumoecia Mulsant & Rey, 1873b: 631
Type species: Aleochara excavata Gyllenhal, 1827
Trochanterella Brundin, 1953: 428
Type species: Atheta cribripennis J. Sahlberg, 1890
Tropatheta Bernhauer, 1927a: 81
Type species: Atheta gestroi Bernhauer, 1927
Umbala Blackwelder, 1952: 402
Type species: Atheta mimetica Cameron, 1939
Stictatheta Cameron, 1939b: 336 [synonymized by Blackwelder 1952: 402]

Type species: Atheta mimetica Cameron, 1939
Xenota Mulsant & Rey, 1873b: 397 [*]
Type species: Homalota myrmecobia Kraatz, 1856
Mycota Mulsant & Rey, 1873b: 502 [Newton & Thayer 2003]
Type species: Homalota humeralis Kraatz, 1856
Xestota Bernhauer, 1908: 361 [*]
Type species: Atheta biarmata Bernhauer, 1908

Characters/Description: Lynch-Arribálzaga 1884: 50 (characters of subgenus Microdota); Fenyes 1920: 181
(characters), 201 (characters as Synaptina), 210 (characters subgenus Atheta), 238 (characters as Iotota), 247
(characters as Rhodeota); Lohse et al. 1990: 125 (characters in key), 126 (characters in key of Bessobia), 126
(characters in key of Dimetrota), 178 (characters of Dimetrota), 188 (characters); Ashe 2002b: 128
(characters in key); Pace 2008a: 281 (characters in key); Aballay et al. 2014: 103 (characters in key).
Key: Fenyes 1920: 182 (subgenera); Pace 1988: 88 (some species of Chile); Pace 1996c: 652 (some species of
Ecuador); Klimaszewski & Peck, 1998: 236 (species of the Galapagos Isl.); Pace 2000a: 428 (some species of
Chile); Pace 2008a: 294 (subgenera of Ecuador and Peru), 302 (species of the subgenus Acrotona of Ecuador
and Peru), 305 (species of the subgenus Xestota of Ecuador and Peru), 307 (species of the subgenus
Microdota of Ecuador and Peru), 310 (species of the subgenus Kladobrachydotina of Ecuador and Peru), 312
(species of the subgenus Datomicra of Ecuador and Peru), 317 (species of the subgenus Dimetrota of Ecuador
and Peru), 319–321 (species of the subgenus Pseudobessobia of Ecuador and Peru); Newton & Peck 2006:
139 (species of the Galapagos Isl.).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, Puerto Rico, US Virgin Isl., British Virgin Isl., Guadeloupe, St.
Lucia, St. Vincent & Grenadines, Grenada.
South America: Colombia, Venezuela, Suriname, French Guiana, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil,
Paraguay, Chile, Argentina.
0305. Brundinia Tottenham, 1949c: 78

Type species: Metaxya apricans Mulsant & Rey, 1873
Metaxya Mulsant & Rey, 1873d: 181 [synonymized by Tottenham 1949c: 78]
Type species: Metaxya apricans Mulsant & Rey, 1873

Characters/Description: Fenyes 1920: 197 (characters as Metaxya).
Distribution.
South America: Peru, Argentina.
0306. Dalotia Casey, 1910a: 106

Type species: Dimetrota pectorina Casey, 1910
Key: Newton & Peck 2006: 139 (species of the Galapagos Isl.).
Distribution.
South America: Ecuador, Galapagos Isl., Bolivia, Brazil, Chile, Argentina.

0307. Dochmonota Thomson, 1859: 40
Type species: Homalota funebris Thomson, 1856

Distribution.
0308. Homia Blackwelder, 1952: 190

Type species: Daya occipitalis Fauvel, 1878
Daya Fauvel, 1878d: 147 [synonymized by Blackwelder 1952: 118]
Type species: Daya occipitalis Fauvel, 1878

Characters/Description: Fenyes 1920: 230 (characters as Daya).
Distribution.
0309. Hydrosmecta Thomson, 1858: 33

Type species: Homalota longula Heer, 1839
Actocharina Bernhauer in Ganglbauer, 1907: 185 [synonymized by Elven et al. 2012: 633]
Type species: Atheta leptotyphloides Bernhauer in Ganglbauer, 1907
Doliponta Blackwelder, 1952: 132 [synonymized by Blackwelder 1952: 132]
Type species: Lipodonta veris Fenyes, 1921
Hydrosmectina Ganglbauer, 1895: 232 [Smetana 2004: 407]
Type species: Homalota subtilissima Kraatz, 1854
Lipodonta Fenyes, 1921a: 24 [synonymized by Blackwelder 1952: 132]
Type species: Lipodonta veris Fenyes, 1921
Thinoecia Mulsant & Rey, 1873d: 184 [synonymized by Blackwelder 1952: 386]
Type species: Thinoecia libitina Mulsant & Rey, 1873
Thinaecia Mulsant & Rey, 1873b: 5 [synonymized by Blackwelder 1952: 386]
Thinaecia Mulsant & Rey, 1874c: 37 [synonymized by Blackwelder 1952: 386]
Thinoecia Mulsant & Rey, 1874a: 38 [synonymized by Blackwelder 1952: 386]
Thinoecia Mulsant & Rey, 1875a: 260 [synonymized by Blackwelder 1952: 386]
Thinoecia Mulsant & Rey, 1875c: 234 [synonymized by Bernhauer & Scheerpeltz 1926: 605]
Characters/Description: Fenyes 1920: 174 (characters as Actocharina), 236 (characters), 237 (characters as
Hydrosmectina); Lohse et al. 1990: 125 (characters in key), 147 (characters); Pace 2008a: 280 (characters in
key).
Distribution.
South America: Ecuador, Chile, Argentina.
0310. Nehemitropia Lohse, 1971: 81
Type species: Staphylinus sordidus Marsham, 1802

Distribution.
South America: Peru, Bolivia, Brazil, Paraguay, Uruguay, Chile, Argentina.
0311. Notothecta Thomson, 1858: 33

Type species: Aleochara flavipes Gravenhorst, 1806
Subgenera:
Kraatzia Saulcy, 1862: 289
Type species: Kraatzia attophila Saulcy, 1862
Notothecta Thomson, 1858: 33
Type species: Aleochara flavipes Gravenhorst, 1806
Notothectina Bernhauer, 1912d: 77 [*]
Type species: Notothecta attae Bernhauer, 1912

Characters/Description: Fenyes 1920: 240 (characters), 241 (characters of subgenera Kraatzia and Notothecta),
242 (characters of subgenus Notothectina).
Key: Fenyes 1920: 241 (subgenera).
Distribution.
Other regions: Palaearctic, Oriental.
0312. Philhygra Mulsant & Rey, 1873d: 160

Type species: Microdota perdubia Mulsant & Rey, 1873
Amphibitherion Notman, 1921: 155 [Seevers 1978: 123]
Type species: Amphibitherion demissum Notman, 1921
Homalotusa Casey, 1906: 340 [Smetana 2004: 390]
Type species: Homalotusa helenica Casey, 1906
Hygroecia Mulsant & Rey, 1873d: 187 [Schülke & Smetana 2015: 547]
Type species: Hygroecia parca Mulsant & Rey, 1873
Phryogora Mulsant & Rey, 1873b: 625 [Schülke & Smetana 2015: 548]
Type species: Homalota hygrobia Thomson, 1856
Plagiarthrina Keys, 1920: 131 [Smetana 2004: 390]
Type species: Metaxya fordhamiana Keys, 1920
Valenusa Casey, 1906: 342 [Newton & Thayer 2003]
Type species: Valenusa parallela Casey, 1906

Characters/Description: Fenyes 1920: 200 (characters); Lohse et al. 1990: 126 (characters in key), 167 (characters).
Distribution.
Other regions: Neartic, Palaearctic, Ethiopian.

0313. Platyola Mulsant & Rey, 1875e: 249
Type species: Homalota fusicornis Mulsant & Rey, 1853
Australoplatyola Scheerpeltz, 1959b: 10 [nomen nudum]
Type species: Platyola antarctica Bernhauer, 1920
Neoplatyola Pace, 1996b: 399 [nomen nudum]
Platyola Mulsant & Rey, 1875d: 411 [synonymized by Blackwelder 1952: 313]

Distribution.
South America: Colombia, Peru, Bolivia, Brazil.
0314. Pycnota Mulsant & Rey, 1873b: 377

Type species: Homalota paradoxa Mulsant & Rey, 1861

Key: Pace, 1988: 86 (some species of Chile).
Distribution.
Other regions: Palaearctic.
0315. Schistoglossa Kraatz, 1856b: 344

Type species: Homalota viduata Erichson, 1837
Hölzelia Scheerpeltz, 1952b: 765 [Smetana 2004: 416]
Type species: Hoelzelia caricis Scheerpeltz, 1952
Protoskiusa Bernhauer, 1900b: 200 [synonymized by Bernhauer & Scheerpeltz 1926: 591]
Type species: Protoskiusa paradoxa Bernhauer, 1900

Distribution.
0316. Stethusa Casey, 1910a: 4

Type species: Atheta klimschi Bernhauer, 1909
3 species are included in this genus from the West Indies and South America.
Characters/Description: Gusarov 2003c: 5 (characters).
Key: Gusarov 2003c: 12 (species of the genus); Newton & Peck 2006: 139 (species of the Galapagos Isl.).
Distribution.
West Indies: US Virgin Isl., Guadeloupe, St. Lucia., Grenada.
South America: Venezuela, French Guiana, Galapagos Isl., Bolivia, Brazil, Paraguay, Uruguay, Argentina.
Other regions: Nearctic, Palaearctic ?.
Subtribe Schistogeniina Fenyes, 1918
0317. Mimacrotona Cameron, 1920a: 268

Type species: Mimacrotona cingulata Cameron, 1920

Distribution.
West Indies: St. Vincent & Grenadines, Grenada.
Other regions: Palaearctic, Ethiopian, Oriental, Australian, Oceanic.
Subtribe Thamiaraeina Fenyes, 1921a
0318. Phoinixiusa Pace, 2011a: 132

Type species: Phoinixiusa ecuadorensis Pace, 2011

Distribution.
0319. Thamiaraea Thomson, 1858: 35

Type species: Aleochara cinnamomea Gravenhorst, 1802
Fusalia Casey, 1911: 145 [synonymized by Gusarov 2003e: 95]
Type species: Sableta brittoni Casey, 1911
Subgenera:
Miatharaea Pace, 1991b: 855
Type species: Thamiaraea smetanai Pace, 1991
Thamiaraea Thomson, 1858: 35 [*]
Type species: Aleochara cinnamomea Gravenhorst, 1802
Characters/Description: Fenyes 1920: 144 (characters); Ashe 2002b: 128 (characters in key).
Distribution.
Incertae sedis in Tribe Athetini Casey, 1910a
0320. Acticola Cameron, 1944a: 618

Type species: Acticola falklandica Cameron, 1944

Distribution.
South America: Falkland Isl.
0321. Apalonia Casey, 1906: 323

Type species: Apalonia seticornis Casey, 1906
Subgenera:
Alopania Pace, 2015b: 133 [*]
Type species: Apalonia dimidiaticornis Pace, 2015
Apalonia Casey, 1906: 323 [*]
Type species: Apalonia seticornis Casey, 1906
Episkiolonia Pace, 2015b: 133 [*]
Type species: Apalonia semiopaca Pace, 2015
79 species are included in this genus from Latin America. Eldredge (2012b: 159) listed 29 species that not belong
to Apalonia Casey and could belong to Myrmedonota Cameron.
Characters/Description: Fenyes 1920: 259 (characters); Eldredge 2012b: 154 (characters).
Key: Pace 2008a: 338 (species of Ecuador and Peru); Pace 2008b: 196 (Brazilian species); Pace 2015b: 132
(subgenera and species of the genus).
Distribution.
West Indies: Guadeloupe, Trinidad-Tobago.
South America: Colombia, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay.
0322. Arecaceopora Pace, 2011a: 126

Type species: Arecaceopora couturieri Pace, 2011

Distribution.
0323. Campbellia Kistner & Jacobson, 1990: 388

Type species: Ecitoxenia lucanoides Campbell, 1973
Key: Kistner & Jacobson 1990: 395 (species of the genus).
Distribution.
0324. Charoxus Sharp, 1883: 224

Type species: Charoxus fodiens Sharp, 1883

Characters/Description: Fenyes 1920: 263 (characters); Kistner 1981: 588 (characters); Ashe 2002b: 127
Key: Kistner 1981: 590 (species of the genus).
Distribution.
South America: Venezuela, Paraguay.
0325. Cosmogastrusa Pace, 1996c: 646
Type species: Cosmogastrusa curticornis Pace, 1996

Distribution.
0326. Dinusella Bernhauer, 1908: 354

Type species: Dinusella brasiliana Bernhauer, 1908

Characters/Description: Fenyes 1920: 267 (characters); Pace 2008a: 280 (characters in key); Pace 2014c: 102
Distribution.
South America: French Guiana, Ecuador, Peru, Brazil, Argentina.
0327. Dityloechusa Pace, 1985: 384

Type species: Dityloechusa franzi Pace, 1985

Distribution.
South America: Venezuela, Paraguay.
0328. Dolerella Pace, 2014c: 101

Type species: Dolerella cayennensis Pace, 2014
Characters/Description: Pace 2014c: 102 (characters in key)
Distribution.
0329. Ecitochara Wasmann, 1887: 404

Type species: Ecitochara fusicornis Wasmann, 1887
Characters/Description: Fenyes 1920: 270 (characters); Seevers 1965: 292 (characters); Kistner & Jacobson 1990:
359 (characters).
Distribution.
0330. Ecitodaemon Reichensperger, 1939b: 292

Type species: Ecitodaemon vagantium Reichensperger, 1939
Characters/Description: Seevers 1965: 292 (characters); Kistner & Jacobson 1990: 433.
Distribution.

0331. Ecitodonia Seevers, 1965: 241
Type species: Ecitopora major Wasmann, 1900
Characters/Description: Seevers 1965: 236 (characters in key).
Distribution.
South America: Guyana, Brazil.
0332. Ecitomorpha Wasmann, 1889: 185 (Fig. 53)

Type species: Ecitomorpha arachnoides Wasmann, 1889
Only one species is known from and is restricted to Latin America.

Characters/Description: Wasman 1900b: 226 (characters); Fenyes 1920: 271 (characters); Seevers 1965: 208
(characters in key), 209 (characters); Kistner & Jacobson 1990: 418 (characters); Ashe 2002b: 128 (characters
in key).
Distribution.
South America: Colombia, Guyana, Ecuador, Peru, Bolivia, Brazil.
0333. Ecitophya Wasmann, 1900b: 224 (Fig. 54)

Type species: Ecitomorpha simulans Wasmann, 1889
Characters/Description: Fenyes 1920: 273 (characters); Seevers 1965: 208 (characters in key), 211 (characters);
Kistner & Jacobson 1990: 396 (characters); Ashe 2002b: 128 (characters in key).
Distribution.
South America: Colombia, Guyana, Suriname, Ecuador, Peru, Bolivia, Brazil.
0334. Ecitopora Wasmann, 1887: 408

Type species: Ecitopora opaca Wasmann, 1887
Characters/Description: Fenyes 1920: 274 (characters); Borgmeier 1949: 130 (characters); Seevers 1965: 236
(characters in key), 236 (characters in key), 239 (characters); Kistner & Mooney 2011: 270 (characters)
Key: Wasman 1900b: 238 (some species of the genus); Kistner & Mooney 2011: 303 (species of the genus).
Distribution.
South America: Guyana, Ecuador, Bolivia, Brazil.
0335. Ecitoschneirla Kistner & Jacobson, 1990: 374

Type species: Ecitoxenia impressa Seevers, 1965
Characters/Description: Ashe 2002b: 128 (characters in key).
Distribution.
0336. Ecitosymbia Bruch, 1923: 182

Type species: Ecitosymbia rufa Bruch, 1923
Ecitoxenides Borgmeier, 1949: 151 [synonymized by Kistner & Jacobson 1990: 341]
Type species: Ecitoxenides cariniceps Borgmeier, 1949
Characters/Description: Borgmeier 1949: 150 (characters); Seevers 1965: 289 (characters); Kistner & Jacobson
1990: 341 (characters).
Distribution.
0337. Ecitoxenia Wasmann, 1900b: 232

Type species: Ecitoxenia mirabilis Wasmann, 1900
Pseudoecitoxenia Costa-Lima, 1932: 59 [synonymized by Blackwelder 1952: 326]
Type species: Pseudoecitoxenia mirabilis Costa-Lima, 1932
Only a single species is included in this genus from South America.

Characters/Description: Fenyes 1920: 334 (characters); Seevers 1965: 289 (characters); Kistner & Jacobson 1990:
351 (characters).
Key: Seevers 1965: 290 (species of the genus); Campbell 1973d: 27 (Colombian species).
Distribution.
0338. Eidmannotherium Scheerpeltz, 1936: 520

Type species: Eidmannotherium attarum Scheerpeltz, 1936

Distribution.
0339. Fernanstiba Pace, 1999c: 171

Type species: Fernanstiba robinsoni Pace, 1999

Distribution.
0340. Gastrosteata Pace, 1999c: 174

Type species: Gastrosteata lapidicola Pace, 1999

Distribution.

0341. Geitonusa Pace, 2008a: 290
Type species: Geitonusa salcedensis Pace, 2008

Characters/Description: Pace 2008a: 280 (characters in key).
Distribution.
0342. Heterostiba Pace, 1983b: 142 (Fig. 59)

Type species: Heterostiba franzi Pace, 1983
Key: Pace 2008a: 284 (species of the Ecuador and Peru).
Distribution.
West Indies: Guadeloupe, St. Lucia.
South America: Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil, Chile.
0343. Idiocolyusa Pace, 2000a: 442

Type species: Idiocolyusa gonzalezi Pace, 2000
Distribution.
0344. Ischnothina Pace, 1999c: 169

Type species: Ischnothina newtoni Pace, 1999

Distribution.
0345. Kaloxypoda Pace, 2008a: 296

Type species: Kaloxypoda ecuadorensis Pace, 2008

Distribution.
0346. Kephamekynella Pace, 2015e: 81

Type species: Kephamekynella peruviana Pace, 2015

Distribution.
0347. Lamprostiba Pace, 1983a: 97 (Fig. 62)
Type species: Lamprostiba franzi Pace, 1983
Key: Pace 2008a: 298 (species of Peru and Ecuador).
Distribution.
South America: Venezuela, French Guiana, Ecuador, Peru, Bolivia, Chile, Argentina.
0348. Leiodota Pace, 1987b: 483

Type species: Leiodota chilensis Pace, 1987

Distribution.
0349. Leptoglossula Eichelbaum, 1915: 112

Type species: Leptoglossa puberula Solsky, 1870
Leptoglossa Solsky, 1870: 260 [synonymized by Eichelbaum 1915: 112]
Neoleptoglossa Bernhauer & Scheerpeltz, 1926: 683 [synonymized by Blackwelder 1952: 216]
Characters/Description: Fenyes 1920: 277 (characters as Leptoglossa); Ashe 2002b: 128 (characters in key).
Distribution.
South America: Brazil, Chile, Argentina.
0350. Leptonia Sharp, 1883: 196 (Fig. 63)

Type species: Leptonia picta Sharp, 1883

Characters/Description: Fenyes 1920: 277 (characters); Paśnik 2007: 1121 (characters as Pseudognypeta); Pace
Key: Pace 2001b: 428 (some species of the genus); Pace 2008a: 332 (species of Ecuador and Peru).
Distribution.
Central America: Mexico, Guatemala, Belize, Nicaragua, Panama.
South America: Colombia, Venezuela, Guyana, Suriname, Ecuador, Peru, Bolivia, Brazil, Argentina.
0351. Onoreusa Pace, 2015e: 104

Type species: Onoreusa ecuadorensis Pace, 2015

Distribution.
0352. Ousipaliaglossa Pace, 1985: 382

Type species: Ousipaliaglossa splendens Pace, 1985

Distribution.
West Indies: St. Vincent & Grenadines.
South America: Venezuela, Ecuador, Peru, Brazil.
0353. Parademosoma Bernhauer, 1929a: 207 (Fig. 69)

Type species: Oxypoda opaciventris Bernhauer, 1929
Acanthaktema Pace, 2008a: 289 [synonymized by Pace 2011b: 59]
Type species: Hydrosmecta ecuadoriana Pace, 2008
Subgenera:
Elixusa Pace, 2014c: 101 [*]
Type species: Parademosoma guyanensis Pace, 2014
Parademosoma Bernhauer, 1929a: 207 [*]
Type species: Oxypoda opaciventris Bernhauer, 1929
Characters/Description: Ashe 2002b: 127 (characters in key); Pace 2008a: 280 (characters in key).
Key: Pace 2008a: 287 (some species of the genus); Pace 2014c: 101 (subgenera of the genus).
Distribution.
South America: Venezuela, French Guiana, Ecuador, Bolivia.
0354. Parapycnota Bernhauer, 1927b: 255

Type species: Atheta weiseriana Bernhauer, 1927

Key: Pace 2008a: 284 (some species of the genus).
Distribution.
0355. Peruusa Pace, 2008a: 293

Type species: Peruusa peruviana Pace, 2008

Key: Pace 2008a: 294 (species of Peru and Ecuador).
Distribution.
South America: Ecuador, Peru, Brazil.
0356. Physetophallia Pace, 1990b: 62

Type species: Physetophallia minima Pace, 1990

Distribution.
South America: Ecuador, Argentina.
0357. Pichinchusa Pace, 2008a: 292
Type species: Pichinchusa miripennis Pace, 2008

Distribution.
0358. Plesioleptonia Pace, 2011b: 59

Type species: Plesioleptonia abdominalis Pace, 2011

Distribution.
0359. Pontomalota Casey, 1885a: 296

Type species: Phytosus opacus LeConte, 1863

Characters/Description: Fenyes 1920: 244 (characters); Ahn & Ashe 1992: 347 (characters); Ashe 2002b: 127
Key: Ahn & Ashe 1992: 350 (species of the genus).
Distribution.
0360. Pseudoleptonia Pace, 2008a: 290

Type species: Pseudoleptonia ecuadorica Pace, 2008
Only one species is known from and restricted to South America.

Distribution.
0361. Pseustantha Pace, 2011b: 61

Type species: Pseustantha escalonai Pace, 2011

Distribution.
0362. Ragabrachydota Pace, 2008a: 296

Type species: Ragabrachydota ecuadorensis Pace, 2008

Distribution.

0363. Retteneciton Kistner & Jacobson, 1990: 381
Type species: Retteneciton elongatus Kistner & Jacobson, 1990

Distribution.
0364. Seeverseciton Kistner & Jacobson, 1990: 367

Type species: Ecitoxenia megalops Seevers, 1965
Distribution.
0365. Serikendusa Pace, 2008a: 381

Type species: Serikendusa peruviana Pace, 2008

Distribution.
0366. Tarphiota Casey, 1893: 332

Type species: Tarphiota pallidipes Casey, 1893

Characters/Description: Fenyes 1920: 254 (characters); Ahn 1996b: 177 (characters); Ashe 2002b: 127 (characters
in key).
Key: Ahn 1996b: 179 (species of the genus); Ahn 1999: 193 (species of the genus).
Distribution.
0367. Tetradonia Wasmann, 1894: 209

Type species: Myrmedonia eppelsheimii Wasmann, 1890
Acanthodonia Bruch, 1923: 184 [synonymized by Borgmeier 1959a: 10]
Type species: Acanthodonia argentina Bruch, 1923
Chlorotusa Casey, 1906: 324 [synonymized by Seevers 1965: 249]
Type species: Chlorotusa megalops Casey, 1906
Characters/Description: Fenyes 1920: 263 (characters as Chlorotusa), 291 (characters as Tetradonia); Seevers
1965: 235 (characters in key), 249 (characters); Jacobson & Kistner 1998: 166 (characters); Ashe 2002b: 127
(characters in key); Santiago-Jiménez 2010: 646 (characters in key); Santiago-Jiménez 2016: 275 (characters
in key).
Key: Borgmeier 1931: 365 (species of Acanthodonia); Scheerpeltz 1972a: 88 (species of Acanthodonia);
Scheerpeltz 1972b: 111 (species of the genus); Jacobson & Kistner 1998: 171 (species of the genus); Beeren et
al. 2016: 13 (species of La Selva Biological Station in Costa Rica).
Distribution.
South America: Ecuador, Peru, Bolivia, Brazil, Paraguay, Argentina.
0368. Thinusa Casey, 1893: 371

Type species: Phytosus maritimus Casey, 1885

Characters/Description: Fenyes 1920: 134 (characters); Ahn 1997: 75 (characters); Ashe 2002b: 129 (characters in
key).
Key: Ahn 1997: 76 (species of the genus).
Distribution.
0369. Xyronarchusa Pace, 2014c: 94 (Fig. 80)

Type species: Xyronarchusa amazonica Pace, 2014

Distribution.
Tribe Autaliini Thomson, 1859
0370. Attonia Wasmann, 1925b: 52

Type species: Attonia hirta Wasmann, 1925

Distribution.
0371. Autalia Leach in Samouelle, 1819: 177

Type species: Staphylinus impressus Olivier, 1795
Hydatobia Gistel, 1834: 10 [synonymized by Bousquet & Bouchard 2017: 120]
Type species: Staphylinus impressus Olivier, 1795
Characters/Description: Hoebeke & Ashe 1994: 193 (characters); Ashe 2002b: 130 (characters in key); J.S. Park &
Ahn 2005: 1 (characters).
Key: Hoebeke & Ashe 1994: 207 (Neotropical species).
Distribution.

0372. Ophioglossa Fauvel, 1866b: 259 (Fig. 67)
Type species: Ophioglossa araucana Fauvel, 1866
Antrogastra Bernhauer, 1912d: 70 [synonymized by Blackwelder 1952: 57]
Type species: Ophioglossa bruchiana Bernhauer, 1912
Ophioglossa Fauvel, 1866a: 10 [Newton & Thayer 2003]
Type species: Ophioglossa araucana Fauvel, 1866
Characters/Description: Fenyes 1920: 134 (characters), 135 (characters as subgenera Ophioglossa, Antrogastra);
Coiffait & Sáiz 1967: 65 (characters in key), 97 (characters); Pace 1986a: 419 (characters in key); Ashe
2002b: 129 (characters in key).
Key: Fenyes 1920: 135 (as subgenera); Pace 1987b: 473 (some species of Chile).
Distribution.
Central America: Mexico, Nicaragua.
South America: Brazil, Paraguay, Chile, Argentina.
Other regions: Oceanic.
0373. Rhopalogastrum Bernhauer, 1912d: 68

Type species: Rhopalogastrum claviventre Bernhauer, 1912

Distribution.
Tribe Cordobanini Bernhauer, 1910
0374. Cordobanus Bernhauer, 1910: 386

Type species: Cordobanus mirabilis Bernhauer, 1910

Distribution.
Tribe Corotocini Fenyes, 1918
Subtribe Abrotelina Seevers, 1957
0375. Abroteles Casey, 1890: 190

Type species: Abroteles beaumonti Casey, 1890
Termitothymus Silvestri, 1901: 1 [synonymized by Borgmeier 1950: 638]
Type species: Termitothymus philetaerus Silvestri, 1901
Characters/Description: Borgmeier 1950: 653 (characters); Seevers 1957: 121 (characters in key), 122 (characters);
Kistner & Jacobson 1976: 29 (characters).
Key: Seevers 1957: 122 (species of the genus).
Distribution.
South America: Colombia, Ecuador, Peru, Bolivia, Brazil, Argentina.
0376. Termitophya Wasmann, 1902a: 95 (Fig. 76)

Type species: Termitophya heyeri Wasmann, 1902

Characters/Description: Fenyes 1920: 346 (characters); Seevers 1957: 71 (characters), 71 (characters in key);
Jacobson et al. 1986: 41 (characters).
Key: Wasmann & Holmgren 1911: 429 (some species of the genus); Seevers 1937: 19 (species of the genus);
Seevers 1957: 72 (species of the genus).
Distribution.
South America: Guyana, Ecuador, Peru, Bolivia, Brazil.
Subtribe Corotocina Fenyes, 1918
0377. Corotoca Schiødte, 1853: 101

Type species: Corotoca melantho Schiødte, 1853
Corotoca Schiødte, 1854: 8 [synonymized by Blackwelder 1952: 108]
Corotoca Schiødte, 1856a: 48 [synonymized by Blackwelder 1952: 108]
Corotoca Schiødte, 1856b: 171 [synonymized by Blackwelder 1952: 108]

Characters/Description: Seevers 1957: 141 (characters in key), 141 (characters); Jacobson et al. 1986: 118
(characters).
Distribution.
0378. Eburniola Mann, 1923: 333

Type species: Eburniola leucogaster Mann, 1923

(characters).
Distribution.
South America: Colombia, Guyana, Peru, Brazil.
0379. Nigriphilus Jacobson et al., 1986: 144

Type species: Thyreoxenus mexicanus Seevers, 1960

Distribution.
0380. Spirachtha Schiødte, 1853: 101

Type species: Spirachtha eurymedusa Schiødte, 1853
Spirachtha Schiødte, 1854: 12 [synonymized by Blackwelder 1952: 356]
Spirachtha Schiødte, 1856a: 52 [synonymized by Blackwelder 1952: 356]
Spirachtha Schiødte, 1856b: 176 [synonymized by Blackwelder 1952: 356]

(characters).
Distribution.
0381. Thyreoxenus Mann, 1923: 329

Type species: Thyreoxenus parviceps Mann, 1923

Characters/Description: Seevers 1957: 141 (characters in key), 156 (characters); Kistner & Jacobson 1976: 33
(characters); Jacobson et al. 1986: 139 (characters).
Key: Seevers 1946: 256 (species of the genus); Seevers 1957: 156 (species of the genus); Kistner & Jacobson 1976:
Distribution.
South America: Colombia, Guyana, Peru, Bolivia, Brazil.
Other regions: Oriental, Oceanic.
Subtribe Eburniogastrina Jacobson et al., 1986
0382. Eburniogaster Seevers, 1938: 424

Type species: Eburniogaster termitocolus Seevers, 1938

(characters); Ashe 2002b: 124 (characters in key).
Distribution.
0383. Termitonidia Seevers, 1938: 428

Type species: Termitonidia lunata Seevers, 1938
Distribution.
Subtribe Termitogastrina Bernhauer & Scheerpeltz, 1926
0384. Neotermitogaster Seevers, 1939: 8

Type species: Neotermitogaster colonus Seevers, 1939

Characters/Description: Seevers 1957: 71 (characters in key), 98 (characters); Jacobson et al. 1986: 71 (characters).
Distribution.
0385. Termitogaster Casey, 1889a: 384 (Fig. 75)

Type species: Termitogaster insolens Casey, 1889

Characters/Description: Fenyes 1920: 345 (characters); Seevers 1939: 1 (characters); Seevers 1957: 71 (characters
in key), 89 (characters); Kistner & Jacobson 1976: 17 (characters).
Key: Seevers 1937: 14 (species of the genus); Seevers 1946: 248 (species of the genus); Seevers 1957: 90 (species
of the genus).
Distribution.
South America: Colombia, Guyana, Ecuador, Peru, Bolivia, Brazil, Argentina.
0386. Termitoides Seevers, 1939: 7

Type species: Termitoides marginatus Seevers, 1939

Characters/Description: Seevers 1957: 71 (characters in key), 99 (characters).
Distribution.
0387. Termitomorpha Wasmann, 1894: 210

Type species: Termitomorpha meinerti Wasmann, 1894
Termitosomus Seevers, 1939: 3 [synonymized by Borgmeier 1950: 639, 640]
Type species: Termitogaster fissipennis Casey, 1890
Thaxteria Fenyes, 1921a: 17 [synonymized by Seevers 1957: 84]
Type species: Thaxteria insularis Fenyes, 1921

Kistner & Jacobson 1976: 8 (characters).
Key: Seevers 1946: 253 (species of Termitosomus); Seevers 1957: 84 (species of the genus).
Distribution.
West Indies: Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Guyana, Peru, Bolivia, Brazil.
0388. Termitonasus Borgmeier, 1959b: 290

Type species: Termitonasus palpalis Borgmeier, 1959

Characters/Description: Kistner & Jacobson 1976: 23 (characters); Ashe 2002b: 124 (characters in key).
Distribution.
Central America: Mexico, Costa Rica.
0389. Termitosyne Seevers, 1957: 81

Type species: Termitosyne platygastra Seevers, 1957
Characters/Description: Seevers 1957: 71 (characters in key).
Distribution.
0390. Termitosynodes Seevers, 1957: 83

Type species: Termitosynodes williamsi Seevers, 1957
Characters/Description: Seevers 1957: 71 (characters in key); Kistner & Jacobson 1976: 3 (characters); Ashe
Distribution.
0391. Trachopeplus Mann, 1923: 353

Type species: Trachopeplus setosus Mann, 1923
Distribution.
South America: Guyana, Ecuador.
0392. Xenogaster Wasmann, 1891: 651 (Fig. 79)

Type species: Xenogaster inflata Wasmann, 1891
Kistner & Jacobson 1976: 27 (characters); Ashe 2002b: 124 (characters in key).
Key: Wasmann & Holmgren 1911: 429 (some species of the genus); Seevers 1957: 101 (species of the genus).
Distribution.
South America: Guyana, Peru, Brazil, Uruguay.
0393. Xenopelta Mann, 1923: 356

Type species: Xenopelta cornuta Mann, 1923
Ceratoxenus Mann, 1923: 360 [synonymized by Emerson 1935: 369]
Type species: Ceratoxenus tricornis Mann, 1923
(characters).
Distribution.
Subtribe Termitoiceina Jacobson, Kistner & Pasteels, 1986
0394. Fonsechellus Silvestri, 1946c: 312

Type species: Fonsechellus diversicolor Silvestri, 1946
Trianellus Silvestri, 1946c: 315 [synonymized by Seevers 1957: 130]
Type species: Fonsechellus bicolor Silvestri, 1946

Characters/Description: Seevers 1957: 130 (characters); Kistner 1994: 201 (characters).
Key: Seevers 1957: 128 (species of the genus); Kistner 1994: 203 (species of the genus).
Distribution.
0395. Mormellus Silvestri, 1946c: 323

Type species: Mormellus bicolor Silvestri, 1946

Characters/Description: Seevers 1957: 134 (characters).
Distribution.
0396. Oecidiophilus Silvestri, 1946c: 331

Type species: Oecidiophilus mimellus Silvestri, 1946


Distribution.
0397. Parvidolum Silvestri, 1946c: 318

Type species: Parvidolum microsomatis Silvestri, 1946

Distribution.
0398. Perlinctus Silvestri, 1946c: 320

Type species: Perlinctus quaesitus Silvestri, 1946

Characters/Description: Seevers 1957: 132 (characters); Jacobson et al. 1986: 93 (characters).
Distribution.
0399. Termitoiceus Silvestri, 1901: 5

Type species: Termitoiceus anastrephoproctus Silvestri, 1901

Characters/Description: Fenyes 1920: 345 (characters); Seevers 1957: 129 (characters); Jacobson et al. 1986: 86
(characters).
Distribution.
Subtribe Termitopithina Jacobson, Kistner & Pasteels, 1986
0400. Termitopithus Seevers, 1957: 80

Type species: Termitopithus crassiusculus Seevers, 1957

Characters/Description: Seevers 1957: 71 (characters in key); Jacobson et al. 1986: 80 (characters).
Distribution.
Subtribe Timeparthenina Fenyes, 1921a
0401. Ptocholellus Silvestri, 1946c: 305

Type species: Ptocholellus mimus Silvestri, 1946
Key: Campbell 1973e: 92 (species of the genus as Ptochellus [sic]).
Distribution.
0402. Reginamimus Kistner, 2000: 191

Type species: Reginamimus milli Kistner, 2000

Distribution.
0403. Termitozophilus Silvestri, 1901: 7

Type species: Termitozophilus laetus Silvestri, 1901
Corymbogaster Mann, 1923: 346 [synonymized by Borgmeier 1950: 638, 640]
Type species: Corymbogaster miranda Mann, 1923
Characters/Description: Fenyes 1920: 348 (characters); Seevers 1957: 65 (characters); Campbell 1973e: 92
(characters in key); Jacobson et al. 1986: 18 (characters).
Key: Campbell 1973e: 92 (species of the genus).
Distribution.
South America: Guyana, Brazil, Paraguay, Argentina.
0404. Termituncula Borgmeier, 1950: 662 (Fig. 77)

Type species: Termituncula gracilipes Borgmeier, 1950

Distribution.
0405. Timeparthenus Silvestri, 1901: 10

Type species: Timeparthenus regius Silvestri, 1901
Autuoria Silvestri, 1946c: 309 [synonymized by Campbell 1973e: 84]
Type species: Autuoria elegantulum Silvestri, 1946

Characters/Description: Seevers 1957: 69 (characters), 68 (characters as Autuoria); Campbell 1973e: 84
(characters), 92 (characters in key); Jacobson et al. 1986: 22 (characters).
Distribution.

Incertae sedis in Tribe Corotocini Fenyes, 1918
0406. Termitosius Silvestri, 1901: 8

Type species: Termitosius pauciseta Silvestri, 1901

Characters/Description: Fenyes 1920: 347 (characters); Seevers 1957: 134 (characters).
Distribution.
Tribe Crematoxenini Mann, 1921
0407. Beyeria Fenyes, 1910: 118

Type species: Beyeria vespa Fenyes, 1910
Acamatoxenus Mann, 1925: 76 [synonymized by Jacobson et al. 1987: 317]
Type species: Acamatoxenus suavis Mann, 1925

Jacobson et al. 1987: 317 (characters); Jacobson & Kistner 1992: 131 (characters); Ashe 2002b: 126
Key: Jacobson et al. 1987: 321 (species of the genus); Jacobson & Kistner 1992: 133 (species of the genus).
Distribution.
0408. Crematoxenus Mann, 1921: 547

Type species: Crematoxenus aenigma Mann, 1921

Characters/Description: Seevers 1965: 207 (characters in key), 217 (characters); Jacobson & Kistner 1992: 159
Distribution.
0409. Cryptomimus Reichensperger, 1926: 114

Type species: Cryptomimus handlirschi Reichensperger, 1926

(characters).
Key: Jacobson & Kistner 1992: 164 (species of the genus).
Distribution.
0410. Diploeciton Wasmann, 1923: lxiii

Type species: Diploeciton constrictum Wasmann, 1923
Acamatoteras Reichensperger, 1936b: 189 [synonymized by Blackwelder 1952: 33]
Type species: Acamatoteras nevermanni Reichensperger, 1936
Characters/Description: Reichensperger 1939a: 134 (characters); Jacobson & Kistner 1992: 150 (characters);
Seevers 1965: 207 (characters in key), 219 (characters).
Distribution.
0411. Ecitosius Seevers, 1965: 272

Type species: Ecitosius gracilis Seevers, 1965

Characters/Description: Seevers 1965: 233 (characters in key); Jacobson & Kistner 1992: 115 (characters); Ashe
Distribution.
0412. Ecitotima Seevers, 1965: 273

Type species: Ecitotima miriventris Seevers, 1965

Characters/Description: Seevers 1965: 233 (characters in key); Jacobson & Kistner 1992: 124 (characters).
Distribution.
0413. Neivaphilus Jacobson & Kistner, 1992: 139

Type species: Neivaphilus watkinsi Jacobson & Kistner, 1992

Distribution.
0414. Philacamatus Bruch, 1933a: 206 (Fig. 70)

Type species: Philacamatus bosqi Bruch, 1933

(characters).
Distribution.
0415. Pulicomorpha Mann, 1924: 87

Type species: Pulicomorpha coecum Mann, 1924

Distribution.
Tribe Diestotini Mulsant & Rey 1871
0416. Diestota Mulsant & Rey, 1870: 194

Type species: Diestota mayeti Mulsant & Rey, 1870
Hypselusa Bernhauer in Bernhauer & Scott, 1931: 592 [synonymized by Pace 1986b: 108]
Type species: Hypselusa scotti Bernhauer, 1931
Ousilusa Cameron, 1920a: 234 [synonymized by Pace 2000b: 131]
Type species: Ousilusa myrmicobia Cameron, 1920
Prosilusa Cameron, 1920a: 236 [synonymized by Blackwelder 1952: 323]
Type species: Prosilusa rufa Cameron, 1920
Subgenera:
Aphaenoglossa Peyerimhoff, 1937: 103
Type species: Aphaenoglossa normandi Peyerimhoff, 1937
Apheloglossa Casey, 1893: 348 [*]
Type species: Apheloglossa rufipennis Casey, 1893
Amenusa Casey, 1906: 349 [synonymized by Seevers 1978: 165]
Type species: Amenusa angustula Casey, 1906
Elachistarthron Notman, 1920: 715 [synonymized by Pace 1986a: 421]
Type species: Elachistarthron ambiguum Notman, 1920
Orthodiatelus Notman, 1920: 716 [synonymized by Pace 1986a: 421]
Type species: Orthodiatelus innotabilis Notman, 1920
Pectusa Casey, 1911: 197 [synonymized by Seevers 1978: 165]
Type species: Pectusa oblonga Casey, 1911
Diestota Mulsant & Rey, 1870: 194
Type species: Diestota mayeti Mulsant & Rey, 1870
Diestota Mulsant & Rey, 1872a: 170 [Newton & Thayer 2003]
Diestota Mulsant & Rey, 1873c: 74 [Newton & Thayer 2003]
Eutrichostygna Pace, 1986a: 421 [*]
Type species: Diestota chavin Pace, 1986

Characters/Description: Pace 1986a: 419 (characters in key); Ashe 2002b: 129 (characters in key); Pace 2008a: 254
(characters in key as subgenera Diestota and Apheloglossa).
Key: Pace 1986a: 419 (subgenera of the genus); Klimaszewski & Peck 1998: 230 (species of the Galapagos Isl.);
Newton & Peck 2006: 137 (species of the Galapagos Isl.); Pace 2008a: 270 (species of the subgenus Diestota
of Ecuador and Peru), 270 (species of the subgenus Apheloglossa of Ecuador and Peru).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Puerto Rico, US Virgin Isl., Saint-Martin, Guadeloupe, St. Vincent &
Grenadines, Grenada.
Central America: Mexico, Guatemala, Nicaragua, Panama.
0417. Eudera Fauvel, 1866b: 257 (Fig. 55)

Type species: Eudera sculptilis Fauvel, 1866
Cajachara Ashe & Leschen, 1995: 86 [synonymized by Pace 2008a: 273]
Type species: Cajachara carltoni Ashe & Leschen, 1995
Eudera Fauvel, 1866a: 8 [Newton & Thayer 2003]
Type species: Eudera sculptilis Fauvel, 1866

Characters/Description: Coiffait & Sáiz 1967: 65 (characters in key), 83 (characters); Pace 1986a: 419 (characters
in key); Pace 2008a: 254 (characters in key).
Key: Coiffait & Sáiz, 1967: 84 (species of Chile); Pace 2008a: 273 (species of the Ecuador and Peru).
Distribution.
West Indies: US Virgin Isl., Guadeloupe.
South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Brazil, Chile, Argentina.
0418. Gansia Sharp, 1883: 282

Type species: Gansia bicolor Sharp, 1883
Characters/Description: Ashe & Lingafelter 1996: 253 (characters); Ashe 2002b: 129 (characters in key); Pace
2008a: 254 (characters in key), 274 (characters in key).
Key: Ashe & Lingafelter 1996: 257 (species of Mexico and Central America).
Distribution.
South America: Colombia, Ecuador, Bolivia, Brazil.
0419. Gansiella Pace, 2008a: 274

Type species: Gansiella peruviana Pace, 2008

Characters/Description: Pace 2008a: 254 (characters in key), 274 (characters in key).
Key: Pace 2014b: 87 (speceis of the genus).
Distribution.
South America: French Guiana, Peru.
0420. Parasilusa Bernhauer, 1908: 338

Type species: Parasilusa iheringi Bernhauer, 1908
Ocyupora Pace, 1986a: 421 [synonymized by Pace 2008a: 254]
Type species: Diestota inca Pace, 1986

Characters/Description: Pace 1996b: 417 (characters in key); Pace 2008a: 254 (characters in key).
Key: Pace 2008a: 254 (species of Ecuador and Peru).

Distribution.
South America: French Guiana, Ecuador, Peru, Brazil.
0421. Plesiomalota Pace, 1983d: 296

Type species: Homalota republicana Bernhauer, 1921
Subgenera:
Chaetopiezusa Pace, 1985: 378 [*]
Type species: Plesiomalota venezuelanorum Pace, 1985
Heteromalota Pace, 1983d: 298 [*]
Type species: Plesiomalota uhligi Pace, 1983
Leptosomaphya Pace, 1983a: 91 [*]
Type species: Leptosomaphya franzi Pace, 1983
Microchara Pace, 1983d: 300 [*]
Type species: Microchara flava Pace, 1983
Ischnosomaops Pace, 1983a: 93 [synonymized by Pace 1986a: 424]
Type species: Leptosomaphya minor Pace, 1983
Microusa Pace, 1983a: 95 [*]
Type species: Microusa franzi Pace, 1983
Plesiomalota Pace, 1983d: 296 [*]
Type species: Homalota republicana Bernhauer, 1920

Characters/Description: Pace 1986a: 419 (characters in key); Pace 2008a: 254 (characters in key).
Key: Pace 1986a: 423 (subgenera of the genus); Pace 2001b: 418 (subgenera of the genus); Pace 2008a: 258
(species of Ecuador and Peru).
Distribution.
Chile, Argentina.
0422. Tachiona Sharp, 1883: 284

Type species: Tachiona deplanata Sharp, 1883

Characters/Description: Fenyes 1920: 127 (characters); Ashe & Wheeler 1988: 177 (characters); Ashe 2002b: 129
Key: Ashe & Wheeler 1988: 177 (some species of the genus); Ashe 1990: 229 (species of the genus); Ashe 1993:
Distribution.
Tribe Diglottini Jacobson, 1909
0423. Diglotta Champion, 1887: 228 (Fig. 51)

Type species: Diglossa mersa Haliday, 1837
Diglossa Haliday, 1837: 252 [synonymized by Bernhauer & Scheerpeltz 1926: 509]
Diglotta Champion, 1899: 265 [synonymized by Blackwelder 1952: 125]
Characters/Description: Ashe 2002b: 129 (characters in key); Caron & Ribeiro-Costa 2008: 56 (characters).
Distribution.
Other regions: Nearctic, Palaearctic, Oceanic.
Tribe Ecitogastrini Fenyes, 1918
0424. Ecitogaster Wasmann, 1900a: 404

Type species: Ecitogaster schmalzi Wasmann, 1900
Ecitogaster Wasmann, 1900b: 222 [synonymized by Blackwelder 1952: 140]
Characters/Description: Borgmeier 1949: 126 (characters); Seevers 1965: 300 (characters); Ashe 2002b: 124
Key: Borgmeier 1949: 127 (species of the genus).
Distribution.
Tribe Falagriini Mulsant & Rey, 1873b
0425. Akrincastiba Pace, 2001b: 421

Type species: Akrincastiba moreti Pace, 2001

Distribution.
0426. Aleodorus Say, 1833: 60 (Fig. 43)

Type species: Aleochara bilobata Say, 1830
Aleodorus Say, 1839: 157 [synonymized by Blackwelder 1952: 45]
Chitalia Sharp, 1883: 235 [synonymized by Bernhauer & Scheerpeltz 1926: 571]
Type species: Chitalia crenata Sharp, 1883
Characters/Description: Fenyes 1920: 148 (characters as Chitalia); Ashe 2002b: 126 (characters in key); Delgado
& Santiago-Jiménez 2009: 67 (characters).
Key: Pace 1990a: 162 (species of the Neotropical region).
Distribution.
South America: Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil, Argentina.
Other regions: Nearctic, Oriental.

0427. Anaulacaspis Ganglbauer, 1895: 256
Type species: Aleochara nigra Gravenhorst, 1802
Euphorbagria Assing, 1997a: 64 [synonymized by Assing 1997a: 65]
Type species: Falagria rotundicollis Lindberg, 1953
Falagriola Reitter, 1909: 74 [synonymized by Blackwelder 1952: 164]
Melagria Casey, 1906: 230 [synonymized by Blackwelder 1952: 237]

Characters/Description: Fenyes 1920: 153 (characters in key), 154 (characters).
Distribution.
South America: Colombia, Venezuela, Brazil, Chile, Argentina.
0428. Borboropora Kraatz in Kraatz & Fuss, 1862: 405

Type species: Borboropora kraatzii Fuss, 1862
Aneurota Casey, 1893: 347 [synonymized by Bernhauer & Scheerpeltz 1926: 573]
Type species: Aneurota sulcifrons Casey, 1893
Orthagria Casey, 1906: 260 [synonymized by Bernhauer & Scheerpeltz 1926: 573]
Type species: Falagria quadriceps LeConte, 1866
Pseudoscopaeus Weise in Reitter, 1877: 8 [synonymized by Bernhauer & Scheerpeltz 1926: 573]
Type species: Pseudoscopaeus reitteri Weise, 1877

Characters/Description: Fenyes 1920: 146 (characters); Delgado & Santiago-Jiménez 2009: 67 (characters).
Key: Delgado & Santiago-Jiménez 2009: 66 (species of the New World).
Distribution.
0429. Bryobiota Casey, 1893: 367

Type species: Phytosus bicolor Casey, 1885

Characters/Description: Fenyes 1920: 130 (characters); Ahn & Ashe 1995: 143 (characters), 151 (characters in
key); Ashe 2002b: 129 (characters in key).
Key: Ahn & Ashe 1995: 145 (species of the genus).
Distribution.
0430. Drepanopora Bernhauer, 1908: 345

Type species: Drepanopora borboroporoides Bernhauer, 1908

Distribution.
0431. Falagria Leach in Samouelle, 1819: 177
Type species: Staphylinus sulcatus Paykull, 1789
Coenobiotes Gistel, 1856: 387 [synonymized by Blackwelder 1952: 102]
Type species: Staphylinus sulcatus Paykull, 1789

Characters/Description: Lynch-Arribálzaga 1884: 29 (characters); Fenyes 1920: 152 (characters); Ashe 2002b: 126
Key: Fenyes 1920: 136 (subgenera); Pace 2008a: 275 (species of Ecuador and Peru).
Distribution.
West Indies: Jamaica, Grenada.
South America: Colombia, Venezuela, Ecuador, Bolivia, Brazil, Paraguay, Argentina.
0432. Falagrioma Casey, 1906: 230

Type species: Falagria thoracica Stephens, 1832

Distribution.
0433. Guajira Bierig, 1938b: 146

Type species: Guajira cubana Bierig, 1938
Distribution.
West Indies: Cuba.
0434. Leptagria Casey, 1906: 249

Type species: Leptagria perexilis Casey, 1906

Distribution.
West Indies: Jamaica, Dominican Republic, US Virgin Isl., British Virgin Isl., St. Lucia, St. Vincent &
South America: Ecuador, Bolivia, Brazil.
Other regions: Nearctic, Madagascan, Oriental.
0435. Lissagria Casey, 1906: 252

Type species: Falagria laeviuscula LeConte, 1866
Omoschema Notman, 1920: 731 [synonymized by Hoebeke 1985: 930]
Type species: Omoschema laticeps Notman, 1920
Key: Eldredge 2012a: 227 (species of the genus).

Distribution.
South America: Venezuela, Ecuador.
0436. Myrmecocephalus W.J. MacLeay, 1871: 134

Type species: Myrmecocephalus cingulatus W.J. MacLeay, 1871
Lorinota Casey, 1906: 238 [synonymized by Blackwelder 1952: 227]
Type species: Falagria cingulata LeConte, 1866
Stenagria Sharp, 1883: 237 [synonymized by Blackwelder 1952: 358]
Type species: Stenagria gracilipes Sharp, 1883
Stilicioides Broun, 1880: 95 [synonymized by Blackwelder 1952: 362]
Type species: Stilicioides micans Broun, 1880

Characters/Description: Fenyes 1920: 158 (characters as Lorinota), 159 (characters as Stenagria); Ashe 2002b: 125
(characters in key); Newton & Peck 2006: 135 (characters in key).
Distribution.
South America: French Guiana, Ecuador, Galapagos Isl., Bolivia, Brazil, Chile, Argentina.
0437. Myrmecopora Saulcy, 1865: 429

Type species: Myrmecopora publicana Saulcy, 1865
Subgenera:
Amechanopora Pace, 2004a: 196
Type species: Myrmecopora vietnamensis Pace, 2004
Anatolagria Assing, 2004a: 48
Type species: Myrmecopora effeminata Assing, 2004
Euphorbiusa Assing, 1997b: 132
Type species: Myrmecopora gravata Peyerimhoff, 1945
Iliusa Mulsant & Rey, 1873b: 6
Type species: Tachyusa fugax Erichson, 1839
Iliusa Mulsant & Rey, 1874c: 38 [Newton & Thayer 2003]
Ilyusa Mulsant & Rey, 1875c: 419 [Newton & Thayer 2003]
Ilyusa Mulsant & Rey, 1875a: 445 [Newton & Thayer 2003]
Lamproxenusa Assing, 1997c: 339
Type species: Tachyusa rufescens Sharp, 1874
Myrmecopora Saulcy, 1865: 429
Type species: Myrmecopora publicana Saulcy, 1865
Paraxenusa Assing, 1997b: 101
Type species: Tachyusa laesa Erichson, 1839
Xenusa Mulsant & Rey, 1873b: 6 [*]
Type species: Tachyusa uvida Erichson, 1840
Xenusa Mulsant & Rey, 1874c: 38 [Newton & Thayer 2003]
Xenusa Mulsant & Rey, 1875a: 436 [Newton & Thayer 2003]
Xenusa Mulsant & Rey, 1875c: 410 [Newton & Thayer 2003]
Characters/Description: Fenyes 1920: 165 (characters), 166 (characters of subgenus Iliusa), 167 (characters
subgenus Xenusa); Ahn & Ashe 1995: 151 (characters in key).
Distribution.
0438. Tropidera Bernhauer, 1908: 349

Type species: Tropidera jenseni Bernhauer, 1908
Mesaraeus Fenyes, 1921a: 21 [synonymized by Blackwelder 1952: 238]
Type species: Mesaraeus laevigatus Fenyes, 1921

Distribution.
South America: French Guiana, Brazil, Paraguay, Argentina.
Tribe Geostibini Seevers, 1978
0439. Alevonota Thomson, 1858: 35

Type species: Homalota atricapilla Mulsant & Rey, 1852
Liota Mulsant & Rey, 1873b: 4 [synonymized by Blackwelder 1952: 222]
Type species: Homalota gracilenta Erichson, 1839
Liota Mulsant & Rey, 1874c: 36 [synonymized by Blackwelder 1952: 222]
Liota Mulsant & Rey, 1875a: 148 [synonymized by Blackwelder 1952: 222]
Liota Mulsant & Rey, 1875c: 122 [synonymized by Blackwelder 1952: 222]

Distribution.
0440. Aloconota Thomson, 1858: 33

Type species: Tachyusa immunita Erichson, 1840
Subgenera:
Aloconota Thomson, 1858: 33 [*]
Type species: Tachyusa immunita Erichson, 1840
Glossola Fowler, 1888: 66 [synonymized by Seevers 1978: 110]
Type species: Homalota gregaria Erichson, 1839
Disopora Thomson, 1859: 39
Type species: Homalota languida Erichson, 1837
Disoporina Fenyes, 1918: 22 [Newton & Thayer 2003]
Type species: Atheta ernestinae Bernhauer, 1898
Disoporina Fenyes, 1920: 205 [synonymized by Bernhauer & Scheerpeltz 1926: 611]
Taphrodota Casey, 1906: 338

Type species: Taphrodota ventralis Casey, 1906
Terasota Casey, 1906: 337
Type species: Terasota brunneipes Casey, 1906
Distribution.
South America: Ecuador, Peru, Chile, Argentina.
0441. Seeversiella Ashe, 1986: 501

Type species: Seeversiella bispinosa Ashe, 1986

Characters/Description: Ashe 1986: 511 (characters in key); Gusarov 2003f: 3 (characters).
Key: Gusarov 2003f: 10 (species of the genus).
Distribution.
Central America: Mexico, Guatemala, Honduras, El Salvador, Costa Rica, Panama.
Tribe Gymnusini Heer, 1839
0442. Adinopsis Cameron, 1919: 242

Type species: Adinopsis rufobrunnea Cameron, 1919
Condylopennis Klimaszewski & Jansen, 1995: 329 [nomen nudum, Newton & Thayer 2003]

Characters/Description: Klimaszewski 1979: 60 (characters); Klimaszewski & Jansen 1995: 327 (characters); Ashe
2002a: 72 (characters in key); Ashe 2002b: 123 (characters in key).
Key: Klimaszewski 1979: 71 (Nearctic and Neotropical species).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, St. Lucia, Trinidad-Tobago.
South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Brazil, Paraguay.
0443. Allodinopsis Ashe, 2002a: 62 (Fig. 44)

Type species: Allodinopsis howdeni Ashe, 2002

Characters/Description: Ashe 2002a: 72 (characters in key).
Distribution.
0444. Metadeinopsis Klimaszewski, 1979: 78
Type species: Metadeinopsis brunnea Klimaszewski, 1979

Characters/Description: Ashe 2002a: 72 (characters in key).
Key: Klimaszewski 1979: 79 (species of the genus); Klimaszewski 1982: 333 (species of the genus); Klimaszewski
& Génier 1985: 63 (species of the genus).
Distribution.
South America: Colombia, French Guiana, Ecuador, Peru, Brazil, Paraguay.
Tribe Homalotini Heer, 1839
Subtribe Bolitocharina Thomson, 1859
0445. Bolitochara Mannerheim, 1830: 75

Type species: Aleochara pulchra Gravenhorst, 1806
Agaribiota Bierig, 1937b: 279 [synonymized by Blackwelder 1952: 41]
Type species: Bolitochara cinctigastra Bierig, 1937
Ditropalia Casey, 1906: 263 [synonymized by Seevers 1978: 164]
Type species: Bolitochara bella Märkel, 1844
Characters/Description: Fenyes 1920: 114 (characters as Ditropalia); Coiffait & Sáiz 1967: 65 (characters in key),
79 (characters); Ashe 1992: 365 (characters in key), 367 (characters).
Key: Coiffait & Sáiz 1967: 80 (Chilean species).
Distribution.
West Indies: Cuba.
Other regions: Palaearctic, Oriental, Australian, Oceanic.
0446. Habramicrada Pace, 2007: 225 (Fig. 58)

Type species: Habramicrada boliviensis Pace, 2007

Distribution.
South America: Peru, Bolivia, Brazil.
0447. Leptusa Kraatz, 1856b: 60

Type species: Bolitochara pulchella Mannerheim, 1830
Subgenera:
Adexiopisalia Pace, 1982c: 585
Type species: Leptusa hummleri Bernhauer, 1910
Adoxopisalia Pace, 1989: 130
Type species: Leptusa opaca Casey, 1893
Afroleptusa Pace, 1983c: 62
Type species: Leptusa sparsa Fauvel, 1898
Akratopisalia Pace, 1996a: 29

Type species: Leptusa cribrata Pace, 1996
Amalopisalia Pace, 1983c: 97
Type species: Leptusa janetscheki Scheerpeltz, 1956
Anisopisalia Pace, 1989: 214
Type species: Leptusa luigionii Bernhauer, 1935
Anomopisalia Pace, 1983c: 82
Type species: Leptusa granulicauda Eppelsheim, 1890
Anosiopisalia Pace, 1995: 247
Type species: Leptusa nemoricultrix Pace, 1995
Aphaireleptusa Pace, 1996a: 30
Type species: Leptusa anmashanensis Pace, 1996
Aleteleptusa Pace, 1997a: 753 [Smetana 2004: 429]
Type species: Leptusa chinensis Pace, 1997
Mimumenepisalia Pace, 1997a: 752 [Smetana 2004: 429]
Type species: Leptusa gonggamontis Pace, 1997
Baldopisalia Pace, 1983e: 130
Type species: Leptusa veronensis Pace, 1983
Boreoleptusa Pace, 1989: 82
Type species: Leptusa norvegica Strand, 1941
Bothrydiopisalia Scheerpeltz, 1966b: 20
Type species: Leptusa bosnica Eppelsheim, 1892
Bryopisalia Pace, 1989: 239
Type species: Leptusa pilatensis Scheerpeltz, 1972
Chondrelytropisalia Scheerpeltz, 1976c: 55
Type species: Leptusa nepalica Scheerpeltz, 1976
Chondridiopisalia Pace, 1983c: 85
Type species: Leptusa granulifera Pace, 1983
Chondridiopisalia Scheerpeltz, 1966b: 19 [nomen nudum, Newton & Thayer 2003]
Type species: Leptusa granulifera Scheerpeltz, 1966
Chondridiopisalia Lohse, 1974: 53 [Newton & Thayer 2003]
Chondropisalia Scheerpeltz, 1966b: 14
Type species: Leptusa schaschli Ganglbauer, 1897
Cidnopisalia Pace, 1989: 235
Type species: Leptusa cavallensis Pace, 1980
Comphopisalia Scheerpeltz, 1966b: 16
Type species: Leptusa brucki Scriba, 1867
Craspedopisalia Scheerpeltz, 1966b: 19
Type species: Leptusa carinata Bernhauer, 1909
Cryptopisalia Pace, 1983c: 88
Type species: Leptusa montisparii Scheerpeltz, 1953
Cryptopisalia Pace, 1983e: 129 [Newton & Thayer 2003]
Cyllopisalia Pace, 1982a: 40
Type species: Leptusa difformis Mulsant & Rey, 1853
Cyllopisalia Pace, 1983c: 79 [Smetana 2004: 431]
Parapisalia Scheerpeltz, 1966b: 18 [synonymized by Pace 1982a: 40]
Sipalia Mulsant & Rey, 1853b: 32 [synonymized by Pace 1982a: 40]
Type species: Homalota difformis Mulsant & Rey, 1853
Dendroleptusa Pace, 1983c: 65
Type species: Leptusa fumida Erichson, 1839
Drepanoleptusa Pace, 1982d: 93
Type species: Leptusa annapurnensis Pace, 1982
Dysleptusa Pace, 1982c: 580
Type species: Oxypoda fuliginosa Aubé, 1850
Ectinopisalia Scheerpeltz, 1966b: 18
Type species: Leptusa baldensis Ganglbauer, 1895
Emmelopisalia Pace, 1983c: 92
Type species: Leptusa norica Pace, 1979
Entomophallopisalia Pace, 1983c: 90
Type species: Leptusa asturiensis Eppelsheim, 1880
Eospisalia Pace, 1982c: 583
Type species: Leptusa ishizuchiensis Pace, 1982
Eremopisalia Pace, 1989: 196
Type species: Leptusa marianii Pace, 1978
Eucryptusa Casey, 1906: 345
Type species: Silusa nanula Casey, 1893
Evaniopisalia Pace, 1982a: 38
Type species: Leptusa ultracollensis Pace, 1982
Exaeretopisalia Pace, 1989: 233
Type species: Leptusa cansigliensis Pace, 1975
Gallopisalia Pace, 1989: 179
Type species: Leptusa lapidicola Brisout de Barneville, 1863
Gatineauleptusa Klimaszewski in Klimaszewski et al., 2018: 363
Type species: Leptusa gatineauensis Klimaszewski & Pelletier in Klimaszewski et al., 2004
Geopisalia Pace, 1984: 214
Type species: Leptusa peyerimhoffi Bernhauer, 1935
Gnopheropisalia Pace, 1983c: 76
Type species: Leptusa granulipennis Eppelsheim, 1880
Habropisalia Pace, 1989: 216
Type species: Leptusa laticeps Scheerpeltz, 1972
Halmaeusa Kiesenwetter in Kiesenwetter & Kirsch, 1877: 160 [*]
Type species: Halmaeusa antarctica Kiesenwetter, 1877
Antarctophytosus Enderlein, 1909: 377 [synonymized by Steel 1964: 366]
Type species: Phytosus atriceps Waterhouse, 1875
Paraphytosus Cameron, 1917c: 125 [synonymized by Steel 1964: 366]
Type species: Phytosus atriceps Waterhouse, 1875
Austromalota Brèthes, 1925: 170 [synonymized by Steel 1964: 366]
Type species: Austromalota rufimixta Brèthes, 1925
Hemipasilia Pace, 1989: 90
Type species: Leptusa oertzeni Eppelsheim, 1888
Heteroleptusa Pace, 1989: 160
Type species: Sipalia frontalis Casey, 1893
Heterotyphlopasilia Pace, 1983c: 66
Type species: Leptusa serbica Scheerpeltz, 1935
Homopisalia Pace, 1982c: 584

Type species: Leptusa bengalensis Pace, 1982
Ischnopisalia Pace, 1989: 226
Type species: Leptusa poggii Pace, 1978
Kochliodepisalia Pace, 1996a: 35
Type species: Leptusa spirarum Pace, 1996
Koilokoryphepisalia Pace, 1999a: 210
Type species: Leptusa argentierensis Pace, 1999
Lasiopisalia Pace, 1983c: 76
Type species: Leptusa crenulata Bernhauer, 1899
Lathripisalia Pace, 1983e: 135
Type species: Leptusa montispasubii Pace, 1975
Leptopasilia Scheerpeltz, 1966b: 13
Type species: Leptusa hopffgarteni Eppelsheim, 1881
Leptusa Kraatz, 1856b: 60
Type species: Bolitochara pulchella Mannerheim, 1830
Sipalia Mulsant & Rey, 1853a: 45 [Newton & Thayer 2003]
Liopisalia Pace, 1989: 177
Type species: Leptusa serullazi Sainte-Claire Deville, 1919
Megacolypisalia Pace, 1983e: 133
Type species: Leptusa laevicauda Scheerpeltz, 1958
Melopisalia Pace, 1983c: 88
Type species: Leptusa pasubiana Scheerpeltz, 1953
Metaplasmopasilia Pace, 1989: 104
Type species: Leptusa apennina Holdhaus, 1924
Microcolypisalia Pace, 1989: 196
Type species: Leptusa occulta Pace, 1975
Micropisalia Scheerpeltz, 1948: 164
Type species: Leptusa alpestris Scheerpeltz, 1935
Myopopisalia Scheerpeltz, 1966b: 14
Type species: Leptusa bonvouloiri Brisout de Barneville, 1867
Nanoglossa Fauvel, 1868a: 379 [*]
Type species: Microglossa chilensis Fauvel, 1866
Microglossa Fauvel, 1866b: 282 [synonymized by Fauvel 1868a: 379]
Microglossa Fauvel, 1866a: 33 [Newton & Thayer 2003]
Pareuryusa Scheerpeltz, 1972d: 131 [synonymized by Pace 1989: 110]
Type species: Euryusa parallela Fairmaire & Germain, 1862
Nanopisalia Pace, 1983c: 94
Type species: Homalota piceata Mulsant & Rey, 1853
Micropisalia Scheerpeltz, 1966b: 20 [synonymized by Pace 1983c: 94]
Nanopisalia Pace, 1983e: 132 [Newton & Thayer 2003]
Neopisalia Scheerpeltz, 1966b: 16
Type species: Leptusa microphthalma Reitter, 1887
Nesopisalia Pace, 1992b: 112
Type species: Leptusa centralis Pace, 1992
Stenoleptusa Scheerpeltz, 1966b: 10 [synonymized by Assing 2009c: 1286]
Type species: Bolitochara laeviuscula Hochhuth, 1849
Oligopisalia Scheerpeltz, 1966b: 20
Type species: Leptusa flavicornis Brancsik, 1874
Oncopisalia Pace, 1982c: 589
Type species: Leptusa monachorum Bernhauer, 1912
Oreopisalia Pace, 1983c: 82
Type species: Leptusa hoelzeli Scheerpeltz, 1948
Pachygastropisalia Scheerpeltz, 1966b: 15
Type species: Leptusa lativentris Sharp, 1873
Pachygluta Thomson, 1858: 34
Type species: Oxypoda ruficollis Erichson, 1839
Parapisalia Scheerpeltz, 1948: 159
Type species: Leptusa puellaris Hampe, 1863
Pasilia Mulsant & Rey, 1871: 226
Type species: Homalota nubigena Kiesenwetter, 1861
Pasilia Mulsant & Rey, 1872b: 316 [Newton & Thayer 2003]
Pasilia Mulsant & Rey, 1873c: 73 [Newton & Thayer 2003]
Pisalia Mulsant & Rey, 1871: 234
Type species: Homalota globulicollis Mulsant & Rey, 1853
Pisalia Mulsant & Rey, 1872b: 324 [Newton & Thayer 2003]
Pisalia Mulsant & Rey, 1873c: 73 [Newton & Thayer 2003]
Tropidiopasilia Scheerpeltz, 1966b: 11 [synonymized by Pace 1989: 190]
Type species: Leptusa tricolor Scriba, 1870
Planepisalia Pace, 1989: 217
Type species: Leptusa kochiana Bernhauer, 1936
Protoleptusa Semenov in Semenov et al., 2001: 156
Type species: Leptusa defuncta Semenov, Perkovsky & Petrenko, 2001
Rhombopisalia Scheerpeltz, 1953: 66
Type species: Leptusa knabli Bernhauer, 1909
Roubaliusa Scheerpeltz, 1966b: 13
Type species: Leptusa storkani Roubal, 1916
Scelopisalia Scheerpeltz, 1966b: 18
Type species: Leptusa gracilipes Krauss, 1899
Scoliophallopisalia Pace, 1989: 232
Type species: Leptusa pascuorum Pace, 1975
Scoliophallopisalia Pace, 1981: 115 [nomen nudum, Newton & Thayer 2003]
Scoliophallopisalia Anonymous, 1984: 249 [Newton & Thayer 2003]
Type species: Leptusa blesioi Pace, 1981
Scolioplatypisalia Pace, 1989: 86
Type species: Leptusa munelensis Apfelbeck, 1907
Stictopisalia Scheerpeltz, 1966b: 16
Type species: Leptusa eximia Kraatz, 1856
Synpisalia Scheerpeltz, 1966b: 16 [synonymized by Pace 1983c: 56]
Type species: Leptusa caucasica Eppelsheim, 1877
Toxophallopisalia Pace, 1983c: 99
Type species: Leptusa austriaca Scheerpeltz, 1935
Trichopasilia Scheerpeltz, 1966b: 12

Type species: Leptusa major Bernhauer, 1900
Typhlopasilia Ganglbauer, 1895: 278
Type species: Leptusa kaufmanni Ganglbauer, 1895
Ulitusa Casey, 1906: 347
Type species: Ulitusa cribratula Casey, 1906
Yunnaleptusa Assing, 2008b: 256
Type species: Leptusa curvata Assing, 2006

Characters/Description: Lynch-Arribálzaga 1884: 87 (characters); Fenyes 1920: 120 (characters), 122 (characters
subgenera Typhlosilia and Leptusa), 123 (characters subgenera Eucryptusa and Pachygluta), 124 (characters
subgenera Oreusa and Pasilia), 125 (characters subgenera Pisalia); Pace 1989: 25 (characters), 110
(characters of the subgenus Nanoglossa), 244 (characters of the subgenus Halmaeusa).
Key: Bernhauer 1900a: 402 (Subgenera and some species); Fenyes 1920: 121 (subgenera); Pace 1989: 25
(subgenera), 110 (species of the subgenus Nanoglossa), 246 (species of the subgenus Halmaeusa); Pace
2001a: 12 (Chilean species).
Distribution.
South America: Chile, Argentina, Falkland Isl.
Other regions: Nearctic, Palaearctic, Ethiopian, Oriental, Australian, Antarctic.
0448. Meoticaphaena Pace, 2015e: 72

Type species: Meotica cordobensis Bernhauer, 1934

Distribution.
0449. Phymatura J. Sahlberg, 1876: 85

Type species: Bolitochara brevicollis Kraatz, 1856
Venusa Casey, 1906: 272 [synonymized by Ashe 1992: 362]
Type species: Venusa picta Casey, 1906

Characters/Description: Fenyes 1920: 116 (characters); Ashe 1992: 366 (characters in key), 371 (characters).
Distribution.
0450. Prosoponotha Pace, 1999c: 146

Type species: Prosoponotha chilensis Pace, 1999

Distribution.
South America: Brazil, Chile.
0451. Tanystetha Pace, 2002: 137

Type species: Tanystetha brasiliensis Pace, 2002
Distribution.
Subtribe Dinardopsina Bernhauer & Scheerpeltz 1926
0452. Dinardopsis Bruch, 1917: 257 (Fig. 52)

Type species: Dinardopsis solenopsidicola Bruch, 1917

Distribution.
Subtribe Gyrophaenina Kraatz, 1856b
0453. Agaricochara Kraatz, 1856b: 361

Type species: Gyrophaena laevicollis Kraatz, 1854
Characters/Description: Seevers 1951: 740 (characters); Ashe 1984: 241 (characters in key), 255 (characters).
Distribution.
West Indies: Martinique, St. Lucia.
South America: Colombia, Ecuador, Peru.
0454. Agaricomorpha Ashe, 1984: 263

Type species: Gyrophaena apacheana Seevers, 1951

Characters/Description: Ashe 1984: 241 (characters in key); Ashe 2002b: 130 (characters in key); Eldredge 2012c:
Distribution.
0455. Ashea Y.H. Kim & Ahn, 2015: 92 (Fig. 47)

Type species: Ashea megacephala Kim & Ahn, 2015

Characters/Description: Y.H. Kim & Ahn 2015: 99 (characters in key).
Distribution.
0456. Brachida Mulsant & Rey, 1871: 4

Type species: Homalota notha Erichson, 1839
Brachida Mulsant & Rey, 1872b: 94 [synonymized by Blackwelder 1952: 84]
Brachida Mulsant & Rey, 1873c: 74 [synonymized by Blackwelder 1952: 84]

Characters/Description: Ashe 1984: 240 (characters in key), 254 (characters).
Key: Pace 2002: 126, 128, 130, 133, 134 (some species of the genus); Pace 2008a: 247 (species of Ecuador and
Peru).
Distribution.
South America: Colombia, Venezuela, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Argentina.
Other regions: Palaearctic, Ethiopian, Madagascan, Oriental, Australian, Oceanic.
0457. Brachycantharus Bierig, 1939a: 16

Type species: Brachycantharus gibber Bierig, 1939
Distribution.
0458. Brachychara Sharp, 1883: 267

Type species: Brachychara crassa Sharp, 1883

Characters/Description: Ashe 1984: 241 (characters in key), 261 (characters); Ashe 2002b: 130 (characters in key).
Distribution.
West Indies: Cuba, St. Vincent & Grenadines.
0459. Dactyloglossa Pace, 1986a: 418

Type species: Dactyloglossa inca Pace, 1986

Distribution.
0460. Eumicrota Casey, 1906: 280

Type species: Gyrophaena corruscula Erichson, 1839

Characters/Description: Seevers 1951: 732 (characters); Ashe 1984: 241 (characters in key), 249 (characters); Ashe
2002b: 130 (characters in key); Y.H. Kim & Ahn 2015: 99 (characters in key).
Key: Pace 2002: 120, 122, 124, 126 (some species of Eumicrota).
Distribution.
West Indies: St. Lucia.
South America: Venezuela, French Guiana, Brazil, Argentina.
0461. Gyrophaena Mannerheim, 1830: 74

Type species: Staphylinus nanus Paykull, 1800
Subgenera:
Agaricophaena Reitter, 1909: 85
Type species: Staphylinus boleti Linnaeus, 1758
Enkentrophaena Eichelbaum, 1913: 139
Type species: Gyrophaena plicata Fauvel, 1898
Gyrophaena Mannerheim, 1830: 74 [*]
Type species: Staphylinus nanus Paykull, 1800
Leptarthrophaena Blackwelder, 1952: 215 [*]
Type species: Gyrophaena affinis Mannerheim, 1830
Leptarthrophaena Scheerpeltz & Höfler, 1948: 170 [synonymized by Blackwelder 1952: 215]
Orphnebioidea Schubert, 1908: 611
Type species: Orphnebioidea rosti Schubert, 1908
Phaenogyra Mulsant & Rey, 1871: 76
Type species: Gyrophaena strictula Erichson, 1839
Phaenogyra Mulsant & Rey, 1872b: 166 [Newton & Thayer 2003]
Phaenogyra Mulsant & Rey, 1873c: 74 [Newton & Thayer 2003]
Razia Blackwelder, 1952: 337
Type species: Zyras abnormalis Bernhauer, 1916
Allocota Bernhauer, 1916b: 428 [synonymized by Blackwelder 1952: 337]
Type species: Zyras abnormalis Bernhauer, 1916

Key: Cameron 1922b: 638 (species of the West Indies); Seevers 1951: 671 (some subgenera); Ashe 1984: 245
(subgenera of Gyrophaena); Pace 2008a: 233 (species of Ecuador and Peru).
Distribution.
West Indies: Cuba, Jamaica, Haiti, US Virgin Isl., Guadeloupe, St. Lucia, Saint Vincent & the Grenadines,
Grenada.
South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Argentina.
0462. Microbrachida Bierig, 1939a: 20

Type species: Microbrachida gilvicornis Bierig, 1939

Distribution.
0463. Neobrachychara Bierig, 1939a: 19

Type species: Neobrachychara horni Bierig, 1939

Distribution.
0464. Phanerota Casey, 1906: 285

Type species: Aleochara fasciata Say, 1834

Subgenera:
Acanthophaena Blackwelder, 1952: 34
Type species: Gyrophaena appendiculata Motschulsky, 1858
Acanthophaena Cameron, 1934: 23 [synonymized by Blackwelder 1952: 34]
Phanerota Casey, 1906: 285 [*]
Type species: Aleochara fasciata Say, 1834

Key: Ashe 1984: 248 (subgenera of Phanerota); Klimaszewski & Peck 1998: 230 (species of the Galapagos Isl.);
Pace 2002: 118 (some Brazilian species of Phanerota); Newton & Peck 2006: 137 (species of the Galapagos
Isl.).
Distribution.
West Indies: St. Lucia, Trinidad-Tobago.
South America: Colombia, French Guiana, Ecuador, Galapagos Isl., Peru, Brazil, Paraguay, Argentina.
Other regions: Nearctic, Ethiopian, Oriental, Australian, Oceanic.
0465. Probrachida Ashe, 1984: 252

Type species: Brachida modesta Sharp, 1883
Characters/Description: Ashe 1984: 240 (characters in key); Ashe 2002b: 130 (characters in key); Eldredge 2012c:
231(characters), 232 (characters in key).
Distribution.
South America: Bolivia, Brazil, Argentina.
Subtribe Homalotina Heer, 1839
0466. Anomognathus Solier, 1849: 337

Type species: Anomognathus filiformis Solier, 1849
Theetura Thomson, 1858: 32 [synonymized by Bernhauer & Scheerpeltz 1926: 547]
Type species: Homalota cuspidata Erichson, 1839
Subgenera:
Anomognathus Solier, 1849: 337 [*]
Type species: Anomognathus filiformis Solier, 1849
Mindoria Cameron, 1941: 401
Type species: Mindoria paradoxa Cameron, 1941

Distribution.
Other regions: Nearctic, Palaearctic, Oriental, Australian, Oceanic.
0467. Cephaloxynum Bernhauer, 1907: 382
Type species: Cephaloxynum gestroi Bernhauer, 1907
Subgenera:
Cephaloxynum Bernhauer, 1907: 382 [*]
Type species: Cephaloxynum gestroi Bernhauer, 1907
Ponticulus Bierig, 1931: 424 [*]
Type species: Cephaloxynum rambouseki Bierig, 1931

Characters/Description: Blackwelder 1943: 394 (characters in key), 458 (characters); Ashe 2002b: 131 (characters
in key).
Key: Blackwelder 1943: 459 (species of the West Indies).
Distribution.
West Indies: Cuba, Jamaica.
South America: Bolivia, Argentina.
0468. Homalota Mannerheim, 1831: 487

Type species: Aleochara plana Gyllenhal, 1810
Epipeda Mulsant & Rey, 1871: 136 [synonymized by Bernhauer & Scheerpeltz 1926: 545]
Type species: Aleochara plana Gyllenhal, 1810
Epipeda Mulsant & Rey, 1872b: 226 [synonymized by Blackwelder 1952: 150]
Lampromalota Cameron, 1920a: 246 [synonymized by Bernhauer & Scheerpeltz 1926: 545]
Type species: Lampromalota brunneicollis Cameron, 1920
Mimomalota Cameron, 1920a: 242 [synonymized by Bernhauer & Scheerpeltz 1926: 545]
Type species: Mimomalota bispina Cameron, 1920
Stonalota Mannerheim, 1846: 508 [synonymized by Blackwelder 1952: 189]
Characters/Description: Lynch-Arribálzaga 1884: 55 (characters); Ashe 2002b: 131 (characters in key).
Distribution.
South America: Venezuela, French Guiana, Ecuador, Brazil, Paraguay, Argentina.
Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Oceanic.
0469. Peliusa Erichson, 1839a: 192

Type species: Peliusa labiata Erichson, 1839

Distribution.
South America: Colombia, Ecuador, Bolivia.
Other regions: Madagascan, Oriental.
0470. Thecturota Casey, 1893: 357 (Fig. 78)

Type species: Thecturota tenuissima Casey, 1893
Hemithecta Casey, 1911: 211 [synonymized by Bernhauer & Scheerpeltz 1926: 564]
Type species: Thecturota ruficollis Casey, 1911

Leptischnusa Pace, 1983d: 308 [synonymized by Pace 1990b: 58]
Type species: Leptischnusa schuberti Pace, 1983
Oligurota Casey, 1893: 361 [synonymized by Bernhauer & Scheerpeltz 1926: 564]
Type species: Oligurota pusio Casey, 1893
Pragensiella Machulka, 1941: 100 [synonymized by Muona 1984: 227, 230]
Type species: Pragensiella magnifica Machulka, 1941
Key: Klimaszewski & Peck 1998: 230 (species of the Galapagos Isl.); Newton & Peck 2006: 137 (species of the
Galapagos Isl.).
Distribution.
South America: Colombia, Galapagos Isl., Bolivia, Brazil, Chile, Argentina.
Subtribe Silusina Fenyes, 1918
0471. Silusa Erichson, 1837: 377 (Fig. 74)

Type species: Silusa rubiginosa Erichson, 1837
Subgenera:
Silusa Erichson, 1837: 377 [*]
Type species: Silusa rubiginosa Erichson, 1837
Stenusa Kraatz, 1856b: 47 [*]
Type species: Silusa rubra Erichson, 1839

Characters/Description: Coiffait & Sáiz 1967: 64 (characters in key), 68 (characters); Ashe 2002b: 129 (characters
in key).
Distribution.
Incertae sedis in Tribe Homalotini Heer, 1839
0472. Anomalophaena Pace, 1982b: 165

Type species: Leptusa weiseri Bernhauer, 1925

Distribution.
0473. Caralepta Pace, 1996b: 435

Type species: Caralepta cotopaxicola Pace, 1996
Distribution.
0474. Coenonica Kraatz, 1857d: 45

Type species: Coenonica puncticollis Kraatz, 1857
Deralia Cameron, 1920a: 238 [synonymized by Pace 2003a: 276]
Type species: Deralia fuscipennis Cameron, 1920
Ruteria Jarrige, 1957: 113 [Y.H. Kim & Ahn 2013: 58]
Type species: Ruteria rufula Jarrige, 1957
Characters/Description: Fenyes 1920: 142 (characters); Y.H. Kim & Ahn 2013: 57 (characters).
Distribution.
West Indies: Cayman Is., Jamaica, Dominican Republic, Guadeloupe, St. Vincent & Grenadines, Grenada.
0475. Cyllogerusa Pace, 1996c: 413

Type species: Homalota rubicula Erichson, 1839

Distribution.
0476. Heterota Mulsant & Rey, 1873b: 162

Type species: Homalota plumbea Waterhouse, 1858
Heterota Mulsant & Rey, 1874c: 194 [synonymized by Blackwelder 1952: 186]
1 species is included in this genus from the West Indies and Central America.
Characters/Description: Frank & Thomas 1984a: 409 (characters)
Distribution.
West Indies: Jamaica, Guadeloupe.
0477. Hoplomicra Sharp, 1883: 273

Type species: Hoplomicra clavicornis Sharp, 1883

Distribution.
0478. Pareudera Coiffait & Sáiz, 1967: 89

Type species: Eudera depressicollis Germain, 1903

Key: Coiffait & Sáiz 1967: 89 (species of Chile).
Distribution.
0479. Polytelusa Pace, 1996c: 416 (Fig. 73)

Type species: Polytelusa colombica Pace, 1996

Characters/Description: Pace 1996b: 417 (characters in key).
Distribution.
0480. Thecturella Cameron, 1922b: 648

Type species: Thecturella insidiosa Cameron, 1922
Distribution.
0481. Thinobiosus Moore & Legner, 1977: 468

Type species: Thinobiosus salinus Moore & Legner, 1977

Distribution.
0482. Thrichidryas Blackwelder, 1952: 389

Type species: Thrichidryas silvestris Bierig, 1939
Thrichidryas Bierig, 1939a: 22 [nomen nudum, validated by Blackwelder 1952: 389]

Distribution.
0483. Trichidryas Pace, 1996b: 416

Type species: Homalota discors Erichson, 1839

Distribution.
0484. Tropocoenonica Pace, 2007: 222

Type species: Tropocoenonica catharinensis Pace, 2007
Distribution.
0485. Xenobiota Bierig, 1938b: 144

Type species: Xenobiota bernhaueri Bierig, 1938
Distribution.
West Indies: Cuba.
Tribe Hoplandriini Casey, 1910
Subtribe Hoplandriina Casey, 1910a
0486. Heliconandria Hanley, 2002a: 121

Type species: Heliconandria peoechma Hanley, 2002

Key: Hanley 2002a: 127 (species of the genus); Hanley 2003a: 88 (characters in key), 108 (characters).
Distribution.
0487. Hoplandria Kraatz, 1857c: 4

Type species: Hoplandria ochracea Kraatz, 1857
Diaboligenus Bierig, 1939a: 26 [synonymized by Pace 1990a: 174]
Type species: Diaboligenus primus Bierig, 1939
Subgenera:
Arrhenandria Génier, 1989: 20
Type species: Hoplandria laeviventris Casey, 1910
Genosema Notman, 1920: 720 [*]
Type species: Genosema sexualis Notman, 1920
Hoplandria Kraatz, 1857c: 4 [*]
Type species: Hoplandria ochracea Kraatz, 1857
Lophomucter Notman, 1920: 722
Type species: Lophomucter laevicollis Notman, 1920
Platonica Sharp, 1883: 214 [*]
Type species: Platonica major Sharp, 1883

Characters/Description: Fenyes 1920: 306 (characters); Génier 1989: 8 (characters); Ashe 2002b: 125 (characters
in key); Hanley 2003a: 86 (characters), 88 (characters in key), 91 (characters subgenus Arrhenandria), 94
(characters subgenus Genosema), 97 (characters subgenus Hoplandria), 100 (characters subgenus
Lophomucter), 103 (characters subgenus Platonica); Pace2008a: 364 (characters in key).
Key: Bernhauer 1935a: 92 (Brazilian species); Génier 1989: 10 (subgenera), 10 (some species of subgenus
Hoplandria), 23 (some species of subgenus Lophomucter); Pace 1990b: 71 (some species of the genus);
Hanley 2003a: 91 (subgenera); Pace 2008a: 367 (some species of Ecuador and Peru).

Distribution.
West Indies: Cuba, Puerto Rico, Guadeloupe, Dominica, St. Vincent & Grenadines, Grenada.
Chile, Argentina.
0488. Leptandria Hanley, 2003b: 2616

Type species: Leptandria ecitophila Hanley, 2003
Leptandria Hanley, 2003a: 106 [Newton & Thayer 2003]
Leptandria Hanley, 2003c: 18 [Newton & Thayer 2003]

Characters/Description: Hanley 2003a: 88 (characters in key), 106 (characters).
Key: Hanley 2003b: 2619 (species of the genus).
Distribution.
Subtribe Platandriina Hanley, 2002c
0489. Ligulata Hanley & Ashe, 1998: 184

Type species: Ligulata hansoni Hanley & Ashe, 1998

Characters/Description: Hanley 2003a: 88 (characters in key), 119 (characters).
Distribution.
0490. Microlia Casey, 1910a: 144 (Fig. 65)

Type species: Dolosota pernix Casey, 1910
Nosora Casey, 1911: 145 [synonymized by Gusarov 2002b: 2]
Type species: Nosora azteca Casey, 1911
Tinotoma Cameron, 1923: 386 [synonymized by Hanley 2003a: 122, 125]
Type species: Tinotoma rufotestacea Cameron, 1923

Characters/Description: Fenyes 1920: 308 (characters as Nosora); Gusarov 2002b: 2 (characters); Ashe 2002b: 125
(characters in key); Hanley 2003a: 88 (characters in key), 122 (characters).
Key: Gusarov 2002b: 8 (species of the genus).
Distribution.
West Indies: Grenada.
Other regions: Nearctic, Australian, Oceanic.
0491. Platandria Casey, 1893: 345
Type species: Platandria mormonica Casey, 1893
Tetrallus Bernhauer, 1905: 252 [synonymized by Hanley 2003a: 114]
Type species: Tetrallus fenyesi Bernhauer, 1905
Characters/Description: Fenyes 1920: 309 (characters), 310 (characters as Tetrallus); Génier & Klimaszewski
1986: 202 (characters); Ashe 2002b: 125 (characters in key); Hanley 2003a: 88 (characters in key), 112
(characters); Pace 2008a: 365 (characters in key).
Key: Génier & Klimaszewski 1986: 205 (species of the genus).
Distribution.
South America: Venezuela, Ecuador, Peru, Bolivia, Argentina.
Other regions: Nearctic, Ethiopian, Oriental, Australian.
Incertae sedis in Tribe Hoplandriini Casey, 1910a
0492. Acantoxyura Pace, 1987a: 194

Type species: Acantoxyura armata Pace, 1987
Characters/Description: Hanley 2003a: 134 (characters).
Distribution.
0493. Bessoglossa Pace, 1986a: 458

Type species: Bessoglossa peruviana Pace, 1986

Characters/Description: Hanley 2003a: 135 (characters); Pace 2008a: 364 (characters in key).
Distribution.
0494. Ecuadorusa Pace, 2008a: 369

Type species: Ecuadorusa andensis Pace, 2008

Key: Pace, 2008a: 370 (species of Ecuador).
Distribution.
0495. Mesoplandria Pace, 2008a: 370

Type species: Mesoplandria loreticola Pace, 2008

Distribution.
0496. Paraplandria Pace, 2008a: 365

Type species: Paraplandria caraorum Pace, 2008
Distribution.
South America: Venezuela, French Guiana, Ecuador, Peru.
0497. Rondonusa Pace, 2015e: 125

Type species: Rondonusa braziliana Pace, 2015

Distribution.
0498. Taplandria Pace, 2015e: 129

Type species: Taplandria flava Pace, 2015
Taplandria Pace, 2015c: 78 [synonymized by Newton 2017: 7]
Type species: Taplandria guyanensis Pace, 2015

Distribution.
South America: French Guiana, Peru, Ecuador.
Tribe Hygronomini Thomson, 1859
Subtribe Hygronomina Thomson, 1859
0499. Caloderella Bernhauer, 1912d: 70 (Fig. 50)

Type species: Caloderella argentina Bernhauer, 1912

Key: Irmler 2011: 169 (Neotropical species); Pace 2008a: 229 (species of Ecuador and Peru).
Distribution.
South America: Colombia, Ecuador, Argentina.
Other regions: Ethiopian, Oriental.
0500. Cryptocompsus Lynch-Arribálzaga, 1884: 96

Type species: Cryptocompsus venustus Lynch-Arribálzaga, 1884

Distribution.
0501. Hygronoma Erichson, 1837: 312
Type species: Aleochara dimidiata Gravenhorst, 1806

Distribution.
South America: French Guiana, Brazil.
Subtribe Saphoglossina Bernhauer & Scheerpeltz, 1926
0502. Barychara Sharp, 1883: 292

Type species: Barychara filicornis Sharp, 1883
Distribution.
West Indies: Haiti.
0503. Saphoglossa Sharp, 1883: 291

Type species: Saphoglossa pictipennis Sharp, 1883

Distribution.
Incertae sedis in Tribe Hygronomini Thomson, 1859
0504. Eymekesina Pace, 2015e: 44

Type species: Eymekesina peruviana Pace, 2015

Distribution.
Tribe Hypocyphtini Laporte, 1835
0505. Callopsenius Wasmann, 1903: 236

Type species: Eupsenius clavicornis Wasmann, 1902
Eupsenius H. Franz, 1989c: 5 [synonymized by Wasmann 1903: 236]
Type species: Eupsenius clavicornis Wasmann, 1902

Characters/Description: Seevers 1941: 341 (characters); Seevers 1957: 283 (characters).
Distribution.

0506. Cypha Leach in Samouelle, 1819: 176
Type species: Tachyporus granulum Gravenhorst, 1806
Eremodromius Iablokoff-Khnzorian, 1961: 148 [synonymized by Assing 2005: 62]
Type species: Eremodromius takhtajani Iablokoff-Khnzorian, 1961
Hypocyphtus Gyllenhal, 1827: 294 [synonymized by Blackwelder 1952: 195]
Type species: Scaphidium longicorne Paykull, 1800
Hypocyphtus Stephens, 1829b: 272 [synonymized by Blackwelder 1952: 195]
Hypocyphtus Mannerheim, 1830: 58 [synonymized by Blackwelder 1952: 195]
Hypocyptus Agassiz, 1846: 191 [synonymized by Blackwelder 1952: 195]
Type species: Scaphidium longicorne Paykull, 1800

Distribution.
West Indies: St. Lucia.
South America: Ecuador, Bolivia.
0507. Holobus Solier, 1849: 335 (Fig. 60)

Type species: Holobus pigmaeus Solier, 1849
Somatium Wollaston, 1854: 563 [Newton & Thayer 2003]
Type species: Somatium anale Wollaston, 1854

Characters/Description: Coiffait & Sáiz 1967: 52 (characters in key), 56 (characters); Ashe 2002b: 131 (characters
in key); Newton & Peck 2006: 135 (characters in key as subgenus of Oligota).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, US Virgin Isl., British Virgin Isl., Antigua & Barbuda, Montserrat,
Guadeloupe, St. Lucia, Barbados, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Galapagos Isl., Peru, Brazil,
Chile, Argentina.
0508. Oligota Mannerheim, 1830: 72

Type species: Aleochara pusillima Gravenhorst, 1806
Deroligota Sharp in Sharp & Scott, 1908: 555 [synonymized by Blackwelder 1952: 121]
Type species: Oligota prolixa Sharp, 1885
Gnatholigota Sharp in Sharp & Scott, 1908: 556 [synonymized by Blackwelder 1952: 172]
Type species: Oligota latifrons Sharp, 1908
Goliota Mulsant & Rey, 1873b: 107 [synonymized by Blackwelder 1952: 174]
Type species: Oligota granaria Erichson, 1837
Goliota Mulsant & Rey, 1874d: 121 [synonymized by Blackwelder 1952: 174]
Logiota Mulsant & Rey, 1873a: 111 [synonymized by Blackwelder 1952: 226]
Logiota Mulsant & Rey, 1873d: 148 [synonymized by Blackwelder 1952: 225]
Type species: Oligota picescens Mulsant & Rey, 1873
Logiota Mulsant & Rey, 1874a: 2 [Newton & Thayer 2003]
Logiota Mulsant & Rey, 1874d: 131 [synonymized by Blackwelder 1952: 226]
Microcera Mannerheim, 1830: 72 [synonymized by Blackwelder 1952: 244]
Type species: Microcera inflata Mannerheim, 1830
Nesoligota Sharp in Sharp & Scott, 1908: 557 [synonymized by Blackwelder 1952: 261]
Type species: Oligota latipennis Sharp, 1908
Oligusa Wasmann, 1897: 267 [synonymized by Pace 2008e: 557]
Type species: Oligusa crematogastris Wasmann, 1897
Paroligota Cameron, 1945a: 159 [synonymized by Williams 1976: 248]
Type species: Paroligota zealandica Cameron, 1945

Characters/Description: Lynch-Arribálzaga 1884: 92 (characters); Coiffait & Sáiz 1967: 51 (characters in key), 52
Key: Coiffait & Sáiz 1967: 53 (species of Chile); Frank 1972: 129 (subgenera), 143 (species of the Caribbean
region).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, US Virgin Isl., British Virgin Isl., Guadeloupe, St. Vincent &
South America: Colombia, French Guiana, Ecuador, Bolivia, Brazil, Chile, Argentina.
Tribe Liparocephalini Fenyes, 1918
0509. Diaulota Casey, 1893: 354

Type species: Diaulota densissima Casey, 1893
Genoplectes Sawada, 1955: 81 [synonymized by Ahn 1996a: 271]
Type species: Genoplectes uenoi Sawada, 1955

Characters/Description: Ahn 1996a: 271 (characters); Ahn 2001: 132 (characters in key); Ashe 2002b: 129
Key: Ahn, 1996a: 274 (species of the genus).
Distribution.
Tribe Lomechusini Fleming, 1821
Subtribe Lomechusina Fleming, 1821
0510. Xenodusa Wasmann, 1894: 205

Type species: Lomechusa cava LeConte, 1863
Pseudolomechusa Mann, 1914: 174 [synonymized by Hoebeke 1976]
Type species: Xenodusa sharpi Wasmann, 1897

Distribution.
Other regions: Nearctic, Oceanic.
Subtribe Myrmedoniina Thomson, 1867a
0511. Amazoncharis Pace, 1990b: 70

Type species: Amazoncharis aspera Pace, 1990

Distribution.
South America: Peru, Brazil.
0512. Asheidium Santiago-Jiménez, 2010: 671

Type species: Asheidium zurquiensis Santiago-Jiménez, 2010

Characters/Description: Santiago-Jiménez 2010: 647 (characters in key); Santiago-Jiménez 2016: 275 (characters
in key).
Distribution.
0513. Delgadoidium Santiago-Jiménez, 2010: 659

Type species: Delgadoidium ecuatorensis Santiago-Jiménez, 2010

in key).
Distribution.
0514. Dromacamatus Bruch, 1933a: 210

Type species: Dromacamatus caviceps Bruch, 1933
Distribution.
0515. Dromeciton Fauvel, 1904b: 282

Type species: Dromeciton wagneri Fauvel, 1904
Characters/Description: Fenyes 1920: 269 (characters); Seevers 1965: 235 (characters in key), 251 (characters).
Distribution.
0516. Drusilla Leach in Samouelle, 1819: 177

Type species: Staphylinus canaliculatus Fabricius, 1787
Anaulax Bernhauer, 1929d: 231 [synonymized by Blackwelder 1952: 51]
Type species: Astilbus semicircularis Bernhauer, 1929
Evansius Bernhauer, 1933c: 297 [synonymized by Blackwelder 1952: 137]
Type species: Astilbus denticollis Bernhauer, 1933
Thoracophagus Kistner, 2003: 561 [synonymized by Hlaváč et al. 2011: 9]
Type species: Thoracophagus congoensis Kistner, 2003
Subgenera:
Apteranopsis Jeannel, 1960b: 208
Type species: Apteranillus lethierryi Fauvel, 1898
Drusilla Leach in Samouelle, 1819: 177
Agaricola Gistel, 1834: 10 [synonymized by Blackwelder 1952: 41]
Astilbus Dillwyn, 1829: 63 [synonymized by Blackwelder 1952: 66]
Camacopalpus Motschulsky, 1858b: 231 [synonymized by Klimaszewski 1982: 421]
Type species: Camacopalpus flavicornis Motschulsky, 1858
Camacopselaphus Gemminger & Harold, 1868: 549 [synonymized by Blackwelder 1952: 92]
Type species: Camacopalpus flavicornis Motschulsky, 1858
Myrmedonia Erichson, 1837: 286 [synonymized by Blackwelder 1952: 254]
Santhota Sharp, 1874: 3 [synonymized by Maruyama 2000: 351]
Type species: Santhota sparsa Sharp, 1874
Tropignorimus Bernhauer, 1915a: 154
Type species: Astilbus carinithorax Bernhauer, 1915
only a single species are included in this genus from South America.
Characters/Description: Fenyes 1920: 136 (characters as Camacopselaphus), 159 (characters as Santhota), 261
(characters as Astilbus), 297 (characters as Myrmedonia).
Distribution.
0517. Ecitana Seevers, 1965: 242

Type species: Ecitana biimpressa Seevers, 1965

Distribution.
0518. Ecitocerus Borgmeier, 1949: 136

Type species: Ecitocerus gracilipes Borgmeier, 1949

Distribution.
0519. Ecitocryptodes Seevers, 1965: 265

Type species: Ecitocryptodes petiolatus Seevers, 1965

Characters/Description: Seevers, 1965: 233 (characters in key).
Distribution.
0520. Ecitocryptus Borgmeier, 1930: 173

Type species: Ecitocryptus sulcatus Borgmeier, 1930
Distribution.
0521. Ecitodiscus Borgmeier, 1949: 134

Type species: Ecitodiscus pedissequus Borgmeier, 1949

Distribution.
0522. Ecitoglossa Borgmeier, 1958: 234

Type species: Ecitoglossa foveicollis Borgmeier, 1958

Characters/Description: Seevers, 1965: 235 (characters in key), 259 (characters); Mooney & Kistner, 2005: 252
(characters).
Key: Seevers 1965: 260 (species of the genus); Mooney & Kistner 2005: 262 (species of the genus).
Distribution.
0523. Ecitonia Wasmann, 1894: 209

Type species: Myrmedonia badariottii Wasmann, 1890

Distribution.
0524. Ecitonilla Wasmann, 1894: 210
Type species: Ecitonilla claviventris Wasmann, 1894

Distribution.
0525. Ecitopelta Borgmeier, 1949: 138

Type species: Ecitopelta reticulata Borgmeier, 1949

Distribution.
0526. Ecitophila Wasmann, 1890: 314

Type species: Ecitophila omnivora Wasmann, 1890

Characters/Description: Fenyes 1920: 273 (characters); Seevers 1965: 245 (characters).
Distribution.
South America: Bolivia, Brazil, Paraguay.
0527. Ecitophiletus Borgmeier, 1932: 399

Type species: Ecitophiletus constrictus Borgmeier, 1932

Distribution.
0528. Ecitophrura Reichensperger, 1939b: 290

Type species: Ecitophrura capito Reichensperger, 1939
Distribution.
0529. Ecitoplectus Borgmeier, 1931: 364

Type species: Ecitoplectus transiens Borgmeier, 1931

Distribution.

0530. Ecitopolites Borgmeier, 1949: 140
Type species: Ecitopolites scopifer Borgmeier, 1949

Distribution.
0531. Ecitotyphlus Borgmeier, 1949: 144

Type species: Ecitotyphlus apterus Borgmeier, 1949

Distribution.
0532. Ecitoxenidia Wasmann, 1909a: 179

Type species: Ecitoxenia brevipes Brues, 1902
Ecitocolax Borgmeier, 1949: 145 [synonymized by Seevers 1965: 246]
Type species: Ecitocolax longicornis Borgmeier, 1949
Ecitoxenidia Wasmann, 1902c: 91 [nomen nudum, noted by Blackwelder 1952: 143]
Kistner et al. 1996: 62 (characters); Ashe 2002b: 126 (characters in key).
Key: Seevers 1959: 76 (species of the genus); Kistner et al. 1996: 66 (species of the genus).
Distribution.
0533. Eurydiotyphla Pace, 1986a: 447

Type species: Eurydiotyphla franzi Pace, 1986

Distribution.
0534. Falagonia Sharp, 1883: 212

Type species: Falagonia mexicana Sharp, 1883

Characters/Description: Fenyes 1920: 275 (characters); Ashe 2002b: 127 (characters in key); Santiago-Jiménez
2010: 646 (characters in key), 647 (characters); Santiago-Jiménez 2016: 275 (characters in key).
Distribution.
0535. Falagonilla Reichensperger, 1939b: 295
Type species: Falagonilla cursor Reichensperger, 1939

Distribution.
0536. Gallardoia Bruch, 1924: 257

Type species: Gallardoia argentina Bruch, 1924

Key: Seevers 1965: 234 (characters in key), 265 (species of the genus).
Distribution.
0537. Labidilla Borgmeier, 1949: 132

Type species: Labidilla dentiguttur Borgmeier, 1949

Distribution.
0538. Labidoculex Reichensperger, 1936a: 234

Type species: Labidoculex fragilis Reichensperger, 1936
Characters/Description: Borgmeier 1958: 230 (characters); Seevers 1965: 235 (characters in key), 252 (characters);
Ashe 2002b: 127 (characters in key); Santiago-Jiménez 2010: 646 (characters in key); Santiago-Jiménez
Distribution.
0539. Macrogerodonia Bernhauer, 1941: 292 (Fig. 64)

Type species: Zyras magnicollis Bernhauer, 1941
Macrogerodonia Bernhauer, 1951: 281 [Newton & Thayer 2003]
Zyrastilbodes Scheerpeltz, 1962b: 200 [synonymized by Pace 1986a: 451]
Type species: Myrmedonia scabripennis Sharp, 1876

Characters/Description: Bernhauer, 1951: 281 (characters).
Key: Scheerpeltz 1962b: 204 (species of Zyrastilbodes); Pace 1986a: 456 (some species of Peru); Pace 2001b: 429
(some species of the genus).
Distribution.
West Indies: Guadeloupe, St. Lucia.
Central America: Guatemala, Honduras, El Salvador, Costa Rica, Panama.

South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay.
0540. Manausdota Pace, 2008b: 193
Type species: Manausdota amazonica Pace, 2008

Key: Pace 2008b: 193 (species of the genus).
Distribution.
0541. Microdonia Casey, 1893: 318

Type species: Microdonia occipitalis Casey, 1893

Kistner et al. 1996: 47 (characters); Ashe 2002b: 126 (characters in key).
Key: Seevers 1959: 67 (species of the genus); Kistner et al. 1996: 52 (species of the genus).
Distribution.
0542. Myrmecoxenia Lynch-Arribálzaga, 1884: 35

Type species: Myrmecoxenia pampeana Lynch-Arribálzaga, 1884

Distribution.
South America: Brazil, Paraguay, Uruguay, Argentina.
0543. Myrmedonota Cameron, 1920a: 272

Type species: Myrmedonota cingulata Cameron, 1920
Characters/Description: Bourguignon & Roisin 2006: 2 (characters); Eldredge 2010: 18 (characters); Mathis &
Eldredge 2014: 96 (characters); Santiago-Jiménez 2014: 50 (characters); Navarro & Caron 2018: 148
(characters).
Key: Santiago-Jiménez 2014: 51 (species of the Nearctic and Neotropics); Navarro & Caron 2018: 148 (species of
the genus).
Distribution.
Other regions: Nearctic, Oriental, Oceanic.
0544. Myrmoecia Mulsant & Rey, 1873: 98

Type species: Myrmedonia tuberiventris Fairmaire, 1855
Myrmoecia Mulsant & Rey, 1874: 130 [Hlaváč et al. 2011: 56]
Nototaphra Casey, 1893: 327 [synonymized by Wasmann 1901b: 252]
Type species: Nototaphra lauta Casey, 1893
Characters/Description: Fenyes 1920: 281 (characters as Nototaphra), 298 (characters).
Distribution.
0545. Newtonidium Santiago-Jiménez, 2010: 707

Type species: Newtonidium magdalenensis Santiago-Jiménez, 2010
in key).
Key: Santiago-Jiménez 2010: 711 (species of the genus).
Distribution.
0546. Orphnebius Motschulsky, 1858b: 263 (Fig. 68)

Type species: Orphnebius ventricosus Motschulsky, 1858
Gastroacanthodonia Scheerpeltz, 1972a: 92 [synonymized by Pace 2008a: 335]
Type species: Acanthodonia nevermanni Scheerpeltz, 1972
Subgenera:
Aulacothoracobius Bernhauer, 1929e: 147
Type species: Orphnebius wasmannianus Bernhauer, 1929
Deroleptus Bernhauer, 1915a: 150
Type species: Astilbus bigladiosus Bernhauer, 1915
Colpoleptus Bernhauer, 1929e: 143
Type species: Astilbus superbus Bernhauer, 1916
Megalocephalobius Bernhauer, 1929e: 146 [synonymized by Assing 2016b: 63)
Type species: Orphnebius falagrioides Bernhauer, 1929
Orphnebiota Cameron, 1920c: 97
Type species: Orphnebiota rufocastanea Cameron, 1920
Mesocephalobius Bernhauer, 1929e: 146
Type species: Orphnebius bakeri Bernhauer, 1929
Microcephalobius Bernhauer, 1929e: 147
Type species: Orphnebius miricornis Bernhauer, 1929
Microrphnebius Pace, 1985: 386 [*]
Type species: Orphnebius ruficollis Pace, 1985
Orphnebius Motschulsky, 1858b: 263 [*]
Type species: Orphnebius ventricosus Motschulsky, 1858
Hygroptera Motschulsky, 1860a: 86
Type species: Hygroptera termitis Motschulsky, 1860
Stenaspidobius Bernhauer, 1929d: 228
Type species: Orphnebius burgeoni Bernhauer, 1929
Strabocephalium Bernhauer, 1911b: 91
Type species: Strabocephalium mirabile Bernhauer, 1911
Thoracobius Bernhauer, 1929e: 147
Type species: Orphnebius brevicollis Bernhauer, 1929

Key: Bernhauer 1929e: 146 (subgenera and some species); Pace 2008a: 335 (species of Ecuador and Peru).
Distribution.
South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Brazil.
Other regions: Ethiopian, Oriental, Oceanic.
0547. Pella Stephens, 1833b: 95

Type species: Staphylinus limbatus Paykull, 1789
Lepla Tottenham, 1939a: 226 [synonymized by Maruyama 2006: 28]
Type species: Aleochara lugens Gravenhorst, 1802
Myrmelia Mulsant & Rey, 1873d: 152 [synonymized by Maruyama 2006: 27]
Type species: Myrmedonia excepta Mulsant & Rey, 1873
Myrmelia Mulsant & Rey, 1873b: 54 [synonymized by Bernhauer & Scheerpeltz 1926: 694]
Myrmelia Mulsant & Rey, 1874a: 6 [Newton & Thayer 2003]
Myrmelia Mulsant & Rey, 1874c: 86 [Newton & Thayer 2003]
Pellochromonia Reitter, 1909: 43 [synonymized by Maruyama 2006: 28]
Type species: Myrmedonia ruficollis Grimm, 1845
Characters/Description: Fenyes 1920: 296 (characters), 298 (characters as Pellochromonia), 299 (characters as
Myrmelia); Klimaszewski et al. 2005: 716 (characters).
Distribution.
South America: Brazil, Ecuador.
0548. Pseudodinusa Bernhauer, 1912d: 78

Type species: Pseudodinusa richteri Bernhauer, 1912

Distribution.
0549. Pseudofalagonia Santiago-Jiménez, 2010: 678

Type species: Falagonia crassiventris Sharp, 1883
in key).
Distribution.
Central America: Mexico, Guatemala, Belize, El Salvador, Costa Rica, Panama.
0550. Pseudothamiaraea Cameron, 1923: 363
Type species: Pseudothamiaraea brunnea Cameron, 1923
Distribution.
West Indies: Cuba, Haiti.
0551. Scotodonia Wasmann, 1894: 210

Type species: Myrmedonia diabolica Wasmann, 1890

Kistner & Berghoff 2008: 619–620 (characters).
Distribution.
0552. Sharpidium Santiago-Jiménez, 2010: 691

Type species: Sharpidium surinamensis Santiago-Jiménez, 2010

in key).
Distribution.
South America: Suriname, French Guiana, Peru.
0553. Termitognathus Borgmeier, 1959b: 304

Type species: Termitognathus caviceps Borgmeier, 1959

Distribution.
0554. Tetradonella Jacobson & Kistner, 1998: 254

Type species: Tetradonella latiarea Jacobson & Kistner, 1998
Distribution.
0555. Tetralophodes Bernhauer, 1922a: 18

Type species: Tetralophodes bruchi Bernhauer, 1922

Distribution.

0556. Thayeridium Santiago-Jiménez, 2010: 666
Type species: Thayeridium darienensis Santiago-Jiménez, 2010

in key).
Distribution.
0557. Typhlonusa Borgmeier, 1958: 231

Type species: Typhlonusa opacicornis Borgmeier, 1958

Distribution.
0558. Wasmannina Mann, 1925: 75

Type species: Wasmannina trapezicollis Mann, 1925
Ecitochlamys Borgmeier, 1933a: 473 [synonymized by Seevers 1965: 183, 268]
Type species: Ecitochlamys comes Borgmeier, 1933
Distribution.
0559. Zyras Stephens, 1833b: 91

Type species: Aleochara haworthi Stephens, 1832
Zyras Stephens, 1835: 430 [Newton & Thayer 2003]
Subgenera:
Acanthocnemidonia Bernhauer, 1936a: 265
Type species: Zyras miricauda Bernhauer, 1936
Acrothoraconia Bernhauer, 1934b: 216
Type species: Zyras mombassana Bernhauer, 1934
Androdonia Bernhauer, 1928a: 69
Type species: Myrmedonia laminata Roth, 1851
Anophthalmodonia Bernhauer, 1936a: 266
Type species: Zyras jordani Bernhauer, 1936
Antronia Bernhauer, 1928a: 54
Type species: Zyras orbicollis Bernhauer, 1928
Thoracodonia Bernhauer, 1928a: 21 [synonymized by Last 1977b: 79]
Type species: Myrmedonia aculeata Eppelsheim, 1885
Aplastonia Bernhauer, 1932: 170
Type species: Zyras rugosissimus Bernhauer, 1932
Apostenonia Bernhauer, 1929b: 201
Type species: Zyras quadrituberculatus Bernhauer, 1929
Apterygodonia Scheerpeltz, 1963a: 100
Type species: Zyras apterus Scheerpeltz, 1963
Aulacodonia Bernhauer, 1928a: 53
Type species: Zyras glaberrimus Bernhauer, 1928
Botsa Blackwelder, 1952: 84
Type species: Zyras tuberculatus Bernhauer, 1932
Watsa Bernhauer, 1932: 171 [synonymized by Blackwelder 1952: 84]
Type species: Zyras tuberculatus Bernhauer, 1932
Callodonia Bernhauer, 1928a: 55
Type species: Zyras politus Bernhauer, 1928
Cameronodonia Dvořák, 1981: 58
Type species: Zyras carinipennis Cameron, 1950
Thoracodonia Cameron, 1950a: 127 [synonymized by Dvořák 1981: 58]
Type species: Zyras carinipennis Cameron, 1950
Camonia Bernhauer, 1928a: 27
Type species: Myrmedonia speciosa Erichson, 1839
Cephalodonia Bernhauer, 1928a: 25
Type species: Zyras bicoloriceps Bernhauer, 1928
Colpodonia Bernhauer, 1929b: 196
Type species: Zyras densithorax Bernhauer, 1929
Craspa Blackwelder, 1952: 110
Type species: Zyras antilope Bernhauer, 1908
Craspedonia Bernhauer, 1928a: 20 [synonymized by Blackwelder 1952: 110]
Type species: Zyras antilope Bernhauer, 1908
Crateodonia Bernhauer, 1928a: 68
Type species: Zyras schoutedeni Bernhauer, 1928
Ctenodonia Wasmann, 1894: 208 [*]
Type species: Ctenodonia inclyta Wasmann, 1894
Dentothalmonia Last, 1956: 209
Type species: Zyras probatus Last, 1956
Diaulaconia Bernhauer, 1928a: 73 [*]
Type species: Zyras biseriatus Bernhauer, 1915
Euryalonia Bernhauer, 1928a: 35
Type species: Zyras capensis Bernhauer & Scheerpeltz, 1926
Eurydonia Bernhauer, 1928a: 20
Type species: Zyras usambarae Bernhauer, 1915
Euryncephalodonia Scheerpeltz, 1974: 346
Type species: Zyras flavomaculatus Scheerpeltz, 1974
Euryndonia Bernhauer, 1928a: 68
Type species: Zyras peringueyi Bernhauer & Scheerpeltz, 1926
Fealina Bernhauer, 1929b: 200
Type species: Zyras insularis Bernhauer, 1929
Glossacantha Gemminger & Harold, 1868: 519 [*]
Type species: Acanthoglossa badia Motschulsky, 1860
Acanthoglossa Motschulsky, 1860a: 88 [synonymized by Gemminger & Harold 1868: 519]
Type species: Acanthoglossa badia Motschulsky, 1860
Grammodonia Bernhauer, 1928a: 55

Type species: Myrmedonia frontalis Erichson, 1839
Isothoracodonia Bernhauer, 1936c: 328
Type species: Zyras crassicornis Bernhauer, 1936
Lastia Dvořák, 1984: 199
Type species: Zyras bundukiensis Last, 1960
Solus Last, 1960b: 52 [synonymized by Dvořák 1984: 199]
Type species: Zyras bundukiensis Last, 1960
Leptodonia Bernhauer, 1928a: 26
Type species: Zyras marshalli Bernhauer, 1928
Macrodonia Wasmann, 1894: 207
Type species: Macrodonia vandepolli Wasmann, 1894
Neotropopella Scheerpeltz, 1976a: 140 [*]
Type species: Zyras brasiliensis Scheerpeltz, 1976
Pachydonia Bernhauer, 1928a: 34
Type species: Zyras dubius Bernhauer, 1928
Paragrammodonia Bernhauer, 1935c: 105
Type species: Zyras overlaeti Bernhauer, 1935
Parophthalmonia Bernhauer, 1928a: 60
Type species: Zyras kristenseni Bernhauer, 1915
Platydonia Bernhauer, 1928a: 21
Type species: Zyras notandus Bernhauer, 1915
Polydonia Bernhauer, 1928a: 21
Type species: Zyras linki Bernhauer, 1915
Pycnodonia Bernhauer, 1928a: 21
Type species: Zyras lioglutoides Bernhauer, 1915
Remionea Blackwelder, 1952: 338
Type species: Zyras escherichi Bernhauer, 1915
Eremonia Bernhauer, 1928a: 19 [synonymized by Blackwelder 1952: 338]
Type species: Zyras escherichi Bernhauer, 1915
Rhopalodonia Cameron, 1939b: 546
Type species: Myrmedonia clavicornis Kraatz, 1859
Rocnema Blackwelder, 1952: 342
Type species: Myrmedonia bangae Cameron, 1926
Blepharonia Bernhauer, 1928a: 24 [synonymized by Blackwelder 1952: 342]
Type species: Myrmedonia bangae Cameron, 1926
Sinozyras Pace, 1999b: 684
Type species: Zyras pygmaeus Pace, 1999
Subversoris Last, 1977a: 943
Type species: Zyras chadensis Last, 1977
Synthoracodonia Scheerpeltz, 1974: 370
Type species: Zyras meridioafricanus Scheerpeltz, 1974
Taprodonia Cameron, 1939b: 516
Type species: Myrmedonia nietneri Kraatz, 1857
Termidonia Motschulsky, 1860a: 87
Type species: Termidonia laminata Motschulsky, 1860
Rhynchodonia Wasmann, 1896b: 620 [Smetana 2004: 461]
Type species: Myrmedonia feae Wasmann, 1896
Termitelia Cameron, 1939b: 517
Type species: Zyras insignis Cameron, 1939
Termitodonia Cameron, 1936: 184
Type species: Zyras flavus Cameron, 1936
Trigonodonia Bernhauer, 1928a: 22
Type species: Zyras panganianus Bernhauer, 1915
Trigonozyras Cameron, 1943: 143
Type species: Zyras sarawakensis Cameron, 1943
Tropidonia Bernhauer, 1928a: 52
Type species: Zyras tubericollis Bernhauer, 1928
Visendor Last, 1960a: 91
Type species: Zyras marukana Cameron, 1933
Zyras Stephens, 1833b: 91 [*]
Type species: Aleochara haworthi Stephens, 1832

Characters/Description: Fenyes 1920: 293 (characters); Seevers 1957: 233 (characters); Ashe 2002b: 127
(characters in key); Klimaszewski et al. 2005: 714 (characters); Hlaváč & Jászay 2009: 52 (characters).
Key: Fenyes 1920: 294 (subgenera); Klimaszewski et al. 2005: 707 (species of the America north of Mexico).
Distribution.
West Indies: Cuba, US Virgin Isl., Saint Vincent & the Grenadines, Grenada.
South America: Colombia, Venezuela, Suriname, Bolivia, Brazil, Paraguay, Argentina.
Subtribe Termitozyrina Seevers, 1957
0560. Iheringocantharus Bernhauer, 1912c: 47

Type species: Iheringocantharus ypiranganus Bernhauer, 1912

Distribution.
0561. Termitonusa Borgmeier, 1950: 660

Type species: Termitonusa sequax Borgmeier, 1950

Distribution.
0562. Termitophagus Silvestri, 1946b: 530

Type species: Termitophagus synterminus Silvestri, 1946

Distribution.

0563. Termitosymbia Seevers, 1957: 245
Type species: Termitosymbia nitida Seevers, 1957

Distribution.
0564. Termitozyras Seevers, 1957: 242

Type species: Termitozyras adamsoni Seevers, 1957
Distribution.
South America: Guyana, Bolivia.
Incertae sedis in Tribe Lomechusini Fleming, 1821
0565. Apalosellia Pace, 2015e: 109

Type species: Apalosellia cotopaxiensis Pace, 2015

Distribution.
Central America: Ecuador.
0566. Aychenonida Pace, 2015a: 90

Type species: Aychenonida guyanensis Pace, 2015

Distribution.
0567. Euryepiskiosa Pace, 2015a: 91 (Fig. 56)

Type species: Euryepiskiosa crassa Pace, 2015

Distribution.
0568. Macrozoque Santiago-Jiménez, 2016: 273

Type species: Macrozoque hoplandrioides Santiago-Jiménez, 2016

Characters/Description: Santiago-Jiménez 2016: 275 (characters in key).
Distribution.
0569. Schizorphnebius Pace, 2015e: 108
Type species: Schizorphnebius osellai Pace, 2015
Distribution.
Tribe Mesoporini Cameron, 1959
0570. Ambracyptus Lundgren, 1984: 163

Type species: Paracyptus minutissima Seevers, 1971
Paracyptus Seevers, 1971: 82 [synonymized by Lundgren 1984: 163]
Type species: Paracyptus minutissima Seevers, 1971
1 species known only from Mexican amber.

Distribution.
0571. Anacyptus Horn, 1877: 87

Type species: Hypocyptus testaceus LeConte, 1863
Microcyptus Horn, 1882: 1 [synonymized by Blackwelder 1952: 244]
Type species: Hypocyptus testaceus LeConte, 1863

Characters/Description: Blackwelder 1943: 532 (characters); Ashe 2002b: 124 (characters in key).
Distribution.
West Indies: Cuba, Montserrat.
Other regions: Nearctic, Oriental.
Tribe Mimecitini Wasmann, 1917
Subtribe Labidopullina Jacobson & Kistner, 1991
0572. Labidopullus Borgmeier, 1958: 240

Type species: Labidopullus appendiculatus Borgmeier, 1958
(characters).
Distribution.
South America: Colombia, Ecuador.

Subtribe Leptanillophilina Fenyes, 1918
0573. Acamatusinella Bruch, 1931: 16

Type species: Acamatusinella globuliventris Bruch, 1931

Characters/Description: Borgmeier 1949: 126 (characters in key); Jacobson & Kistner 1991: 31 (characters).
Distribution.
0574. Ecitomerus Borgmeier, 1933b: 371

Type species: Ecitomerus impressifrons Borgmeier, 1933

Characters/Description: Borgmeier 1949: 125 (characters in key); Seevers 1965: 209 (characters in key), 224
(characters); Jacobson & Kistner 1991: 42 (characters)
Distribution.
0575. Ecitophanes Borgmeier, 1930: 165

Type species: Ecitophanes crassicornis Borgmeier, 1930

Distribution.
0576. Ecitosoma Borgmeier, 1939: 458

Type species: Ecitosoma lamellatum Borgmeier, 1939

Distribution.
0577. Leptanillophilus Holmgren, 1908: 340

Type species: Leptanillophilus similis Holmgren, 1908
Acamatusina Bruch, 1930b: 18 [synonymized by Bruch 1933c: 18]
Type species: Acamatusina inopinata Bruch, 1930

(characters); Jacobson & Kistner 1991: 17 (characters).
Distribution.
South America: Peru, Bolivia, Argentina.
0578. Mimacamatus Bruch, 1933c: 16
Type species: Mimacamatus mirabilis Bruch, 1933
Ecitomimus Borgmeier, 1949: 124 [synonymized by Blackwelder 1952: 141]
Type species: Mimacamatus fraterculus Bruch, 1933
Mimacamatus Borgmeier, 1933b: 374 [nomen nudum, synonymized by Blackwelder 1952: 247]
Type species: Mimacamatus mirabilis Borgmeier, 1933
Synacamatus Bruch, 1933b: 351 [nomen nudum, synonymized by Seevers 1965: 223]
Type species: Mimacamatus fraterculus Bruch, 1933
Characters/Description: Borgmeier 1949: 126 (characters in key as Mimacamatus and Ecitomimus); Seevers 1965:
208 (characters in key), 223 (characters); Jacobson & Kistner 1991: 25 (characters)
Distribution.
Subtribe Mimecitina Wasmann, 1917
0579. Labidoglobus Reichensperger, 1933: 179

Type species: Labidoglobus nevermanni Reichensperger, 1933
(characters)
Distribution.
0580. Labidosphaerula Reichensperger, 1939b: 282

Type species: Labidosphaerula schmidti Reichensperger, 1939

(characters).
Distribution.
0581. Mimeciton Wasmann, 1893: 97

Type species: Mimeciton pulex Wasmann, 1893
(characters).
Distribution.

South America: Ecuador, Brazil.
0582. Paramimeciton Reichensperger, 1935: 210
Type species: Mimeciton coeci Reichensperger, 1935
(characters).
Distribution.
0583. Pseudomimeciton Heikertinger, 1926: 614

Type species: Mimeciton zikani Wasmann, 1923
Metamimeciton Reichensperger, 1936a: 232 [synonymized by Blackwelder 1952: 240, 248]
Type species: Mimeciton antennatus Mann, 1926
Characters/Description: Seevers 1965: 208 (characters in key as subgenus of Mimeciton); Jacobson & Kistner
1991: 88 (characters); Ashe 2002b: 123 (characters in key).
Distribution.
Subtribe Mimonillina Bernhauer & Scheerpeltz, 1926
0584. Labidomimus Wasmann, 1923: lxiii

Type species: Labidomimus petiolatus Wasmann, 1923
Labidomimus Wasmann, 1925b: 126 [synonymized by Blackwelder 1952: 207]
Characters/Description: Jacobson & Kistner 1991: 113 (characters); Seevers 1965: 231 (characters).
Distribution.
0585. Mimonilla Wasmann, 1913: 380

Type species: Mimonilla ecitonis Wasmann, 1913
(characters).
Distribution.
Tribe Myllaenini Ganglbauer, 1895
0586. Amazonopora Pace, 1996d: 38 (Fig. 45)

Type species: Amazonopora manausensis Pace, 1996

Characters/Description: Pace 1996d: 38 (characters in key); Ahn & Ashe 1999: 132 (characters).
Key: Ahn & Ashe 1999: 131 (species of the genus); Pace 2014a: 62 (species of the genus).
Distribution.
South America: French Guiana, Peru, Brazil.
0587. Bryothinusa Casey, 1904: 312

Type species: Bryothinusa catalinae Casey, 1904
Anopsisus Bernhauer, 1929c: 187 [synonymized by Haghebaert 1990: 33]
Type species: Phytosus microphthalmus Bernhauer, 1929
Halesthenus Sawada, 1955: 83 [synonymized by Sawada 1971a: 81]
Type species: Halesthenus minutus Sawada, 1955

Characters/Description: Fenyes 1920: 131 (characters); Pace 1996d: 38 (characters in key); Ahn 1998: 333
Distribution.
0588. Myllaena Erichson, 1837: 382

Type species: Aleochara dubia Gravenhorst, 1806
Centroglossa Matthews, 1838: 194 [synonymized by Blackwelder 1952: 96]
Type species: Centroglossa conuroides Matthews, 1838

Characters/Description: Lynch-Arribálzaga 1884: 98 (characters); Fenyes 1920: 136 (characters); Coiffait & Sáiz
1967: 59 (characters); Ahn & Ashe 1999: 132 (characters); Ashe 2002b: 129 (characters in key); Newton &
Peck 2006: 135 (characters in key); Caron & Klimaszewski 2008: 356 (characters).
Key: Coiffait & Sáiz 1967: 61 (species of Chile); Pace 2008a: 230 (species of Ecuador and Peru).
Distribution.
West Indies: Cayman Is., Jamaica, Haiti, Guadeloupe, St. Lucia, St. Vincent & Grenadines, Grenada.
South America: Colombia, Venezuela, French Guiana, Ecuador, Galapagos Isl., Peru, Brazil, Chile, Argentina.
0589. Rothium Moore & Legner, 1977: 460

Type species: Rothium sonorensis Moore & Legner, 1977
Characters/Description: Ahn & Ashe 1996: 235 (characters); Klimaszewski & Peck 1998: 227 (characters); Ashe
2002b: 128 (characters in key); Newton & Peck 2006: 135 (characters in key).

Key: Ahn & Ashe 1996: 239 (species of the genus); Newton & Peck 2006: 136 (species of the Galapagos Isl.).
Distribution.
South America: Ecuador, Galapagos Isl., Peru.
Tribe Oxypodini Thomson, 1859
Subtribe Dinardina Mulsant & Rey, 1873a
0590. Blepharhymenus Solier, 1849: 339

Type species: Blepharhymenus sulcicollis Solier, 1849
Subgenera:
Blepharhymenus Solier, 1849: 339
Blepharrhymenus Gemminger & Harold, 1868: 505 [Blackwelder 1952: 78]
Colusa Casey, 1885a: 288 [synonymized by Bernhauer & Scheerpeltz 1926: 732]
Type species: Colusa gracilis Casey, 1885
Echidnoglossa Wollaston, 1864: 530 [synonymized by Bernhauer & Scheerpeltz 1926: 732]
Type species: Echidnoglossa constricta Wollaston, 1864
Blepharrhymorphus Ihssen, 1934: 215
Type species: Blepharhymenus mirandus Fauvel, 1899
Syntomenus Blackwelder, 1952: 367
Type species: Echidnoglossa ventricosus Quedenfeldt, 1881
Syntomenus Bernhauer, 1939c: 601 [nomen nudum, validated by Blackwelder 1952: 367]

Key: Pace 1987b: 491 (some species of Chile).
Distribution.
Other regions: Nearctic, Palaearctic, Ethiopian, Australian.
0591. Euthorax Solier, 1849: 345

Type species: Euthorax ruficornis Solier, 1849
Myrmecochara Kraatz, 1857c: 40 [synonymized by Blackwelder 1952: 254]
Type species: Myrmecochara pictipennis Kraatz, 1857
Campoporus Lynch-Arribálzaga, 1884: 64 [synonymized by Blackwelder 1952: 93]
Type species: Campoporus elegantulus Lynch-Arribálzaga, 1884
Eurynotida Casey, 1906: 343 [synonymized by Blackwelder 1952: 160]
Type species: Eurynotida ornata Casey, 1906
Dinusina Bernhauer, 1909b: 249 [synonymized by Blackwelder 1952: 128]
Type species: Dinusina gestroi Bernhauer, 1909
Characters/Description: Fenyes 1920: 360 (characters), 387 (characters as Dinusina); Ashe 2002b: 125 (characters
in key).
Distribution.
0592. Thiasophila Kraatz, 1856b: 69

Type species: Aleochara angulata Erichson, 1837
Myrmecodelus Motschulsky, 1858b: 239 [synonymized by Blackwelder 1952: 254]
Thyasophila Fairmaire & Laboulbène, 1856: 461 [synonymized by Blackwelder 1952: 389]
Thiasophilomorpha Horion, 1967: 359 [synonymized by Zerche 1987: 92]
Type species: Thiasophilomorpha granulifera Horion, 1967
Subgenera:
Apatusa Pace, 1997b: 41 [*]
Type species: Thiasophila pulchricornis Pace, 1997
Thiasophila Kraatz, 1856b: 69

Key: Pace 1997b: 41 (some subgenera of the genus).
Distribution.
Subtribe Meoticina Seevers, 1978
0593. Alisalia Casey, 1911: 219

Type species: Alisalia brevipennis Casey, 1911

Distribution.
West Indies: Haiti.
0594. Apimela Mulsant & Rey, 1873b: 4

Type species: Homalota macella Erichson, 1839
Apimela Mulsant & Rey, 1874c: 36 [synonymized by Blackwelder 1952: 58]
Type species: Homalota macella Erichson, 1839
Apimela Mulsant & Rey, 1875a: 74 [synonymized by Blackwelder 1952: 58]
Apimela Mulsant & Rey, 1875c: 48 [synonymized by Blackwelder 1952: 58]
Gyronychina Casey, 1911: 218 [synonymized by Blackwelder 1952: 177]
Type species: Calodera attenuata Casey, 1885

Gampsonycha Bernhauer, 1912b: 109 [synonymized by Blackwelder 1952: 167]
Type species: Homalota pallens Mulsant & Rey, 1851
Characters/Description: Fenyes 1920: 316 (characters in key), 317 (characters), 318 (characters as Gampsonycha),
319 (characters as Gyronychina).
Distribution.
South America: Bolivia, Brazil, Chile, Argentina.
0595. Bamona Sharp, 1883: 287

Type species: Bamona gracilis Sharp, 1883

Distribution.
0596. Gyronycha Casey, 1893: 372

Type species: Gyronycha valens Casey, 1893
Distribution.
0597. Meotica Mulsant & Rey, 1873d: 174

Type species: Meotica parasita Mulsant & Rey, 1873
Cryptusa Mulsant & Rey, 1873d: 176 [synonymized by Blackwelder 1952: 113]
Type species: Meotica capitalis Mulsant & Rey, 1873
Cryptusa Mulsant & Rey, 1873b: 4 [synonymized by Blackwelder 1952: 113]
Cryptusa Mulsant & Rey, 1874a: 30 [synonymized by Blackwelder 1952: 113]
Cryptusa Mulsant & Rey, 1874c: 36 [synonymized by Blackwelder 1952: 113]
Cryptusa Mulsant & Rey, 1875a: 100 [synonymized by Blackwelder 1952: 113]
Cryptusa Mulsant & Rey, 1875c: 74 [synonymized by Blackwelder 1952: 113]
Meotica Mulsant & Rey, 1873b: 4 [synonymized by Blackwelder 1952: 238]
Meotica Mulsant & Rey, 1874a: 28 [Newton & Thayer 2003]
Meotica Mulsant & Rey, 1874c: 36 [synonymized by Blackwelder 1952: 238]
Sipaliella Casey, 1911: 159 [synonymized by Gusarov 2002a: 85]
Type species: Sipalia filaria Casey, 1911
Characters/Description: Fenyes 1920: 252 (characters as Sipaliella), 319 (characters).
Distribution.
South America: Peru, Argentina.
Other regions: Nearctic, Palaearctic, Madagascan, Oriental.
Subtribe Microglottina Fenyes, 1918
0598. Haploglossa Kraatz, 1856b: 78

Type species: Aleochara villosula Stephens, 1832
Microglotta Kraatz, 1862a: 300 [synonymized by Blackwelder, 1952: 245]
Type species: Aleochara pulla Gyllenhal, 1827

Characters/Description: Fenyes 1920: 391 (characters as Microglotta); Klimaszewski & Ashe 1991: 410
Distribution.
Subtribe Oxypodina Thomson, 1859
0599. Ambodina Sharp, 1883: 157

Type species: Ambodina granulata Sharp, 1883

Distribution.
0600. Calodera Mannerheim, 1830: 85

Type species: Calodera nigrita Mannerheim, 1830
Callidera Agassiz, 1846: 60 [synonymized by Blackwelder 1952: 89]
Callidera Gistel, 1856: 387 [synonymized by Blackwelder 1952: 89]
Caloderella Bernhauer, 1921b: 182 [synonymized by Scheerpeltz 1934: 1672]
Type species: Calodera nigerrima Bernhauer, 1921
Caloderodema Scheerpeltz, 1974: 375 [Newton & Thayer 2003]
Type species: Calodera basutoensis Scheerpeltz, 1974
Caloderona Bernhauer, 1921f: 176 [synonymized by Blackwelder 1952: 91]
Type species: Calodera dilatata Bernhauer, 1921
Caloderopsis Scheerpeltz, 1934: 1672 [synonymized by Blackwelder 1952: 91]
Type species: Calodera nigerrima Bernhauer, 1921
Ityocara Thomson, 1867a: 239 [synonymized by Assing 2003: 225]
Type species: Calodera rubens Erichson, 1837
Ityocara Thomson, 1867b: 46 [synonymized by Blackwelder 1952: 206]
Type species: Calodera rubens Erichson, 1837
Nasirema Casey, 1893: 307 [synonymized by Assing 2002: 196]
Type species: Nasirema humilis Casey, 1893
Pentaulacodera Bernhauer, 1943: 179 [synonymized by Blackwelder 1952: 298]
Type species: Calodera grandipennis Bernhauer, 1943

Triaulacodera Bernhauer, 1943: 180 [synonymized by Blackwelder 1952: 393]
Type species: Calodera minima Bernhauer, 1943
Characters/Description: Lynch-Arribálzaga 1884: 85 (characters); Fenyes 1920: 329 (characters), 338 (characters
as Ityocara), 339 (characters as Nasirema); Pace 1987c: 168 (characters in key).
Distribution.
South America: Brazil, Paraguay, Chile, Argentina.
0601. Cousya Mulsant & Rey, 1875e: 258

Type species: Calodera nigrata Fairmaire & Laboulbène, 1856
Chilomorpha Krása, 1914: 146 [synonymized by Assing & Schülke 2007: 34]
Type species: Chilomorpha bernhaueri Krása, 1914
Cousya Mulsant & Rey, 1875d: 420 [synonymized by Blackwelder 1952: 109]

Key: Pace 2001b: 431 (some species of the genus).
Distribution.
West Indies: Grenada.
0602. Ocalea Erichson, 1837: 298

Type species: Ocalea castanea Erichson, 1837
Subgenera:
Ocalea Erichson, 1837: 298
Type species: Ocalea castanea Erichson, 1837
Tetrocalea Cameron, 1939b: 576
Type species: Ocalea rufobrunnea Cameron, 1939
Characters/Description: Lynch-Arribálzaga 1884: 79 (characters); Fenyes 1920: 340 (characters); Ashe 2002b: 125
Key: Lynch-Arribálzaga 1884: 80 (species of Buenos Aires province, Argentina).
Distribution.
South America: Bolivia, Argentina.
0603. Oxypoda Mannerheim, 1830: 69

Type species: Oxypoda spectabilis Märkel, 1844
Ancillota Casey, 1910a: 165 [synonymized by Gusarov 2003e: 15]
Type species: Ancillota sollemnis Casey, 1910
Moluciba Casey, 1911: 156 [synonymized by Gusarov 2003e: 15]
Type species: Moluciba grandipennis Casey, 1911
Oxeopus Gistel, 1856: 387 [synonymized by Blackwelder 1952: 280]
Subgenera:
Atlantoxypoda Zerche, 1996: 304
Type species: Oxypoda assingi Zerche, 1996
Baeoglena Thomson, 1867a: 248
Type species: Oxypoda praecox Erichson, 1839
Bessopora Thomson, 1859: 37
Type species: Oxypoda testacea Erichson, 1839
Dromyusa Mulsant & Rey, 1875e: 192 [Newton & Thayer 2003]
Type species: Oxypoda picta Mulsant & Rey, 1875
Dromyusa Mulsant & Rey, 1875d: 354 [Newton & Thayer 2003]
Canaroxypoda Zerche, 1996: 296
Type species: Oxypoda palmensis Zerche, 1996
Cyrtonychochaeta Scheerpeltz, 1947: 347
Type species: Cyrtonychochaeta hoelzeli Scheerpeltz, 1947
Demosoma Thomson, 1859: 37
Type species: Oxypoda formiceticola Märkel, 1841
Deropoda Bernhauer, 1902: 134
Type species: Oxypoda amicta Erichson, 1839
Baptopoda Bernhauer, 1902: 133 [synonymized by Assing 2004b: 70]
Type species: Oxypoda magnicollis Fauvel, 1878
Disochara Thomson, 1858: 34
Type species: Oxypoda elongatula Aubé, 1850
Mycetodrepa Thomson, 1859: 37
Type species: Aleochara alternans Gravenhorst, 1802
Oxypoda Mannerheim, 1830: 69 [*]
Type species: Oxypoda spectabilis Märkel, 1844
Paroxypoda Ganglbauer, 1895: 64
Type species: Oxypoda lugubris Kraatz, 1856
Podoxya Mulsant & Rey, 1875e: 135 [*]
Type species: Oxypoda lentula Erichson, 1837
Podoxya Mulsant & Rey, 1875d: 297 [Newton & Thayer 2003]
Sedomoma Tottenham, 1939a: 226
Type species: Oxypoda soror Thomson, 1855
Sphenoma Mannerheim, 1830: 68 [*]
Type species: Sphenoma abdominale Mannerheim, 1830
Thliboptera Thomson, 1859: 37 [Newton & Thayer 2003]
Type species: Oxypoda togata Erichson, 1839
31 species (1 in Mexican amber) are included in this genus from Central and South America.
Characters/Description: Fenyes 1920: 219 (characters as Moluciba), 363 (characters); Lohse et al. 1990: 125
(characters in key), 126 (characters); Ashe 2002b: 125 (characters in key); Gusarov 2003e: 15 (characters).
Distribution.
South America: Colombia, Brazil, Chile, Argentina.
0604. Polylobus Solier, 1849: 354 (Fig. 72)

Type species: Polylobus maculipennis Solier, 1849

Characters/Description: Fenyes 1920: 377 (characters); Klimaszewski & Sturm 1991: 3 (characters); Pace 1997b:
43 (characters in key); Pace 2000a: 400 (characters); Ashe 2002b: 125 (characters in key).
Key: Klimaszewski & Sturm 1991: 3 (Colombian species associate to Espeletia flowers of the Paramo region).
Distribution.
South America: Colombia, Ecuador, Peru, Brazil, Chile, Argentina.
0605. Tectusa Bernhauer, 1899: 18

Type species: Leptusa difficilis Eppelsheim, 1881
Parocyusa Bernhauer, 1902: 223 [synonymized by Assing & Schülke 2007: 32]
Type species: Ocyusa holdhausi Bernhauer, 1902

Characters/Description: Fenyes 1920: 353 (characters in key as Parocyusa), 356 (characters as Parocyusa), 358
(characters); Ashe 2002b: 125 (characters in key as Parocyusa).
Distribution.
0606. Xenomma Wollaston, 1854: 543

Type species: Xenomma planifrons Wollaston, 1854

Distribution.
0607. Zoosetha Mulsant & Rey, 1873b: 4

Type species: Homalota inconspicua Erichson, 1839
Ocyusida Bernhauer, 1900b: 197 [synonymized by Assing 1998: 74]
Type species: Ocyusida skalitzkyi Bernhauer, 1900

Distribution.
Sbtribu Phloeoporina Thomson, 1859
0608. Phloeopora Erichson, 1837: 311

Type species: Aleochara corticalis Gravenhorst, 1802
Phloeodroma Kraatz, 1856b: 338 [synonymized by Lohse 1989: 223]
Type species: Phloeodroma concolor Kraatz, 1856

Characters/Description: Fenyes 1920: 321 (characters as Phloeodrama), 322 (characters); Pace 1987c: 168
(characters in key); Lohse et al. 1990: 125 (characters in key), 140 (characters); Ashe 2002b: 125 (characters
in key).
Distribution.
West Indies: Cuba, Haiti, Puerto Rico, US Virgin Isl., Guadeloupe.
South America: Colombia, Chile, Argentina.
Incertae sedis in Tribe Oxypodini Thomson, 1859
0609. Abrophaena Pace, 1983d: 313

Type species: Abrophaena boliviana Pace, 1983

Distribution.
0610. Acrocyusa Bernhauer, 1930: 202

Type species: Ocyusa grandicornis Bernhauer, 1930

Distribution.
South America: French Guiana, Peru, Brazil.
0611. Akanthinoknemusa Pace, 2015e: 124

Type species: Akanthinoknemusa cerroblancoicola Pace, 2015

Distribution.
0612. Alogotheke Pace, 2015c: 77

Type species: Alogotheke guyanense Pace, 2015

Distribution.
0613. Andinostiba Scheerpeltz, 1972d: 152

Type species: Atheta eleganticornis Scheerpeltz, 1972

Characters/Description: Pace 1987c: 168 (characters in key).
Distribution.
0614. Anocalea Fenyes, 1921a: 27

Type species: Anocalea thaxteri Fenyes, 1921

Distribution.
0615. Chilodera Cameron, 1944a: 619

Type species: Chilodera falklandica Cameron, 1944

Distribution.
South America: Falkland Isl.
0616. Chiloestiba Pace, 2015d: 28

Type species: Chiloestiba zanettii Pace, 2015

Distribution.
0617. Cnemostyla Pace, 1988: 94

Type species: Cnemostyla mirabilis Pace, 1988

Characters/Description: Pace 1988: 94 (characters).
Distribution.
0618. Cornellusa Pace, 2015e: 120

Type species: Cornellusa peruviana Pace, 2015

Distribution.
0619. Ctenopeuca Bernhauer, 1915b: 299

Type species: Ctenopeuca heynei Bernhauer, 1915

Distribution.
0620. Dasymera Fauvel, 1866b: 290
Type species: Dasymera chillana Fauvel, 1866
Dasymera Fauvel, 1866a: 41 [Newton & Thayer 2003]
Type species: Dasymera chillana Fauvel, 1866

Distribution.
South America: Bolivia, Paraguay, Chile,Argentina.
0621. Demoglena Pace, 1987b: 505

Type species: Demoglena andensis Pace, 1987

Distribution.
0622. Diacanthochara Pace, 1983b: 145

Type species: Diacanthochara franzi Pace, 1983
Haplochara Pace, 1985: 388 [synonymized by Pace 2015e: 121]
Type species: Haplochara franzi Pace, 1985

Key: Pace 2008a: 352 (species of Ecuador and Peru as Haplochara).
Distribution.
South America: Venezuela, French Guiana, Ecuador, Peru, Brazil.
0623. Ecitodulus Wasmann, 1900b: 235

Type species: Ecitodulus crassicornis Wasmann, 1900

Distribution.
0624. Feluva Blackwelder, 1952: 165

Type species: Brachyglossa varicolor Fauvel, 1866
Brachyglossa Fauvel, 1866a: 276 [synonymized by Blackwelder 1952: 165]
Brachyglossa Fauvel, 1866b: 27 [synonymized by Blackwelder 1952: 165]

Characters/Description: Fenyes 1920: 317 (characters as Brachyglossa); Newton & Peck 2006: 135 (characters in
key).
Distribution.

South America: Colombia, French Guiana, Ecuador, Galapagos Isl., Peru, Brazil, Chile, Argentina.
0625. Gastrorhopalus Solier, 1849: 333
Type species: Gastrorhopalus niger Solier, 1849

Characters/Description: Fenyes 1920: 336 (characters); Pace 1988: 94 (characters).
Key: Pace 1987b: 485 (some species of Chile); Pace 1999c: 180 (some species of Chile); Pace 2008a: 332 (some
species of the genus).
Distribution.
0626. Glossarhymenus Pace, 2008a: 357

Type species: Glossarhymenus otonganus Pace, 2008

Distribution.
0627. Gnypetosoma Cameron, 1922a: 127

Type species: Gnypetosoma calocera Cameron, 1922
Distribution.
0628. Idiostiba Pace, 1990b: 74

Type species: Idiostiba franzi Pace, 1990

Distribution.
South America: Colombia, Ecuador, Brazil, Argentina.
0629. Kaweshkarusa Pace, 1992a: 359

Type species: Kaweshkarusa daccordii Pace, 1992

Distribution.
0630. Kochlifeluva Pace, 2011b: 63

Type species: Kochlifeluva ecuadorensis Pace, 2011

Distribution.
0631. Meoticaops Pace, 1983b: 150
Type species: Meoticaops franzi Pace, 1983
Distribution.
South America: Venezuela, Ecuador, Peru, Bolivia.
0632. Myrmigaster Sharp, 1876a: 50

Type species: Myrmigaster singularis Sharp, 1876
Fauvelia Wasmann, 1896a: 174 [synonymized by Ashe & Kistner 2005: 223]
Type species: Fauvelia permira Wasmann, 1896
Pseudodinarda Wasmann, 1896a: 174 [synonymized by Ashe & Kistner 2005: 223]
Type species: Fauvelia permira Wasmann, 1896
Characters/Description: Fenyes 1920: 280 (characters); Ashe & Kistner 2005: 223 (characters).
Distribution.
0633. Neocalodera Pace, 1987c: 168

Type species: Bolitochara arcuata Fauvel, 1866

Distribution.
0634. Newtonusa Pace, 1999c: 180

Type species: Newtonusa chilensis Pace, 1999

Distribution.
0635. Nordenskjoldella Enderlein, 1912: 65

Type species: Nordenskjoldella flavitarsis Enderlein, 1912

Distribution.
0636. Ocyusa Kraatz, 1856b: 156

Type species: Oxypoda maura Erichson, 1837
Deubelia Bernhauer, 1899: 15 [synonymized by Assing & Schülke 2001: 170]
Type species: Deubelia diabolica Bernhauer, 1899

Characters/Description: Fenyes 1920: 349 (characters as Deubelia), 352 (characters); Lohse et al. 1990: 125

Distribution.
0637. Oligonotus Lynch-Arribálzaga, 1884: 89

Type species: Oligonotus exiguus Lynch-Arribálzaga, 1884

Distribution.
0638. Parabainusa Pace, 1997b: 41

Type species: Parabainusa tropica Pace, 1997

Distribution.
0639. Parahydrosmecta Scheerpeltz, 1972d: 149

Type species: Atheta quemquemtreuana Scheerpeltz, 1972

Distribution.
0640. Pnephorusa Pace, 2000a: 459

Type species: Pnephorusa decorata Pace, 2000

Distribution.
0641. Poikilnotha Pace, 2000a: 473

Type species: Poikilnotha valdiviana Pace, 2000

Distribution.
0642. Polyacanthode Bernhauer, 1939e: 148

Type species: Polyacanthode kochianus Bernhauer, 1939
Distribution.
0643. Polylobinus Bernhauer, 1908: 370
Type species: Polylobinus brasiliensis Bernhauer, 1908

Characters/Description: Fenyes 1920: 377 (characters); Pace 2000a: 400 (characters).
Distribution.
0644. Prymnorhopala Bernhauer, 1922a: 21

Type species: Prymnorhopala opaca Bernhauer, 1922

Distribution.
0645. Pseudomyllaena Pace, 1986a: 463

Type species: Pseudomyllaena franzi Pace, 1986

Distribution.
0646. Somasterochara Pace, 1986a: 458

Type species: Somasterochara peruviana Pace, 1986

Distribution.
0647. Spanioda Blackwelder, 1952: 354

Type species: Calodera fairmairei Bernhauer, 1927
Spaniodera Bernhauer, 1927b: 262 [synonymized by Blackwelder 1952: 354]
Type species: Calodera fairmairei Bernhauer, 1927

Characters/Description: Pace 1987c: 168 (characters in key); Pace 1992a: 361 (characters in key).
Key: Pace 1987b: 498 (some species of Chile); Pace 2000a: 455 (some species of Chile).
Distribution.
South America: Peru, Brazil, Chile, Argentina.
0648. Sphaenidatoma Pace, 1987c: 169

Type species: Tachyusa elegantula Scheerpeltz, 1972

Distribution.
0649. Teratanytoma Pace, 1999c: 182
Type species: Teratanytoma newtoni Pace, 1999

Distribution.
0650. Tomocoryphusa Pace, 2000a: 453

Type species: Phloeopora pacaorum Pace, 1987

Distribution.
0651. Tricolpochila Bernhauer, 1908: 368

Type species: Tricolpochila kraatzi Bernhauer, 1908

Characters/Description: Fenyes 1920: 381 (characters); Pace 2000a: 400 (characters).
Distribution.
South America: Peru, Bolivia, Brazil, Paraguay, Chile, Argentina.
0652. Troglocyphodas Pace, 1986c: 496

Type species: Troglocyphodas decui Pace, 1986

Distribution.
Tribe Paglini Newton & Thayer, 1992
0653. Pagla Blackwelder, 1952: 287

Type species: Hoplandria anthracina Fairmaire & Germain, 1862
Euryglossa Fauvel, 1866b: 326 [synonymized by Fauvel 1868a: 379]
Euryglossa Fauvel, 1866a: 77 [synonymized by Blackwelder 1952: 287]
Pachyglossa Fauvel, 1868a: 379 [synonymized by Blackwelder 1952: 287]
Subgenera:
Pagla Blackwelder, 1952: 287 [*]
Parallelopagla Pace, 2007: 227 [*]
Type species: Pagla levis Pace, 2007
Key: Pace 2007: 227 (subgenera of the genus).
Distribution.
0654. Paglhoplasymma Pace, 2015e: 58
Type species: Paglhoplasymma heliconiae Pace, 2015

Distribution.
Tribe Paradoxenusini Bruch, 1937
0655. Paradoxenusa Bruch, 1937: 354

Type species: Paradoxenusa silvestrii Bruch, 1937

Distribution.
Tribe Philotermitini Seevers, 1957
0656. Pseudophilotermes Bernhauer, 1934c: 511

Type species: Pseudophilotermes bruchi Bernhauer, 1934
Neophilotermes Seevers, 1957: 257 [synonymized by Kistner & Jacobson 1976: 49]
Type species: Philotermes laxicornis Sharp, 1883
Characters/Description: Seevers 1957: 251 (characters in key); Kistner 1971: 17 (characters, Neophilotermes);
Ashe 2002b: 128 (characters in key).
Key: Kistner 1971: 18 (Neophilotermes); Kistner & Jacobson 1976: 49 (species of the genus).
Distribution.
South America: Venezuela, Peru, Argentina.
Tribe Phytosini Thomson, 1867a
0657. Cameronium C. Koch, 1936: 202

Type species: Phytosus obockianus Fauvel, 1905

Distribution.
Other regions: Palaearctic, Ethiopian, Madagascan.
0658. Phytosus Curtis, 1838: pl. 718

Type species: Phytosus spinifer Curtis, 1838

Subgenera:
Actosus Mulsant & Rey, 1871: 300
Type species: Myrmedonia nigriventris Chevrolat, 1843
Euphytosus Bernhauer & Scheerpeltz, 1926: 552
Type species: Phytosus schenklingi Bernhauer, 1922
Paraphytosus Bernhauer, 1922b: 236 [synonymized by Bernhauer & Scheerpeltz 1926: 552]
Type species: Phytosus schenklingi Bernhauer, 1922
Phytosus Curtis, 1838: pl. 718 [*]
Type species: Phytosus spinifer Curtis, 1838

Distribution.
Tribe Placusini Mulsant & Rey, 1871
0659. Euvira Sharp, 1883: 278

Type species: Euvira nigra Sharp, 1883
Crimalia Casey, 1911: 206 [synonymized by Seevers 1978: 168]
Type species: Crimalia quadriceps Casey, 1911

Characters/Description: Fenyes 1920: 118 (characters as Crimalia); Cameron 1948b: 233 (characters); Ashe &
Kistner 1989: 86 (characters); Ashe 2002b: 130 (characters in key); Newton & Peck 2006: 135 (characters in
key); Santiago-Jiménez & Santiago-Navarro 2016: 49 (characters in key).
Distribution.
West Indies: Cuba, Guadeloupe, Saint Vincent & the Grenadines.
Central America: Mexico, Guatemala, Nicaragua, Costa Rica.
South America: Colombia, Venezuela, Ecuador, Galapagos Isl., Peru, Brazil, Argentina.
0660. Kyrtusa Pace, 2008a: 252

Type species: Kyrtusa curticornis Pace, 2008
Kirtusa Pace, 2008a: 225, 252 [synonymized by Newton 2017: 18]

Characters/Description: Santiago-Jiménez & Santiago-Navarro 2016: 49 (characters in key).
Distribution.
0661. Placukorna Santiago-Jiménez in Santiago-Jiménez & Santiago-Navarro, 2016: 46

Type species: Placukorna ipsa Santiago-Jiménez in Santiago-Jiménez & Santiago-Navarro, 2016

Characters/Description: Santiago-Jiménez & Santiago-Navarro 2016: 49 (characters in key).
Distribution.
0662. Placusa Erichson, 1837: 370 (Fig. 71)

Type species: Aleochara pumilio Gravenhorst, 1802
Subgenera:
Calpusa Mulsant & Rey, 1871: 108
Type species: Placusa adscita Erichson, 1839
Calpusa Mulsant & Rey, 1872b: 198 [synonymized by Blackwelder 1952: 92]
Calpusa Mulsant & Rey, 1873c: 74 [synonymized by Blackwelder 1952: 92]
Placusa Erichson, 1837: 370
Type species: Aleochara pumilio Gravenhorst, 1802
Pelekoglossa Cameron, 1920a: 226 [Smetana 2004: 492]
Type species: Pelekoglossa cingulata Cameron, 1920

Characters/Description: Coiffait & Sáiz 1967: 64 (characters in key), 65 (characters); Klimaszewski et al. 2001: 6
(characters); Ashe 2002b: 131 (characters in key); Santiago-Jiménez & Santiago-Navarro 2016: 49
(characters).
Distribution.
West Indies: Saint Vincent & the Grenadines, Guadeloupe.
South America: Colombia, Venezuela, Suriname, Peru, Bolivia, Brazil, Chile, Argentina.
Tribe Pronomaeini Mulsant & Rey, 1873b
0663. Perupora Pace, 2015e: 47

Type species: Perupora huanucensis Pace, 2015

Distribution.
0664. Pronomaea Erichson, 1837: 378

Type species: Pronomaea rostrata Erichson, 1837
Cephalomaea Bernhauer in Bernhauer & Paulian, 1942: 368 [synonymized by Blackwelder 1952: 96]
Type species: Pronomaea kamerunensis Bernhauer, 1942

Distribution.
West Indies: Jamaica, Guadeloupe.
0665. Pseudomniophila Pace, 1985: 372

Type species: Pseudomniophila unidentata Pace, 1985

5 species are known from Central and South America.
Characters/Description: Klimaszewski & Ashe 1993: 79 (characters).
Key: Klimaszewski & Ashe 1993: 81 (some species of the genus).
Distribution.
South America: Venezuela, Ecuador, Peru.
Tribe Sceptobiini Seevers, 1978
0666. Dinardilla Wasmann, 1901a: 145

Type species: Dinardilla liometopi Wasmann, 1901

Characters/Description: Fenyes 1920: 265 (characters); Danoff-Burg 1994: 39 (characters in key); Ashe 2002b:
Key: Danoff-Burg 1994: 39 (species of the genus).
Distribution.
0667. Sceptobius Sharp, 1883: 211

Type species: Sceptobius dispar Sharp, 1883
Apteronina Wasmann, 1901a: 146 [synonymized by Danoff-Burg 1994: 36]
Type species: Apteronina schmitti Wasmann, 1901
Symbiochara Fenyes, 1909: 325 [synonymized by Danoff-Burg 1994: 33]
Type species: Symbiochara lativentris Fenyes, 1909

Characters/Description: Fenyes 1920: 260 (characters as Apteronina), 285 (characters), 288 (characters as
Symbiochara); Danoff-Burg 1994: 32 (characters), 39 (characters in key); Ashe 2002b: 127 (characters in
key).
Key: Danoff-Burg 1994: 39 (species of the genus).
Distribution.
Tribe Tachyusini Thomson, 1859
0668. Gnypeta Thomson, 1858: 33

Type species: Homalota labilis Erichson, 1839
Euliusa Casey, 1906: 215 [synonymized by Blackwelder 1952: 173]
Type species: Gnypeta lucens Bernhauer, 1905
Gnypetoma Casey, 1906: 196 [synonymized by Blackwelder 1952: 173]
Type species: Tachyusa baltifera LeConte, 1863
Characters/Description: Fenyes 1920: 162 (characters); Ashe 2002b: 128 (characters in key); Klimaszewski et al.
2008: 15 (characters); Paśnik 2010: 37 (characters in key), 71 (characters).
Distribution.
West Indies: Jamaica, St. Lucia, Grenada.
South America: Colombia, French Guiana, Ecuador, Peru, Bolivia, Brazil, Chile, Argentina.
0669. Ischnopoda Stephens, 1833b: 91 (Fig. 61)

Type species: Staphylinus leucopus Marsham, 1802
Amanota Casey, 1905: 189 [synonymized by Paśnik 2006b: 13]
Type species: Amanota capensis Casey, 1906
Calischnopoda Reitter, 1909: 73 [synonymized by Paśnik 2006b: 13]
Type species: Drusilla exarata Mannerheim, 1830
Cathusya Mulsant & Rey, 1873b: 6 [synonymized by Paśnik 2006b: 12]
Type species: Tachyusa scitula Erichson, 1837
Cathusya Mulsant & Rey, 1874c: 38 [synonymized by Blackwelder 1952: 95]
Cathusya Mulsant & Rey, 1875a: 409 [synonymized by Blackwelder 1952: 95]
Cathusya Mulsant & Rey, 1875c: 383 [synonymized by Blackwelder 1952: 95]
Ischnopoda Stephens, 1835: 430 [Newton & Thayer 2003]
Leucopus Bertolini, 1873: 48 [Newton & Thayer, 2003]
Pischnopoda Tottenham, 1939a: 226 [rejected by ICZN 1961: 241]
Type species: Staphylinus leucopus Marsham, 1802
Rechota Sharp, 1883: 228 [synonymized by Paśnik 2006b: 13]
Type species: Rechota impressa Sharp, 1883
Characters/Description: Fenyes 1920: 160 (characters as Amanota), 168 (characters as Rechota), 170 (characters in
key as Calischnopoda and Cathusya), 171 (characters), 171 (characters as Calischnopoda), 172 (characters as
Cathusya); Ashe 2002b: 128 (characters in key as Rechota); Paśnik 2006b: 12 (characters); Paśnik 2010: 37
Key: Paśnik 2006b: 17 (species of the genus).
Distribution.
South America: French Guiana, Ecuador, Bolivia, Brazil, Chile, Argentina.
0670. Meronera Sharp, 1887a: 779

Type species: Falagria venustula Erichson, 1839
Merona Sharp, 1883: 229 [synonymized by Sharp 1887a: 779]
Type species: Falagria venustula Erichson, 1839

Characters/Description: Fenyes 1920: 164 (characters); Ashe 2002b: 126 (characters in key); Paśnik 2010: 39
Key: Pace 1990a: 158 (species of the Neotropical region); Pace 2008a: 278 (species of Ecuador).
Distribution.
West Indies: Cuba, Jamaica, Guadeloupe, Grenada.

South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Bolivia, Brazil, Chile, Argentina.
0671. Neolara Sharp, 1883: 231
Type species: Neolara centralis Sharp, 1883

Characters/Description: Casey 1906: 188 (characters); Fenyes 1920: 167 (characters); Ashe 2002b: 126 (characters
in key); Paśnik 2010: 39 (characters in key), 105 (characters).
Distribution.
West Indies: Cuba, Grenada.
South America: Colombia, Venezuela, French Guiana, Ecuador, Bolivia, Brazil, Paraguay, Argentina.
0672. Pseudognypeta Cameron, 1923: 365

Type species: Pseudognypeta picta Cameron, 1923
Characters/Description: Paśnik 2007: 1121 (characters); Paśnik 2010: 38 (characters in key), 95 (characters).
Distribution.
0673. Tachyusa Erichson, 1837: 307

Type species: Tachyusa constricta Erichson, 1837
Caliusa Mulsant & Rey, 1873b: 6 [synonymized by Blackwelder 1952: 88]
Caliusa Mulsant & Rey, 1874c: 38 [synonymized by Blackwelder 1952: 88]
Caliusa Mulsant & Rey, 1875a: 409 [synonymized by Blackwelder 1952: 88]
Caliusa Mulsant & Rey, 1875c: 383 [synonymized by Paśnik 2006a: 19]
Type species: Tachyusa balteata Erichson, 1839
Chyusata Tottenham, 1945: 70 [synonymized by ICZN 1961: 241]
Type species: Tachyusa constricta Erichson, 1837
Tachyusilla Casey, 1906: 213 [synonymized by Paśnik 2006a: 19]
Type species: Tachyusa balteata Erichson, 1839
Tachyusota Casey, 1906: 213 [synonymized by Paśnik 2006a: 19]
Type species: Tachyusa gemma Casey, 1906
1 species is included in this genus from Central. Paśnik (2006a: 148) in his revision of Tachyusa, concluded that
Tachyusa gratiosa Bernhauer, 1912d: 73, should be assigned to tribe Bolitocharini Thomson.
Characters/Description: Fenyes 1920: 169 (characters), 170 (characters in key as Caliusa), 172 (characters), 173
(characters as Caliusa); Ashe 2002b: 128 (characters in key); Paśnik 2006a: 19 (characters); Paśnik 2010: 36
Key: Paśnik 2006a: 23 (species of the genus).
Distribution.
0674. Teliusa Casey, 1906: 203
Type species: Teliusa alutacea Casey, 1906

Characters/Description: Fenyes 1920: 174 (characters); Paśnik 2005: 734 (characters); Paśnik 2010: 36 (characters
in key), 75 (characters).
Key: Paśnik 2005: 735 (species of the genus).
Distribution.
West Indies: Cuba, Jamaica, St. Lucia, Grenada.
Tribe Taxicerini Lohse in Lohse & Lucht, 1989
0675. Halobrecta Thomson, 1858: 35

Type species: Homalota puncticeps Thomson, 1852
Exatheta Cameron, 1920a: 265 [synonymized by Sawada 1985: 106]
Type species: Exatheta cingulata Cameron, 1920
Glaphya Mulsant & Rey, 1873d: 172 [Newton & Thayer 2003]
Type species: Dinaraea pubes Mulsant & Rey 1873
Glaphya Mulsant & Rey, 1873b: 646 [synonymized by Bernhauer & Scheerpeltz 1926: 621]
Glaphya Mulsant & Rey, 1874a: 26 [Newton & Thayer, 2003]
Glaphya Mulsant & Rey, 1874c: 678 [synonymized by Blackwelder 1952: 170]

Characters/Description: Fenyes 1920: 185 (characters); Gusarov 2004: 2 (characters).
Key: Gusarov 2004: 8 (Nearctic species).
Distribution.
Tribe Termitohospitini Seevers, 1941
Subtribe Termitohospitina Seevers, 1941
0676. Blapticoxenus Mann, 1923: 362

Type species: Blapticoxenus brunneus Mann, 1923
Termitocolonus Seevers, 1941: 336 [synonymized by Seevers 1957: 199]
Type species: Termitocolonus ericiogaster Seevers, 1941

Distribution.
0677. Neotermitosocius Kanao et al., 2012: 70 (Fig. 66)

Type species: Neotermitosocius bolivianus Kanao, Eldredge & Maruyama, 2012

Distribution.
0678. Paratermitosocius Seevers, 1941: 340
Type species: Perinthus vestitus Mann, 1923

Distribution.
0679. Termitohospes Seevers, 1941: 333

Type species: Termitohospes miricorniger Seevers, 1941

(characters).
Distribution.
South America: Guyana, Brazil, Peru.
0680. Termitosocius Seevers, 1941: 338

Type species: Termitosocius microps Seevers, 1941

Distribution.
0681. Termitosodalis Seevers, 1941: 337

Type species: Termitosodalis barticae Seevers, 1941
Termitomus Silvestri, 1947: 138 [synonymized by Seevers 1957: 200]
Type species: Termitomus fasciatus Silvestri, 1947

Distribution.
Tribe Termitonannini Fenyes, 1918
Subtribe Perinthina Bernhauer & Scheerpeltz, 1926
0682. Macrognathellus Silvestri, 1946a: 20

Type species: Macrognathellus paraguayensis Silvestri, 1946
1 species is known from and is restricted to South America.
Characters/Description: Seevers 1957: 165 (characters in key), 174 (characters); Kistner 2004: 100 (characters).
Distribution.
0683. Perinthus Casey, 1890: 192
Type species: Perinthus dudleyanus Casey, 1890

(characters); Ashe 2002b: 131 (characters in key); Kistner 2004: 10 (characters).
Key: Seevers 1937: 10 (species of the genus); Kistner 2004: 16 (species of the genus).
Distribution.
South America: Colombia, Guyana, Ecuador, Peru, Bolivia, Brazil, Paraguay.
0684. Poduroides Mann, 1926: 151

Type species: Poduroides bovingi Mann, 1926

Distribution.
0685. Termitocola Seevers, 1937: 7

Type species: Termitocola cylindricornis Seevers, 1937
(characters); Ashe 2002b: 131 (characters in key); Kistner 2004: 35 (characters).
Key: Kistner & Jacobson 1976: 48 (species of the genus); Kistner 2004: 39 (species of the genus).
Distribution.
Central America: Mexico, Nicaragua, Panama.
0686. Termitonicus Mann, 1926: 153

Type species: Termitonicus mahout Mann, 1926

Distribution.
South America: Guyana, Brazil, Argentina.
0687. Termitopelta Borgmeier, 1950: 656

Type species: Termitopelta fulgens Borgmeier, 1950

Distribution.

Subtribe Termitonannina Fenyes, 1918
0688. Chaetonannus Borgmeier, 1959b: 295

Type species: Chaetonannus fragilicornis Borgmeier, 1959

Distribution.
0689. Eunannodes Silvestri, 1946a: 15

Type species: Eunannodes reconditi Silvestri, 1946

Distribution.
0690. Macrotrichurus Silvestri, 1946a: 11

Type species: Macrotrichurus brasiliensis Silvestri, 1946

Characters/Description: Seevers, 1957: 189 (characters).
Distribution.
0691. Nannellus Silvestri, 1946a: 17

Type species: Nannellus anoplotermitis Silvestri, 1946

Distribution.
0692. Nannusa Borgmeier, 1959b: 298

Type species: Nannusa longicornis Borgmeier, 1959

Distribution.
0693. Termitocomes Seevers, 1941: 342

Type species: Termitocomes wasmanni Seevers, 1941
Distribution.
0694. Termitonannus Wasmann, 1902b: 2
Type species: Termitonannus schmalzi Wasmann, 1902
Tetraphilus Silvestri, 1946a: 2 [synonymized by Blackwelder 1952: 378]
Type species: Termitonannus brachycerus Silvestri, 1946
Characters/Description: Seevers 1957: 182 (characters); Kistner 2006: 633 (characters).
Distribution.
South America: Guyana, Peru, Bolivia, Brazil, Paraguay, Argentina.
0695. Termitonilla Borgmeier, 1950: 651

Type species: Termitonilla luteola Borgmeier, 1950

Distribution.
Tribe Termitusini Fenyes, 1918
Subtribe Termitospectrina Seevers, 1957
0696. Termitospectrum Mann, 1926: 154

Type species: Termitospectrum thoracicum Mann, 1926

Distribution.
Tribe Trichopseniini LeConte & Horn, 1883
0697. Megaxenistusa Seevers, 1945: 66

Type species: Megaxenistusa rhinotermitis Seevers, 1945

Distribution.
0698. Prorhinopsenius Pasteels & Kistner, 1971: 353

Type species: Hamitopsenius micronesiae Seevers, 1957
Palaeopsenius Seevers, 1971: 78 [synonymized by Kistner 1998: 53]

Type species: Palaeopsenius mexicanus Seevers, 1971
4 species are included in this genus from the West Indies and Central America, all in amber.
Key: Kistner, 1998: 62 (species of the genus).
Distribution.
Other regions: Australian, Oceanic.
0699. Rhinotermopsenius Seevers, 1941: 329

Type species: Rhinotermopsenius saltatorius Seevers, 1941

Distribution.
0700. Termitopsenius Wasmann, 1902b: 4

Type species: Termitopsenius limulus Wasmann, 1902
Pulicipsenius Seevers, 1941: 327 [synonymized by Seevers 1957: 275]
Type species: Pulicipsenius acanthoscelis Seevers, 1941

Characters/Description: Seevers 1941: 331 (characters); Seevers 1957: 267 (characters in key), 275 (characters).
Distribution.
South America: Guyana, Brazil, Argentina.
Incertae sedis in Subfamily Aleocharinae
0701. Salinamexus Moore & Legner, 1977: 463

Type species: Salinamexus browni Moore & Legner, 1977
Biophytosus Moore & Legner, 1977: 465 [synonymized by Ahn 1996c: 204]
Type species: Biophytosus reticulatus Moore & Legner, 1977

Characters/Description: Ahn 2001: 131 (characters in key); Ashe 2002b: 129 (characters in key); Maruyama 2011:
106 (characters).
Distribution.
Subfamily SCAPHIDIINAE Latreille, 1807
Tribe Cypariini Achard, 1924
0702. Cyparium Erichson, 1845: 3 (Fig. 81)

Type species: Cyparium palliatum Erichson, 1845
Yparicum Achard, 1920: 126 [synonymized by Löbl 1992: 498]
Type species: Yparicum yunnanum Achard, 1920

Characters/Description: Matthews 1888: 165 (characters); Leschen & Löbl 1995: 448 (characters in key), 448
(characters); Hwang & Ahn 2001: 369 (characters); Navarrete-Heredia et al. 2002: 201 (characters in key);
Fierros-Lopéz 2002: 8 (characters).
Key: Matthews 1888: 166 (species of Central America); Fierros-Lopéz 2002: 10 (Mexican species).
Distribution.
West Indies: Cuba, Jamaica, Dominican Republic.
South America: Colombia, Suriname, Ecuador, Peru, Bolivia, Brazil.
Tribe Scaphidiini Latreille, 1806
0703. Scaphidium Olivier, 1790: 20.1 (Figs. 82–83)

Type species: Scaphidium quadrimaculatum Olivier, 1790
Ascaphidium Pic, 1915a: 24 [synonymized by Leschen & Löbl 1995: 453]
Type species: Ascaphidium sikorai Pic, 1915
Cribroscaphium Pic, 1920: 93 [synonymized by Leschen & Löbl 1995: 453]
Type species: Scaphidium irregulare Pic, 1920
Falsoascaphidium Pic, 1923: 16 [synonymized by Leschen & Löbl 1995: 453]
Type species: Scaphidium subdepressum Pic, 1921
Hemiscaphium Achard, 1922: 12 [synonymized by Leschen & Löbl 1995: 453]
Type species: Scaphidium striatipenne Gestro, 1879
Hyposcaphidium Achard, 1922: 12 [synonymized by Leschen & Löbl 1995: 453]
Type species: Scaphidium rufopygum Lewis, 1893
Isoscaphidium Achard, 1922: 12 [synonymized by Leschen & Löbl 1995: 453]
Type species: Scaphidium quadriguttatum Say, 1823
Pachyscaphidium Achard, 1922: 12 [synonymized by Leschen & Löbl 1995: 453]
Type species: Scaphidium arrowi Achard, 1920
Parascaphium Achard, 1923: 97 [synonymized by Löbl 1968: 386]
Type species: Scaphium optabile Lewis, 1893
Scaphidiolum Achard, 1922: 12 [synonymized by Leschen & Löbl 1995: 453]
Type species: Scaphidium basale Laporte, 1840
Scaphidopsis Achard, 1922: 12 [synonymized by Leschen & Löbl 1995: 453]
Type species: Scaphidium pardale Laporte, 1840
Characters/Description: Matthews 1888: 159 (characters); Leschen & Löbl 1995: 448 (characters in key), 453
(characters); Hwang & Ahn 2001: 369 (characters); Navarrete-Heredia et al. 2002: 201 (characters in key);
Fierros-Lopéz 2005: 12 (characters).
Key: Matthews 1888: 159 (species of Central America); Fierros-Lopéz 2005: 14 (species of Central America).
Distribution.
Central America: Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama.
South America: Colombia, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Oceanic.
Tribe Scaphisomatini Casey, 1893
0704. Alexidia Reitter, 1880: 43

Type species: Alexidia rogenhoferi Reitter, 1880
Characters/Description: Löbl & Leschen 2003: 316 (characters); Leschen & Löbl 2006: 19 (characters in key).
Key: Löbl & Leschen 2003: 316 (species of the genus).
Distribution.
South America: Colombia, Ecuador, Brazil.
0705. Amalocera Erichson, 1845: 4

Type species: Amalocera picta Erichson, 1845

Characters/Description: Löbl 1974: 39 (characters); Leschen & Löbl 2006: 19 (characters in key).
Key: Löbl 1974: 39 (species of the genus).
Distribution.
0706. Baeocera Erichson, 1845: 4

Type species: Baeocera falsata Achard, 1920
Amaloceroschema Löbl, 1967: 1 [synonymized by Ogawa & Löbl 2013: 302]
Type species: Baeocera freudei Löbl, 1967
Cyparella Achard, 1924: 28 [synonymized by Löbl 1987: 856]
Type species: Scaphisoma rufoguttatum Fairmaire, 1898
Eubaeocera Cornell, 1967: 2 [synonymized by Löbl 1977b: 5]
Type species: Baeocera abdominalis Casey, 1900
Sciatrophes Blackburn, 1903: 100 [synonymized by Löbl 1977a: 102]
Type species: Sciatrophes latens Blackburn, 1903

Characters/Description: Matthews 1888: 168 (characters); Navarrete-Heredia et al. 2002: 202 (characters in key);
Leschen & Löbl 2006: 19 (characters in key).
Key: Matthews 1888: 168 (species of Central America); Löbl 1983: 162 (Chilean species); Fierros-Lopéz 2010:
206 (Mexican species)); Löbl 2018: 1312 (Chilean species).
Distribution.
Central America: Mexico, Guatemala, Nicaragua.
South America: Colombia, Galapagos Isl., Brazil, Chile, Argentina.
0707. Scaphisoma Leach, 1815: 89 (Fig. 84)

Type species: Silpha agaricina Linnaeus, 1758
Caryoscapha Ganglbauer, 1899: 343 [synonymized by Leschen & Löbl 2006: 24]
Type species: Scaphisoma limbatum Erichson, 1845
Macrobaeocera Pic, 1925: 195 [synonymized by Löbl 1975a: 271]
Type species: Scaphosoma phungi Pic, 1922
Macroscaphosoma Löbl, 1970b: 128 [synonymized by Leschen & Löbl 2006: 25]
Type species: Macroscaphosoma collarti Löbl, 1970
Metalloscapha Löbl, 1975b: 384 [synonymized by Löbl 2002: 182]
Type species: Metalloscapha papua Löbl, 1975
Mimoscaphosoma Pic, 1928: 49 [synonymized by Löbl 1997: 83]
Type species: Scaphosoma bruchi Pic, 1928
Pseudoscaphosoma Pic, 1915b: 31 [synonymized by Löbl 1975a: 270]
Type species: Pseudoscaphosoma testaceomaculatum Pic, 1915
Scaphella Achard, 1924: 29 [synonymized by Löbl 1970a: 730]
Type species: Scaphosoma antennatum Achard, 1920
Scaphiomicrus Casey, 1900: 58 [synonymized by Fall 1910: 119]
Type species: Scaphisoma pusilla LeConte, 1860
Scaphosoma Agassiz, 1846: 332 [synonymized by Löbl 1997: 82]
Scaphosomum Gistel, 1856: 36 [Newton & Thayer 2003]
Scutoscaphosoma Pic, 1916: 3 [synonymized by Löbl 1981: 156]
Type species: Scaphosoma rouyeri Pic, 1916

Leschen & Löbl 2006: 18 (characters in key), 24 (characters).
Key: Matthews 1888: 171 (species of Central America); Fierros-Lopéz 2006: 67 (Mexican species).
Distribution.
West Indies: Cuba.
South America: Colombia, Venezuela, Peru, Brazil, Argentina.
0708. Toxidium LeConte, 1860: 324 (Fig. 85)

Type species: Toxidium gammaroides LeConte, 1860
Leschen & Löbl 2006: 19 (characters in key).
Key: Matthews 1888: 178 (species of Central America).
Distribution.
Subfamily PIESTINAE Erichson, 1839a

0709. Hypotelus Erichson, 1839a: 31 (Figs. 86–87)
Type species: Hypotelus pusillus Erichson, 1840
Antropiestus Bernhauer, 1917a: 45 [synonymized by Caron et al. 2012: 550]
Type species: Piestus andinus Bernhauer, 1917
Eccoptopiestus Scheerpeltz, 1952a: 295 [synonymized by Caron et al. 2012: 550]
Type species: Piestus laevis Solsky, 1872

Characters/Description: Erichson 1840: 840 (characters); Blackwelder 1943: 41 (characters in key and characters);
Navarrete-Heredia et al. 2002: 207 (characters in key); Caron et al. 2012: 550 (characters); Bortoluzzi et al.
Key: Bortoluzzi et al. 2017: 458 (species of the genus).
Distribution.
West Indies: Cuba, Jamaica, Dominican Republic, St. Vincent & Grenadines, Grenada.
South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Bolivia, Brazil.
0710. Piestus Gravenhorst, 1806: 223 (Figs. 88–90)

Type species: Piestus sulcatus Gravenhorst, 1806
Elytropiestus Scheerpeltz, 1952a: 294 [synonymized by Caron et al. 2012: 500]
Type species: Piestus paradoxus Bernhauer, 1917
Trachypiestus Scheerpeltz, 1952a: 291 [synonymized by Caron et al. 2012: 500]
Type species: Piestus angularis Fauvel, 1864
Trichocoryne Gray in Griffith & Pidgeon, 1831: 306 [synonymized by Laporte 1835: 126]
Type species: Zirophorus penicillatus Dalman, 1821
Zirophorus Dalman, 1821: 372 [synonymized by Laporte 1835: 126]
Type species: Zirophorus fronticornis Dalman, 1821
Characters/Description: Blackwelder 1943: 41 (characters in key), 43 (characters); Navarrete-Heredia et al. 2002:
207 (characters in key); Caron et al. 2012: 500 (characters); Pérez et al. 2017: 422 (characters).
Key: Blackwelder 1943: 44 (species of the West Indies); Scheerpeltz 1951: 5 (species of Zirophorus); Scheerpeltz
1952a: 282 (subgenera and species); Newton & Peck 2006: 141 (species of the Galapagos Isl.); Caron et al.
2012: 504 (species of the genus); Caron et al. 2013: 129 (species of the genus); Pérez et al. 2017: 422 (species
of the genus).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, Saint-Barthelemy, Guadeloupe, Dominica,
Martinique, St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
Bolivia, Brazil, Paraguay, Uruguay, Argentina.
0711. Siagonium Kirby & Spence, 1815: pl.1 (Fig. 91)

Type species: Siagonium quadricorne Kirby & Spence, 1815

Characters/Description: Navarrete-Heredia et al. 2002: 207 (characters in key).
Distribution.
Subfamily OSORIINAE Erichson, 1839a
Tribe Eleusinini Sharp, 1887a
0712. Eleusima Irmler Irmler, 2017b: 277

Type species: Eleusima platysoma Irmler, 2017

Characters/Description: Irmler 2017b: 277 (characters in key).
Distribution.
0713. Eleusis Laporte, 1835: 131 (Figs. 96–98)

Type species: Eleusis tibialis Laporte, 1835
Chasolium Laporte, 1835: 132 [synonymized by Fauvel 1865: 35]
Type species: Chasolium ernestini Laporte, 1835
Isomalus Erichson, 1839a: 31 [synonymized by Gemminger & Harold 1868: 675]
Type species: Isomalus humilis Erichson, 1840
Lissopiestus Scheerpeltz, 1952a: 295 [synonymized by Caron et al. 2012: 564]
Type species: Isomalus interruptus Erichson, 1840

Characters/Description: Blackwelder 1943: 120 (characters in key), 155 (characters); Navarrete-Heredia et al.
2002: 211 (characters in key); Irmler 2017b: 277 (characters), 278 (characters in key).
Key: Irmler 2017b: 278 (groups of species), 302 (species of genus).
Distribution.
West Indies: Cuba, Trinidad-Tobago.
Argentina.
0714. Renardia Motschulsky, 1865: 583 (Fig. 114)

Type species: Renardia jubilaea Motschulsky, 1865
Eumalus Sharp, 1887a: 732 [synonymized by Hammond 1970: 70]
Type species: Eumalus nigriceps Sharp, 1887

Characters/Description: Navarrete-Heredia et al. 2002: 211 (characters in key); Irmler 2017b: 278 (characters in
key).
Distribution.
0715. Zeoleusis Steel, 1950a: 215

Type species: Eleusis virgula Fauvel, 1889

Distribution.
Other regions: Oriental, Oceanic, Australian.
Tribe Leptochirini Sharp, 1887a
0716. Leptochirus Germar, 1823: 35 (Figs. 103–104)

Type species: Leptochirus scoriaceus Germar, 1823
Subgenera:
Leptochirus Germar, 1823: 35 [*]
Type species: Leptochirus scoriaceus Germar, 1823
Mesochirus Bernhauer, 1903a: 120 [*]
Type species: Cucujus maxillosus Fabricius, 1801
Strongylochirus Bernhauer, 1903a: 120
Type species: Leptochirus laevis Laporte, 1840
Tropiochirus Bernhauer, 1903a: 120 [*]
Type species: Leptochirus proteus Fauvel, 1864

Characters/Description: Bernhauer 1903a: 117 (characters); Blackwelder 1943: 120 (characters in key), 162
(characters); Navarrete-Heredia et al. 2002: 212 (characters in key); Wu & Zhou 2007: 81 (characters in key);
Asenjo & Ribeiro-Costa 2013: 1258 (characters as subgenus Tropiochirus).
Key: Bernhauer 1903a: 120 (subgenera and species); Asenjo & Ribeiro-Costa 2013: 1283 (species of the subgenus
Tropiochirus).
Distribution.
South America: Colombia, Venezuela, Guyana, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay,
Argentina.
0717. Plastus Bernhauer, 1903a: 142

Type species: Leptochirus convexus Laporte, 1835
Subgenera:
Barychirus Greenslade, 1971: 135
Type species: Priochirus colossalis Bernhauer, 1927
Eutriacanthus Jakobson, 1908: 466
Type species: Leptochirus unicolor Laporte, 1835
Triacanthochirus Bernhauer, 1923: 63 [synonymized by Greenslade, 1971: 137]
Triacanthus Bernhauer, 1903: 136, 158 [synonymized by Bernhauer, 1923b: 63]
Exochirus Greenslade, 1971: 139
Type species: Leptochirus beccarii Fauvel, 1878
Leiochirus Greenslade, 1971: 134
Type species: Priochirus latifrons Greenslade, 1971
Plastus Bernhauer, 1903: 142
Type species: Leptochirus convexus Laporte, 1835
Sinumandibulus Wu & Zhou, 2007: 81
Type species: Priochirus japonicus Sharp, 1889
Stigmatochirus Bernhauer, 1903: 141, 159
Type species: Priochirus dohrni Fauvel, 1902
Syncampsochirus Bernhauer, 1903: 141, 159 [*]
Type species: Leptochirus samoensis Blanchard, 1853

Characters/Description: Wu & Zhou 2007: 81 (characters, characters in key).
Distribution.
Other regions: Palaearctic, Madagascan, Oriental, Australian, Oceanic.
0718. Priochirus Sharp, 1887a: 740 (Fig. 112)

Type species: Priochirus haemorrhous Sharp, 1887
Subgenera:
Cephalomerus Bernhauer, 1903a: 139, 159
Type species: Leptochirus pygmaeus Kraatz, 1859
Euleptarthrus Jakobson, 1908: 466
Type species: Leptochirus longicornis Fauvel, 1864
Leptarthrus Bernhauer, 1903a: 141, 159 [synonymized by Scheerpeltz 1933: 1004]
Neoleptarthrus Scheerpeltz, 1933: 1004 [synonymized by Greenslade 1971: 132]
Priochirus Sharp, 1887a: 740 [*]
Type species: Priochirus haemorrhous Sharp, 1887
Paraborolinus Nakane & Sawada, 1960: 121
Type species: Borolinus bicornis Nakane & Sawada, 1960
Peucodontus Bernhauer, 1903a: 140, 159
Type species: Leptochirus mandibularis Kraatz, 1859
Characters/Description: Bernhauer 1903a: 135 (characters); Navarrete-Heredia et al. 2002: 212 (characters in key);
Wu & Zhou 2007: 81 (characters in key), 89 (characters).
Key: Bernhauer 1903a: 136 (subgenera and species of the genus); Wu & Zhou 2007: 81 (subgenera of the genus).
Distribution.
Central America: Mexico, Guatemala, Honduras, Belize, Costa Rica, Panama.
Tribe Osoriini Erichson, 1839a
Subtribe Osoriina Erichson, 1839a

0719. Anancosorius Bernhauer, 1908: 292
Type species: Anancosorius klimschi Bernhauer, 1908

Characters/Description: Irmler 2010b: 368 (characters in key); Irmler 2015a: 177 (characters in key).
Distribution.
0720. Antillosorius Irmler, 2010b: 366

Type species: Osorius salutator Darlington, 1937
Characters/Description: Irmler 2010b: 368 (characters in key); Irmler 2015a: 177 (characters in key).
Key: Irmler 2015b: 158 (Cuban species).
Distribution.
West Indies: Bahamas, Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico.
0721. Craspedus Bernhauer, 1908: 296

Type species: Craspedus iheringi Bernhauer, 1908
key); Irmler 2015a: 176 (characters in key).
Distribution.
0722. Holotrochopsis Fagel, 1955b: 58

Type species: Holotrochus hulstaerti Bernhauer, 1935

Characters/Description: Irmler 2015a: 174 (characters), 176 (characters in key).
Key: Irmler 2015a: 187 (species of the genus).
Distribution.
South America: Guyana, Peru, Bolivia, Brazil.
0723. Holotrochus Erichson, 1839a: 30 (Figs. 101–102)

Type species: Holotrochus volvulus Erichson, 1840
Neotrochus Blackwelder, 1943: 164 [synonymized by Irmler 1982b: 379]
Type species: Holotrochus cylindrus Erichson, 1840
Subgenera:
Holotrochus Erichson, 1839a: 30 [*]
Type species: Holotrochus volvulus Erichson, 1840
Scapulotrochus Coiffait, 1979: 26
Type species: Holotrochus scapularis Coiffait, 1979
Characters/Description: Erichson 1840: 757 (characters); Blackwelder 1943: 119 (characters in key), 119
(characters in key, Neotrochus), 169 (characters), 164 (characters as Neotrochus); Irmler 1987: 81 (characters
in key); Newton & Peck 2006: 142 (characters in key); Irmler 2010b: 368 (characters in key); Navarrete-
Heredia et al. 2002: 212 (characters in key); Irmler 2015a: 176 (characters in key).
Key: Blackwelder 1943: 170 (species of the West Indies), 165 (species of Neotrochus of the West Indies); Irmler
1982b: 380 (Neotropical species); Irmler 1987: 81 (Neotropical species); Irmler 2005c: 11 (species of
Holotrochus neotropicus group), 17 (species of Holotrochus brasiliensis group [actually in the genus
Paratorchus McColl]); Newton & Peck 2006: 144 (species of the Galapagos Isl.); Irmler 2007b: 136 (species
of Holotrochus syntheticus group); Irmler 2010c: 51 (species of Holotrochus simplex group); Irmler 2013:
161 (species of Holotrochus pubescens group); Irmler 2015b: 158 (Cuban species).
Distribution.
West Indies: Bahamas, Cuba, Cayman Is., Jamaica, Puerto Rico, St. Vincent & Grenadines, Grenada, Trinidad-
Tobago.
Bolivia, Brazil, Paraguay, Argentina.
0724. Mimogonia Coiffait, 1978b: 177

Type species: Mimogonia europaea Coiffait, 1978

Characters/Description: Irmler 1981a: 144 (characters in key); Irmler 2010b: 368 (characters in key); Irmler 2015a:
Key: Irmler 1981a: 144; Irmler 2005d: 54 (species of the genus); Irmler 2007c: 63 (species of the genus); Irmler
2010a: 492 (species with large prominent eyes); Irmler 2013: 160 (species of Brazil).
Distribution.
South America: French Guiana, Peru, Bolivia, Brazil, Paraguay.
Other regions: Palaearctic [in doubt].
0725. Mimogonus Fauvel, 1903: 261

Type species: Osorius fumator Fauvel, 1889

2002: 212 (characters in key); Irmler 2010b: 368 (characters in key); Irmler 2015a: 176 (characters in key).
Key: Irmler 1981a: 144 (some species of the genus); Irmler 2010a: 487 (Neotropical species); Irmler 2015b: 158
(Cuban species).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, Guadeloupe.
Other regions: Palaearctic, Ethiopian, Madagascan, Oriental, Oceanic.
0726. Mimotrochus Irmler, 1987: 107

Type species: Mimotrochus pecki Irmler, 1987

Characters/Description: Irmler 1987: 81 (characters in key); Irmler 2010b: 368 (characters in key); Irmler 2015a:
Key: Irmler 1987: 81 (species of the genus).
Distribution.
0727. Molosoma Say, 1831: 48 (Fig. 106)

Type species: Oxytelus latipes Gravenhorst, 1806
Osoriellus Fagel, 1959: 186 [synonymized by Irmler 2016b: 139]
Type species: Osorius linearis Bernhauer, 1915

Characters/Description: Irmler 2003a: 7 (characters); Irmler 2010b: 368 (characters in key); Irmler 2015a: 177
Key: Irmler 2014: 233 (species group of the genus); Irmler 2014: 235 (species of Osoriellus oculatus group), 236
(species of Osoriellus laeviusculus group), 244 (species of Osoriellus bicornis group), 248 (species of
Osoriellus hirtulus group), 250 (species of Osoriellus granarius group), 256 (species of Osoriellus guiananus
group), 291 (species of Osoriellus osoriellus group), 308 (some species); Irmler 2015b: 158 (Cuban species);
Irmler 2016b: 144 (some species of French Guiana).
Distribution.
West Indies: Bahamas, Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl.,
Montserrat, St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil,
Paraguay, Uruguay, Argentina.
Other regions: Nearctic, Ethiopian.
0728. Neorhabdopsis Irmler, 2015a: 174

Type species: Neorhabdopsis lutzei Irmler, 2015
Characters/Description: Irmler 2015a: 177 (characters in key).
Distribution.
0729. Neosorius Fagel, 1959: 119

Type species: Osorius burgeoni Cameron, 1928
Characters/Description: Irmler 2003a: 8 (characters); Irmler 2010b: 368 (characters in key); Irmler 2015a: 175
Distribution.
Other regions: Ethiopian, Madagascan.
0730. Oryssomma Notman, 1925: 4
Type species: Oryssomma schwarzi Notman, 1925

Key: Irmler 2015a: 182 (species).
Distribution.
South America: Venezuela, French Guiana, Ecuador, Peru, Brazil.
0731. Osoriosetus Irmler, 2015a: 175

Type species: Osoriosetus hanagarthi Irmler, 2015

Distribution.
0732. Osorius Guérin-Méneville, 1829: pl. 9 (Fig. 110)

Type species: Osorius brasiliensis Guérin-Ménéville, 1829

Characters/Description: Lynch-Arribálzaga 1884: 345 (characters); Notman 1925: 6 (characters); Blackwelder
1943: 120 (characters in key), 174 (characters); Navarrete-Heredia et al. 2002: 212 (characters in key);
Newton & Peck 2006: 142 (characters in key); Irmler 2010b: 368 (characters in key), 370 (characters); Irmler
Key: Bernhauer 1933c: 332 (species of the Argentina); Darlington 1937: 285 (species of the West Indies);
Blackwelder 1943: 175 (species of the West Indies); Irmler 2010b: 370 (Neotropical species).
Distribution.
0733. Ouloglene Notman, 1925: 3

Type species: Ouloglene barberi Notman, 1925
Distribution.
0734. Paratorchus McColl, 1985: 355

Type species: Paratrochus hamatus McColl, 1982

Paratrochus McColl, 1982: 31 [synonymized by McColl 1985: 355]
Type species: Paratrochus hamatus McColl, 1982
Key: Irmler 2005c: 17 (some species as Holotrochus brasiliensis group); Irmler 2015a: 185 (species of the genus).
Distribution.
South America: Colombia, Venezuela, Ecuador, Bolivia, Brazil, Chile.
0735. Tavakilianidia Orousset, 1985: 60

Type species: Tavakilianidia anophtalma Orousset, 1985

Distribution.
0736. Tumboecus Fagel, 1959: 62

Type species: Tumboecus nitidus Fagel, 1959

Distribution.
South America: French Guiana, Peru.
Tribe Thoracophorini Reitter, 1909
Subtribe Clavilispinina Newton & Thayer, 1992
0737. Allotrochus Fagel, 1955b: 78 (Fig. 92)

Type species: Holotrochus curticollis Fauvel, 1898

Characters/Description: Navarrete-Heredia et al. 2002: 213 (characters in key); Newton 2017: 23 (characters).
Key: Irmler 2000a: 247 (Neotropical species); Naomi & Irmler 2012: 106 (species of the genus); Irmler 2015b: 158
(Cuban species).
Distribution.
West Indies: Cuba.
South America: Venezuela, Ecuador, Peru, Brazil.
Other regions: Ethiopian, Oriental, Australian, Oceanic, Madagascan.
0738. Clavilispinus Bernhauer, 1926: 255 (Fig. 94)

Type species: Paralispinus siargaoanus Bernhauer, 1926
Ancaeus Fauvel, 1864: 56 [synonymized by Cameron 1930: 42]
Type species: Ancaeus megacephalus Fauvel, 1864
Neolispinodes Bernhauer, 1937: 579 [synonymized by Herman 2001d: 1232]
Paralispinus Bernhauer, 1921a: 67 [synonymized by Blackwelder 1952: 291]

Characters/Description: Bernhauer 1921a: 67 (characters); Blackwelder 1943: 120 (characters in key as
Paralispinus), 156 (characters as Paralispinus); Navarrete-Heredia et al. 2002: 213 (characters in key);
Newton & Peck 2006: 142 (characters in key).
Key: Blackwelder 1943: 157 (species of the West Indies as Paralispinus); Irmler 1991: 85 (Neotropical species);
Irmler 2003b: 350 (Neotropical species of Clavilispinus exiguus group); Irmler 2015b: 158 (Cuban species).
Distribution.
West Indies: Cuba, Jamaica, Dominican Republic, Puerto Rico, US Virgin Isl., Guadeloupe, St. Lucia, St.
Vincent & Grenadines, Grenada, Trinidad-Tobago, Barbados.
Bolivia, Brazil, Paraguay.
0739. Geotrochopsis Irmler, 2016a: 109
Type species: Geotrochopsis pubescens Irmler, 2016
Key: Irmler, 2016a: 114 (key to species).
Distribution.
Subtribe Glyptomina Newton & Thayer, 1992
0740. Arborilispinus Irmler, 2010d: 230

Type species: Arborilispinus longulus Irmler, 2010

Distribution.
0741. Espeson L.W. Schaufuss, 1882a: 168

Type species: Espeson moratus L.W. Schaufuss, 1882

2002: 213 (characters in key); Newton & Peck 2006: 142 (characters in key); Irmler 2012b: 333 (characters in
key).
Key: Blackwelder 1943: 144 (species of the West Indies); Scheerpeltz 1970b: 116 (species of the genus); Irmler
2012b: 333 (Neotropical species of the genus); Irmler 2015b: 158 (Cuban species).
Distribution.

West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Montserrat, Guadeloupe,
Dominica, Martinique, St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
Central America: Mexico, Guatemala, El Salvador, Nicaragua, Costa Rica, Panama.
South America: Colombia, Venezuela, Guyana, French Guiana, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil,
Paraguay, Argentina.
Other regions: Nearctic, Ethiopian, Madagascan, Oriental, Australian.
0742. Geomitopsis Scheerpeltz, 1931: 388

Type species: Cylindropsis zariquieyi Dodero, 1918
Aschnaosorius Makhan, 2008: 1 [synonymized by Irmler 2016a: 114]
Type species: Cubanotyphlus rishwani Makhan, 2007
Subgenera:
Geomitopsis Scheerpeltz, 1931: 388 [*]
Type species: Cylindropsis zariquieyi Dodero, 1918
Libanotyphlus Coiffait in Vandel et al., 1954: 155 [synonymized by Coiffait 1959: 218]
Type species: Libanotyphlus dubertreti Coiffait, 1954
Pseudogeomitopsis Orousset, 1985: 63 [*]
Type species: Geomitopsis remilleti Orousset, 1985

Characters/Description: Coiffait & Sáiz 1965c: 577 (characters); Orousset 1985: 63 (characters); Navarrete-
Heredia et al. 2002: 213 (characters in key); Irmler 2016a: 114 (characters).
Key: Sáiz 1973b: 21 (Chilean species); Orousset 1985: 64 (subgenera); Irmler 2016a: 115 (species of the
Neotropical region).
Distribution.
South America: Suriname, French Guiana, Peru, Brazil, Chile.
0743. Glyptoma Erichson, 1839a: 32 (Fig. 100)

Type species: Glyptoma crassicorne Erichson, 1840
Calocerus Fauvel, 1891: 88 [synonymized by Blackwelder 1943: 141]
Type species: Glyptoma crassicorne Erichson, 1840

Characters/Description: Erichson 1840: 908 (characters); Blackwelder 1943: 120 (characters in key), 141
(characters); Navarrete-Heredia et al. 2002: 213 (characters in key); Irmler 2015c: 298 (characters).
Key: Blackwelder 1943: 142 (species of the West Indies); Scheerpeltz 1970c: 130 (species of Calocerus).
Distribution.
West Indies: Guadeloupe, Trinidad-Tobago.
Central America: Mexico, Guatemala, Belize, El Salvador, Nicaragua, Costa Rica, Panama.
0744. Lispinomimus Irmler, 2003a: 11

Type species: Lispinomimus atavus Irmler, 2003
1 species known only from Dominican amber.

Distribution.
0745. Pseudespeson Lecoq, 1994: 299

Type species: Pseudespeson rossii Lecoq, 1994
Characters/Description: Irmler 2012b: 333 (characters in key).
Key: Irmler 2012b: 333 (Neotropical species of the genus).
Distribution.
West Indies: US Virgin Isl., Montserrat, Guadeloupe, Martinique, St. Lucia, St. Vincent & Grenadines, Grenada,
Trinidad-Tobago.
South America: Venezuela, Peru, Brazil.
Other regions: Palaearctic, Ethiopian, Madagascan, Oriental.
Subtribe Lispinina Bernhauer & Schubert, 1910
0746. Liberiana Blackwelder, 1942: 82 (Fig. 105)

Type species: Pseudolispinodes femoralis Blackwelder, 1942

Characters/Description: Irmler 2003c: 86 (characters in key); Irmler 2003a: 5 (characters).
Distribution.
0747. Lispinuncus Irmler, 2005b: 173

Type species: Lispinuncus pulcher Irmler, 2005

Distribution.
0748. Lispinus Erichson, 1839a: 31

Type species: Lispinus attenuatus Erichson, 1840
Subgenera:
Aulacolispinus Scheerpeltz, 1957a: 223, 239, 240 [*]
Type species: Lispinus unistriatus Fauvel, 1878
Lispinus Erichson, 1839a: 31 [*]
Type species: Lispinus attenuatus Erichson, 1840
Paralispinus Eichelbaum, 1913: 117
Type species: Lispinus amaniensis Eichelbaum, 1913
Pseudolispinodes Bernhauer, 1926: 258 [*]
Type species: Holosus madurensis Bernhauer, 1914
Spinilus Blackwelder, 1942: 83 [synonymized by Blackwelder 1952: 223]
Type species: Lispinodes bistriatus Fauvel, 1895
Stictolispinus Scheerpeltz, 1957a: 241 [*]

Type species: Lispinus quadripunctulus Fauvel, 1864

Characters/Description: Erichson 1840: 828 (characters); Blackwelder 1942: 83 (characters); Blackwelder 1943:
120 (characters in key), 120 (characters in key of subgenus Pseudolispinodes), 129 (characters); Irmler 2003c:
86 (characters in key); Navarrete-Heredia et al. 2002: 212 (characters in key); Newton & Peck 2006: 142
Key: Blackwelder 1942: 77 (subgenera); Blackwelder 1943: 130 (species of the West Indies); Irmler 1994: 54
(Neotropical species); Irmler 2000b: 82 (some Neotropical species); Irmler 2001a: 9, 10 (some Neotropical
species); Irmler 2006a: 3 (species of Costa Rica); Irmler 2009: 55 (Peruvian species); Irmler 2012a: 223
(species of Ecuador); Irmler 2015b: 158 (Cuban species).
Distribution.
West Indies: Bahamas, Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Dominica,
Montserrat, Guadeloupe, St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
0749. Nacaeus Blackwelder, 1942: 82 (Figs. 107–108)

Type species: Lispinus planellus Sharp, 1887
Subgenera:
Liberiana Blackwelder, 1942: 82
Type species: Pseudolispinodes femoralis Blackwelder, 1942
Liberiella Blackwelder, 1942: 81
Type species: Pseudolispinodes cooki Blackwelder, 1942
Nacaeus Blackwelder, 1942: 82 [*]
Type species: Lispinus planellus Sharp, 1887
Rumeba Blackwelder, 1942: 82
Type species: Pseudolispinodes lispinoides Blackwelder, 1942
Characters/Description: Navarrete-Heredia et al. 2002: 212 (characters in key); Irmler 2003a: 4 (characters); Irmler
2003c: 86 (characters in key); Newton & Peck 2006: 142 (characters in key).
Key: Irmler 2003c: 98 (Neotropical species); Irmler 2006b: 212 (species similar to Nacaeus sculpturatus), 214
(species with light red or yellow elytra); Irmler 2015b: 158 (Cuban species).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, Montserrat, Guadeloupe, St. Lucia, St.
Vincent & Grenadines, Grenada.
0750. Neolosus Blackwelder, 1942: 85 (Fig. 109)

Type species: Holosus tachyporiformis Motschulsky, 1858
Subgenera:
Neolosus Blackwelder, 1942: 85
Type species: Holosus tachyporiformis Motschulsky, 1858
Osholus Blackwelder, 1952: 278 [synonymized by Herman 2001d: 1286]
Type species: Holosus tachiniformis Motschulsky, 1858
Holosus Motschulsky, 1858a: 496 [synonymized by Herman 2001d: 1286]
Type species: Holosus tachiniformis Motschulsky, 1857
Relinda Blackwelder, 1942: 85
Type species: Holosus mycetoporiformis Motschulsky, 1858

Characters/Description: Blackwelder 1942: 84 (characters as Holosus); Irmler 2003c: 86 (characters in key).
Key: Irmler 1999: 268 (South American species); Irmler 2001a: 11 (species of Neolosus zischkai group).
Distribution.
South America: Colombia, Ecuador, Peru, Bolivia, Brazil.
Other regions: Palaearctic, Ethiopian, Oriental, Australian.
0751. Tannea Blackwelder, 1952: 373 (Figs. 115–116)

Type species: Lispinus tenellus Erichson, 1840

Characters/Description: Irmler 2003c: 86 (characters in key); Navarrete-Heredia et al. 2002: 212 (characters in
key).
Key: Irmler 2003c: 86 (Neotropical species); Irmler 2005e: 153 (Tannea amazonica species group), 155 (Tannea
bierigii species group), 156 (Tannea longicornis species group); Irmler 2007a: 72 (species of Costa Rica);
Irmler 2012c: 220 (species of Ecuador); Irmler 2015b: 158 (Cuban species).
Distribution.
St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
Argentina.
Subtribe Thoracophorina Reitter, 1909
0752. Aneucamptus Sharp, 1887a: 725 (Fig. 93)

Type species: Thoracophorus excisicollis Motschulsky, 1860

Characters/Description: Irmler 1985: 41 (characters in key); Navarrete-Heredia et al. 2002: 213 (characters in key).
Key: Irmler 1985: 41 (species of the genus).
Distribution.
0753. Dirocephalus Silvestri, 1938: 251 (Fig. 95)

Type species: Dirocephalus myrmecophilus Silvestri, 1938
Diplopsis Fauvel, 1902a: 33 [synonymized by Blackwelder 1952: 165]

Type species: Diplopsis multicostata Fauvel, 1902
Fauva Blackwelder, 1952: 165 [synonymized by Irmler 2005a: 103]
Type species: Diplopsis multicostata Fauvel, 1902
Characters/Description: Fauvel 1902a: 33 (characters as Fauva); Irmler 2001b: 121 (characters as Fauva);
Navarrete-Heredia et al. 2002: 212 (characters in key); Irmler 2005a: 104 (characters in key).
Key: Irmler 2001b: 126 (species of Fauva); Irmler 2005a: 104 (species of the genus).
Distribution.
South America: Venezuela, Guyana, Ecuador, Peru, Bolivia, Brazil, Paraguay.
0754. Euctenopsia Bruch, 1942: 137 (Fig. 99)

Type species: Euctenopsia ogloblini Bruch, 1942

Distribution.
South America: Paraguay, Argentina.
0755. Mesotrochus Wasmann, 1890: 317

Type species: Mesotrochus paradoxus Wasmann, 1890

Characters/Description: Notman 1925: 3 (characters); Irmler 2003a: 2 (characters).
Distribution.
0756. Pardirocephalus Bruch, 1942: 130 (Fig. 111)

Type species: Pardirocephalus cordobensis Bruch, 1942

Distribution.
0757. Pselaphomimus Bruch, 1942: 134 (Fig. 113)

Type species: Pselaphomimus amphiphilus Bruch, 1942

Distribution.
0758. Quadricephalus Irmler, 2005a: 105

Type species: Quadricephalus obstrusus Irmler, 2005
Distribution.
0759. Rhopalopherus Bernhauer, 1909b: 227

Type species: Rhopalopherus gestroi Bernhauer, 1909

Characters/Description: Irmler 2003a: 2 (characters).
Distribution.
0760. Thoracophorus Motschulsky, 1837: 98 (Figs. 117–118)

Type species: Thoracophorus corticinus Motschulsky, 1837
Thoraxophorus Motschulsky, 1837: 98
Subgenera:
Leipophorus Bernhauer, 1926: 261
Type species: Thoracophorus minutissimus Bernhauer, 1926
Stilbogastrus Bernhauer, 1908: 286 [*]
Type species: Thoracophorus nitidus Bernhauer, 1908
Thoracophorus Motschulsky, 1837: 98 [*]
Type species: Thoracophorus corticinus Motschulsky, 1837
Characters/Description: Blackwelder 1943: 120 (characters in key), 148 (characters); Coiffait & Sáiz 1968: 460
(characters in key), 460 (characters); Irmler 1985: 41 (characters in key); Navarrete-Heredia et al. 2002: 213
(characters in key); Irmler 2003a: 2 (characters); Newton & Peck 2006: 142 (characters in key); Irmler 2015c:
298 (characters); Ferro 2015: 2 (characters).
Key: Blackwelder 1943: 149 (species of the West Indies); Irmler 1985: 41 (species of the genus); Irmler 2015b:
158 (Cuban species); Ferro 2015: 4 (species of America north of Mexico).
Distribution.
West Indies: Bahamas, Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Montserrat,
Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
Bolivia, Brazil, Paraguay, Chile, Argentina.
0761. Verhaaghiella Irmler, 2005a: 105

Type species: Fauva becki Irmler, 2001

Characters/Description: Irmler, 2005a: 104 (characters in key).
Key: Irmler, 2005a: 104 (species).
Distribution.

Subfamily OXYTELINAE Fleming, 1821
Tribe Blediini Ádám, 2001
0762. Bledius Leach in Samouelle, 1819: 174 (Figs. 120–121)

Type species: Staphylinus armatus Panzer, 1799
Microbledius Herman, 1972a: 118 [synonymized by Herman 1986: 295]
Type species: Microbledius playanus Herman, 1972
Subgenera:
Astycops Thomson, 1859: 43
Type species: Oxytelus talpa Gyllenhal, 1810
Bargus Schiødte, 1866: 145, 148
Type species: Oxytelus pallipes Gravenhorst, 1806
Belidus Mulsant & Rey, 1878a: 657
Type species: Bledius angustus Mulsant & Rey, 1861
Bledius Leach in Samouelle, 1819: 174
Type species: Staphylinus armatus Panzer, 1799
Dicarenus Gistel, 1834: 9
Type species: Staphylinus arenarius Paykull, 1800
Cotysops Tottenham, 1939a: 225 [synonymized by Herman 1970a: 375]
Type species: Staphylinus arenarius Paykull, 1800
Elbidus Mulsant & Rey, 1878a: 572 [synonymized by Herman 1970a: 375]
Type species: Oxytelus bicornis Germar, 1823
Neobledius M. Abdullah & Qadri, 1968: 394 [synonymized by Herman 1986: 295]
Type species: Neobledius karachiensis Abdullah & Qadri, 1968
Euceratobledius Znojko, 1929: 203
Type species: Oxytelus furcatus Olivier, 1811
Hesperophilus Stephens, 1829a: 24
Type species: Staphylinus fracticornis Paykull, 1790
Blediodes Mulsant & Rey, 1878a: 576 [synonymized by Herman 1970a: 375]
Tadunus Schiødte, 1866: 144, 147 [synonymized by Gemminger & Harold 1868: 643]
Psamathobledius Herman, 1972a: 136
Type species: Bledius punctatissimus LeConte, 1877
Pucerus Mulsant & Rey, 1878a: 654
Type species: Bledius verres Erichson, 1840
Teratobledius Ádám, 2001: 218
Type species: Bledius tibialis Heer, 1839
71 species are included in this genus from Latin America. It is one of the super-large genera of the subfamily, its
taxonomy is resolved only to species groups, keys to species level mostly missing [György Makranczy,
personal communication].
Characters/Description: Lynch-Arribálzaga 1884: 351 (characters); Blackwelder 1943: 57 (characters in key), 112
(characters); Coiffait & Sáiz 1968: 415 (characters in key), 423 (characters); Herman 1970a: 361 (characters
in key), 375 (characters); Herman 1972a: 153 (characters); Sáiz 1973c: 229 (characters); Navarrete-Heredia et
al. 2002: 230 (characters in key); Makranczy 2006: 103 (characters in key); Newton & Peck 2006: 145
Key: Lynch-Arribálzaga 1884: 353 (species of Buenos Aires province, Argentina); Bernhauer 1927b: 232 (species
of the Argentina); Blackwelder 1943: 113 (species of the West Indies); Coiffait & Sáiz 1968: 424 (Chilean
species); Herman 1972a: 154 (species groups of the genus); Sáiz 1973c: 229 (Chilean species); Herman 1976:
77 (species groups of the genus); Herman 1983: 8 (species groups of the genus); Herman 1986: 107 (species
groups); Newton & Peck 2006: 145 (species of the Galapagos Isl.); Caron & Ribeiro-Costa 2007: 453
(seashore of southern Brazil); Castro et al. 2016: 146 (Brazilian seashore species).
Distribution.
Antigua & Barbuda, Montserrat, Guadeloupe, Dominica, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, French Guiana, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil, Uruguay,
Chile, Argentina.
Tribe Coprophilini Heer, 1839
0763. Homalotrichus Solier, 1849: 321

Type species: Homalotrichus striatus Solier, 1849
Sharpia Fauvel, 1878c: 488 [synonymized by Fauvel 1900b: 189]
Type species: Sharpia banksi Fauvel, 1878
Toxoderus Fauvel, 1900b: 189 [synonymized by Herman 1970a: 368]
Type species: Sharpia banksi Fauvel, 1878
8 species are included in this genus from South America. As with other genera of Coprophilini, most of the species
are lacking strong internal sclerites in the aedeagi and species-specific female genitalia, therefore are very
difficult to separate [György Makranczy, personal communication].
Characters/Description: Coiffait & Sáiz 1968: 415 (characters in key); Herman 1970a: 360 (characters in key), 368
(characters); Makranczy 2006: 103 (characters in key); Pérez et al. 2018: 385 (characters).
Distribution.
South America: Peru, Chile, Argentina.
Tribe Deleasterini Reitter, 1909
0764. Deleaster Erichson, 1839b: 610

Type species: Anthophagus dichrous Gravenhorst, 1802

Characters/Description: Herman 1970a: 360 (characters in key), 363 (characters); Navarrete-Heredia et al. 2002:
229 (characters in key); Makranczy 2006: 102 (characters in key).
Distribution.

Tribe Euphaniini Reitter, 1909
0765. Euphanias Fairmaire & Laboulbène, 1856: 657

Type species: Euphanias insignicornis Fairmaire & Laboulbène, 1856
Pholidus Mulsant & Rey, 1856: 7 [synonymized by Redtenbacher 1857: 991]
Type species: Pholidus insignis Mulsant & Rey, 1856

Characters/Description: Makranczy 2006: 102 (characters in key).
Distribution.
Tribe Oxytelini Fleming, 1821
0766. Anotylus Thomson, 1859: 44

Type species: Oxytelus sculpturatus Gravenhorst, 1806
Anotylops Fagel, 1957b: 8 [synonymized by Hammond 1976: 145]
Type species: Anotylops seydeli Fagel, 1957
Boettcherinus Bernhauer, 1936b: 82 [synonymized by Herman 1970a: 414]
Type species: Oxytelus planaticollis Bernhauer, 1936
Delopsis Fauvel, 1895a: 198 [synonymized by Blackwelder 1952: 342]
Type species: Delopsis cornuta Fauvel, 1895
Emopotylus Bernhauer, 1910: 359 [synonymized by Herman 1970a: 414]
Type species: Oxytelus cuernavacanus Bernhauer, 1910
Metoxytelus Coiffait & Sáiz, 1968: 422 [synonymized by Hammond 1976: 145]
Type species: Oxytelus sulcicollis Gemminger & Harold, 1868
Microxytelus Fagel, 1956: 272 [synonymized by Herman 1970a: 414]
Type species: Oxytelus nitidifrons Wollaston, 1871
Neoplatystethus M. Abdullah & Qadri, 1970: 127 [synonymized by Hammond 1976: 146]
Type species: Neoplatystethus hameedi Abdullah & Qadri, 1970
Neopyctocraerus M. Abdullah & Qadri, 1970: 126 [synonymized by Hammond 1976: 146]
Type species: Neopyctocraerus shafqati Abdullah & Qadri, 1970
Oncoparia Bernhauer in Bernhauer & Jeannel, 1936: 214 [synonymized by Hammond 1976: 145]
Type species: Oncoparia parasita Bernhauer, 1936
Oxytelodes Bernhauer, 1908: 290 [synonymized by Herman 1970a: 414]
Type species: Oxytelodes holdhausi Bernhauer, 1908
Oxytelops Fagel, 1956: 273 [synonymized by Herman 1970a: 414]
Type species: Staphylinus tetracarinatus Block, 1799
Oxytelopsis Fauvel, 1895a: 199 [synonymized by Makranczy 2006: 83]
Type species: Oxytelopsis cimicoides Fauvel, 1895
Oxytelosus Cameron, 1950b: 92 [synonymized by Fagel 1956: 274]
Oxytelosus Fagel, 1956: 274 [synonymized by Herman 1970a: 414]
Type species: Oxytelus abnormalis Cameron, 1938
Paracaccoporus Steel, 1948: 188 [synonymized by Herman 1970a: 414]
Type species: Oxytelus ocularis Fauvel, 1877
Pseudodelopsis Fagel, 1957b: 3 [synonymized by Herman 1970a: 414]
Type species: Pseudodelopsis scotti Fagel, 1957
Pseudoplatystethus M. Abdullah & Qadri, 1970: 129 [synonymized by Hammond 1976: 146]
Type species: Neoplatystethus meccii Abdullah & Qadri, 1970
Pseudopyctocraerus M. Abdullah & Qadri, 1970: 125 [synonymized by Hammond 1976: 146]
Type species: Platystethus mahmoodi Abdullah & Qadri, 1970
Rimba Blackwelder, 1952: 342 [synonymized by Makranczy 2006: 83]
Type species: Delopsis cornuta Fauvel, 1895
Styloxys Gozis, 1886: 15 [synonymized by Blackwelder 1943: 91]
Type species: Staphylinus rugosus Fabricius, 1775
58 species are included in this genus from Latin America. A very large genus, extremely poorly known considering
its great Neotropical diversity; the majority of the species in the region are undescribed [György Makranczy,
personal communication].
Characters/Description: Bernhauer 1908: 290 (characters); Coiffait & Sáiz 1968: 415 (characters in key), 415
(characters in key as Metoxytelus); Herman 1970a: 362 (characters in key), 362 (characters in key,
Oxytelopsis), 414 (characters); Navarrete-Heredia et al. 2002: 230 (characters in key); Makranczy 2006: 105
(characters in key); Aballay et al. 2014: 103 (characters in key).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Antigua &
Barbuda, Montserrat, Guadeloupe, Dominica, St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Bolivia, Brazil, Uruguay, Chile,
Argentina.
0767. Apocellus Erichson, 1839a: 30 (Fig. 119)

Type species: Lathrobium sphaericollis Say, 1831
Ocaleomorpha Fleischer, 1921b: 114 [synonymized by Fleischer 1921a: 181]
Type species: Ocaleomorpha lacoi Fleischer, 1921
Pheidoloxenides Wasmann, 1925a: 8 [synonymized by Herman 1970a: 411]
Type species: Pheidoloxenides dampfi Wasmann, 1925
Pheidoloxenides Wasmann, 1925b: 122 [synonymized by Herman 1970a: 411]
Type species: Pheidoloxenides dampfi Wasmann, 1925
36 species are included in this genus from Latin America. A medium-sized New World genus most frequently
mentioned because of its myrmecoid appearance and possible myrmecophily; also well-known for multiple
introductions to Europe with soil [György Makranczy, personal communication].
Characters/Description: Lynch-Arribálzaga 1884: 373 (characters); Erichson 1840: 812 (characters); Blackwelder
1943: 57 (characters in key), 88 (characters); Coiffait & Sáiz 1968: 415 (characters in key), 447 (characters);
Herman 1970a: 362 (characters in key), 411 (characters); Navarrete-Heredia et al. 2002: 230 (characters in
key); Makranczy 2006: 105 (characters in key); Newton & Peck 2006: 145 (characters in key).
Key: Blackwelder 1943: 89 (species of the West Indies); Chani-Posse & Scheibler 2013: 198 (species of the
southwestern of the South America).
Distribution.

West Indies: Cayman Is., Puerto Rico, US Virgin Isl., Montserrat, Guadeloupe, Dominica, Martinique, Grenada,
Trinidad-Tobago.
South America: Colombia, Venezuela, Bolivia, Brazil, Paraguay, Chile, Argentina.
0768. Ecitoclimax Borgmeier, 1934: 452

Type species: Ecitoclimax tarsalis Borgmeier, 1934
Only one species is known from and is restricted to South America. Formerly monotypical genus, with a recently
discovered second species (yet undescribed). Myrmecophilous habits possible [György Makranczy, personal
communication].
Characters/Description: Seevers 1965: 339 (characters); Herman 1970a: 362 (characters in key), 403 (characters);
Makranczy 2006: 105 (characters in key).
Distribution.
0769. Oxytelus Gravenhorst, 1802: 101 (Fig. 122)

Type species: Staphylinus piceus Linnaeus, 1767
Anisopsidius Fagel, 1960: 8 [synonymized by Makranczy 2006: 82]
Type species: Anisopsis quadricollis Bernhauer, 1932
Anisopsis Fauvel, 1904a: 108 [synonymized by Makranczy 2006: 82]
Type species: Anisopsis flexuosa Fauvel, 1904
Anisopsodes Fagel, 1960: 11 [synonymized by Herman 1970a: 404]
Type species: Anisopsodes ornatus Fagel, 1960
Hoplitodes Fauvel, 1904a: 109 [synonymized by Makranczy 2006: 82]
Type species: Hoplitodes echidne Fauvel, 1904
Paranisopsis Cameron, 1938: 3 [synonymized by Herman 1970a: 404]
Type species: Paranisopsis dorylina Cameron, 1938
Paroxytelopsis Cameron, 1933b: 36 [synonymized by Makranczy 2006: 82]
Type species: Paroxytelopsis dorylinus Cameron, 1933
Subgenera:
Epomotylus Thomson, 1859: 43
Type species: Oxytelus sculptus Gravenhorst, 1806
Oxytelus Gravenhorst, 1802: 101
Basilewskyorus Fagel, 1957a: 41 [synonymized by Herman 1970a: 408]
Type species: Oxytelus rugegensis Cameron, 1956
Caccoporus Thomson, 1859: 43 [synonymized by Gemminger & Harold 1868: 648]
Tanycraerus Thomson, 1859: 43
Type species: Oxytelus luteipennis Erichson, 1839
12 species are included in this genus from Latin America. A mostly Old World genus with a small diversity in the
Neotropics [György Makranczy, personal communication].
(characters); Coiffait & Sáiz 1968: 415 (characters in key of subgenus Epomotylus); Herman 1970a: 362
(characters in key), 408 (characters); Navarrete-Heredia et al. 2002: 230 (characters in key); Makranczy 2006:
105 (characters in key); Newton & Peck 2006: 145 (characters in key).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Saint-Kitts &
Nevis, Antigua & Barbuda, Montserrat, Guadeloupe, St. Lucia, St. Vincent & Grenadines, Barbados,
Grenada, Trinidad-Tobago.
South America: Colombia, Guyana, Galapagos Isl., Peru, Bolivia, Brazil, Chile, Argentina.
0770. Platystethus Mannerheim, 1830: 46 (Fig. 124)

Type species: Staphylinus morsitans Paykull, 1792
Subgenera:
Craetopycrus Tottenham, 1939a: 225
Type species: Oxytelus cornutus Gravenhorst, 1802
Platystethus Mannerheim, 1830: 46
Pyctocraerus Thomson, 1859: 43 [synonymized by Fauvel 1872: 178]

(characters); Herman 1970a: 361 (characters in key), 401 (characters); Navarrete-Heredia et al. 2002: 230
(characters in key); Makranczy 2006: 105 (characters in key); Newton & Peck 2006: 145 (characters in key).
Key: Lynch-Arribálzaga 1884: 359 (species of Buenos Aires province, Argentina).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Antigua & Barbuda,
Guadeloupe, Dominica, St. Lucia, St. Vincent & Grenadines, Barbados, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil, Uruguay, Argentina.
Tribe Thinobiini J. Sahlberg, 1876
0771. Carpelimus Leach in Samouelle, 1819: 174

Type species: Oxytelus fuliginosus Gravenhorst, 1802
Batychrus Gistel, 1834: 9 [synonymized by Blackwelder 1952: 74]
Type species: Oxytelus corticinus Gravenhorst, 1806
Boopinus Klima, 1904: 43, 46, 56 [synonymized by Bernhauer & Schubert 1911: 97]
Type species: Trogophloeus memnonius Erichson, 1840
Bucephalinus C. Koch, 1934: 42 [synonymized by Blackwelder 1952: 87]
Bucephalinus Blackwelder, 1952: 87 [synonymized by Blackwelder 1952: 87]
Type species: Trogophloeus priesneri C. Koch, 1934
Glomus Gistel, 1848: xi [synonymized by Blackwelder 1952: 170]
Type species: Aleochara pusilla Gravenhorst, 1802
Myopinus Blackwelder, 1952: 253 [synonymized by Herman 1970a: 390]

Type species: Trogophloeus elongatulus Erichson, 1839
Myopinus Scheerpeltz, 1937b: 116 [synonymized by Blackwelder 1952: 253]
Nanolobus Cameron, 1933a: 74 [synonymized by Herman 1970a: 389]
Type species: Nanolobus pacificus Cameron, 1933
Oxytrogus Wendeler, 1930: 183 [synonymized by Herman 1970a: 389]
Type species: Oxytrogus oculatus Wendeler, 1930
Paraboopinus Scheerpeltz, 1937b: 109 [synonymized by Herman 1970a: 389]
Type species: Trogophloeus nitidus Baudi di Selve, 1848
Paratrogophloeus Hatch, 1957: 91 [synonymized by Herman 1970a: 390]
Type species: Carpelimus bilineatus Stephens 1834
Taenosoma Mannerheim, 1830: 10, 50 [synonymized by Lacordaire 1854: 118]
Type species: Aleochara pusilla Gravenhorst, 1802
Thoracoplatynus Scheerpeltz, 1937b: 109 [synonymized by Herman 1970a: 390]
Troginus Mulsant & Rey, 1878a: 758 [synonymized by Scheerpeltz 1933: 1089]
Type species: Trogophloeus exiguus Erichson, 1839
Trogophloeus Mannerheim, 1830: 9, 49 [synonymized by Tottenham 1939b: 227]
Type species: Oxytelus corticinus Gravenhorst, 1806
Subgenera:
Carpelimus Leach in Samouelle, 1819: 174 [*]
Stenoderophloeus Scheerpeltz, 1972d: 66 [*]
Type species: Trogophloeus guttifer Scheerpeltz, 1972
107 species are included in this genus from Latin America. A very large, Cosmopolitan genus with hundreds of
species in the Neotropics; lacking any recent work. In the total absence of illustrations the species cannot be
identified. A recent catalogue (Herman, 2001d) misplaced a high number of Thinodromus species under
Carpelimus because of the misinterpretation of the name Paracarpalimus Scheerpeltz 1937b (see also under
Thinodromus) [György Makranczy, personal communication].
Characters/Description: Lynch-Arribálzaga 1884: 368 (characters of Trogophloeus); Blackwelder 1943: 57
(characters in key), 58 (characters); Coiffait & Sáiz 1968: 415 (characters in key of Trogophloeus), 429
(characters of Trogophloeus); Herman 1970a: 361, 362 (characters in key), 389 (characters); Navarrete-
Heredia et al. 2002: 230 (characters in key); Makranczy 2006: 63 (characters), 105 (characters in key);
Newton & Peck 2006: 145 (characters in key); Makranczy 2018: 75 (characters of genus).
Key: Lynch-Arribálzaga 1884: 369 (species of Buenos Aires province, Argentina); Blackwelder 1943: 59 (species
of the West Indies); Coiffait & Sáiz 1968: 430 (Chilean species of Trogophloeus); Newton & Peck 2006: 145
(species of the Galapagos Isl.).
Distribution.
Antigua & Barbuda, Guadeloupe, Martinique, St. Lucia, St. Vincent & Grenadines, Barbados, Grenada,
Trinidad-Tobago.
South America: Ecuador, Galapagos Isl., Peru, Bolivia, Brazil, Paraguay, Uruguay, Chile, Argentina.
0772. Crassodemus Herman, 1968: 1

Type species: Crassodemus foraminosus Herman, 1968
Only one species is known from and is restricted to South America. A formerly monotypical genus with a few,
recently discovered (still undescribed) species. Appears to be widely distributed in the tropical parts of South
America [György Makranczy, personal communication].
Characters/Description: Herman 1970a: 360 (characters in key), 372 (characters); Makranczy 2006: 104
Distribution.
0773. Neoxus Herman, 1970a: 394

Type species: Thinobius crassicornis Casey, 1889
2 species are included in this genus from Central America. A widely distributed but small genus, containing several
undescribed species. Similar habits to those of Thinobius; because of its small size and fragile body rarely
turning up in collections [György Makranczy, personal communication].
Characters/Description: Herman 1970a: 361 (characters in key); Navarrete-Heredia et al. 2002: 230 (characters in
key); Makranczy 2006: 105 (characters in key).
Distribution.
0774. Parosus Sharp, 1887a: 704 (Fig. 123)

Type species: Parosus hilaris Sharp, 1887
21 species are known from and are restricted to Latin America. The only Neotropical oxyteline genus with a recent
revision (Makranczy 2014). The expected diversity is even greater than documented there, and may rise to
about 40 species. Biology became better known recently, with discovery of the possible larva collected with
adults in bracts of Gesneriaceae [György Makranczy, personal communication].
Characters/Description: Blackwelder 1943: 57 (characters in key), 103 (characters); Herman 1970a: 361, 362
(characters in key), 399 (characters); Makranczy 2006: 105 (characters in key); Makranczy 2014: 83
(characters).
Key: Blackwelder 1943: 103 (species of the West Indies); Makranczy 2014: 83 (species of the genus); López-
García & Marín-Gómez 2018: 564 (Colombian species).
Distribution.
West Indies: Guadeloupe, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil.
0775. Sciotrogus Sharp, 1887a: 707 (Fig. 125)

Type species: Sciotrogus opacus Sharp, 1887
2 species are known from and are restricted to the West Indies and Central America. Genus with doubtful status
currently containing only 2 species, difficult separation from Thinobius, may be synonymous with that
[György Makranczy, personal communication].
Characters/Description: Herman 1970a: 361 (characters in key), 395 (characters); Navarrete-Heredia et al. 2002:
229 (characters in key); Makranczy 2006: 105 (characters in key).
Distribution.

0776. Teropalpus Solier, 1849: 330
Type species: Teropalpus suturalis Solier, 1849
Nesoteropalpus Coiffait & Saiz, 1968: 432 [Nomem nudum, designated by Herman 2001d: 1734]
Type species: Trogophloeus skottsbergii Bernhauer, 1921
Trogolinus Sharp, 1900: 231 [synonymized by Fauvel 1900b: 182]
Type species: Trogophloeus unicolor Sharp, 1900
4 species are included in this genus from South America. Small genus known for its introduction to several
Holarctic locations [György Makranczy, personal communication].
Characters/Description: Herman 1970a: 361 (characters in key), 381 (characters); Makranczy 2006: 53
Distribution.
0777. Thinobius Kiesenwetter, 1844: 355

Type species: Thinobius ciliatus Kiesenwetter, 1844
Subgenera:
Bracharthrothinophilus Scheerpeltz, 1959a: 60
Type species: Thinobius nitens Fauvel, 1871
Oedarthrothinophilus Scheerpeltz, 1959a: 60
Type species: Thinobius nodicornis Eppelsheim, 1884
Platyderothinophilus Scheerpeltz, 1959a: 58
Type species: Thinobius major Kraatz, 1857
Myopothinophilus Scheerpeltz, 1959a: 58 [synonymized by Schülke & Smetana 2015: 790]
Type species: Thinobius klimai Bernhauer, 1902
Thinobius Kiesenwetter, 1844: 355
Type species: Thinobius ciliatus Kiesenwetter, 1844
Thinobiellus Bernhauer, 1909a: 198 [synonymized by Schülke 2009: 829]
Type species: Thinobius rossicus Bernhauer, 1909
Torrentomus Bierig, 1934b: 213 [synonymized by Herman 1970a: 395]
Type species: Torrentomus torrei Bierig, 1934
Yosiityphlus Sawada, 1971b: 327 [synonymized by Gusarov & Makranczy 2004: 1]
Type species: Yosiityphlus kuroshio Sawada, 1971
Thiphonilus Tottenham, 1939a: 225
Type species: Thinobius linearis Kraatz, 1857
Thinophilus Mulsant & Rey, 1878a: 764 [synonymized by Tottenham 1939a: 225]
Type species: Thinophilus linearis Kraatz, 1857
25 species are included in this genus from Latin America. Rather large and widespread genus but with very poorly
resolved taxonomy and a few conspicuous forms in the Neotropics. Most of the species are undescribed and
because of the small size and fragile body are rarely preserved in collections, but eventually the species count
in the New World could rise to a hundred [György Makranczy, personal communication].
Characters/Description: Blackwelder 1943: 57 (characters in key), 57 (characters in key as Torrentomus), 87
(characters as Torrentomus), 105 (characters); Coiffait & Sáiz 1968: 415 (characters in key), 427 (characters);
Herman 1970a: 361 (characters in key), 395 (characters); Sáiz 1973c: 233 (characters); Navarrete-Heredia et
al. 2002: 229 (characters in key); Makranczy 2006: 105 (characters in key).
Key: Blackwelder 1943: 87 (species of Torrentomus of the West Indies), 105 (species of the West Indies); Coiffait
& Sáiz 1968: 428 (Chilean species); Sáiz 1973c: 234 (Chilean species); Scheibler & Chani-Posse 2009: 257
(species from south-western South America).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, Montserrat, Dominica, St. Vincent &
South America: Bolivia, Paraguay, Chile, Argentina.
0778. Thinodromus Kraatz, 1857b: 866

Type species: Trogophloeus dilatatus Erichson, 1839
Apocellagria Cameron, 1920b: 143 [synonymized by Makranczy 2006: 65]
Type species: Apocellagria indica Cameron, 1920
Warburtonia Oke, 1933: 104 [synonymized by Herman 1970a: 385]
Type species: Warburtonia inflatipes Oke, 1933
Subgenera:
Amisammus Gozis, 1886: 15
Type species: Carpelimus arcuatus Stephens, 1834
Carpaliaceus Gildenkov, 2000: 45
Type species: Thinodromus thoracicus Gildenkov, 2000
Mendaxinus Gildenkov, 2004: 9
Type species: Trogophloeus schoutedeni Cameron, 1928
Mendaxinus Gildenkov, 2001: 54 [nomem nudum, Gildenkov 2004: 9]
Type species: Trogophloeus schoutedeni Cameron, 1928
Paracarpalimus Scheerpeltz, 1937b: 105 [*]
Type species: Homalotrichus luteipes Solier, 1849
Thinodromus Kraatz, 1857b: 866
Type species: Trogophloeus dilatatus Erichson, 1839
45 species are included in this genus from Latin America. A large genus with many Neotropical species, which
when properly known could number well over a hundred species. Some groups are unique to the New World,
but in the context of the great disparity of Thinodromus should be maintained as species groups. A number of
species are catalogued in Herman (2001b) under Carpelimus [György Makranczy, personal communication].
Characters/Description: Herman 1970a: 361, 362 (characters in key), 385 (characters); Navarrete-Heredia et al.
2002: 230 (characters in key); Makranczy 2006: 65 (characters), 104 (characters in key); Makranczy 2018: 75
(characters of genus), 77 (characters of Thinodromus ferrugineus species group), 93 (characters of
Thinodromus circulus species group), 96 (characters of Thinodromus diffusus species group).
Distribution.
South America: Venezuela, Ecuador, Peru, Bolivia, Brazil, Paraguay, Chile, Argentina.
0779. Trogactus Sharp, 1887a: 702

Type species: Trogactus championi Sharp, 1887

8 species are known from and are restricted to the West Indies and Central America. A characteristic-looking
Neotropical genus, however, similar forms in the genus Thinodromus also exist (with extremely elongate
antennae). Most frequently encountered in leafpacks near streams, rare in collections, although good fliers.
Species count will rise significantly after revision (in preparation) [György Makranczy, personal
communication].
Characters/Description: Herman 1970a: 387 (characters); Navarrete-Heredia et al. 2002: 230 (characters in key);
Makranczy 2006: 67 (characters), 104 (characters in key).
Distribution.
West Indies: US Virgin Isl., Guadeloupe, St. Lucia, Saint Vincent & the Grenadines, Barbados, Grenada,
Grenada.
Incertae sedis in Subfamily Oxytelinae
0780. Dolichoxenus Engel & Chatzimanolis, 2009: 426

Type species: Dolichoxenus newtoni Engel & Chatzimanolis, 2009
1 species known only from Dominican amber.

Distribution.
Subfamily OXYPORINAE Fleming, 1821
0781. Oxyporus Fabricius, 1775: 267 (Figs. 126–127)

Type species: Staphylinus rufus Linnaeus, 1758
Characters/Description: Campbell 1969: 228 (characters).
Key: Scheerpeltz 1960: 78 (Neotropical species); Campbell 1969: 229 (New World species); Campbell 1974: 157
(some species); Campbell 1990: 214 (species of the Mexico and Central America); Navarrete-Heredia &
Novelo-Gutiérrez 1990: 231 (Mexican species); Márquez et al. 2005: 7 (Mexican species); Márquez & Asiain
2006: 58 (Mexican species).
Distribution.
0782. Pseudoxyporus Nakane & Sawada, 1956: 116, 120

Type species: Oxyporus hoplites Sharp, 1889

Characters/Description: Campbell 1969: 229 (characters in key), 230 (characters); Makranczy 2012: 113
(characters); M.H. Kim et al. 2016: 344 (characters).
Distribution.
Subfamily MEGALOPSIDIINAE Leng, 1920
0783. Megalopinus Eichelbaum, 1915: 104 (Figs. 128–131)

Type species: Oxyporus caelatus Gravenhorst, 1802
Aulacotrachelus L. Benick, 1920: 1 [synonymized by L. Benick 1922: 1]
Megalops Erichson, 1839a: 30 [synonymized by Scheerpeltz 1933: 1142]
Megalopsidia Leng, 1918: 204 [synonymized by Blackwelder 1952: 235]
Subgenera:
Gata Blackwelder, 1943: 202 [*]
Type species: Megalops praeditus Sharp, 1886
Perostylus L. Benick, 1917: 190 [synonymized by Blackwelder 1943: 202]
Type species: Megalops praeditus Sharp, 1886
Megalopinus Eichelbaum, 1915: 104 [*]
Megalopsidiella Bernhauer, 1933c: 333 [*]
Type species: Megalopsidia ogloblini Bernhauer, 1933
Polycyrtopsidia Scheerpeltz, 1972d: 95 [*]
Type species: Megalopsidia sanguinitriguttata Scheerpeltz, 1972
Stylopodus L. Benick, 1917: 190 [*]
Type species: Megalops cephalotes Erichson, 1840

Characters/Description: Coiffait & Sáiz 1968: 395 (characters, Megalopsidia).
Key: Bernhauer 1933c: 333 (Megalopsidia of the Argentina); Blackwelder 1943: 203 (species of the subgenus
Stylopodus of the West Indies); Sáiz 1970d: 3 (species of Chile); Puthz 1972: 119 (Megalopinus of the Chile);
Puthz 1994a: 17 (species of Megalopinus peploides group); Puthz 1994b: 425 (species of incisicollis
complex); Puthz 2012: 620 (Neotropical species and species group), 674 (species of the subgroup pulcher of
caelatus group), 682 (species of cephalotes group), 690 (species of incisicollis group), 711 (species the frons-
complex of punctatus group).
Distribution.
West Indies: Cuba, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
Subfamily SCYDMAENINAE Leach, 1815
Supertribe Cephenniitae Reitter, 1882d
Tribe Cephenniini Reitter, 1882d

0784. Cephazteca Jałoszyński, 2011b: 229
Type species: Cephazteca xilitla Jałoszyński, 2011

Distribution.
Central America: Mexico, Nicaragua, Costa Rica.
0785. Furcodes Jałoszyński, 2012b: 461

Type species: Furcodes apicalis Jałoszyński, 2012

Characters/Description: Jałoszyński 2012b: 470 (characters in key).
Distribution.
Central America: Mexico, Honduras.
0786. Monstrophennium Jałoszyński, 2012b: 458

Type species: Cephennium spinicolle L.W. Schaufuss, 1867

Distribution.
0787. Paracephennium O'Keefe, 1999: 74 (Fig. 137)

Type species: Paracephennium monteverde O'Keefe, 1999

Characters/Description: Jałoszyński 2012b: 460 (characters), 470 (characters in key); Jałoszyński 2012c: 249
(characters).
Key: O’Keefe 1999: 76 (species of Costa Rica).
Distribution.
0788. Pseudocephennium Reitter, 1883a: 385

Type species: Pseudocephennium integricolle Reitter, 1883

Characters/Description: Jałoszyński 2012b: 463 (characters), 470 (characters in key).
Distribution.
South America: Venezuela, Guyana, Suriname, Peru, Bolivia.
0789. Shyri Jałoszyński, 2012b: 453

Type species: Shyri pichincha Jałoszyński, 2012

Distribution.
Tribe Eutheiini Casey, 1897
0790. Eutheia Stephens, 1830: 115 (Fig. 134)

Type species: Eutheia scydmaenoides Stephens, 1830
Euthia Agassiz, 1846: 152 [synonymized by Newton & H. Franz 1998: 142]
Type species: Eutheia scydmaenoides Stephens, 1830

Characters/Description: O’Keefe 1999: 72 (characters); Jałoszyński 2011a: 122 (characters); Jałoszyński 2013f:
169 (characters).
Distribution.
Supertribe Mastigitae Fleming, 1821
Tribe Clidicini Casey, 1897
0791. Leptochromus Motschulsky, 1855: 12 (Fig. 135)

Type species: Leptochromus fulvescens Motschulsky, 1855
Hecotus Sharp, 1887b: 70 [synonymized by Csiki 1919: 87]
Type species: Hecotus mexicanus Sharp, 1887
6 species (1 in Mexican amber) are known from and are restricted to Central and South America.
Characters/Description: O'Keefe 1998: 263 (characters); O’Keefe 1999: 68 (characters); O’Keefe 2002: 214
(characters); Lord et al. 2014: 119 (characters).
Key: O’Keefe 1999: 70 (species of Costa Rica); O’Keefe 2002: 218 (species of the genus); Lord et al. 2014: 122
Distribution.
0792. Papusus Casey, 1897: 542

Type species: Papusus macer Casey, 1897

Characters/Description: H. Franz 1985c: 184 (characters); O'Keefe 1998: 263 (characters); O'Keefe 2003: 266
(characters).
Key: O'Keefe 2003: 271 (species of the genus).
Distribution.
Supertribe Scydmaenitae Leach, 1815

Tribe Chevrolatiini Reitter, 1882d
0793. Chevrolatia Jacquelin du Val, 1850: 45

Type species: Chevrolatia insignis Jacquelin du Val, 1850

Characters/Description: O'Keefe 1997: 166 (characters).
Key: O'Keefe 1997: 168 (species of the genus).
Distribution.
Tribe Glandulariini L.W. Schaufuss, 1889
0794. Afroeudesis H. Franz, 1963: 16

Type species: Pseudoeudesis basilewskyi H. Franz, 1963
Subgenera:
Afroeudesis H. Franz, 1963: 16
Type species: Pseudoeudesis basilewskyi H. Franz, 1963
Eburneus Jałoszyński, 2016e: 165
Type species: Microscydmus trianguliceps H. Franz, 1980
Eudesoides Jałoszyński, 2016e: 169 [*]
Type species: Pseudoeudesis sulcata H. Franz, 1967
Nanoscydmus Jałoszyński, 2009: 173
Type species: Nanoscydmus baoshanus Jałoszyński, 2009

Distribution.
0795. Alloraphes H. Franz, 1980c: 212 (Fig. 132)

Type species: Alloraphes jamaicae H. Franz, 1980

Characters/Description: H. Franz 1980c: 223 (characters in key); H. Franz 1989b: 277 (characters); Jałoszyński
2013a: 551 (characters).
Key: H. Franz 1980c: 216 (species of the genus).
Distribution.
South America: Venezuela, Peru, Bolivia, Brazil.
0796. Amimoscydmus Jałoszyński, 2013c: 259

Type species: Mimoscydmus baruerii H. Franz, 1980
Distribution.
0797. Archiconnus H. Franz, 1980c: 189

Type species: Archiconnus huallaganus H. Franz, 1980
Characters/Description: Jałoszyński 2013b: 525 (characters); Jałoszyński 2018c: 292 (characters).
Distribution.
0798. Euconnomorphus H. Franz, 1980c: 207 (Fig. 133)

Type species: Euconnomorphus pilosissimus H. Franz, 1980

Characters/Description: H. Franz 1980c: 222 (characters in key); Jałoszyński 2012a: 69 (characters); Jałoszyński
2013e: 143 (characters).
Distribution.
South America: Peru, Bolivia.
0799. Euconnus Thomson, 1859: 61

Type species: Pselaphus hirticollis Illiger, 1798
Subgenera:
Allonapochus H. Franz, 1972: 3
Type species: Euconnus mauritii H. Franz, 1972
Allonapochus H. Franz, 1986c: 199 [synonymized by Newton & H. Franz 1998: 145]
Type species: Euconnus perineti H. Franz, 1986
Androconnus H. Franz, 1986a: 21 [*]
Type species: Euconnus bechyneae H. Franz, 1986
Austroconophron H. Franz, 1971: 69
Type species: Euconnus tindoui H. Franz, 1971
Borneoconnus H. Franz, 1992: 900
Type species: Euconnus laticlava H. Franz, 1992
Cladoconnus Reitter, 1909: 226
Type species: Scydmaenus mothoulskii Sturm, 1838
Dimorphoconnus H. Franz, 1975: 177
Type species: Scydmaenus insigniventris Lea, 1912
Euconnus Thomson, 1859: 61 [*]
Type species: Pselaphus hirticollis Illiger, 1798
Diarthroconnus Ganglbauer, 1900: 261 [synonymized by Jałoszyński 2017d: 192]
Type species: Euconnus birnbacheri Ganglbauer, 1900
Glaphostoma L.W. Schaufuss, 1889: 2 [synonymized by Csiki 1919: 45]
Type species: Glaphostoma cribricolle L.W. Schaufuss, 1889
Scopophus Casey, 1897: 365 [synonymized by Jałoszyński 2017b: 459]
Type species: Euconnus affinis Casey, 1897
Euconophron Reitter, 1909: 226
Type species: Euconnus promptus Coquerel, 1860

Ursoconnus H. Franz, 1963: 42 [synonymized by Jałoszyński 2017f: 538]
Type species: Euconnus basilewskyi H. Franz, 1962
Eupentarius Reitter, 1907: 300
Type species: Euconnus grandiclavis Reitter, 1907
Filonapochus H. Franz, 1986c: 202
Type species: Euconnus grylloides H. Franz, 1986
Glabriconnus Jałoszyński, 2016b: 573
Type species: Euconnus banari Jałoszyński, 2016
Heteroconnus H. Franz, 1963: 40
Type species: Euconnus impressicollis H. Franz, 1963
Microconnus H. Franz, 1994a: 81 [*]
Type species: Euconnus canaimanus H. Franz, 1994
Myrmecodytes Croissandeau, 1898: 160
Type species: Scydmaenus pharaonis Motschulsky, 1851
Napochus Thomson, 1859: 61 [*]
Type species: Scydmaenus claviger Müller & Kunze, 1822
Cephaloconnus H. Franz in Newton & H. Franz, 1998: 145 [synonymized by Jałoszyński 2017f: 534]
Type species: Euconnus hypertrophicus H. Franz, 1962
Cerviconnus H. Franz, 1962: 368 [synonymized by Jałoszyński 2017f: 534]
Type species: Euconnus bundukii H. Franz, 1962
Connophron Casey, 1897: 390 [synonymized by Csiki 1919: 58]
Type species: Connophron repletum Casey, 1897
Glandularia L.W. Schaufuss, 1889: 3 [synonymized by H. Franz 1970: 548]
Type species: Glandularia fricatoris L.W. Schaufuss, 1889
Neonapochus Machulka, 1929: 21 [synonymized by Jałoszyński 2016g: 40]
Type species: Scydmaenus maklinii Mannerheim, 1844
Paranapochus H. Franz, 1986c: 192 [synonymized by Jałoszyński 2016g: 44]
Type species: Euconnus marojejyanus H. Franz, 1986
Protoscydmus H. Franz, 1992: 948 [synonymized by Jałoszyński 2016g: 40]
Type species: Protoscydmus sabahensis H. Franz, 1992
Smicrophus Casey, 1897: 467 [synonymized by Jałoszyński 2017a: 427]
Type species: Smicrophus leviceps Casey, 1897
Nodoconnus H. Franz, 1986c: 258
Type species: Euconnus gomyanus H. Franz, 1986
Paratetramelus H. Franz, 1963: 25
Type species: Euconnus kenyae H. Franz, 1963
Psomophus Casey, 1897: 366
Type species: Euconnus callidus Casey, 1897
Scydmaenites Croissandeau, 1898: 165 [synonymized by Jałoszyński 2017g: 570]
Type species: Euconnus kraatzii Reitter, 1881
Spanioconnus Ganglbauer, 1899: 46 [synonymized by H. Franz 1985c: 165]
Type species: Scydmaenus wetterhallii Gyllenhal, 1813
Xestophus Casey, 1897: 365 [synonymized by H. Franz 1985c: 167, Jałoszyński 2017g: 574]
Type species: Scydmaenus salinator LeConte, 1852
Pycnophus Casey, 1897: 385 [*]
Type species: Scydmaenus rasus LeConte, 1852
Nudatoconnus H. Franz, 1980c: 116 [synonymized by H. Franz 1985c: 167]
Type species: Euconnus sphaerocephalus H. Franz, 1980
Rhomboconnus H. Franz, 1986a: 20 [*]
Type species: Euconnus perplexus H. Franz, 1986
Tetramelus Motschulsky, 1869: 257 [*]
Type species: Scydmaenus oblongus Sturm, 1838
Drastophus Casey, 1897: 389 [synonymized by Jałoszyński 2017a: 426]
Type species: Drastophus laevicollis Casey, 1897
Euconnoides Croissandeau, 1898: 154 [synonymized by Csiki 1919: 64]
Type species: Scydmaenus styriacus Grimmer, 1841

Characters/Description: H. Franz 1980c: 86 (characters of subgenus Napochus), 116 (characters of subgenus
Nudatoconnus), 133 (characters of subgenus Tetramelus), 223 (characters in key); H. Franz 1986a: 19
(characters in key of subgenus Rhomboconnus), 19 (characters in key of subgenus Nudatoconnus), 19
(characters in key of subgenus Napochus), 20 (characters in key of subgenus Androconnus); H. Franz 1993b:
75 (characters in key of subgera Androconnus and Pycnophus); O’Keefe 1998: 264 (characters), 265
(characters of subgenus Noctophus), 265 (characters of subgenus Smicrophus), 266 (characters of subgenus
Xestophus), 266 (characters of subgenus Drastophus), 268 (characters of subgenus Napochus); Jałoszyński
2012a: 69 (characters); Jałoszyński 2015b: 10 (characters of subgenus Napochus); Jałoszyński 2015c: 485
(characters of subgenus Rhomboconnus); Jałoszyński 2015e: 249 (characters of subgenus Tetramelus), 292
(characters of subgenus Paratetramelus), 297 (characters of subgenus Heteroconnus); Jałoszyński 2017e: 81
(characters of subgenus Rhomboconnus); Jałoszyński & Newton 2017: 403 (characters of subgenus
Austroconophron); Jałoszyński 2018a: 588 (characters of subgenus Rhomboconnus).
Key: Reitter & Croissandeau 1890: 216 (some Brazilian species); H. Franz 1967: 616 (subgenera), 620 (species of
subgenus Napochus), 627 (species of subgenus Tetramelus), 656 (species incertae sedis in subgenera of
Euconnus), 688 (species of subgenus Napochus from tropical region of South America), 695 (some species of
subgenus Tetramelus), 715 (for some species); H. Franz 1980c: 113 (some species of subgenus Napochus),
129 (species of subgenus Nudatoconnus), 139 (species of subgenus Tetramelus), 150 (some species), 160
(some species), 187 (species of Jamaica and Caribbean islands), 238 (species of subgenus Tetramelus from
south of Chile), 259 (some species from south of Chile); H. Franz 1983a: 218 (species of Northeast of Brazil);
H. Franz 1983b: 136 (species of Suriname); H. Franz 1984: 28 (species from Caribbean Islands); H. Franz
1985b: 130 (species from Suriname); H. Franz 1986a: 19 (subgenera), 40 (Venezuelan species of subgenus
Androconnus); H. Franz 1988: 69 (Venezuelan species), 74 (Venezuelan species); H. Franz 1989a: 10
(Venezuelan species of subgenus Pycnophus), 52 (some Venezuelan species); H. Franz 1991: 45 (Cuban
species); H. Franz 1993b: 75 (South American species); H. Franz 1993c: 226 (Southern Brazilian species); H.
Franz 1994b: 374 (Mexican species); H. Franz 1995: 76 (species of Central America).
Distribution.
West Indies: Cuba, Jamaica, Dominican Republic, Puerto Rico, US Virgin Isl., Guadeloupe, Dominica, Grenada,
Trinidad-Tobago, Curacao.
South America: Colombia, Venezuela, Suriname, Ecuador, Peru, Bolivia, Brazil, Paraguay, Uruguay, Chile,
Argentina.
0800. Heteroscydmus H. Franz, 1980c: 208

Type species: Heteroscydmus yapacariensis H. Franz, 1980

Characters/Description: H. Franz 1980c: 223 (characters in key); Jałoszyński 2013c: 252 (characters).

Distribution.
0801. Homoconnus Sharp, 1887b: 64

Type species: Scydmaenus batesi L.W. Schaufuss, 1866
17 species (1 in Dominican amber) are known from and are restricted to Latin America.
Characters/Description: H. Franz 1980c: 222 (characters in key).
Key: H. Franz 1980c: 194 (species of the genus); H. Franz 1994a: 68 (species of the alluvial plains of the Orinoco
and Amazon river).
Distribution.
West Indies: Cuba, Dominican Republic.
South America: Venezuela, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil.
0802. Madagassoconnus H. Franz, 1986c: 112

Type species: Cyrtoscydmus kolbei L.W. Schaufuss, 1890
Madagassoconnus H. Franz, 1977: 351 [nomen nudum, synonymized by Newton & H. Franz 1998: 147]

Distribution.
Other regions: Madagascan.
0803. Magellanoconnus H. Franz, 1967: 628

Type species: Euconnus laurisilvae H. Franz, 1967
Subgenera:
Anthicimimus H. Franz, 1993a: 100 [*]
Type species: Euconnus anthiciformis H. Franz, 1967
Magellanoconnus H. Franz, 1967: 628 [*]
Type species: Euconnus laurisilvae H. Franz, 1967

Characters/Description: H. Franz 1993a: 100 (characters); Jałoszyński 2015a: 446 (characters as Anthicimimus);
Chandler & Newton 2017: 407 (characters), 411 (characters), 411(characters in key as subgenera
Anthicimimus and Magellanoconnus).
Key: H. Franz 1967: 644 (species of subgenus Magellanoconnus); H. Franz 1980c: 133 (species of the subgenus
Magellanoconnus from tropical area of South America), 251 (species of subgenus Magellanoconnus from
south of Chile).
Distribution.
Central America: Colombia, Ecuador, Bolivia, Chile, Argentina.
Other regions: Ethiopian, Australian.
0804. Meridaphes Jałoszyński, 2015d: 258

Type species: Scydmoraphes venezolanus H. Franz, 1988

Distribution.
0805. Mexiconnus Jałoszyński, 2013d: 530

Type species: Archiconnus mexicanus H. Franz, 1994

Distribution.
0806. Microraphes H. Franz, 1980c: 222

Type species: Microraphes pygmaeus H. Franz, 1980

Distribution.
0807. Microscydmus Saulcy & Croissandeau, 1893: 225

Type species: Scydmaenus nanus Schaum, 1844
Subgenera:
Microscydmus Saulcy & Croissandeau, 1893: 225 [*]
Type species: Scydmaenus nanus Schaum, 1844
Nepaloconnus H. Franz, 1979: 242 [synonymized by Jałoszyński 2016d: 470]
Type species: Euconnus khumbuensis H. Franz, 1979
Neoscydmus H. Franz, 1980c: 200 [*]
Type species: Microscydmus amazonicus H. Franz, 1980
Opresus Casey, 1897: 494
Type species: Scydmaenus misellus LeConte, 1852
Parastenichnus H. Franz, 1970: 574
Type species: Parastenichnus singaporensis H. Franz, 1970
Scydmomicrus Jałoszyński, 2014d: 11
Type species: Microscydmus australiensis H. Franz, 1975
Valdivioconnus H. Franz, 1980c: 252 [*]
Type species: Euconnus stenichniformis H. Franz, 1967
Characters/Description: H. Franz 1967: 659 (characters); H. Franz 1980c: 224 (characters in key); H. Franz 1993a:
100 (characters of subgenus Valdivioconnus); Jałoszyński 2014d: 4 (characters), 11 (characters of subgenus
Scydmomicrus).
Key: H. Franz 1967: 660 (species of the genus); H. Franz 1980c: 254 (species of the subgenus Valdivioconnus); H.
Franz 1984: 30 (species of the Caribbean Islands); H. Franz 1993c: 226 (Southern Brazilian species).
Distribution.
West Indies: Jamaica, Dominican Republic, Puerto Rico, US Virgin Isl., Dominica.
South America: Venezuela, Peru, Bolivia, Brazil, Chile, Argentina.
0808. Mimoscydmus H. Franz, 1980c: 220

Type species: Mimoscydmus brasiliensis H. Franz, 1980

Distribution.
0809. Napochomorphus H. Franz, 1980c: 206

Type species: Napochomorphus extraneus H. Franz, 1980

Distribution.
0810. Napoconnus H. Franz, 1957: 31

Type species: Euconnus kubienai H. Franz, 1957

Characters/Description: H. Franz 1980c: 75 (characters); Jałoszyński 2016d: 464 (characters).
Key: H. Franz 1980c: 84 (Sudamerican species of Napoconnus).
Distribution.
South America: Venezuela, Peru, Bolivia, Brazil.
0811. Neuraphanax Reitter, 1883a: 385

Type species: Eumicrus dux L.W. Schaufuss, 1867

Characters/Description: H. Franz 1967: 718 (characters); H. Franz 1980c: 222 (characters in key); H. Franz 1986a:
11 (characters).
Key: H. Franz 1967: 720 (species of the genus); H. Franz 1980c: 74 (species of the genus); H. Franz 1986a: 17
(Venezuelan species).
Distribution.
South America: Colombia, Venezuela, Peru.
0812. Neuraphes Thomson, 1859: 61

Type species: Scydmaenus angulatus Müller & Kunze, 1822
Neuraphes Thomson, 1862: 80 [synonymized by Newton & H. Franz 1998: 140, 148]
Subgenera:
Neuraphes Thomson, 1859: 61
Type species: Scydmaenus angulatus Müller & Kunze, 1822
Tropithorax Croissandeau, 1894: 359 [synonymized by Ganglbauer 1899: 25]
Type species: Neuraphes fauveli Croissandeau, 1894
Pararaphes Reitter, 1891b: 131
Type species: Neuraphes capellae Reitter, 1881
Rhynchoraphes Apfelbeck, 1918: 101 [Schülke & Smetana 2015: 882]
Type species: Neuraphes triangularis Apfelbeck, 1918
1 species from Domincan amber.
Distribution.
0813. Obesoconnus Jałoszyński, 2014b: 482 (Fig. 136)

Type species: Obesoconnus guyanensis Jałoszyński, 2014
Characters/Description: Jałoszyński 2018c: 292 (characters).
Distribution.
0814. Parapseudoconnus H. Franz, 1980c: 218

Type species: Pseudoconnus aberrans H. Franz, 1980
Subgenera:
Neuraphomimus H. Franz, 1986d: 44 [*]
Type species: Pseudoconnus simulator H. Franz, 1980
Pseudoconnus H. Franz, 1980c: 216 [synonymized by H. Franz 1986d: 44]
Vetusteconnus H. Franz, 1993a: 101 [synonymized by Newton & H. Franz 1998: 148]
Parapseudoconnus H. Franz, 1980c: 218 [*]
Type species: Pseudoconnus aberrans H. Franz, 1980

Characters/Description: H. Franz 1980c: 223 (characters in key, Pseudoconnus); Jałoszyński 2013b: 534
(characters), 536 (characters of subgenus Parapseudoconnus), 540 (characters of subgenus Neuraphomimus).
Distribution.
0815. Perumicrus Jałoszyński, 2015d: 264

Type species: Scydmoraphes peruanus H. Franz, 1980

Distribution.
0816. Plaumanniola Costa-Lima, 1962: 415 (Figs. 138–139)

Type species: Plaumanniola sanctaecatharinae Costa-Lima, 1962

Characters/Description: Lawrence & Reichardt 1966: 39 (characters); Jałoszyński 2013d: 318 (characters);
Jałoszyński 2016a: 362 (characters).
Distribution.
South America: French Guiana, Bolivia, Brazil.

0817. Protandroconnus H. Franz, 1989c: 636
Type species: Protandroconnus peruensis H. Franz, 1989
Distribution.
0818. Protoconnus H. Franz, 1967: 716

Type species: Protoconnus andicola H. Franz, 1967

Characters/Description: Jałoszyński 2018b: 43 (characters).
Key: H. Franz 1980c: 199 (species of the genus).
Distribution.
South America: Venezuela, Ecuador, Peru, Bolivia, Paraguay.
0819. Pseudoraphes H. Franz, 1980c: 209

Type species: Pseudoraphes huallagae H. Franz, 1980

Distribution.
0820. Sciacharis Broun, 1893: 1062

Type species: Sciacharis fulva Broun, 1893
Subgenera:
Maorinus H. Franz, 1980b: 291
Type species: Phagonophana alacer Broun, 1915
Maoria H. Franz, 1975: 14 [synonymized by H. Franz 1980b: 291]
Type species: Phagonophana alacer Broun, 1915
Palaeoconnus H. Franz, 1995: 53 [*]
Type species: Scydmaenus brevipes Sharp, 1887
Sciacharis Broun, 1893: 1062
Type species: Sciacharis fulva Broun, 1893
Allomaoria H. Franz, 1975: 49 [synonymized by H. Franz 1980b: 262]
Type species: Phagonophana lanosa Broun, 1885
Chathamaenus H. Franz, 1980b: 254 [synonymized by Jałoszyński 2014a: 224]
Type species: Chathamaenus chathamensis H. Franz, 1980

Characters/Description: H. Franz 1980b: 262 (characters); H. Franz 1993a: 99 (characters), 100 (characters of
subgenus Sciacharis); Jałoszyński & Newton 2017: 407 (characters of subgenus Sciacharis); Jałoszyński
2018d: 573 (characters of subgenus Palaeoconnus), 573 (characters in key of subgenera Palaeoconnus,
Sciacharis, and Maorinus).
Key: H. Franz 1980b: 263 (subgenera); H. Franz 1993a: 100 (subgenera); Jałoszyński 2018d: 573 (subgenera).
Distribution.
South America: Bolivia, Chile.
0821. Stenichnaphes H. Franz, 1980b: 255
Type species: Stenichnaphes urbanus H. Franz, 1980

Characters/Description: H. Franz, 1989b: 277 (characters).
Distribution.
0822. Stenichnoconnus H. Franz, 1985b: 114

Type species: Stenichnoconnus huijbregtsensis H. Franz, 1985

Distribution.
South America: Suriname.
0823. Venezolanoconnus H. Franz, 1988: 81

Type species: Venezolanoconnus andinus H. Franz, 1988
Characters/Description: Jałoszyński 2012a: 74 (characters); Jałoszyński 2017c: 293 (characters).
Key: H. Franz 1988: 83 (Venezuelan species).
Distribution.
Tribe Scydmaenini Leach, 1815
0824. Scydmaenus Latreille, 1802: 116

Type species: Pselaphus hellwigii Paykull, 1800 nec Herbst, 1792
Subgenera:
Afroscydmaenus H. Franz, 1963: 5
Type species: Scydmaenus mariusi H. Franz, 1963
Agathelor L.W. Schaufuss, 1884: 420
Type species: Agathelor deplanatum L.W. Schaufuss, 1884
Androscydmaenus H. Franz, 1980d: 691
Type species: Scydmaenus pilosissimus H. Franz, 1961
Androscydmus Newton in Newton & H. Franz, 1998: 152
Type species: Scydmaenus densepunctatus H. Franz, 1992
Androscydmaenus H. Franz, 1992: 870 [synonymized by Newton in Newton & H. Franz 1998: 152]
Type species: Scydmaenus densepunctatus H. Franz, 1992
Archiscydmaenus H. Franz in Newton & H. Franz, 1998: 152 [*]
Type species: Scydmaenus huijbregtsi H. Franz, 1985
Palaeoscydmaenus H. Franz, 1985b: 109 [synonymized by H. Franz in Newton & H. Franz 1998: 152]

Type species: Scydmaenus huijbregtsi H. Franz, 1985
Armatoscydmaenus H. Franz, 1971: 100 [*]
Type species: Scydmaenus regularis L.W. Schaufuss, 1889
Breviclavatoscydmaenus H. Franz, 1994a: 63 [*]
Type species: Scydmaenus kavacanus H. Franz, 1994
Choleropsis H. Franz, 1975: 291
Type species: Heterognathus gracilis King, 1864
Cholerus Thomson, 1859: 62 [*]
Type species: Pselaphus hellwigii Herbst, 1792
Mastigus Illiger, 1802: 296 [synonymized by Newton in Newton & H. Franz 1998: 152]
Type species: Pselaphus hellwigii Herbst, 1792
Conoscydmaenus H. Franz, 1980c: 68 [*]
Type species: Scydmaenus unicornis H. Franz, 1980
Corbulifer H. Franz, 1975: 286
Type species: Scydmaenus tamborinensis H. Franz, 1975
Eustemmoides H. Franz, 1958a: 3
Type species: Scydmaenus alluvialis H. Franz, 1958
Eustemmus Reitter, 1882c: 582
Type species: Scydmaenus antidotus Schaum, 1841
Geoscydmaenus H. Franz, 1961: 177
Type species: Scydmaenus pullus H. Franz, 1961
Heteroeustemmus H. Franz, 1971: 97
Type species: Eumicrus atrorufus Fauvel, 1889
Heteromicrus H. Franz, 1975: 300 [*]
Type species: Scydmaenus mirus H. Franz, 1975
Kingius H. Franz in Newton & H. Franz, 1998: 153
Type species: Heterognathus carinatus King, 1864
Heterognathus King, 1864: 96 [synonymized by H. Franz in Newton & H. Franz 1998: 153]
Type species: Heterognathus carinatus King, 1864
Mascarensia H. Franz, 1972: 11
Type species: Scydmaenus spinipes H. Franz, 1972
Mimoscydmaenus H. Franz, 1986b: 967
Type species: Scydmaenus chiangmaii H. Franz, 1986
Nepaloscydmaenus H. Franz, 1973: 118
Type species: Scydmaenus ghokarnae H. Franz, 1973
Paraeustemmus H. Franz, 1955: 1051
Type species: Scydmaenus vividus Reitter, 1887
Parageoscydmaenus H. Franz, 1980c: 69 [*]
Type species: Scydmaenus baloghi H. Franz, 1980
Parallomicrus H. Franz in Newton & H. Franz, 1998: 153
Type species: Scydmaenus rufus Müller & Kunze, 1822
Allomicrus H. Franz, 1975: 284 [synonymized by H. Franz in Newton & H. Franz 1998: 153]
Type species: Scydmaenus rufus Müller & Kunze, 1822
Parascydmaenus Jeannel & Paulian, 1945: 125
Type species: Scydmaenus spathifer Reitter, 1907
Pseudeumicrus Jeannel & Paulian, 1945: 126
Type species: Scydmaenus tachyoryctidis Jeannel & Paulian, 1945
Scottiscydmaenus H. Franz, 1975: 277
Type species: Scydmaenus lodoiceae Scott, 1922
Scydmaenus Latreille, 1802: 116 [*]
Type species: Pselaphus hellwigii Paykull, 1800 nec Herbst, 1792
Acholerops Casey, 1897: 535 [synonymized by H. Franz 1985c: 182]
Type species: Acholerops retrusa Casey, 1897
Alaudula L.W. Schaufuss, 1889: 31 [synonymized by Vít 2008: 265]
Type species: Alaudula rectispina L.W. Schaufuss, 1889
Clavigeroscydmus H. Franz, 1986: 374 [synonymized by Jałoszyński 2016c: 462]
Type species: Clavigeroscydmus corticariaeformis H. Franz, 1986
Eumicrus Laporte, 1832: 395 [synonymized by Reitter 1885d: 58]
Type species: Scydmaenus tarsatus Müller & Kunze, 1822
Ensimus Thomson, 1859: 62 [Newton & H. Franz 1998: 154]
Microstemma Motschulsky, 1858c: 57 [Newton & H. Franz 1998: 154]
Pseudomicrus Motschulsky, 1863: 425 [synonymized by Newton & H. Franz 1998: 154]
Type species: Scydmaenus alatus Nietner, 1856
Seychellinus H. Franz, 1986c: 359
Type species: Scydmaenus insularum Scott, 1922
Termitoscydmaenus H. Franz, 1980a: 55
Type species: Scydmaenus kistneri H. Franz, 1980
Trapezoscydmaenus H. Franz, 1986c: 366
Type species: Scydmaenus cavipennis Peyerimhoff, 1939

Characters/Description: Casey 1897: 470 (characters); H. Franz 1980c: 69 (characters subgenus Scydmaenus), 70
(characters in key of subgenera Parageoscydmaenus and Scydmaenus); Jałoszyński 2016c: 455 (characters);
Jałoszyński 2016f: 135 (characters), 139 (characters in key).
Key: Reitter & Croissandeau 1890: 219 (some Brazilian species); H. Franz 1980c: 70 (species of the Latin
America); H. Franz 1986a: 10 (Venezuelan species); H. Franz 1993b: 28 (South American species); H. Franz
1993c: 226 (Southern Brazilian species); H. Franz 1994a: 65 (species of subgenus Breviclavatoscydmaenus of
the alluvial plains of Orinoco and Amazon river); H. Franz 1994b: 334 (Mexican species); H. Franz 1995: 52
(species of the Central America).
Distribution.
West Indies: Cuba, Domincan Republic, US Virgin Isl., Montserrat, Guadeloupe, Martinique.
South America: Colombia, Venezuela, Suriname, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil, Paraguay, Chile,
Argentina.
Subfamily STENINAE W.S. MacLeay, 1825
0825. Stenus Latreille, 1797: 77 (Figs. 140–141)

Type species: Staphylinus juno Paykull, 1789
Subgenera:
Hemistenus Motschulsky, 1860b: 557 [*]
Type species: Stenus gilvipes Motschulsky, 1858

Mesostenus Rey, 1884: 326 [synonymized by Bernhauer & Schubert 1911: 178]
Type species: Stenus impressus Germar, 1823
Parastenus Heyden, 1905: 262 [synonymized by Bernhauer & Schubert 1911: 178]
Type species: Stenus impressus Germar, 1823
Hypostenus Rey, 1884: 390 [*]
Type species: Stenus kiesenwetteri Rosenhauer, 1856
Areus Casey, 1884c: 150 [synonymized by Horn, 1885: 113]
Type species: Stenus flavicornis Erichson, 1840
Astenus Lynch-Arribálzaga, 1884: 341 [synonymized by Bernhauer & Schubert 1911: 171]
Type species: Stenus speculifrons Fauvel, 1877
Stenosidotus Lynch-Arribálzaga, 1884: 338 [synonymized by Bernhauer & Schubert 1911: 171]
Type species: Stenus aenescens Lynch-Arribálzaga, 1884
Systenus Eichelbaum, 1913: 124 [synonymized by Scheerpeltz 1933: 1172]
Type species: Stenus amaniensis Eichelbaum, 1913
Metatesnus Ádám in Lázló & Gábor, 2001: 126
Type species: Stenus binotatus Ljungh, 1804
Adamostenus Özdikmen & Darilmaz, 2008: 303
Mestastenus Ádám, 1987a: 135 [synonymized by Ádám in Lázló & Gábor 2001: 126]
Nestus Rey, 1884: 246
Type species: Stenus boops Ljungh, 1810
Stenus Latreille, 1797: 77 [*]
Zolmaenus Stephens, 1829b: 291 [synonymized by Blackwelder 1952: 409]
Tesnus Rey, 1884: 315 [*]
Type species: Stenus opticus Gravenhorst, 1806
Mutinus Casey, 1884c: 146 [synonymized by Bernhauer & Schubert 1911: 169]
Type species: Stenus dispar Casey, 1884

Characters/Description: Lynch-Arribálzaga 1884: 332 (characters); Blackwelder 1943: 208 (characters); Coiffait &
Sáiz 1968: 398 (characters).
Key: Lynch-Arribálzaga 1884: 334 (species of Buenos Aires province, Argentina); L. Benick 1928: 43, 45 (some
species); L. Benick 1938b: 146 (subgenera), 154 (some Brazilian species), 162 (some species of the Central
and South America); L. Benick 1938a: 256 (subgenera), 259 (species of the Central America); Blackwelder
1943: 209 (species of the West Indies); L. Benick 1949: 566 (New World species); Coiffait & Sáiz 1968: 399
(Chilean species); Puthz 1972: 120 (species of Chile); Puthz 1977a: 161 (species of Stenus aculeatus group);
Puthz 1984: 88 (species of Stenus podagricus group), 177 (species of Stenus aeneas group); Puthz 2000: 124
(species apterous of subgenus Hemistenus of Mexico); Puthz 2006a: 663 (Neotropical species of the subgenus
Tesnus); Puthz 2015: 753 (Neotropical groups and species).
Distribution.
West Indies: Bahamas, Cuba, Jamaica, Haiti, Dominican Republic, Guadeloupe, Grenada, Trinidad-Tobago.
Paraguay, Uruguay, Chile, Argentina.
Subfamily EUAESTHETINAE Thomson, 1859
Tribe Alzadaesthetini Scheerpeltz, 1974
0826. Alzadaesthetus Kistner, 1961a: 217

Type species: Alzadaesthetus chilensis Kistner, 1961

Characters/Description: Sáiz, 1972: 173 (characters in key), 174 (characters).
Key: Sáiz 1972: 175 (key to species).
Distribution.
Tribe Austroesthetini Cameron, 1944b
0827. Chilioesthetus Sáiz, 1968a: 73 (Fig. 142)

Type species: Chilioesthetus lorenae Sáiz, 1968

Characters/Description: Sáiz, 1972: 174 (characters in key).
Key: Sáiz 1968a: 74 (some species); Puthz 2008: 23 (species of the genus).
Distribution.
0828. Nothoesthetus Sáiz, 1970a: 295

Type species: Nothoesthetus coiffaiti Sáiz, 1969

Characters/Description: Sáiz 1972: 173 (characters in key), 176 (characters).
Key: Sáiz 1972: 177 (key to species).
Distribution.
Tribe Euaesthetini Thomson, 1859
0829. Edaphus Motschulsky, 1857: 7 (Figs. 143–144)

Type species: Edaphus nitidus Motschulsky, 1857
Edaphellus Fauvel, 1878a: 220 [synonymized by Székessy 1939: 98]
Type species: Edaphellus novaeguineae Fauvel, 1878
Edaphus LeConte, 1861: 67 [synonymized by Blackwelder 1952: 144]
Type species: Edaphus nitidus LeConte, 1861
Hawkeswoodedaphus Makhan, 2007: 1 [synonymized by Puthz 2011a: 18]
Type species: Edaphus aschnaae Makhan, 1995

Microphthartus Blattný, 1925: 185 [synonymized by Puthz 1974: 912]
Type species: Microphthartus luridus Blattný, 1925
Rhenanus Wüsthoff, 1935: 48 [synonymized by Holdhaus 1936: 10]
Type species: Rhenanus rosskotheni Wüsthoff, 1935
Tetrameres L.W. Schaufuss, 1877a: 460 [synonymized by L.W. Schaufuss 1877b: 24]
Type species: Tetratarsus plicatulus L.W. Schaufuss, 1877
Tetratarsus L.W. Schaufuss, 1877b: 24 [synonymized by Reitter 1882b: 211]
Type species: Tetratarsus plicatulus L.W. Schaufuss, 1877

Key: Puthz 1973: 51 (Neotropical species); Puthz 2001c: 26 (Mexican species); Puthz 2006a: 84 (species of
ventralis group); Puthz 2014: 814 (species of the New World).
Distribution.
Paraguay, Chile.
0830. Euaesthetus Gravenhorst, 1806: 201

Type species: Euaesthetus scaber Gravenhorst, 1806
Eristhetus Leach in Samouelle, 1819: 174 [synonymized by Lacordaire 1854: 109]
Type species: Euaesthetus scaber Gravenhorst, 1806
Key: Puthz 2001b: 32 (Neotropical species).
Distribution.
West Indies: Cuba.
Central America: Mexico, Nicaragua, Costa Rica.
0831. Octavius Fauvel, 1873b: 62

Type species: Octavius pyrenaeus Fauvel, 1873
Exoctavius Bierig, 1934b: 221 [synonymized by Puthz 1977b: 106]
Type species: Exoctavius bermudezi Bierig, 1934
Brinckiellus Scheerpeltz, 1974: 98, 105 [synonymized by Puthz 1980: 27]
Type species: Brinckiellus capensis Scheerpeltz, 1974
Doletica Cameron, 1938: 1 [synonymized by Kistner 1962: 101]
Type species: Doletica bicolor Cameron, 1938
Nepalopeplus Coiffait, 1982: 129 [synonymized by Puthz 1987: 453]
Type species: Nepalopeplus himalayicus Coiffait, 1982
Thaxterius Bernhauer, 1935b: 213 [synonymized by Kistner 1960a: 117]
Type species: Thaxterius sulcicollis Bernhauer, 1935
Traegardhius Scheerpeltz, 1974: 93, 105 [synonymized by Puthz 1980: 27]
Type species: Traegardhius natalensis Scheerpeltz, 1974
Turellus Sharp, 1876a: 423 [synonymized by Puthz 1980: 26]
Type species: Turellus batesi Sharp, 1876

Characters/Description: Fauvel 1873a: 220 (characters); Puthz 1977b: 106 (characters); Navarrete-Heredia et al.
Key: Puthz 1977b: 108 (some Neotropical species); Puthz 2001a: 27 (Neotropical species).
Distribution.
West Indies: Cuba.
Central America: Mexico, Belize, Costa Rica, Panama.
South America: Venezuela, Suriname, French Guiana, Ecuador, Peru, Brazil, Paraguay.
0832. Phaenoctavius Pace, 1986d: 45 (Fig. 145)

Type species: Phaenoctavius brasiliensis Pace, 1986

Distribution.
0833. Schatzmayrina C. Koch, 1934: 63

Type species: Schatzmayrina oxyclypea C. Koch, 1934

Characters/Description: Puthz 2007: 84 (characters).
Key: Puthz 2007: 85 (some species of the genus).
Distribution.
0834. Tamotus L.W. Schaufuss, 1872: 248

Type species: Tamotus femoratus L.W. Schaufuss, 1874
Paredaphus Bernhauer, 1915b: 291 [synonymized by Puthz 1973: 60]
Type species: Paredaphus tropicus Bernhauer, 1915

Characters/Description: L.W. Schaufuss 1882a: 170 (characters); Bernhauer 1915b: 291 (characters); Puthz 1973:
60 (characters).
Key: Puthz 1973: 62 (species of the genus); Puthz 2002: 123 (species of the genus).
Distribution.
West Indies: Cuba.
South America: Suriname, Ecuador, Peru, Brazil, Paraguay.
Tribe Stenaesthetini Bernhauer & Schubert, 1911
0835. Stenaesthetus Sharp, 1874: 79

Type species: Stenaesthetus sunioides Sharp, 1874

Aulacosthaetus Bernhauer, 1939d: 211 [synonymized by Puthz 1980: 20]
Type species: Aulacosthaetus rambouseki Bernhauer, 1939
Gerhardia Kistner, 1960b: 32 [synonymized by Puthz 2011b: 795]
Type species: Stenaesthetus africanus Bernhauer, 1915
Rishwanedaphus Makhan, 2013a: 1 [synonymized by Puthz 2013: 105]
Type species: Rishwanedaphus amrishi Makhan, 2013
22 species are included in this genus from The West Indies and South America.
Characters/Description: Orousset 1990: 11 (characters).
Key: Puthz 1973: 67 (Neotropical species); Orousset 1990: 19 (group of species), 19 (species of the group
Stenaesthetus illatus), 37 (species of the group Stenaesthetus mrazi).
Distribution.
South America: Suriname, Colombia, Ecuador, Peru, Brazil, Paraguay.
Other regions: Palaearctic, Ethiopian, Madagascan, Oriental.
Subfamily SOLIERIINAE Newton & Thayer, 1992
0836. Solierius Bernhauer, 1921a: 68 (Fig. 146)

Type species: Physognathus obscurus Solier, 1849
Physognathus Solier, 1849: 303 [synonymized by Bernhauer 1921a: 68]
Type species: Physognathus obscurus Solier, 1849

Distribution.
Subfamily LEPTOTYPHLINAE Fauvel, 1874b
Tribe Metrotyphlini Coiffait, 1963a
0837. Apotyphlus Coiffait & Sáiz, 1965a: 129

Type species: Apotyphlus relictus Coiffait & Sáiz, 1965

Characters/Description: Coiffait & Sáiz 1965a: 135 (characters in key).
Distribution.
0838. Venezillotyphlus Decu, 1990: 37
Type species: Venezillotyphlus pecki Decu, 1990

Distribution.
Tribe Neotyphlini Coiffait, 1963a
0839. Chiliotyphlus Coiffait, 1963a: 378

Type species: Chiliotyphlus aculeensis Coiffait, 1963

Characters/Description: Coiffait 1963a: 372 (characters in key); Coiffait & Sáiz 1965a: 136 (characters in key).
Key: Coiffait 1963a: 378 (Chilean species).
Distribution.
0840. Cubanotyphlus Coiffait & Decou, 1972: 131 (Fig. 147)

Type species: Cubanotyphlus jimenezi Coiffait & Decou, 1972
Characters/Description: Frank & Thomas 1984c: 1412 (characters).
Distribution.
West Indies: Cuba.
0841. Eutyphlops Coiffait, 1983: 345

Type species: Eutyphlus ornatus Coiffait, 1963
Eutyphlus Coiffait, 1963a: 375 [synonymized by Coiffait 1983: 345]
Type species: Eutyphlus ornatus Coiffait, 1963
Characters/Description: Coiffait & Sáiz 1965a: 133 (characters as Eutyphlus), 135 (characters in key as Eutyphlus).
Key: Coiffait & Sáiz 1965a: 133 (species of Chile as Eutyphlus).
Distribution.
0842. Macrotyphlus Coiffait & Sáiz, 1965a: 131

Type species: Macrotyphlus sexualis Coiffait & Sáiz, 1965

Characters/Description: Coiffait & Sáiz 1965a: 136 (characters in key).
Key: Coiffait & Sáiz 1965a: 133 (species of Chile); Sáiz 1976: 7 (some species of the group bicornis), 8 (some
species of the group politus).
Distribution.
0843. Mayatyphlus Gusarov, 2003d: 2

Type species: Mayatyphlus carltoni Gusarov, 2003

Distribution.

Central America: Belize.
0844. Oreinotyphlus Coiffait, 1963a: 372

Type species: Oreinotyphlus castrii Coiffait, 1963

Characters/Description: Coiffait 1963a: 371 (characters in key); Coiffait & Sáiz 1965a: 135 (characters in key).
Key: Coiffait 1963a: 373 (species of Chile).
Distribution.
0845. Paramacrotyphlus Sáiz, 1975: 52

Type species: Paramacrotyphlus valdiviensis Sáiz, 1975

Distribution.
Subfamily PSEUDOPSINAE Ganglbauer, 1895
0846. Pseudopsis Newman, 1834: 313 (Figs. 148–150)

Type species: Pseudopsis sulcata Newman, 1834
Subgenera:
Pseudopsiella Bernhauer, 1939d: 204
Type species: Pseudopsis arrowi Bernhauer, 1939
Chiliopseudopsis Coiffait & Sáiz, 1968: 458 [synonymized by Herman 1975: 257]
Type species: Pseudopsis adustipennis Fairmaire & Germaine, 1862
Pseudopsis Newman, 1834: 313 [*]
Type species: Pseudopsis sulcata Newman, 1834
Characters/Description: Coiffait & Sáiz 1968: 457 (characters), 458 (characters, Chiliopseudopsis); Herman 1975:
257 (characters).
Key: Herman 1975: 262 (species of the genus); Asenjo & Ribeiro-Costa 2010: 231 (South American species).
Distribution.
South America: Colombia, Venezuela, Ecuador, Peru, Chile, Argentina.
Subfamily PAEDERINAE Fleming, 1821
Tribe Cylindroxystini Bierig, 1943
Subtribe Cylindroxystina Bierig, 1943
0847. Cylindroxystus Bierig, 1943: 158
Type species: Cylindroxystus longulus Bierig, 1943

Characters/Description: Herman 1991: 12 (characters); Ramírez & Asenjo 2017: 438 (characters).
Key: Herman 1991: 15 (species of the genus); Ramírez & Asenjo 2017: 439 (species of the genus).
Distribution.
West Indies: Dominican Republic, Puerto Rico.
South America: Colombia, Venezuela, Guyana, Ecuador, Peru, Brazil.
0848. Neolindus Scheerpeltz, 1933: 1219 (Fig. 163)

Type species: Lindus religans Sharp, 1876
Lindus Sharp, 1876a: 281 [synonymized by Scheerpeltz 1933: 1219]
Type species: Lindus religans Sharp, 1876
Characters/Description: Sharp 1876a: 281 (characters); Herman 1991: 33 (characters).
Key: Irmler 1981b: 209 (species of the genus); Herman 1991: 35, 38 (species of the genus); Asenjo 2011: 64
Distribution.
South America: Colombia, Venezuela, French Guiana, Ecuador, Peru, Bolivia, Brazil.
Tribe Lathrobiini Laporte, 1835
Subtribe Astenina Hatch, 1957
0849. Astenus Dejean, 1833: 65 (Fig. 152)

Type species: Staphylinus angustatus Paykull, 1789
Subgenera:
Astenopleuritus Coiffait, 1982: 98
Type species: Astenus flavipennis Cameron, 1931
Astenus Dejean, 1833: 65 [*]
Astenognathus Reitter, 1909: 150 [synonymized by Assing 2008a: 1241]
Type species: Sunius bimaculatus Erichson, 1840
Thoobia Gistel, 1856: 389 [synonymized by Blackwelder 1952: 388]
Eurysunius Reitter, 1909: 149
Type species: Sunius paradoxus Eppelsheim, 1878
Mecognathus Wollaston, 1854: 595
Type species: Mecognathus chimaera Wollaston, 1854
Suniogaster Reitter, 1909: 151 [synonymized by Blackwelder 1943: 365]
Type species: Sunius ampliventris Reitter, 1900
Neognathus Sharp, 1874: 69
Type species: Neognathus angulatus Sharp, 1874

Characters/Description: Stephens 1833a: 275 (characters); Blackwelder 1943: 231 (characters in key), 365
(characters); Blackwelder 1939: 101 (characters in key); Navarrete-Heredia et al. 2002: 267 (characters in
key); Newton & Peck 2006: 148 (characters in key).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Dominican Republic, St. Lucia, St. Vincent & Grenadines, Grenada,
Trinidad-Tobago.
South America: Colombia, Venezuela, Galapagos Isl., Brazil, Uruguay, Argentina.
Subtribe Echiasterina Casey, 1905
0850. Amrishronetus Makhan, 2013b: 1

Type species: Amrishronetus rishwani Makhan, 2013

Distribution.
South America: Suriname.
0851. Echiaster Erichson, 1839a: 29 (Fig. 156)

Type species: Echiaster longicollis Erichson, 1840
Subgenera:
Echiaster Erichson, 1839a: 29 [*]
Type species: Echiaster longicollis Erichson, 1840
Leptogenius Casey, 1886b: 214
Type species: Leptogenius brevicornis Casey, 1886
Polyasterellus Bernhauer, 1925: 34 [*]
Type species: Echiaster bruchi Bernhauer, 1925

Characters/Description: Lynch-Arribálzaga 1884: 277 (characters); Blackwelder 1939: 101 (characters in key);
Blackwelder 1943: 231 (characters in key), 369 (characters); Coiffait & Sáiz 1968: 380 (characters in key),
383 (characters); Navarrete-Heredia et al. 2002: 267 (characters in key); Newton & Peck 2006: 148
of the West Indies); Scheerpeltz 1969b: 86 (species of the genus).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Antigua & Barbuda, Montserrat, Martinique, St.
Vincent & Grenadines, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil, Paraguay, Chile,
Argentina.
0852. Haplonazeris Coiffait & Sáiz, 1968: 380
Type species: Mecognathus sculptilis Fairmaire & Germain, 1862
Characters/Description: Sáiz 1970c: 41 (characters).
Key: Sáiz 1970c: 45 (species of the genus).
Distribution.
0853. Myrmecosaurus Wasmann, 1909b: 766 (Fig. 162)

Type species: Echiaster myrmecophilus Holmgren, 1908
Myrmescopaeus Brèthes, 1916: 431 [synonymized by Bruch 1918: 187]
Type species: Myrmescopaeus gallardoi Brèthes, 1916

Characters/Description: Blackwelder 1939: 100 (characters in key); Coiffait & Sáiz 1968: 379 (characters in key),
382 (characters).
Distribution.
South America: Bolivia, Brazil, Chile, Argentina.
Other regions: Nearctic [adventive].
0854. Pseudastenus Bernhauer, 1933b: 520

Type species: Pseudastenus barretoi Bernhauer, 1933

Distribution.
0855. Ronetus Blackwelder, 1943: 364

Type species: Ronetus ortinolus Blackwelder, 1943

Key: Makhan, 2013c: 2 (species of Surinam).
Distribution.
South America: Colombia, Venezuela, Suriname.
0856. Santiagonius Bruch, 1930a: 31 (Fig. 171)

Type species: Santiagonius gomezi Bruch, 1930

Distribution.
0857. Sphaeronum Sharp, 1876a: 224

Type species: Sphaeronum pallidum Sharp, 1876

Characters/Description: Blackwelder 1939: 100 (characters in key); Blackwelder 1943: 231 (characters in key),
374 (characters); Navarrete-Heredia et al. 2002: 267 (characters in key).
Distribution.
West Indies: Cuba.
0858. Termitosaurus Silvestri, 1946b: 525

Type species: Termitosaurus insinuatus Silvestri, 1946

Distribution.
0859. Zonaster Sharp, 1886: 595

Type species: Zonaster optatus Sharp, 1886

Distribution.
Subtribe Lathrobiina Laporte, 1835
0860. Attaxenus Wasmann, 1925b: 157

Type species: Attaxenus horridus Wasmann, 1925
Distribution.
0861. Dacnochilus LeConte, 1861: 66 (Fig. 154)

Type species: Dacnochilus laetus LeConte, 1863
Acalophaena Sharp, 1886: 554 [synonymized by Jiménez-Sánchez & Galián 2013: 82]
Type species: Calophaena basilis Lynch-Arribálzaga, 1884
Calophaena Lynch-Arribálzaga, 1884: 267 [synonymized by Sharp 1886: 554]
Type species: Calophaena basalis Lynch-Arribálzaga, 1884
Characters/Description: Blackwelder 1939: 98 (characters in key as Acalophaena), 98 (characters in key);
Navarrete-Heredia et al. 2002: 268 (characters in key as Acalophaena), 268 (characters in key).
Distribution.
South America: Colombia, Venezuela, Guyana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Argentina.
0862. Lathrobium Gravenhorst, 1802: 51

Type species: Staphylinus elongatus Linnaeus, 1767
Glyptomerodes Scheerpeltz, 1976c: 31 [nomen nudum, Newton & Thayer 2003]
Glyptomerodoschema Scheerpeltz, 1976c: 29 [synonymized by Assing 2012e: 164]
Type species: Lathrobium janetscheki Scheerpeltz, 1976
Lathrobius Billberg, 1820: 16 [synonymized by Blackwelder 1952: 211]
Subgenera:
Abletobium Casey, 1905: 79
Type species: Abletobium pallescens Casey, 1905
Apteralium Casey, 1905: 77 [*]
Type species: Lathrobium brevipenne LeConte, 1863
Type species: Lathrobium krniense Joseph, 1868
Glyptomerus H. Müller, 1856: 308
Type species: Glyptomerus cavicola Müller, 1856
Typhlobium Kraatz, 1856a: 625 [synonymized by Kraatz 1857a: 668, 669]
Type species: Typhlobium stagophilum Kraatz, 1856
Lathrobioma Casey, 1905: 98
Type species: Lathrobium tenue LeConte, 1863
Lathrobiopsis Casey, 1905: 97
Type species: Lathrobiopsis texana Casey, 1905
Lathrobium Gravenhorst, 1802: 51 [*]
Bathrolium Gozis, 1886: 14 [Newton & Thayer, 2003]
Type species: Staphylinus punctatus Fourcroy, 1785
Centrocnemiella Strand, 1934: 276 [synonymized by Assing 2008a: 1270]
Type species: Lathrobium krniense Joseph, 1868
Centrocnemis Joseph, 1868: 366 [synonymized by Strand 1934: 276]
Hypophylladobius Fauvel, 1885: 34 [synonymized by Blackwelder 1952: 196]
Type species: Lathrobium anophthalmum Fauvel, 1885
Litolathra Casey, 1905: 93 [synonymized by Blackwelder 1939: 108]
Type species: Litolathra suspecta Casey, 1905
Lathrolepta Casey, 1905: 103
Type species: Lathrobium debile LeConte, 1880
Palaeolobrathium M. Abdullah & A. Abdullah, 1968: 2
Type species: Lathrobium whitei Abdullah & Abdullah, 1968
Sucoca Blackwelder, 1952: 365 [*]
Type species: Heterosoma dohrni Bernhauer, 1903
Heterosoma Bernhauer, 1903b: 33 [synonymized by Blackwelder 1952: 365]
Type species: Heterosoma dohrni Bernhauer, 1903

Characters/Description: Lynch-Arribálzaga 1884: 235 (characters); Blackwelder 1939: 97 (characters in key,
Glyptomerus); Blackwelder 1939: 98 (characters in key); Blackwelder 1943: 230 (characters in key), 307
(characters); Navarrete-Heredia et al. 2002: 268 (characters in key); Newton & Peck 2006: 149 (characters in
key); Assing 2012d: 51 (characters).
Key: Lynch-Arribálzaga 1884: 237 (species of Buenos Aires province, Argentina); Blackwelder 1939: 98
(subgenera); Blackwelder 1943: 308 (species of the West Indies).
Distribution.
West Indies: Cuba, Jamaica, Dominican Republic, Puerto Rico, Trinidad-Tobago.
South America: Brazil, Paraguay, Uruguay, Argentina.

0863. Lobrathium Mulsant & Rey, 1878b: 78
Type species: Lathrobium multipunctum Gravenhorst, 1802
Subgenera:
Eulathrobium Casey, 1905: 114
Type species: Lathrobium grande LeConte, 1863
Lathrotropis Casey, 1905: 115 [synonymized by Blackwelder 1939: 109]
Type species: Lathrobium jacobinum LeConte, 1863
Lobrathium Mulsant & Rey, 1878b: 78 [*]
Type species: Lathrobium multipunctum Gravenhorst, 1802
Lathrobiella Casey, 1905: 133 [synonymized by Blackwelder 1939: 109]
Type species: Lathrobium collaris Erichson, 1840
Lathrotaxis Casey, 1905: 122 [synonymized by Blackwelder 1939: 109]
Type species: Lathrobium longiusculum Gravenhorst, 1802
Paralobrathium Bordoni, 1999: 119 [synonymized by Assing 2007b: 753]
Type species: Lathrobium apicale Baudi di Selve, 1857
Ponthrobium Korge, 1971: 27
Type species: Lathrobium heinzi Korge, 1971
Allobrathium Coiffait, 1972a: 134 [synonymized by Coiffait 1981c: 53]
Type species: Lathrobium lethierryi Reiche, 1869
9 species are included in this genus from West and South America.
Characters/Description: Blackwelder 1939: 98 (characters in key); Blackwelder 1943: 230 (characters in key), 311
(characters).
Key: Blackwelder 1939: 98 (subgenera); Blackwelder 1943: 312 (species of the West Indies); Assing 2012d: 51
(characters).
Distribution.
West Indies: Cuba, Cayman Is., Haiti, Dominican Republic, US Virgin Isl.
0864. Paederopsis Wasmann, 1912: 98

Type species: Paederopsis myrmecophila Wasmann, 1912

Distribution.
0865. Platybrathium Bierig, 1933: 478

Type species: Platybrathium panamense Bierig, 1933

Distribution.
0866. Pseudolathra Casey, 1905: 129
Type species: Lathrobium anale LeConte, 1880
Subgenera:
Allolathra Coiffait, 1972a: 132
Type species: Lathrobium methneri Bernhauer, 1915
Madecalobrathium Scheerpeltz, 1961: 251
Type species: Lathrobium keiseri Scheerpeltz, 1961
Pseudolathra Casey, 1905: 129 [*]
Type species: Lathrobium anale LeConte, 1880
Lathrobidium Portevin, 1929: 382 [Newton & Thayer 2003]
Type species: Lathrobium lusitanicum Erichson, 1840
Linolathra Casey, 1905: 131 [synonymized by Blackwelder 1939: 110]
Type species: Linolathra filitarsis Casey, 1905
Microlathra Casey, 1905: 142 [synonymized by Blackwelder 1939: 110]
Type species: Lathrobium pallidulum LeConte, 1880
Paralathra Casey, 1905: 130 [synonymized by Blackwelder 1939: 110]
Type species: Paralathra filicornis Casey, 1905

Characters/Description: Coiffait & Sáiz 1968: 380 (characters in key), 390 (characters); Assing 2012d: 51
(characters).
Distribution.
West Indies: Cuba, Haiti, Dominican Republic, US Virgin Isl., St. Lucia.
South America: Colombia, Venezuela, Bolivia, Brazil, Paraguay, Uruguay, Chile, Argentina.
0867. Stereocephalus Lynch-Arribálzaga, 1884: 231 (Fig. 174)

Type species: Stereocephalus seriatipennis Lynch-Arribálzaga, 1884

Characters/Description: Lynch-Arribálzaga 1884: 231 (characters); Herman 1979: 2 (characters); Matějíček 2018:
65 (characters).
Key: Herman 1979: 4 (species of the genus); Matějíček 2018: 68 (species of the genus).
Distribution.
South America: Venezuela, Brazil, Paraguay, Uruguay, Argentina.
0868. Tetartopeus Czwalina, 1888: 349

Type species: Lathrobium terminatum Gravenhorst, 1802
Deratopeus Casey, 1905: 112 [synonymized by Watrous 1980: 313]
Type species: Deratopeus parvipennis Casey, 1905
Only one species is included in this genus from West Indies.

Distribution.
West Indies: Cuba.
Other regions: Palaearctic, Oriental, Australian.

Subtribe Medonina Casey, 1905
0869. Achenomorphus Motschulsky, 1858d: 647

Type species: Achenomorphus columbicus Motschulsky, 1858
Subgenera:
Achenomorphus Motschulsky, 1858d: 647
Type species: Achenomorphus columbicus Motschulsky, 1858
Aderocharis Sharp, 1886: 552 [*]
Type species: Paederus corticinus Gravenhorst, 1802
Dorocharis Blackwelder, 1939: 99 [*]
Type species: Aderocharis chapini Blackwelder, 1939

Characters/Description: Blackwelder 1943: 230 (characters in key of Aderocharis), 250 (characters of
Aderocharis); Navarrete-Heredia et al. 2002: 268 (characters in key).
Key: Blackwelder 1943: 251 (species of the subgenus Aderocharis of the West Indies).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Puerto Rico, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
South America: Colombia, Guyana, French Guiana, Bolivia, Brazil.
0870. Argoderus Bierig, 1933: 498 (Fig. 151)

Type species: Argoderus panamensis Bierig, 1933
Distribution.
0871. Deroderus Sharp, 1886: 577

Type species: Deroderus vestitus Sharp, 1886
Lypomedon Blackwelder, 1952: 228 [synonymized by Frania 1986: 474]
Type species: Lithocharis tabacina Casey, 1884
Polymedon Casey, 1905: 156 [synonymized by Blackwelder 1952: 228]
Type species: Lithocharis tabacina Casey, 1884

Characters/Description: Frania 1986: 472 (characters); Frania 1987: 250 (characters of larvae); Navarrete-Heredia
et al. 2002: 268 (characters in key).
Distribution.
0872. Ecitocleptis Borgmeier, 1949: 115

Type species: Ecitocleptis socia Borgmeier, 1949
Distribution.
0873. Ecitomedon Bernhauer, 1925: 35
Type species: Ecitomedon bruchi Bernhauer, 1925
Distribution.
0874. Eusclerus Sharp, 1886: 575

Type species: Eusclerus rugifrons Sharp, 1886

Distribution.
0875. Hypomedon Mulsant & Rey, 1878b: 152

Type species: Lithocharis debilicornis Wollaston, 1857
Asteria Fauvel, 1889: 120 [synonymized by Blackwelder 1952: 65]
Type species: Medon effluens Fauvel, 1889
Chloëcharis Lynch-Arribálzaga, 1884: 257 [synonymized by Blackwelder 1939: 103]
Type species: Chloecharis rufula Lynch-Arribálzaga, 1884
Euastenus Fiori, 1915b: 10 [synonymized by Blackwelder 1939: 103]
Type species: Euastenus pallidus Fiori, 1915
Hemimedon Casey, 1905: 160 [synonymized by Fauvel 1889: 120]
Type species: Hemimedon angustus Casey, 1905
Lena Casey, 1886b: 211 [synonymized by Blackwelder 1939: 103]
Type species: Lena testacea Casey, 1886
Only one species is included in this genus from Latin America.

Characters/Description: Blackwelder 1939: 99 (characters in key).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, US Virgin Isl., Antigua & Barbuda, Guadeloupe, St. Lucia, Saint
Vincent & the Grenadines, Grenada, Trinidad-Tobago.
South America: Galapagos Isl., Peru, Brazil, Argentina.
0876. Labrocharis Bierig, 1933: 494

Type species: Labrocharis obsoleta Bierig, 1933
Subgenera:
Labrocharis Bierig, 1933: 494 [*]
Type species: Labrocharis obsoleta Bierig, 1933
Labroporus Bierig, 1933: 496 [*]
Type species: Labrocharis imitatrix Bierig, 1933

Distribution.

0877. Leiporaphes Bernhauer, 1912c: 37
Type species: Medon attarum Bernhauer, 1912

Distribution.
0878. Lithocharis Dejean, 1833: 65

Type species: Paederus ochraceus Gravenhorst, 1802
Metaxyodonta Casey, 1886a: 29 [synonymized by Bernhauer & Schubert 1912: 231]
Type species: Metaxyodonta alutacea Casey, 1886

Blackwelder 1943: 230 (characters in key), 239 (characters); Coiffait & Sáiz 1968: 379 (characters in key);
Navarrete-Heredia et al. 2002: 268 (characters in key); Newton & Peck 2006: 149 (characters in key).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Saint-Kitts &
Nevis, Antigua & Barbuda, Montserrat, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent &
Grenadines, Barbados, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Galapagos Isl., Peru, Bolivia, Brazil, Paraguay, Chile, Argentina.
0879. Lypeticus Sharp, 1886: 556

Type species: Lithocharis munda Sharp, 1876
Distribution.
0880. Medon Stephens, 1833a: 273

Type species: Medon ruddii Stephens, 1833
Subgenera:
Cryptoporus Motschulsky, 1858d: 654
Type species: Cryptoporus flavipes Motschulsky, 1858
Medon Stephens, 1833a: 273 [*]
Type species: Medon ruddii Stephens, 1833
Oxymedon Casey, 1905: 177 [synonymized by Blackwelder 1939: 104]
Type species: Oxymedon rubrum Casey, 1905
Medonodonta Casey, 1905: 176
Type species: Medonodonta alutacea Casey, 1905
Mespalerus Sharp, 1886: 560 [*]
Type species: Mespalerus debilis Sharp, 1886
Nitimedon Hatch, 1957: 158
Type species: Medon quadratum Hatch, 1957
Platymedon Casey, 1889b: 184
Type species: Platymedon laticolle Casey, 1889
Paramedon Casey, 1905: 166 [synonymized by Blackwelder 1939: 104]
Type species: Paramedon arizonicum Casey, 1905
Tetramedon Casey, 1905: 178
Type species: Tetramedon rufipenne Casey, 1905

(characters); Coiffait & Sáiz 1968: 380 (characters in key), 386 (characters); Frania 1987: 2553 (characters,
larvae); Navarrete-Heredia et al. 2002: 268 (characters in key).
Key: Blackwelder 1939: 99 (subgenera); Blackwelder 1943: 270 (species of the West Indies); Coiffait & Sáiz
1968: 387 (Chilean species).
Distribution.
West Indies: Cuba, Haiti, Dominican Republic, Dominica, St. Lucia, Trinidad-Tobago.
South America: Peru, Bolivia, Brazil, Chile, Argentina.
0881. Monocharis Sharp, 1886: 569

Type species: Lithocharis vestita Sharp, 1876
Distribution.
0882. Myrmecomedon Bernhauer, 1912c: 35

Type species: Myrmecomedon bruchi Bernhauer, 1912

Distribution.
0883. Neomedon Sharp, 1886: 557

Type species: Neomedon princeps Sharp, 1886

Characters/Description: Blackwelder 1939: 99 (characters in key); Navarrete-Heredia et al. 2002: 268 (characters
in key).
Distribution.
0884. Ophiomedon Sharp, 1886: 567

Type species: Ophiomedon stipes Sharp, 1886

Distribution.
Central America: Guatemala, Nicaragua, Costa Rica, Panama.
Other regions: Australian, Oceanic.
0885. Pseudomedon Mulsant & Rey, 1878b: 166

Type species: Lathrobium obsoletum Nordmann, 1837
Euphonus Fauvel, 1902c: 181 [synonymized by Assing 2009a: 1183]
Type species: Euphonus pallidus Fauvel, 1902
Pseudomedon Mulsant & Rey, 1878c: 166 [synonymized by Blackwelder 1952: 327–328]
Ramona Casey, 1886b: 213
Type species: Ramona capitulum Casey, 1886
Distribution.
Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Australian.
0886. Sciocharis Lynch-Arribálzaga, 1884: 260

Type species: Sciocharis castanoptera Lynch-Arribálzaga, 1884
Sciocharella Casey, 1905: 158 [synonymized by Scheerpeltz 1970a: 229, 244]
Type species: Sciocharella delicatula Casey, 1905

Characters/Description: Lynch-Arribálzaga 1884: 260 (characters); Bernhauer 1921d: 35 (characters).
Key: Lynch-Arribálzaga 1884: 263 (species of Buenos Aires province, Argentina); Scheerpeltz 1970a: 230
(Central and South American species).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Puerto Rico, Antigua & Barbuda, Dominica, Dominican Republic, St. Lucia,
St. Vincent & the Grenadines, Grenada, Trinidad-Tobago.
Central America: Mexico, Guatemala, El Salvador, Costa Rica, Nicaragua, Panama.
South America: Colombia, Venezuela, Suriname, Peru, Bolivia, Brazil, Paraguay, Argentina.
0887. Scioporus Sharp, 1886: 561 (Fig. 172)

Type species: Scioporus brunneus Sharp, 1886
Distribution.
Central America: Guatemala, Nicaragua, Panama.
South America: Colombia, Venezuela, Peru, Brazil.
0888. Scopobium Blackwelder, 1939: 97
Type species: Ophiomedon anthracinum Cameron, 1913
(characters); Navarrete-Heredia et al. 2002: 268 (characters in key).
Distribution.
West Indies: St. Lucia, St. Vincent & Grenadines, Grenada.
0889. Stilocharis Sharp, 1886: 576

Type species: Stilocharis longula Sharp, 1886

Characters/Description: Frania 1986: 475 (characters); Frania 1987: 2553 (characters, larvae); Navarrete-Heredia
Distribution.
0890. Stilomedon Sharp, 1886: 565

Type species: Lithocharis convexa Sharp, 1876

Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Guadeloupe, Dominica, St.
Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
Central America: Mexico, Guatemala, Belize, Costa Rica, Panama.
0891. Suniotrichus Sharp, 1886: 587

Type species: Suniotrichus capillaris Sharp, 1886
in key).
Distribution.
0892. Sunius Stephens, 1829a: 24

Type species: Paederus melanocephalus Fabricius, 1792
Tetracanthognathus Scheerpeltz, 1963b: 432 [synonymized by Assing 2010: 1052]
Type species: Tetracanthognathus kuehnelti Scheerpeltz, 1963
Subgenera:
Caloderma Casey, 1886a: 5 [*]
Type species: Caloderma rugosum Casey, 1886
Sunius Stephens, 1829a: 24

Type species: Paederus melanocephalus Fabricius, 1792
Medonella Casey, 1905: 178 [synonymized by Blackwelder 1939: 103]
Type species: Medonella minuta Casey, 1905
Micromedon Casey, 1905: 155 [synonymized by Blackwelder 1939: 103]
Type species: Lithocharis seminigra Fairmaire, 1860
Oligopterus Casey, 1886a: 12 [synonymized by Blackwelder 1944: 118]
Type species: Oligopterus cuneicollis Casey, 1886
Xenocharis Bierig, 1934c: 328 [synonymized by Blackwelder 1994: 118]
Type species: Xenocharis occipitalis Bierig, 1934
Trachysectus Casey, 1886a: 32
Type species: Lathrobium confluentum Say, 1834

Characters/Description: Lynch-Arribálzaga 1884: 297 (characters); Blackwelder 1939: 99 (characters in key,
Hypomedon); Blackwelder 1943: 230 (characters in key), 259 (characters); Navarrete-Heredia et al. 2002: 268
Distribution.
West Indies: Cuba, Haiti, Dominican Republic, Puerto Rico, St. Lucia, St. Vincent & Grenadines, Grenada,
Trinidad-Tobago.
South America: Colombia, Venezuela, Galapagos Isl.
0893. Thinocharis Kraatz, 1859a: 142

Type species: Thinocharis pygmaea Kraatz, 1859

Distribution.
Subtribe Scopaeina Mulsant & Rey, 1878c
0894. Euscopaeus Sharp, 1886: 548

Type species: Euscopaeus crassitarsis Sharp, 1886

Characters/Description: Coiffait 1960b: 284 (characters in key).
Distribution.
0895. Micranops Cameron, 1913: 350

Type species: Micranops brunneus Cameron, 1913
Microscopaeus Coiffait, 1981a: 19 [synonymized by Frisch et al. 2002: 46]
Type species: Scopaeus microphthalmus Eppelsheim, 1888
Nivorus Herman, 1965a: 119 [synonymized by Frisch et al. 2002: 46]
Type species: Orus cameroni Blackwelder, 1943
Characters/Description: Blackwelder 1939: 98 (characters in key); Frisch et al. 2002: 28 (characters).
Key: Herman 1965a: 120 (species of Nivorus); Herman 1965b: 86 (species of Nivorus).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Grenada.
South America: Suriname, Brazil, Argentina.
0896. Orus Casey, 1885b: 136 (Fig. 166)

Type species: Orus punctatus Casey, 1885
Subgenera:
Leucorus Casey, 1905: 192 [*]
Type species: Leucorus rubens Casey, 1905
Orus Casey, 1885b: 136
Type species: Orus punctatus Casey, 1884
Pycnorus Casey, 1905: 194
Type species: Scopaeus dentiger LeConte, 1880
(characters); Herman 1965b: 74 (characters); Navarrete-Heredia et al. 2002: 268 (characters in key); Frisch et
al. 2002: 28 (characters).
Key: Blackwelder 1939: 98 (subgenera); Blackwelder 1943: 277 (species of the West Indies as Orus); Herman
1965a: 113 (subgenera), 114 (species of the subgenus Leucorus); Herman 1965b: 74 (species of the subgenera
Orus and Pycnorus).
Distribution.
0897. Scopaeomerus Sharp, 1886: 538

Type species: Scopaeomerus chiriquensis Sharp, 1886

Distribution.
0898. Scopaeus Erichson, 1839a: 29

Type species: Paederus laevigatus Gyllenhal, 1827
Alloscopaeus Coiffait, 1968: 414 [synonymized by Frisch et al. 2002: 46]
Type species: Scopaeus didymus Erichson, 1840
Anomoscopaeus Coiffait, 1968: 426 [synonymized by Frisch et al. 2002: 46]
Type species: Xantholinus gracilis Sperk, 1835
Euscopaeus Coiffait, 1960b: 285 [synonymized by Coiffait 1968: 414]
Type species: Scopaeus didymus Erichson, 1840
Geoscopaeus Coiffait, 1960b: 284 [synonymized by Frisch et al. 2002: 46]

Type species: Scopaeus baudrimonti Coiffait, 1952
Heteroscopaeus Coiffait, 1960b: 285 [synonymized by Coiffait 1968: 418]
Type species: Scopaeus sericans Mulsant & Rey, 1855
Hyposcopaeus Coiffait, 1960b: 285 [synonymized by Frisch et al. 2002: 46]
Type species: Scopaeus scitulus Baudi di Selve, 1857
Stilpon Coiffait, 1952: 6 [synonymized by Coiffait 1960b: 284]
Type species: Scopaeus baudrimonti Coiffait, 1952
Subgenera:
Asiascopaeus Coiffait, 1984: 176
Type species: Scopaeus asiaticus Bernhauer, 1915
Hyperscopaeus Coiffait, 1984: 174
Type species: Scopaeus spathiferus Coiffait, 1970
Scopaeodera Casey, 1886b: 220 [*]
Type species: Echiaster nitidus LeConte, 1863
Scopaeoma Casey, 1905: 211
Type species: Scopaeus rotundiceps Casey, 1886
Scopaeopsis Casey, 1905: 214 [*]
Type species: Echiaster opacus LeConte, 1863
Scopaeus Erichson, 1839a: 29, 604 [*]
Type species: Paederus laevigatus Gyllenhal, 1827
Leptorus Casey, 1886b: 220 [synonymized by Blackwelder 1939: 105]
Type species: Scopaeus exiguus Erichson, 1840
Polyodontus Solier, 1849: 310 [synonymized by Blackwelder 1939: 105]
Type species: Polyodontus angustatus Solier, 1849
Pseudorus Casey, 1910b: 190 [synonymized by Blackwelder 1939: 105]
Type species: Pseudorus prolixipennis Casey, 1910
Scoponeus Motschulsky, 1858d: 641 [synonymized by Kraatz 1862b: 61]
Type species: Scoponeus testaceus Motschulsky, 1858
Typhloscopaeus Jarrige, 1951: 333
Type species: Scopaeus pauliani Jarrige, 1951

Characters/Description: Erichson 1840: 604 (characters); Lynch-Arribálzaga 1884: 245 (characters); Blackwelder
1939: 100 (characters in key); Blackwelder 1943: 230 (characters in key), 279 (characters); Coiffait 1960b:
284 (characters in key as Geoscopaeus), 284 (characters in key of Scopaeus); Coiffait & Sáiz 1968: 380
(characters in key), 389 (characters); Navarrete-Heredia et al. 2002: 267 (characters in key); Frisch et al.
2002: 28 (characters); Newton & Peck 2006: 148 (characters in key).
Key: Lynch-Arribálzaga 1884: 246 (species of Buenos Aires province, Argentina); Blackwelder 1939: 100
(subgenera); Blackwelder 1943: 280 (species of the West Indies); Coiffait & Sáiz 1968: 389 (Chilean
species).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Guadeloupe,
St. Lucia, St. Vincent & the Grenadines, Barbados, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Galapagos Isl., Peru, Brazil, Chile, Argentina.
Subtribe Stilicina Casey, 1905
0899. Eustilicus Sharp, 1886: 579 (Fig. 159)

Type species: Eustilicus crassidens Sharp, 1886
Omostilicus Casey, 1905: 229 [synonymized by Blackwelder 1939: 106]
Type species: Omostilicus sonorinus Casey, 1905
Stilicolina Casey, 1905: 228 [synonymized by Frania 1986: 472]
Type species: Stilicus tristis Melsheimer, 1844
Trochoderus Sharp, 1886: 580 [*] [synonymized by Frania 1986: 472]
Type species: Trochoderus dubius Sharp, 1886

Characters/Description: Frania 1986: 472 (characters); Frania 1987: 2544 (characters of larvae); Navarrete-Heredia
Key: Herman 1970b: 8 (species of Stilicolina).
Distribution.
0900. Rugilus Leach in Samouelle, 1819: 173

Type species: Staphylinus orbiculatus Paykull, 1789
Subgenera:
Afrostilicus Fagel, 1953: 284
Type species: Stilicus confusus Fagel, 1951
Allostilicus Fagel, 1953: 285
Type species: Stilicus uniformis Bernhauer, 1915
Cephalostilicus Fagel, 1950: 1
Type species: Stilicus schimperi Fauvel, 1907
Eurystilicus Fagel, 1953: 284
Type species: Stilicus rubelloides Fagel, 1951
Fossulostilicus Fagel, 1955a: 37
Type species: Stilicus punctipennis Bernhauer, 1915
Eustilicus Fagel, 1953: 285 [synonymized by Fagel 1955a: 37]
Type species: Stilicus punctipennis Bernhauer, 1915
Hemistilicus Fagel, 1951: 228
Type species: Stilicus inermis Fagel, 1951
Heterostilicus Fagel, 1953: 284
Type species: Stilicus dubiosus Fagel, 1953
Mirostilicus Fagel, 1952: 13
Type species: Stilicus anommatophilus Wasmann, 1904
Parastilicus Jeannel & Paulian, 1945: 72
Type species: Stilicus nidicola Bernhauer, 1944
Pedistilicus Fagel, 1951: 223
Type species: Stilicus pholoeophilus Fagel, 1951
Rugilus Leach in Samouelle, 1819: 173
Sepedomorphus Gistel, 1834: 9 [synonymized by Blackwelder 1952: 350]

Stilicosoma Casey, 1905: 219 [synonymized by Löbl & Smetana 2004: 619]
Type species: Rugilus rufipes Germar, 1836
Stilicus Berthold, 1827: 331 [synonymized by Blackwelder 1952: 362]
Stilidentus Coiffait, 1978a: 129 [synonymized by Rougemont 1987: 211]
Type species: Stilicus dorjulensis Coiffait, 1978
Tetragnathostilicus Scheerpeltz, 1976b: 118 [synonymized by Assing 2012c: 118]
Type species: Stilicus gracilis Eppelsheim, 1895
Scabrostilicus Fagel, 1953: 283
Type species: Stilicus crassus Fagel, 1951

Characters/Description: Lynch-Arribálzaga 1884: 272 (characters as Stilicus); Bernhauer 1921b: 173 (characters as
Stilicus); Blackwelder 1939: 100 (characters in key as Stilicus); Blackwelder 1943: 230 (characters in key),
299 (characters); Coiffait & Sáiz 1968: 380 (characters in key as Stilicus), 385 (characters, Stilicus); Frania
1986: 476 (characters); Frania 1987: 2554 (characters of larvae); Navarrete-Heredia et al. 2002: 267
Key: Lynch-Arribálzaga 1884: 273 (as Stilicus of Buenos Aires province, Argentina); Bernhauer 1921b: 173 (as
Stilicus); Bernhauer 1939b: 245 (as Stilicus of Argentina); Blackwelder 1943: 300 (species of the West Indies).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Puerto Rico, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, French Guiana, Peru, Bolivia, Brazil, Uruguay, Chile, Argentina.
Subtribe Stilicopsina Casey, 1905
0901. Dibelonetes R.F. Sahlberg, 1847: 791 (Fig. 155)

Type species: Dibelonetes biplagiatus R.F. Sahlberg, 1847
Sunides Motschulsky, 1858d: 638 [synonymized by Bernhauer & Schubert 1912: 212]
Type species: Sunides boreophiloides Motschulsky, 1858
Subgenera:
Brachynetes Bernhauer, 1922a: 12 [*]
Type species: Dibelonetes apterus Bernhauer, 1922
Apteronetes Bierig, 1933: 516 [synonymized by Blackwelder 1952 : 60]
Type species: Dibelonetes apterus Bernhauer, 1921
Dibelonetes R.F. Sahlberg, 1847: 791
Type species: Dibelonetes biplagiatus R.F. Sahlberg, 1847
Heteronetes Bierig, 1933: 511 [*]
Type species: Dibelonetes vulcanus Bierig, 1933
Melanetes Bierig, 1933: 515 [*]
Type species: Dibelonetes laticeps Sharp, 1886
in key).
Distribution.
Other regions: Oriental, Australian, Oceanic.
0902. Dibelophacis Bierig, 1933: 508

Type species: Dibelophacis horni Bierig, 1933

Distribution.
0903. Stamnoderus Sharp, 1886: 607

Type species: Stamnoderus championi Sharp, 1886

Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Guadeloupe,
Dominica, St. Vincent & Grenadines, Grenada.
0904. Stilicopsis Sachse, 1852: 144

Type species: Stilicopsis paradoxa Sachse, 1852
One species is included in this genus from Galapagos Isl.

in key); Newton & Peck 2006: 148 (characters in key).
Distribution.
South America: Galapagos Isl.
0905. Stiliphacis Bierig, 1938b: 142

Type species: Stiliphacis occipitalis Bierig, 1938
356 (characters).
Distribution.
West Indies: Cuba, Jamaica, Puerto Rico, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
0906. Stilosaurus Blackwelder, 1943: 348

Type species: Stilosaurus rulomus Blackwelder, 1943

Distribution.
West Indies: Cuba, Puerto Rico, Dominican Republic.
0907. Suniocharis Sharp, 1886: 586

Type species: Suniocharis modesta Sharp, 1886
Subgenera:
Parasuniocharis Bernhauer, 1933b: 519 [*]
Type species: Suniocharis boxi Bernhauer, 1933
Suniocharis Sharp, 1886: 586 [*]
Type species: Suniocharis modesta Sharp, 1886

Distribution.
Other regions: Madagascan.
0908. Suniophacis Blackwelder, 1943: 345

Type species: Suniophacis concolor Blackwelder, 1943
Distribution.
West Indies: Cuba, Jamaica, Haiti, Antigua & Barbuda.
0909. Suniosaurus Bierig, 1938b: 139

Type species: Suniosaurus cuadriceps Bierig, 1938
Characters/Description: Blackwelder 1943: 231 (characters in key), 365 (characters).
Distribution.
West Indies: Cuba.
0910. Xenasterides Newton, 2017: 17

Type species: Xenaster plaumanni Bierig, 1939
Xenaster Bierig, 1939b: 179 [synonymized by Newton 2017: 17]
Type species: Xenaster plaumanni Bierig, 1939

Distribution.
Tribe Paederini Fleming, 1821
Subtribe Cryptobiina Casey, 1905
0911. Aderobium Casey, 1905: 23

Type species: Cryptobium angustifrons Sharp, 1876

Characters/Description: Casey 1905: 28 (characters); Blackwelder 1939: 95 (characters in key).
Distribution.
0912. Biocrypta Casey, 1905: 51

Type species: Cryptobium prospiciens LeConte in Schwarz, 1878

Characters/Description: Casey 1905: 26 (characters in key); Blackwelder 1939: 97 (characters in key);
Blackwelder 1943: 230 (characters in key), 335 (characters); Navarrete-Heredia et al. 2002: 266 (characters in
key).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, Antigua & Barbuda, Montserrat,
Guadeloupe, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Ecuador, Brazil, Argentina.
0913. Homaeotarsus Hochhuth, 1851: 34

Type species: Homaeotarsus chaudoirii Hochhuth, 1851
Spirosoma Motschulsky, 1858b: 206 [synonymized by Blackwelder 1939: 113]
Type species: Spirosoma fulvescens Motschulsky, 1858
Subgenera:
Eucryptina Casey, 1905: 28 [*]
Type species: Cryptobium opacum Sharp, 1876
Gastrolobium Casey, 1905: 31 [*]
Type species: Lathrobium bicolor Gravenhorst, 1802
Hesperobium Casey, 1886a: 33
Type species: Cryptobium tumidum LeConte in Schwarz, 1878
Homaeotarsus Hochhuth, 1851: 34
Type species: Homaeotarsus chaudoirii Hochhuth, 1851
Homoeobium Blackwelder, 1939: 96
Type species: Homaeotarsus bakerianus Blackwelder, 1939
Nemoeotus Blackwelder, 1939: 96
Type species: Cryptobium rubiginosum Bernhauer, 1912

Characters/Description: Casey 1905: 24 (characters in key as Homaeotarsus and Eucryptina), 28 (characters as
Homaeotarsus and Eucryptina); Blackwelder 1939: 95 (characters in key); Blackwelder 1943: 230 (characters

in key), 325 (characters); Navarrete-Heredia et al. 2002: 266 (characters in key); Newton & Peck 2006: 148
Key: Blackwelder 1939: 96 (subgenera); Blackwelder 1943: 326 (species of the West Indies).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, St. Vincent & Grenadines,
South America: Colombia, Venezuela, Galapagos Isl., Bolivia, Brazil, Paraguay, Argentina.
Other regions: Nearctic, Palaearctic, Madagascan, Oriental.
0914. Ochthephilum Stephens, 1829a: 24 (Fig. 164)

Type species: Paederus fracticornis Paykull, 1800
Subgenera:
Ababactus Sharp, 1885: 533 [*]
Type species: Ababactus depressus Sharp, 1886
Astenobium Bernhauer, 1911a: 411 [*]
Type species: Cryptobium excellens Bernhauer, 1911
Cryptobiella Casey, 1905: 29 [*]
Type species: Cryptobiella colonica Casey, 1905
Neobactus Blackwelder, 1939: 96
Type species: Cryptobium nunenmacheri Blackwelder, 1939
Ochthephilum Stephens, 1829a: 24
Cryptobium Mannerheim, 1830: 38 [synonymized by Blackwelder 1943: 331]
Epimachus Gistel, 1834: 8 [synonymized by Blackwelder 1952: 150]

Characters/Description: Casey 1905: 29 (characters as Cryptobiella); Lynch-Arribálzaga 1884: 202 (characters as
Cryptobium); Blackwelder 1939: 96 (characters in key as Cryptobium); Blackwelder 1943: 230 (characters in
key), 331 (characters); Navarrete-Heredia et al. 2002: 266 (characters in key).
Distribution.
West Indies: Cuba, Guadeloupe, Trinidad-Tobago.
South America: Colombia, Venezuela, French Guiana, Peru, Bolivia, Brazil, Paraguay, Uruguay, Chile,
Argentina.
0915. Opithes Blackwelder, 1952: 277

Type species: Ophites versatilis Erichson, 1840
Ophites Erichson, 1839a: 29 [synonymized by Blackwelder 1952: 277]
Type species: Ophites versatilis Erichson, 1840
Characters/Description: Lynch-Arribálzaga 1884: 228 (characters); Erichson 1840: 627 (characters); Blackwelder
1939: 95 (characters in key as Ophites); Blackwelder 1943: 230 (characters in key as Ophites), 343
(characters as Ophites); Drugmand 2000: 17 (characters); Navarrete-Heredia et al. 2002: 266 (characters in
key).
Key: Drugmand 2000: 21 (species of the genus).
Distribution.
South America: Colombia, Brazil, Argentina.
0916. Pycnocrypta Casey, 1905: 25 (Fig. 170)

Type species: Cryptobium maxilosum Guérin-Méneville, 1844
Characters/Description: Casey 1905: 28 (characters); Bierig 1933: 476 (characters); Blackwelder 1939: 97
(characters in key); Navarrete-Heredia et al. 2002: 266 (characters in key).
Distribution.
0917. Scopaeodes Sharp, 1876a: 208

Type species: Scopaeodes gracilis Sharp, 1876

Characters/Description: Casey 1905: 55 (characters in key); Blackwelder 1939: 95 (characters in key).
Distribution.
Subtribe Dicaxina Schomann & Solodovnikov, 2017
0918. Baryopsis Fairmaire & Germain, 1862: 442 (Fig. 153)

Type species: Baryopsis brevipennis Fairmaire & Germain, 1862

Characters/Description: Fauvel 1868b: 7 (characters); Coiffait & Sáiz 1968: 379 (characters in key), 393
(characters); Sáiz 1973a: 131 (characters); Assing 2012a: 335 (characters).
Key: Coiffait & Sáiz 1968: 394 (Chilean species); Sáiz 1979: 125 (Chilean species); Assing 2012a: 344 (Peruvian
species).
Distribution.
South America: Peru, Bolivia, Chile.
0919. Pseudocryptobium Bernhauer, 1921a: 74

Type species: Latona spinolae Guérin-Méneville, 1844
Latona Guérin-Méneville, 1844: 13 [synonymized by Bernhauer 1921a: 74]
Type species: Latona spinolae Guérin-Méneville, 1844

Characters/Description: Guérin-Méneville 1844: 13 (characters as Latona).
Distribution.
Subtribe Dolicaonina Casey, 1905
0920. Acaratopus Herman, 1981: 370

Type species: Acaratopus edenus Herman, 1981

Key: Herman 1981: 359 (species of the genus).
Distribution.
0921. Gnathymenus Solier, 1849: 326 (Fig. 160)

Type species: Gnathymenus apterus Solier, 1849
Lithocaon Sharp, 1886: 555 [synonymized by Herman 1981: 373]
Type species: Lithocaon sparsus Sharp, 188
Litozoon Bierig, 1939b: 176 [synonymized by Herman 1981: 373]
Type species: Litozoon progenitor Bierig, 1939
Macrognathymenus Coiffait & Sáiz, 1968: 374 [synonymized by Herman 1981: 373]
Type species: Gnathymenus obesus Fauvel, 1868
Xanthornobium Scheerpeltz, 1967: 237 [synonymized by Herman 1981: 373]
Type species: Xanthornobium vogelsangi Scheerpeltz, 1967
79c species are known from and are restricted to Central and South America.
Characters/Description: Coiffait & Sáiz 1968: 370 (characters); Herman 1981: 373 (characters); Navarrete-Heredia
Key: Coiffait & Sáiz 1968: 373 (subgenera of Chile), 373 (Chilean species of Gnathymemus), 374 (species of Chile
as Macrognathymenus), 375 (subgenera of Chile), 375 (Chilean species of Gnathymemus); Herman 1981: 359
(species of the genus); Bortoluzzi & Caron 2018: 169 (species of the genus).
Distribution.
South America: Colombia, Venezuela, Ecuador, Bolivia, Brazil, Chile.
0922. Stenopholea Herman, 1969: 2 (Fig. 173)

Type species: Stenopholea reddelli Herman, 1969
Characters/Description: Herman 1981: 495 (characters); Navarrete-Heredia et al. 2002: 266 (characters in key).
Distribution.
Subtribe Paederina Fleming, 1821
0923. Eupaederus Scheerpeltz, 1957a: 269

Type species: Paederus cyanipennis Guérin-Méneville, 1830
Only a single species is included in this genus from South America.

Distribution.
South America: Peru, Chile.
Other regions: Oriental.
0924. Paederus Fabricius, 1775: 268 (Fig. 167)

Type species: Staphylinus riparius Linnaeus, 1758
Subgenera:
Anomalopaederus Scheerpeltz, 1966a: 400
Type species: Paederus lacordairei Perroud, 1864
Eopaederus Scheerpeltz, 1957b: 458
Type species: Paederus caligatus Erichson, 1840
Gnathopaederus Chapin, 1927: 75
Type species: Gnathopaederus szechuanus Chapin, 1927
Harpopaederus Scheerpeltz, 1957b: 459
Type species: Paederus schoenherri Czwalina, 1889
Oedopaederus Scheerpeltz, 1957b: 466 [synonymized by Assing 2015b: 184]
Type species: Paederus chinensis Bernhauer, 1931
Heteropaederus Scheerpeltz, 1957b: 460 [*]
Type species: Paederus fuscipes Curtis, 1826
Neopaederus Blackwelder, 1939: 97 [*]
Type species: Paederus morio Mannerheim, 1830
Nepalopaederus Scheerpeltz, 1976b: 111
Type species: Paederus nepalensis Scheerpeltz, 1976
Oreinopaederus Scheerpeltz, 1976b: 112
Type species: Paederus vastus Scheerpeltz, 1976
Paederognathus Wendeler, 1928: 37 [*]
Type species: Paederus turrialbanus Wendeler, 1927
Gnathopaederus Wendeler, 1927: 1 [synonymized by Wendeler 1928: 37]
Paederognathus Wendeler, 1931: 43 [synonymized by Blackwelder 1952: 286]
Paederus Fabricius, 1775: 268 [*]
Geopaederus Gistel, 1848: x [synonymized by Blackwelder 1952: 169]
Leucopaederus Casey, 1905: 67 [synonymized by Blackwelder 1939: 111]
Type species: Paederus ustus LeConte, 1858
Paederillus Casey, 1905: 62 [synonymized by Blackwelder 1939: 111]
Type species: Paederus littorarius Gravenhorst, 1806
Poederomorphus Gautier des Cottes, 1862: 75
Type species: Poederomorphus pedoncularius Gautier des Cottes, 1862
Dioncopaederus Scheerpeltz, 1957b: 464 [synonymized by Herman 2003: 5]

Type species: Paederus littoralis Gravenhorst, 1802
Pseudopaederus Bernhauer, 1915a: 137
Type species: Paederus nigerrimus Bernhauer, 1915

Blackwelder, 1943: 230 (characters in key), 321 (characters); Navarrete-Heredia et al. 2002: 266 (characters
in key).
of the West Indies); Scheerpeltz 1965a: 214 (Venezuelan species); Sanderson 1967: 59 (species of the Greater
Antillean); Vieira et al. 2014: 246 (Brazilian species of medical importance).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Martinique, St. Vincent & Grenadines, Trinidad-Tobago.
Uruguay, Argentina.
Incertae sedis in Tribe Paederini Fleming, 1821
0925. Bolbophites Fauvel, 1904b: 278

Type species: Bolbophites aspericeps Fauvel, 1904

Distribution.
0926. Ecitobium Wasmann, 1923: lxiii

Type species: Ecitobium zikani Wasmann, 1923

Characters/Description: Fischer 1943: 256 (characters); Seevers 1965: 320 (characters).
Distribution.
0927. Ecitonides Wasmann, 1894: 212 (Fig. 157)

Type species: Ecitonides tuberculosus Wasmann, 1894

Characters/Description: Wasmann 1896a: 150 (characters); Wasmann 1900b: 247 (characters); Borgmeier 1949:
121 (characters); Seevers 1965: 317 (characters).
Key: Wasmann 1900b: 248 (species of the genus); Borgmeier 1949: 122 (species of the genus).
Distribution.
South America: Peru, Brazil, Paraguay, Argentina.
0928. Ecitosaurus Fischer, 1943: 259
Type species: Labidosaurus lujae Wasmann, 1925
Labidosaurus Wasmann, 1925b: 49 [synonymized by Fischer 1943: 259]
Type species: Labidosaurus lujae Wasmann, 1925

Characters/Description: Borgmeier 1931: 355 (characters as Labidosaurus); Seevers 1965: 321 (characters).
Distribution.
0929. Ecitotropis Borgmeier, 1936: 297 (Fig. 158)

Type species: Ecitotropis carinata Borgmeier, 1936
Distribution.
0930. Labidophites Borgmeier, 1956: 14

Type species: Labidophites cariniceps Borgmeier, 1956

Distribution.
0931. Mimophites Fauvel, 1904b: 280

Type species: Mimophites bouvieri Fauvel, 1904
Characters/Description: Borgmeier 1949: 116 (characters); Borgmeier 1956: 1 (characters); Seevers 1965: 316
Key: Borgmeier 1949: 116 (species of the genus); Borgmeier 1956: 6 (species of the genus).
Distribution.
0932. Monista Sharp, 1876a: 271 (Fig. 161)

Type species: Monista typica Sharp, 1876

Blackwelder 1943: 230 (characters in key), 296 (characters); Navarrete-Heredia et al. 2002: 267 (characters in
key).
Distribution.
West Indies: Grenada, Dominica, Trinidad-Tobago.

South America: Colombia, Venezuela, Peru, Bolivia, Brazil, Argentina.
0933. Synecitonides Reichensperger, 1936a: 236

Type species: Synecitonides phasma Reichensperger, 1936

Characters/Description: Seevers 1965: 319 (characters); Navarrete-Heredia et al. 2002: 267 (characters in key).
Distribution.
Tribe Pinophilini Nordmann, 1837
Subtribe Pinophilina Nordmann, 1837
0934. Araeocerus Nordmann, 1837: 7, 157

Type species: Araeocerus niger Nordmann, 1837
Scotocerus Bernhauer, 1918: 67 [synonymized by Blackwelder 1943: 386]
Type species: Araeocerus curtipennis Bernhauer, 1918
Characters/Description: Blackwelder 1943: 229 (characters in key), 386 (characters); Abarbanell & Ashe 1989: 1-
Distribution.
West Indies: Cuba.
South America: Brazil, Uruguay, Argentina.
Other regions: Nearctic, Australian.
0935. Lathropinus Sharp, 1886: 628

Type species: Lathropinus parrallelus Sharp, 1886
Characters/Description: Abarbanell & Ashe 1989: 1–2 (characters); Navarrete-Heredia et al. 2002: 266 (characters
in key).
Distribution.
South America: Colombia, Ecuador, Peru, Brazil, Paraguay, Uruguay, Argentina.
0936. Pinophilus Gravenhorst, 1802: 201

Type species: Pinophilus latipes Gravenhorst, 1802
Heteroleucus Sharp, 1886: 629 [synonymized by Blackwelder 1952: 184]
Type species: Heteroleucus marginatus Sharp, 1886
Pityophilus Brullé in Audouin, 1837: 75 [synonymized by Bernhauer & Schubert 1912: 191]
Type species: Pinophilus latipes Gravenhorst, 1802
Characters/Description: Sharp 1883: 620 (characters); Lynch-Arribálzaga 1884: 305 (characters); Blackwelder
1943: 229 (characters in key), 376 (characters); Abarbanell & Ashe 1989: 3 (characters); Navarrete-Heredia et
al. 2002: 266 (characters in key).
of the West Indies); Abarbanell & Ashe 1989: 4 (species of the America north of Mexico).
Distribution.
West Indies: Bahamas, Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, Guadeloupe, St.
Vincent & the Grenadines, Trinidad-Tobago.
South America: Colombia, Venezuela, French Guiana, Brazil, Paraguay, Uruguay, Argentina.
0937. Pinostygus Campbell & Peck, 1989: 397 (Fig. 169)

Type species: Pinostygus galapagoensis Campbell & Peck, 1989
Only one species is known from and is restricted to Galapagos Isl.

Characters/Description: Newton & Peck 2006: 148 (characters in key).
Distribution.
0938. Taenodema Laporte, 1835: 120 (Fig. 175)

Type species: Paederus semicyaneus Perty, 1830
Gymnurus Nordmann, 1837: 158 [synonymized by Bernhauer & Schubert 1912: 196]
Type species: Gymnurus cyanescens Nordmann, 1837
Taenodemiella Bernhauer, 1923a: 51 [synonymized by Blackwelder 1952: 372]
Type species: Taenodema corumbanum Bernhauer, 1923
Characters/Description: Sharp 1886: 618 (characters); Erichson 1840: 679 (characters); Navarrete-Heredia et al.
Distribution.
South America: Colombia, Venezuela, Suriname, French Guiana, Peru, Bolivia, Brazil, Paraguay.
Subtribe Procirrina Bernhauer & Schubert, 1912
0939. Oedichirus Erichson, 1839a: 29 (Fig. 165)

Type species: Oedichirus paederinus Erichson, 1840
Elytrobaeus R.F. Sahlberg, 1847: 801 [synonymized by Fauvel 1875: xvii]
Type species: Elytrobaeus geniculatus R.F. Sahlberg, 1847
Oedichiranus Reitter, 1906: 263 [synonymized by Herman 2010: 38]
Type species: Oedichirus dimidiatus Reitter, 1906
Characters/Description: Erichson 1840: 684 (characters); Navarrete-Heredia et al. 2002: 266 (characters in key);
Herman 2010: 36 (characters); Herman 2013: 9 (characters in key), 10 (characters).

Distribution.
South America: Ecuador, Peru, Bolivia, Brazil, Argentina.
0940. Oedodactylus Fairmaire & Germain, 1862: 441

Type species: Oedodactylus fuscobrunneus Fairmaire & Germain, 1862
Characters/Description: Coiffait & Sáiz 1968: 369 (characters); Navarrete-Heredia et al. 2002: 266 (characters in
key); Herman 2010: 46 (characters); Herman 2013: 9 (characters in key).
Distribution.
0941. Palaminus Erichson, 1839a: 29 (Fig. 168)

Type species: Palaminus pilosus Erichson, 1840
Parapalaminus Bierig, 1943: 155 [synonymized by Herman 2010: 52]
Type species: Palaminus symphylus Bierig, 1943

Characters/Description: Erichson 1840: 681 (characters); Lynch-Arribálzaga 1884: 327 (characters); Blackwelder
1943: 229 (characters in key), 388 (characters); Navarrete-Heredia et al. 2002: 266 (characters in key);
Herman 2010: 50 (characters); Herman 2013: 9 (characters in key).
Key: Notman 1929: 2 (species of the genus).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Puerto Rico, US Virgin Isl., Guadeloupe, St. Vincent & Grenadines, Grenada,
Trinidad-Tobago.
South America: Colombia, Venezuela, Suriname, French Guiana, Peru, Bolivia, Brazil, Argentina.
0942. Stylokyrtus Herman, 2010: 66

Type species: Oedodactylus errans Sharp, 1876

Characters/Description: Herman 2013: 9 (characters in key).
Distribution.
Subfamily STAPHYLININAE Latreille, 1802
Tribe Diochini Casey, 1906
0943. Antarctothius Coiffait & Sáiz, 1969: 279

Type species: Othius antarcticus Fairmaire, 1885
Key: Coiffait & Sáiz 1969: 281 (species of Tierra del Fuego, Chile).
Distribution.
0944. Diochus Erichson, 1839a: 300 (Fig. 188)

Type species: Diochus nanus Erichson, 1839
Rhegmatocerus Motschulsky, 1858d: 657 [synonymized by Gemminger & Harold 1868: 608]
Type species: Rhegmatocerus punctipennis Motschulsky, 1858

(characters); Smetana 1982: 27 (characters); Navarrete-Heredia et al. 2002: 295 (characters in key); Newton
& Peck 2006: 152 (characters in key).
Key: Irmler 2017a: 4 (Neotropical species groups of the genus), 57 (males of the Neotropical species), 59 (females
of the Neotropical species).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Dominican Republic, Puerto Rico, British Virgin Isl., Guadeloupe,
Dominica, Saint Vincent & the Grenadines, Barbados, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Guyana, French Guiana, Suriname, Ecuador, Galapagos Isl., Peru, Bolivia,
Brazil, Paraguay, Argentina.
Tribe Platyprosopini Lynch-Arribálzaga, 1884
0945. Platyprosopus Mannerheim, 1830: 8, 36 (Fig. 219)

Type species: Platyprosopus elongatus Mannerheim, 1830
Megaprosopoda Strand, 1935: 297 [synonymized by Bordoni 2015: 57]
Type species: Platyprosopus beduinus Nordmann, 1837
Megaprosopus Reitter, 1908: 104 [synonymized by Strand 1935: 297]
Type species: Platyprosopus beduinus Nordmann, 1837
Characters/Description: Lynch-Arribálzaga 1884: 166 (characters); Smetana 1982: 22 (characters); Navarrete-
Key: Irmler 1977: 58 (Neotropical species of the genus).
Distribution.
South America: Peru, Brazil, Paraguay, Argentina.
Tribe Staphylinini Latreille, 1802
Subtribe Acylophorina Outerelo & Gamarra, 1985
0946. Acylophorus Nordmann, 1837: 127

Type species: Acylophorus ahrensii Nordmann, 1837
Rhygmacera Motschulsky, 1845: 40 [synonymized by Motschulsky 1858d: 656]
Type species: Rhygmacera nitida Motschulsky, 1845
Subgenera:
Acylophorus Nordmann, 1837: 127
Type species: Acylophorus ahrensii Nordmann, 1837
Indoacylophorus Bierig, 1938a: 123
Type species: Acylophorus microcephalus Cameron, 1932
Neoacylophorus Bierig, 1938a: 123 [*]
Type species: Acylophorus oculifer Bierig, 1938
Palpacylophorus Smetana, 1971: 247, 268 [*]
Type species: Acylophorus leechi Smetana, 1971
Paracylophorus Bierig, 1938a: 123
Type species: Acylophorus schmidti Bierig, 1938

Characters/Description: Blackwelder 1943: 395 (characters in key), 466 (characters); Smetana 1971: 246
(characters), 250 (characters of the subgenus Acylophorus), 268 (characters of the subgenus
Palpacylophorus); Smetana 1988: 181 (characters in key), 337 (characters); Navarrete-Heredia et al. 2002:
298 (characters in key); Schillhammer & Brunke 2018: 311 (characters in key), 317 (characters).
Key: Blackwelder 1943: 467 (species of the West Indies); Smetana 1971: 247 (subgenera), 250 (species of the
subgenus Acylophorus); Smetana 1988: 339 (species of the genus).
Distribution.
West Indies: Cuba, Dominican Republic, Trinidad-Tobago.
South America: Colombia, French Guiana, Brazil, Paraguay, Argentina.
Subtribe Amblyopinina Seevers, 1944
0947. Amblyopinodes Seevers, 1955: 247

Type species: Amblyopinus gahani Fauvel, 1901
Subgenera:
Amblyopinodes Seevers, 1955: 247 [*]
Type species: Amblyopinus gahani Fauvel, 1901
Streptopinodes Martínez et al., 1970: 133 [*]
Type species: Amblyopinus longus E. Franz, 1930

Characters/Description: Seevers 1955: 247 (characters); Machado-Allison 1962a: 312 (characters); Machado-
Allison 1963: 380 (characters in key), 382 (characters); Ashe & Timm 1988: 49 (characters in key).
Key: Machado-Allison 1962a: 313 (Brazilian species of the genus); Machado-Allison 1962b: 82 (species of the
genus); Machado-Allison 1963: 384 (species of the genus).
Distribution.
South America: Venezuela, Peru, Bolivia, Brazil, Paraguay, Uruguay, Argentina.
0948. Amblyopinus Solsky, 1875: 10
Type species: Amblyopinus ielskii Solsky, 1875
Omaloxenus Notman, 1923: 1 [synonymized by Costa-Lima 1936: 58]
Type species: Omaloxenus bequaerti Notman, 1923
Characters/Description: Seevers 1955: 221 (characters); Coiffait & Sáiz 1968: 366 (characters in key); Ashe &
Timm 1988: 48 (characters in key); Ashe & Timm 1995: 375 (characters); Navarrete-Heredia et al. 2002: 298
Key: Seevers 1944: 160 (species of the genus); Seevers 1955: 224 (group of species); Machado-Allison 1963: 380
(characters in key); Ashe & Timm 1995: 376 (species of the Central American).
Distribution.
South America: Colombia, Venezuela, Guyana, Ecuador, Peru, Brazil, Chile, Argentina.
0949. Cheilocolpus Solier, 1849: 320 (Figs. 178–180)

Type species: Staphylinus pyrostoma Solier, 1849

Characters/Description: Coiffait & Sáiz 1966: 385 (characters in key), 386 (characters); Sáiz 1971b: 365
(characters); Smetana 1981: 354 (characters in key).
Key: Coiffait & Sáiz 1966: 387 (Chilean species); Sáiz 1971b: 365 (Chilean species).
Distribution.
0950. Chilamblyopinus Ashe & Timm, 1988: 49 (Fig. 181)

Type species: Chilamblyopinus piceus Ashe & Timm, 1988

Characters/Description: Ashe & Timm 1988: 49 (characters in key).
Distribution.
0951. Devilleferus Jenkins-Shaw & Solodovnikov in Jenkins-Shaw et al., 2017: 717

Type species: Devilleferus brunkei Jenkins-Shaw & Solodovnikov in Jenkins-Shaw et al., 2017

Distribution.
0952. Edrabius Fauvel, 1900a: 65

Type species: Edrabius philippianus Fauvel, 1900

Characters/Description: Seevers 1955: 254 (characters); Machado-Allison 1963: 380 (characters in key); Coiffait
& Sáiz 1968: 365 (characters in key); Ashe & Timm 1988: 48 (characters in key).
Key: Coiffait & Sáiz 1968: 366 (Chilean species); Scheerpeltz 1957c: 219 (species of the genus).
Distribution.
South America: Peru, Bolivia, Uruguay, Chile, Argentina.
0953. Heterothops Stephens, 1829a: 23

Type species: Staphylinus binotatus Gravenhorst, 1802
Trichopygus Nordmann, 1837: 137 [synonymized by Erichson 1839b: 480]
Type species: Tachyporus dissimilis Gravenhorst, 1802
Characters/Description: Lynch-Arribálzaga 1884: 115 (characters in key), 122 (characters); Blackwelder 1943: 395
(characters in key), 464 (characters); Smetana 1971: 14 (characters); Smetana 1988: 318 (characters);
Navarrete-Heredia et al. 2002: 298 (characters in key).
Key: Blackwelder 1943: 464 (species of the West Indies); Smetana 1971: 15 (species of the genus); Smetana 1988:
Distribution.
West Indies: Cuba, Jamaica, Dominican Republic, Trinidad-Tobago.
South America: Colombia, Venezuela, Ecuador, Peru, Bolivia, Paraguay, Brazil, Argentina.
0954. Loncovilius Germain, 1903: 439

Type species: Quedius semiflavus Fairmaire & Germain, 1862
Subgenera:
Lienturius Coiffait & Sáiz, 1966: 402 [*]
Type species: Staphylinus leiocephalus Solier, 1849
Apoquedius Scheerpeltz, 1972d: 24, 25 [synonymized by Herman 2001a: 24]
Type species: Quedius aeneipennis Fairmaire & Germain, 1862
Loncovilius Germain, 1903: 439 [*]
Type species: Quedius semiflavus Fairmaire & Germain, 1862

Characters/Description: Coiffait & Sáiz 1966: 385 (characters in key), 401 (characters); Smetana 1981: 354
Key: Coiffait & Sáiz 1966: 402 (subgenera), 402 (species of the subgenus Lienturius), 408 (species of subgenus
Loncovilius); Sáiz 1971b: 365 (species of the subgenus Lienturius).
Distribution.
0955. Megamblyopinus Seevers, 1955: 251

Type species: Amblyopinus mniszechi Solsky, 1875

Characters/Description: Seevers 1955: 251 (characters); Machado-Allison 1963: 380 (characters in key); Coiffait
& Sáiz 1968: 366 (characters in key); Ashe & Timm 1988: 48 (characters in key).
Distribution.
South America: Peru, Brazil, Chile.
0956. Mimosticus Sharp, 1884: 327 (Fig. 201)

Type species: Mimosticus viridipennis Sharp, 1884
Characters/Description: Brunke & Solodovnikov 2014b: 57 (characters).
Key: Brunke & Solodovnikov 2014b: 60 (species of the genus).
Distribution.
South America: Colombia, Venezuela, Ecuador, Peru, Bolivia.
0957. Philonthellus Bernhauer, 1939a: 13

Type species: Philonthus chilenus Bernhauer, 1939

Distribution.
0958. Rolla Blackwelder, 1952: 342

Type species: Leptoparius paradoxus Bernhauer, 1917
Leptoparius Bernhauer, 1917d: 87 [synonymized by Blackwelder 1952: 217]
Type species: Leptoparius paradoxus Bernhauer, 1917

Distribution.
Subtribe Anisolinina Hayashi, 1993
0959. Misantlius Sharp, 1885: 393 (Figs. 202–203)

Type species: Misantlius carinulatus Sharp, 1885

Characters/Description: Navarrete-Heredia et al. 2002: 301 (characters in key); Asiain & Márquez 2003: 39
(characters); Schillhammer 2004a: 264 (characters).
Key: Asiain & Márquez 2003: 40 (species of the genus).
Distribution.
0960. Tympanophorus Nordmann, 1837: 9

Type species: Tympanophorus canaliculatus Nordmann, 1837
Diaphoetus Waterhouse, 1884: 213 [synonymized by Fauvel 1902b: 41]
Type species: Diaphoetes rugosus C. Waterhouse, 1884
Characters/Description: Moore & Legner 1979: 36 (characters); Sharp 1883: 342 (characters); Fauvel 1902b: 42
(characters in key); Navarrete-Heredia et al. 2002: 299 (characters in key); Schillhammer 2004a: 267
(characters); Brunke et al. 2017: 1022 (characters), 1024 (characters in key); Janák 2018: 295 (characters in
key).
Distribution.

Subtribe Cyrtoquediina Brunke & Solodovnikov in Brunke et al., 2016
0961. Bolitogyrus Chevrolat in d’Orbigny, 1842: 641 (Fig. 176)

Type species: Quedius buphthalmus Erichson, 1840
Cyrtothorax Kraatz, 1858: 366 [synonymized by Smetana 1988: 314]
Type species: Cyrtothorax sallei Kraatz, 1858
Characters/Description: Smetana 1988: 180 (characters in key), 314 (characters); Navarrete-Heredia et al. 2002:
298 (characters in key); Brunke & Solodovnikov 2014a: 11 (characters), 21 (Buphthalmus group), 38
(Bullatus lineage), 45 (Ashei group), 51 (Cornutus group), 63 (Divisus group), 70 (Strigifrons group).
Key: Brunke & Solodovnikov 2014a: 17 (Neotropical species).
Distribution.
0962. Cyrtoquedius Bernhauer, 1917d: 92

Type species: Quedius basiventris Sharp, 1884
Characters/Description: Scheerpeltz 1955: 171 (characters); Brunke et al. 2016: 446 (characters).
Key: Scheerpeltz 1955: 172 (Neotropical species).
Distribution.
West Indies: Domincan Republic.
South America: Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil, Paraguay, Argentina.
0963. Parisanopus Brèthes, 1900: 216

Type species: Parisanopus castaneipennis Brèthes, 1900

Distribution.
0964. Sedolinus Solodovnikov, 2006: 276 (Fig. 225)

Type species: Sedolinus quediosimilis Solodovnikov, 2006
Distribution.
Subtribe Erichsoniina Brunke & Solodovnikov in Brunke et al., 2016
0965. Erichsonius Fauvel, 1874a: 201

Type species: Staphylinus cinerascens Gravenhorst, 1802
Actobius Fauvel, 1875: xxix [synonymized by Blackwelder 1943: 440]
Subgenera:
Erichsonius Fauvel, 1874a: 201
Sectophilonthus Tottenham, 1949a: 358 [*]
Type species: Philonthus biparamerosus Tottenham, 1949
Parerichsonius Coiffait, 1963c: 9 [synonymized by Frank 1981b: 103]
Type species: Philonthus signaticornis Mulsant & Rey, 1853

Characters/Description: Blackwelder 1943: 395 (characters in key), 440 (characters); Coiffait 1963c: 7 (characters
in key), 9 (characters in key for subgenera); Frank 1975: 179 (characters); Smetana 1995: 50 (characters in
key); Staniec 1999: 31 (characters); Navarrete-Heredia et al. 2002: 300 (characters in key).
Key: Frank 1975: 180 (species of the New World).
Distribution.
Subtribe Hyptiomina Casey, 1906
0966. Holisus Erichson, 1839a: 298 (Fig. 194)

Type species: Holisus analis Erichson, 1839
Hyptioma Casey, 1906: 362 [synonymized by Casey 1910c: 106]
Type species: Hyptioma cubensis Casey, 1906
Piestomorphus Motschulsky, 1858d: 666 [synonymized by Gemminger & Harold 1868: 596]
Type species: Piestomorphus ater Motschulsky, 1858
2002: 298 (characters in key); Newton & Peck 2006: 152 (characters in key); Chani-Posse 2014a: 74
Distribution.
West Indies: Bahamas, Cuba, Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., Antigua &
Barbuda, St. Lucia, St. Vincent & Grenadines.
South America: Colombia, Venezuela, Peru, Bolivia, Brazil, Argentina.
Other regions: Nearctic, Ethiopian.
Subtribe Philonthina Kirby in Richardson, 1837
0967. Atopocentrum Bernhauer, 1906a: 327

Type species: Atopocentrum mirabile Bernhauer, 1906

Characters/Description: Chani-Posse 2014a: 74 (characters in key).
Distribution.
0968. Belonuchus Nordmann, 1837: 129

Type species: Staphylinus haemorrhoidalis Fabricius, 1801
Musicoderus Sharp, 1885: 455 [synonymized by Blackwelder 1943: 420]
Type species: Musicoderus cephalotes Sharp, 1885
Trapezinotus Motschulsky, 1870: 49 [synonymized by Fauvel 1895b: 266]
Type species: Trapeziderus bicolor Motschulsky, 1860

(characters); Bernhauer 1916a: 265 (characters); Coiffait 1963c: 8 (characters in key); Smetana 1995: 50
2006: 153 (characters in key); Aballay et al. 2014: 104 (characters in key); Chani-Posse 2014a: 75 (characters
in key).
Key: Blackwelder 1943: 421 (species of the West Indies); Smetana 1995: 725 (species and species group of the
genus, America north of Mexico); Márquez et al. 2004: 24 (some Mexican species).
Distribution.
Antigua & Barbuda, Montserrat, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent & Grenadines,
South America: Colombia, Venezuela, French Guiana, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil, Paraguay,
Uruguay, Argentina.
0969. Bisnius Stephens, 1829a: 23

Type species: Staphylinus cephalotes Gravenhorst, 1802
Gefyrobius Thomson, 1859: 24 [synonymized by Blackwelder 1943: 399]
Type species: Staphylinus nitidulus Gravenhorst, 1802
Characters/Description: Smetana 1995: 51 (characters in key), 513 (characters); Navarrete-Heredia et al. 2002: 301
(characters in key); Chani-Posse 2014a: 75 (characters in key).
Key: Smetana 1995: 517 (species and species group of the genus, America north of Mexico).
Distribution.
South America: Colombia, Venezuela, Chile, Argentina.
0970. Cafius Stephens, 1829a: 23 (Fig. 177)

Type species: Staphylinus xantholoma Gravenhorst, 1806
Subgenera:
Bryonomus Casey, 1885a: 313 [*]
Type species: Philonthus canescens Mäklin, 1852
Cafius Stephens, 1829a: 23
Type species: Staphylinus xantholoma Gravenhorst, 1806
Euremus Blackwelder, 1943: 435 [*]
Type species: Philonthus bistriatus Erichson, 1840
Ifacus Blackwelder, 1952: 198
Type species: Cafius sabulosus Fauvel, 1877
Philonthopsis Blackwelder, 1943: 435 [synonymized by Blackwelder 1952: 198]
Type species: Cafius sabulosus Fauvel, 1877
Platygonium Motschulsky, 1845: 40
Type species: Platygonium sculticeps Motschulsky, 1858
Suborthidus Coiffait, 1963c: 20 [synonymized by Gusarov 1991: 8]
Type species: Philonthus cicatricosus Erichson, 1840
Pseudoremus Blackwelder, 1943: 435 [*]
Type species: Philonthus lithocharinus LeConte, 1863

Characters/Description: Blackwelder 1943: 395 (characters in key), 435 (characters); Coiffait 1963c: 8 (characters
in key); Coiffait & Sáiz 1968: 348 (characters in key), 361 (characters); Coiffait 1974: 5 (characters in key);
Orth & Moore 1980: 181 (characters); Smetana 1995: 50 (characters in key); Navarrete-Heredia et al. 2002:
301 (characters in key); Jeon & Ahn 2005: 261(characters, larva); Newton & Peck 2006: 153 (characters in
key); Chani-Posse 2014a: 76 (characters in key).
Key: Blackwelder 1943: 436 (species of the West Indies); Coiffait 1963c: 20 (subgenera); Coiffait & Sáiz 1968:
361 (Chilean species).
Distribution.
West Indies: Bahamas, Cuba, Jamaica, Dominican Republic, Puerto Rico, US Virgin Isl., Saint-Kitts & Nevis,
Antigua & Barbuda, Montserrat, Guadeloupe, Dominica, Saint Lucia, Saint Vincent & the Grenadines,
Barbados, Grenada, Trinidad-Tobago.
South America: Venezuela, Galapagos Isl.?, Brazil, Chile.
0971. Chroaptomus Sharp, 1885: 437 (Fig. 182)

Type species: Philonthus flagrans Erichson, 1840
Characters/Description: Navarrete-Heredia et al. 2002: 301 (characters in key); Chani-Posse 2006a: 363
(characters); Chani-Posse 2014a: 75 (characters in key).
Key: Chani-Posse 2006a: 366 (species of the genus).
Distribution.
Central America: Mexico, Guatemala, Nicaragua, Honduras, Costa Rica, Panama.
0972. Delgadobius Chani-Posse & Couturier, 2012: 82

Type species: Delgadobius amazonensis Chani-Posse & Couturier, 2012

Distribution.
0973. Ecitophytes Wasmann, 1923: lxii

Type species: Ecitophytes coniceps Wasmann, 1923

Characters/Description: Seevers 1965: 326 (characters); Chani-Posse 2014a: 75 (characters in key).
Distribution.
0974. Flohria Sharp, 1884: 391

Type species: Flohria laticornis Sharp, 1884

Characters/Description: Navarrete-Heredia et al. 2002: 301 (characters in key); Chani-Posse 2014a: 75 (characters
in key); Chani-Posse & Newton 2015: 373 (characters).
Distribution.
Central America: Mexico, Honduras, Nicaragua, Costa Rica.
0975. Gabrius Stephens, 1829a: 23

Type species: Staphylinus aterrimus Gravenhorst, 1802
Characters/Description: Coiffait 1963c: 7 (characters in key); Coiffait & Sáiz 1968: 347 (characters in key);
Smetana 1995: 51 (characters in key), 603 (characters); Navarrete-Heredia et al. 2002: 301 (characters in
key); Chani-Posse 2011: 3 (characters); Chani-Posse 2014a: 75 (characters in key).
Key: Coiffait 1963a: 9 (group of species); Chani-Posse 2011: 3 (species of the southern of South American).
Distribution.
Central America: Mexico, Guatemala, El Salvador, Costa Rica, Panama.
South America: Colombia, Peru, Chile, Argentina.
0976. Gabronthus Tottenham, 1955: 178

Type species: Gabrius maritimus Motschulsky, 1858
Characters/Description: Coiffait 1963c: 7 (characters in key); Smetana 1995: 49 (characters in key), 478
(characters); Navarrete-Heredia et al. 2002: 301 (characters in key); Chani-Posse 2014a: 76 (characters in
key).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Puerto Rico, US Virgin Isl., Saint-Kitts & Nevis, Antigua & Barbuda,
Guadeloupe.
0977. Hesperus Fauvel, 1874a: 200
Type species: Staphylinus rufipennis Gravenhorst, 1806
Onthostygnus Sharp, 1884: 392 [synonymized by Navarrete-Heredia et al. 2002: 338]
Type species: Onthostygnus fasciatus Sharp, 1884
Subgenera:
Hesperotropis Gridelli, 1924a: 196
Type species: Hesperus perfoliatus Gridelli, 1924
Hesperus Fauvel, 1874a: 200 [*]
Type species: Staphylinus rufipennis Gravenhorst, 1802
Characters/Description: Coiffait 1963c: 8 (characters in key); Moore & Legner 1979: 22 (characters); Smetana
1995: 50 (characters in key), 486 (characters); Navarrete-Heredia et al. 2002: 301 (characters in key); Li et al.
2010: 521 (characters); Chani-Posse 2014a: 76 (characters in key).
Key: Scheerpeltz 1936: 488 (some Neotropical species); Scheerpeltz 1971b: 153 (species of the genus).
Distribution.
South America: Bolivia, Brazil, Uruguay, Argentina.
0978. Laetulonthus Moore & Legner, 1973: 213

Type species: Staphylinus laetulus Say, 1830

Characters/Description: Moore & Legner 1979: 23 (characters); Smetana 1995: 50 (characters in key), 505
(characters); Navarrete-Heredia et al. 2002: 301 (characters in key); Chani-Posse 2014a: 76 (characters in
key).
Distribution.
0979. Leptopeltoides Chani-Posse & Asenjo, 2013: 450

Type species: Leptopeltoides ecuatoriensis Chani-Posse & Asenjo, 2013

Key: Chani-Posse & Asenjo 2013: 451 (species of the genus).
Distribution.
0980. Leptopeltus Bernhauer, 1906a: 337 (Fig. 198)

Type species: Philonthus flavipennis Erichson, 1840

Key: Scheerpeltz 1960: 114 (species of the genus); Chani-Posse & Asenjo 2013: 445 (species of the genus).
Distribution.
South America: Colombia, Venezuela, Ecuador, Peru, Bolivia, Argentina.

0981. Linoderus Sharp, 1885: 452 (Fig. 199)
Type species: Linoderus gracilipes Sharp, 1885

Characters/Description: Chani-Posse 2014a: 75 (characters in key); Chani-Posse & Solodovnikov 2017: 87
(characters).
Distribution.
0982. Neobisnius Ganglbauer, 1895: 464

Type species: Gabrius villosulus Stephens, 1833

Characters/Description: Coiffait 1963c: 8 (characters in key); Coiffait & Sáiz 1968: 348 (characters in key); Moore
& Legner 1979: 24 (characters); Frank 1981a: 5 (characters); Smetana 1995: 50 (characters in key);
Navarrete-Heredia et al. 2002: 301 (characters in key); Chani-Posse 2014a: 76 (characters in key).
Key: Coiffait 1963c: 18 (group of species); Frank 1981a: 5 (species of the America north of Mexico), 6 (species of
Mexico and Central America), 7 (species of the West Indies), 7 (species of South American).
Distribution.
West Indies: Cuba, Cayman Is., Jamaica, Haiti, Dominican Republic, Puerto Rico, US Virgin Isl., St. Lucia, St.
Vincent & Grenadines, Barbados, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Bolivia, Brazil, Paraguay, Uruguay, Chile, Argentina.
0983. Neopescolinus Chani-Posse, 2014b: 249

Type species: Neopescolinus nevermanni Chani-Posse, 2014

Distribution.
0984. Odontolinus Sharp, 1885: 454 (Fig. 204)

Type species: Odontolinus fasciatus Sharp, 1885

Characters/Description: Chani-Posse 2014a: 76 (characters in key); Chani-Posse & Solodovnikov 2017: 91
(characters).
Distribution.
Central America: Honduras, Nicaragua, Costa Rica, Panama.
0985. Ophionthus Bernhauer, 1908: 328

Type species: Ophionthus serpentinus Bernhauer, 1908

Distribution.
0986. Paederallus Sharp, 1885: 456
Type species: Paederallus fragilis Sharp, 1885

Distribution.
0987. Paederomimus Sharp, 1885: 438

Type species: Paederomimus difformiceps Sharp, 1885
58 species are known from and are restricted to Latin America. Except for one species Paederomimus chinensis
(Cameron) from China, probably it is am wrong labeling (Schillhammer 2004b).
2002: 301 (characters in key); Chani-Posse 2014a: 76 (characters in key).
Distribution.
West Indies: Puerto Rico, US Virgin Isl., St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Peru, Brazil, Paraguay, Uruguay.
0988. Pescolinus Sharp, 1885: 453 (Fig. 206)

Type species: Pescolinus palmatus Sharp, 1885

Characters/Description: Chani-Posse 2014a: 76 (characters in key); Chani-Posse 2014b: 239 (characters).
Key: Chani-Posse 2014b: 241 (species of the genus).
Distribution.
0989. Phileciton Wasmann, 1894: 211

Type species: Phileciton badariottii Wasmann, 1894

Distribution.
0990. Philonthus Stephens, 1829a: 23 (Figs. 208–209)

Type species: Staphylinus splendens Fabricius, 1792
Cephalonthus Blackwelder, 1952: 96 [synonymized by Smetana in Löbl & Smetana 2004: 34]
Type species: Philonthus caffer Boheman, 1848
Endeius Coiffait & Sáiz, 1968: 355 [synonymized by Chani-Posse 2013: 404]
Type species: Endeius loensis Coiffait & Sáiz, 1968
Kirschenblatia Bolov & Kryzhanovskij in Bolov, 1969: 512 [synonymized by Schülke & Uhlig 1989: 247]
Type species: Kirschenblatia kabardensis Bolov & Kryzhanovskij, 1969
Kenonthus Coiffait, 1960a: 361 [synonymized by Smetana in Löbl & Smetana 2004: 34]
Type species: Philonthus montivagus Heer, 1839

Laxobates Gistel, 1834: 8 [synonymized by Smetana 1958: 143]
Metagabrius Coiffait, 1974: 84, 86 [synonymized by Smetana 1995: 51]
Type species: Philonthus juvenilis Peyerimhoff, 1858
Palaeophilonthus Coiffait, 1972b: 50 [synonymized by Smetana 1995: 51]
Type species: Sectophilonthus rossicus Coiffait, 1965
Paragabrius Coiffait, 1963c: 12 [synonymized by Smetana 1995: 51]
Type species: Staphylinus micans Gravenhorst, 1802
Paralionthus Ádám, 1996: 236 [synonymized by Assing & Schülke 1999: 14]
Type species: Staphylinus punctus Gravenhorst, 1802
Philonthopsis Cameron, 1932: 261 [synonymized by Schillhammer 1995: 74]
Type species: Philonthopsis antennalis Cameron, 1932
Spatulonthus Tottenham, 1955: 154 [synonymized by Smetana 1995: 51]
Type species: Philonthus longicornis Stephens, 1832
Trionthus Coiffait, 1963c: 14 [synonymized by Smetana 1995: 51]
Type species: Staphylinus lepidus Gravenhorst, 1802
Subgenera:
Glaphyronthus Scheerpeltz, 1978: 220
Type species: Philonthus tenellicornis Scheerpeltz, 1978
Onychophilonthus Neresheimer & H. Wagner, 1924: 156
Type species: Staphylinus marginatus Ström, 1768
Philonthus Stephens, 1829a: 23 [*]
Piezarthrus Lecoq, 1998: 63
Type species: Philonthus pollux Fauvel, 1905
Raucalius Tottenham, 1949a: 304
Type species: Philonthus peripateticus Tottenham, 1949
Heterogaster Cameron, 1951: 406 [synonymized by Verdcourt 1952: 176]
Type species: Heterogaster niger Cameron, 1951
Xanthopygoides Verdcourt, 1952: 176 [synonymized by Solodovnikov 2009: 77]
Type species: Heterogaster niger Cameron, 1951
Characters/Description: Lynch-Arribálzaga 1884: 146 (characters); Moore & Legner 1979: 28 (characters);
Blackwelder 1943: 395 (characters in key), 398 (characters); Coiffait 1963c: 7 (characters in key as
Paragabrius), 8 (characters in key as Spatulonthus); Coiffait & Sáiz 1968: 348 (characters in key as Endeius),
348 (characters in key as Spatulonthus), 348 (characters in key of Philonthus), 350 (characters), 355
(charactes as Endeius); Smetana 1995: 50 (characters in key), 51 (characters); Chani-Posse 2008: 382
(characters as Endeius); Chani-Posse 2010a: 6 (characters); Navarrete-Heredia et al. 2002: 298, 301
(characters in key); Newton & Peck 2006: 153 (characters in key of Philonthus), 153 (characters in key as
Endeius); Aballay et al. 2014: 104 (characters in key); Chani-Posse 2014a: 76 (characters in key).
of the West Indies); Coiffait 1963c: 14 (subgenera); Coiffait & Sáiz 1968: 351 (group of species for Chile),
351 (species of Chile of the group intermedius), 351 (species of Chile of the group rectangulus), (species of
Chile of the group fumarius), 352 (Chilean species of Philonthus), 356 (Chilean species as Endeius); Sáiz
1971a: 343 (Chilean species as Endeius); Smetana 1991: 227 (some species of Mexico and Central America);
Smetana 1995: 56 (species and species group of the genus, America north of Mexico); Navarrete-Heredia
2003: 6 (Mexican species); Chani-Posse 2006b: 9 (larvae associated with cattle dung in Tucumán, Argentina);
Newton & Peck 2006: 156 (species of Galapagos Isl. as Endeius), 157 (species of Galapagos Isl. as
Philonthus); Chani-Posse 2008: 386 (Endeius); Chani-Posse 2010a: 9 (species from southern South
American).
Distribution.
Saint-Kitts & Nevis, Antigua & Barbuda, Montserrat, Guadeloupe, Dominica, St. Lucia, St. Vincent &
Grenadines, Barbados, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, Guyana, French Guiana, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil,
0991. Pridonius Blackwelder, 1952: 321

Type species: Quedius iheringi Bernhauer, 1907
Prionidus Bernhauer, 1907: 287 [synonymized by Blackwelder 1952: 321]
Type species: Quedius iheringi Bernhauer, 1907

Characters/Description: Chani-Posse et al. 2018a: 34.
Distribution.
0992. Prionopedinus Scheerpeltz, 1972c: 36

Type species: Trigonopselaphus elvirae Bernhauer, 1921
Prionopus Bernhauer, 1921e: 20 [synonymized by Scheerpeltz 1972c: 36]
Type species: Trigonopselaphus elvirae Bernhauer, 1921

Distribution.
0993. Proxenobius Seevers, 1965: 326

Type species: Proxenobius borgmeieri Seevers, 1965

Characters/Description: Navarrete-Heredia et al. 2002: 298 (characters in key); Chani-Posse 2014a: 75 (characters
in key).
Distribution.
0994. Pterygolaetus Bierig, 1937a: 194

Type species: Pterygolaetus williamsi Bierig, 1937

Distribution.

0995. Remus Holme, 1837: 64
Type species: Remus sericeus Holme, 1837
Pseudidus Mulsant & Rey, 1876: 574 [synonymized by Blackwelder 1943: 435]
Type species: Remus sericeus Holme, 1837
1 species is included in this genus from the West Indies. Remus pruinosus (Erichson, 1840) apparently reported in
error from Cuba (Chani-Posse et al., 2018b: 82).
Characters/Description: Coiffait 1963c: 8 (characters in key); Coiffait 1974: 5 (characters in key); Chani-Posse
Distribution.
West Indies: Cuba.
0996. Xanthodermus Bernhauer, 1912c: 40

Type species: Belonuchus vestitus R.F. Sahlberg, 1847

Distribution.
0997. Xenobius Borgmeier, 1931: 358

Type species: Xenobius rotundiceps Borgmeier, 1931

Characters/Description: Seevers 1965: 324 (characters); Chani-Posse 2014a: 75 (characters in key).
Distribution.
Subtribe Philothalpina Chatzimanolis & Brunke in Chani-Posse et al., 2018a
0998. Philothalpus Kraatz, 1857a: 540 (Fig. 210)

Type species: Philonthus fervidus Erichson, 1840
Allostenopsis Bernhauer, 1921a: 74 [synonymized by Chatzimanolis & Ashe 2005: 64]
Eugastus Sharp, 1876a: 139 [synonymized by Herman 2001a: 10]
Type species: Eugastus mundus Sharp, 1876
Stenopsis Bernhauer, 1907: 286 [synonymized by Chatzimanolis & Ashe 2005: 66]
Type species: Stenopsis antennaria Bernhauer, 1907
Characters/Description: Blackwelder 1943: 394 (characters in key), 451 (characters); Chatzimanolis & Ashe 2005:
66 (characters); Torres-Rodríguez et al. 2012: 228 (characters), 236 (characters in key).
Key: Chatzimanolis & Ashe 2005: 113 (species of the genus); Asenjo & Ribeiro-Costa 2009: 56 (species of the
genus).
Distribution.
Central America: Nicaragua, Costa Rica, Panama.
Subtribe Quediina Kraatz, 1857a
0999. Quedionuchus Sharp, 1884: 336

Type species: Quedius impunctus Solsky, 1868

Characters/Description: Smetana 1971: 147 (characters as subgenus of Quedius); Smetana 1976a: 223 (characters).
Key: Smetana 1971: 147 (species the genus, as subgenus of Quedius); Smetana 1976a: 224 (Central American
species as subgenus of Quedius).
Distribution.
1000. Quedius Stephens, 1829a: 22 (Fig. 223)

Type species: Staphylinus levicollis Brullé, 1832
Aemulus Gistel, 1834: 8 [synonymized by Blackwelder 1952: 40]
Type species: Staphylinus fuliginosus Gravenhorst, 1802
Anastictodera Casey, 1915: 421 [synonymized by Smetana 1971: 38]
Type species: Quedius compransor Fall, 1912
Arphirus Tottenham, 1945: 70 [synonymized by Smetana 1958: 328]
Type species: Staphylinus semiobscurus Marsham, 1802
Ediquus Mulsant & Rey, 1876: 615, 616 [synonymized by Scheerpeltz 1933: 1428]
Type species: Quedius microps Gravenhorst, 1847
Quediochrus Casey, 1915: 398, 420 [synonymized by Smetana 1971: 38]
Type species: Quedius spelaeus Horn, 1871
Quediellus Casey, 1915: 398, 402 [synonymized by Smetana 1971: 39]
Type species: Quedius debilis Horn, 1878
Thanatomanes Gistel, 1856: 388 [synonymized by Blackwelder 1952: 384]
Type species: Staphylinus impressus Panzer, 1796
Subgenera:
Distichalius Casey, 1915: 398, 404
Type species: Staphylinus capucinus Gravenhorst, 1806
Euryquedius Reitter, 1909: 108 [*]
Type species: Quedius curtus Erichson, 1840
Farus Blackwelder, 1952: 165
Type species: Quedius przewalskii Reitter, 1887
Ediquus Reitter, 1887a: 211 [synonymized by Blackwelder 1952: 165]
Type species: Quedius przewalskii Reitter, 1887
Megaquedius Casey, 1915: 421 [*]
Type species: Quedius explanatus LeConte, 1858
Microsaurus Dejean, 1833: 61
Type species: Staphylinus lateralis Gravenhorst, 1802
Tenebrobius Rambousek, 1915a: 27, 29 [synonymized by Gridelli 1924b: 18]
Type species: Quedius bernhaueri Rambousek, 1915
Paraquedius Casey, 1915: 397, 400
Type species: Quedius puncticeps Horn, 1878
Quediopsis Portevin, 1929: 337

Type species: Quedius riparius F. Kellner, 1843
Quedius Stephens, 1829a: 22 [*]
Type species: Staphylinus levicollis Brullé, 1832
Raphirus Stephens, 1829a: 23 [*]
Type species: Staphylinus attenuatus Gravenhorst, 1802
Microquedius Coiffait, 1967: 419 [synonymized by Smetana 1993: 50]
Type species: Philonthus alpestris Heer, 1839
Quediops Coiffait, 1963b: 373 [synonymized by Smetana 1993: 50]
Type species: Quedius plancus Erichson, 1840
Quedionthus Bordoni, 1973: 53 [synonymized by Coiffait 1978c: 9]
Type species: Quedius riparius F. Kellner, 1843
Sauridus Mulsant & Rey, 1876: 616, 700 [synonymized by Blackwelder 1952: 344]
Type species: Staphylinus picipes Mannerheim, 1830
Velleius Leach in Samouelle, 1819: 172
Type species: Staphylinus dilatatus Fabricius, 1787
Laverna Gistel, 1829: 1129 [synonymized by Blackwelder 1952: 212]
Type species: Staphylinus dilatatus Fabricius, 1787
Characters/Description: Lynch-Arribálzaga 1884: 114 (characters in key), 119 (characters); Coiffait & Sáiz 1966:
385 (characters in key), 411 (characters); Smetana 1971: 38 (characters), 42 (characters of the subgenus
Megaquedius), 49 (characters of the subgenus Microsaurus), 128 (characters of the subgenus Quedius),153
(characters of the subgenus Distichalius), 183 (characters of the subgenus Raphirus), 235 (characters of the
subgenus Paraquedius); Smetana 1981: 354 (characters in key); Smetana 1988: 180 (characters in key), 181
(characters); Staniec 1999: 48(characters of pupa); Navarrete-Heredia et al. 2002: 298 (characters in key).
Key: Scheerpeltz 1955: 172 (Cyrtoquedius); Smetana 1971: 41 (subgenera of Quedius), 42 (species of the
subgenus Megaquedius), 49 (species of the subgenus Microsaurus), 129 (species of the subgenus Quedius),
154 (species of the subgenus Distichalius), 184 (species of the subgenus Raphirus); Smetana 1975a: 37
(Mexican species of the subgenus Megaquedius); Smetana 1975b: 314 (species of caseyi group in subgenus
Microsaurus), 322 (species of aenescens group in subgenus Raphirus); Smetana 1976b: 113 (Mexican species
of the subgenus Microsaurus); Smetana 1988: 184 (subgenera), 186 (species of the subgenus Microsaurus),
227 (species of the subgenus Distichalius), 233 (species of the subgenus Raphirus); Gusarov 2001: 376
Distribution.
South America: Colombia, Venezuela, Peru, Bolivia, Paraguay, Chile, Argentina.
Subtribe Staphylinina Latreille, 1802
1001. Creophilus Leach in Samouelle, 1819: 172 (Figs. 183–185)

Type species: Staphylinus maxillosus Linnaeus, 1758
Saprophilus Streubel, 1839: 136 [synonymized by Blackwelder 1952: 344]
Type species: Staphylinus maxillosus Linnaeus, 1758
Characters/Description: Moore & Legner 1979: 33 (chracters); Lynch-Arribálzaga 1884: 128 (characters);
Blackwelder 1943: 394 (characters in key), 447 (characters); Smetana & Davies 2000: 18 (characters in key),
21 (characters); Navarrete-Heredia et al. 2002: 300 (characters in key); Newton & Peck 2006: 152 (characters
in key); Clarke 2011: 755 (characters); Aballay et al. 2014: 104 (characters in key).
Key: Coiffait & Sáiz 1968: 364 (Chilean species); Clarke 2011: 758 (species of the genus).
Distribution.
West Indies: Cuba, Jamaica, Dominican Republic.
Central America: Mexico, Guatemala, Honduras.
South America: Galapagos Isl., Peru, Bolivia, Brazil, Paraguay, Uruguay, Chile, Argentina.
1002. Dinothenarus Thomson, 1858: 29 (Fig. 187)

Type species: Staphylinus pubescens DeGeer, 1774
Subgenera:
Dinothenarus Thomson, 1858: 29 [*]
Type species: Staphylinus pubescens DeGeer, 1774
Protabemus Scheerpeltz, 1966c: 114, 115 [synonymized by Smetana 1992: 187]
Type species: Staphylinus xanthocephalus Kraatz, 1859
Parabemus Reitter, 1909: 118 [*]
Type species: Staphylinus fossor Scopoli, 1772
Hypabemus Scheerpeltz, 1966c: 112, 116 [synonymized by Smetana & Davies 2000: 26]
Type species: Staphylinus chrysocomus Mannerheim, 1830
Parocypus Bernhauer, 1915c: 52 [synonymized by Smetana & Davies 2000: 26]
Type species: Staphylinus dehradunensis Bernhauer, 1915

Characters/Description: Smetana & Davies 2000: 20 (characters in key), 26 (characters); Navarrete-Heredia et al.
2002: 300 (characters in key); Santiago-Jiménez 2008: 51 (characters).
Key: Santiago-Jiménez 2008: 56 (key to species of Dinothenarus s. str. of America).
Distribution.
1003. Hadrotes Mannerheim, 1852: 313

Type species: Staphylinus crassus Mannerheim, 1846

Characters/Description: LeConte 1861: 64 (characters in key); Horn 1879: 199 (characters); Hatch 1957: 212
(characters); Moore & Legner 1979: 34 (characters); Navarrete-Heredia et al. 2002: 300 (characters in key).
Distribution.
Other regions: Nearctic, Australian.
1004. Leistotrophus Perty, 1830: 32 (Fig. 197)

Type species: Leistotrophus gravenhorsti Perty, 1830
Discocephalus Nordmann, 1837: 3, tab. 1, fig. 8 [synonymized by Gemminger & Harold 1868: 576]
Type species: Staphylinus versicolor Gravenhorst, 1806
Schizochilus Gray in Griffith & Pidgeon, 1831: 304 [synonymized by Laporte 1835: 111]

Type species: Schizochilus brasiliensis Gray, 1832
1 species is included in this genus from Central and South America.

Characters/Description: Smetana 1959: 394 (characters in key) ; Navarrete-Heredia et al. 2002: 300 (characters in
key); Márquez & Asiain 2004: 256 (characters).
Distribution.
1005. Ocypus Leach in Samouelle, 1819: 172

Type species: Staphylinus cyaneus Paykull, 1789
Subgenera:
Angulephallus Khachikov, 2005: 23
Type species: Staphylinus brunnipes Fabricius, 1781
Matidus Motschulsky, 1860b: 569
Type species: Matidus forficularius Motschulsky, 1860
Ocypus Leach in Samouelle, 1819: 172 [*]
Type species: Staphylinus cyaneus Paykull, 1789
Goerius Westwood, 1827: 58 [synonymized by Smetana & Davies 2000: 31]
Type species: Staphylinus olens O. Müller, 1764
Isopterum Gistel, 1856: 388, 424 [synonymized by Blackwelder 1952: 205]
Type species: Staphylinus cyaneum Paykull, 1789
Nudabemus Coiffait, 1982: 74 [synonymized by Löbl & Smetana 2004: 674]
Type species: Nudabemus caerulescens Coiffait, 1982
Pseudocypus Mulsant & Rey, 1876: 291
Type species: Staphylinus mus Brullé, 1832
Fortunocypus Coiffait, 1964: 82 [synonymized by Coiffait 1974: 483]
Type species: Ocypus fortunatarum Wollaston, 1871

Characters/Description: Smetana & Davies 2000: 21 (characters in key), 31 (characters); Navarrete-Heredia et al.
Distribution.
1006. Ontholestes Ganglbauer, 1895: 417 (Fig. 205)

Type species: Staphylinus murinus Linnaeus, 1758
Trichoderma Stephens, 1833b: 98 [synonymized by Blackwelder 1952: 394]
Type species: Staphylinus murinus Linnaeus, 1758

Characters/Description: Smetana 1959: 394 (characters in key) ; Moore & Legner 1979: 27 (chracters); Smetana &
Davies 2000: 19 (characters in key), 22 (characters).
Distribution.
South America: Suriname, Guyana, Peru, Bolivia, Brazil, Paraguay, Argentina.
1007. Platydracus Thomson, 1858: 29 (Figs. 211–218)

Type species: Staphylinus stercorarius Olivier, 1795
Subgenera:
Chaetodracus G. Müller, 1926: 27
Type species: Staphylinus patricius Bernhauer, 1915
Chitocompsus Bernhauer, 1913: 232
Type species: Staphylinus polyphemus Bernhauer, 191
Nesiolinus Bernhauer, 1915e: 122
Type species: Staphylinus bakeri Bernhauer, 1915
Paraplatydracus Levasseur, 1980: 369
Type species: Platydracus girardi Levasseur, 1980
Platydracus Thomson, 1858: 29 [*]
Type species: Staphylinus stercorarius Olivier, 1795
Amichorus Sharp, 1884: 390 [synonymized by Navarrete-Heredia et al. 2002: 323]
Type species: Amichorus fauveli Sharp, 1884
Bemasus Mulsant & Rey, 1876: 259 [synonymized by Bernhauer & Schubert 1914: 374]
Type species: Staphylinus lutarius Gravenhorst, 1806
Chapmaniella Bernhauer, 1934a: 145 [synonymized by Newton 2017: 28]
Type species: Chapmaniella miranda Bernhauer, 1934
Euryolinus Bernhauer, 1915b: 297 [synonymized by Newton 2017: 28]
Type species: Staphylinus semicyaneus Bernhauer, 1915
Glenothorax Bierig, 1937a: 199 [synonymized by Newton 2017: 28]
Type species: Glenothorax viridipennis Bierig, 1937
Neotasgius G. Müller, 1925: 41 [synonymized by Smetana & Davies 2000: 24, 39]
Type species: Ocypus brevicornis Weise, 1877
Plesiolinus Bernhauer, 1917c: 93 [synonymized by Newton 2017: 28]
Type species: Staphylinus costaricensis Bernhauer, 1916
Poikilodracus Scheerpeltz, 1974: 168
Type species: Staphylinus brincki Scheerpeltz, 1974
Tropoplatydracus Levasseur, 1980: 367
Type species: Platydracus lecordieri Levasseur, 1980

Characters/Description: Lynch-Arribálzaga 1884: 135 (characters as Staphylinus); Blackwelder 1943: 394
(characters in key as Staphylinus); Smetana & Davies 2000: 18 (characters in key), 24 (characters); Navarrete-
Heredia et al. 2002: 300 (characters in key); Aballay et al. 2014: 104 (characters in key); Newton 2017: 30
(characters).
Key: Lynch-Arribálzaga 1884: 137 (species of Buenos Aires province, Argentina); Márquez et al. 2004: 28 (some
Mexican species).
Distribution.

1008. Thinopinus LeConte, 1852: 215
Type species: Thinopinus pictus LeConte, 1852
Trichocanthus Motschulsky, 1853: 78 [synonymized by Gemminger & Harold 1868: 576]
Type species: Trichocanthus variegatus Motschulsky, 1853

Characters/Description: Moore & Legner 1979: 35 (chracters); Navarrete-Heredia et al. 2002: 300 (characters in
key).
Distribution.
Subtribe Tanygnathinina Reitter, 1909
1009. Atanygnathus Jakobson, 1909: 521

Type species: Tanygnathus terminalis Erichson, 1839
Tanygnathinus Reitter, 1909: 105 [synonymized by Bernhauer & Schubert 1916: 439]
Tanygnathus Erichson, 1839b: 417 [synonymized by Bernhauer & Schubert 1916: 439]
(characters); Smetana 1988: 368 (characters); Navarrete-Heredia et al. 2002: 298 (characters in key); Newton
& Peck 2006: 152 (characters in key).
Key: Blackwelder 1943: 471 (species of the West Indies); Smetana 1988: 369 (species of the genus).
Distribution.
West Indies: Cuba, Dominican Republic, Puerto Rico, Antigua & Barbuda, St. Vincent & Grenadines, Grenada,
Trinidad-Tobago.
Subtribe Xanthopygina Sharp, 1884
1010. Darwinilus Chatzimanolis, 2014a: 32 (Fig. 186)

Type species: Darwinilus sedarisi Chatzimanolis, 2014

Distribution.
1011. Dysanellus Bernhauer, 1911a: 419 (Fig. 189)

Type species: Dysanellus bruchi Bernhauer, 1911
Characters/Description: Scheerpeltz 1965b: 277 (characters); Navarrete-Heredia et al. 2002: 299 (characters in
key); Chatzimanolis & Ashe 2009: 3 (characters); Chatzimanolis 2012: 267 (characters); Torres-Rodríguez et
al. 2012: 219 (characters), 235 (characters in key); Chatzimanolis 2015c: 17 (characters); Chatzimanolis
Distribution.
1012. Elecatopselaphus Scheerpeltz, 1972c: 39

Type species: Trigonopselaphus violaceus Sharp, 1876

Distribution.
South America: Colombia, Peru, Brazil.
1013. Elmas Blackwelder, 1952: 146 (Fig. 190)

Type species: Selma modesta Sharp, 1876
Selma Sharp, 1876b: 426 [synonymized by Blackwelder 1952: 146]
Type species: Selma modesta Sharp, 1876
Characters/Description: Ashe & Chatzimanolis 2003: 3 (characters); Torres-Rodríguez et al. 2012: 220
Key: Ashe & Chatzimanolis 2003: 33 (species of the genus); Ashe & Chatzimanolis 2006: 28 (species of the
genus).
Distribution.
South America: Colombia, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil.
1014. Gastrisus Sharp, 1876a: 136 (Fig. 191)

Type species: Gastrisus laevigatus Sharp, 1876
Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Chatzimanolis & Ashe 2009: 3
(characters).
Distribution.
South America: Colombia, Venezuela, Peru, Brazil, Uruguay, Argentina.
1015. Glenus Kraatz, 1857a: 541

Type species: Staphylinus biplagiatus Perty, 1830
Characters/Description: Sharp 1876a: 147 (characters); Navarrete-Heredia et al. 2002: 299 (characters in key);
Torres-Rodríguez et al. 2012: 221 (characters), 235 (characters in key).
Distribution.
South America: Colombia, Brazil, Peru, Bolivia, Paraguay, Argentina.
1016. Haematodes Laporte, 1835: 112

Type species: Haematodes bicolor Laporte, 1835
Platycnemus Nordmann, 1837: 135 [synonymized by Lacordaire 1854: 72]
Type species: Platycnemus lateritius Nordmann, 1837
Platytoma Chevrolat in d’Orbigny, 1847: 263 [synonymized by Lacordaire 1854: 72]
Weiserianum Bernhauer, 1927b: 247 [synonymized by Brunke & Chatzimanolis 2018: 5 ]
Type species: Weiserianum woltersi Bernhauer, 1927

Characters/Description: Lynch-Arribálzaga 1884: 114 (characters in key), 115 (characters); Erichson 1839a: 340
(characters); Scheerpeltz 1937c: 340 (characters as Weiserianum); Chatzimanolis 2016: 285 (characters);
Brunke & Chatzimanolis 2018: 5 (characters).
Key: Scheerpeltz 1937c: 351 (species as Weiserianum); Brunke & Chatzimanolis 2018: 5 (species of the genus).
Distribution.
South America: Venezuela?, Brazil, Paraguay, Uruguay, Argentina.
1017. Isanopus Sharp, 1876a: 141 (Fig. 196)

Type species: Isanopus tenuicornis Sharp, 1876
Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Chatzimanolis 2008: 1769
(characters).
Key: Chatzimanolis 2008: 1789 (species of the genus).
Distribution.
South America: Ecuador, Peru, Bolivia, Brazil.
1018. Nausicotus Sharp, 1884: 368

Type species: Trigonopselaphus opacipennis Sharp, 1876
Characters/Description: Chatzimanolis & Ashe 2009: 3 (characters).
Distribution.
Central America: Nicaragua, Costa Rica.
South America: Brazil, Uruguay, Argentina.
1019. Nordus Blackwelder, 1952: 262

Type species: Brachydirus xanthocerus Nordmann, 1837
Brachydirus Nordmann, 1837: 131 [synonymized by Erichson 1839a: 353]
Type species: Brachydirus xanthocerus Nordmann, 1837
Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Chatzimanolis 2004: 6
(characters); Torres-Rodríguez et al. 2012: 223 (characters), 236 (characters in key); Aballay et al. 2014: 104
Key: Scheerpeltz 1971a: 94 (Brachydirus); Chatzimanolis 2004: 47 (species of the genus); Navarrete-Heredia et al.
2006: 105 (species of the genus).
Distribution.
1020. Ocyolinus Sharp, 1884: 362

Type species: Ocyolinus rugatus Sharp, 1884
Characters/Description: Chatzimanolis & Ashe 2009: 3 (characters); Chatzimanolis 2017b: 806 (characters).
Key: Chatzimanolis & Ashe 2009: 22 (species of the genus); Chatzimanolis 2017b: 808 (species of the genus).
Distribution.
Central America: Guatemala, Nicaragua, Costa Rica, Panama.
South America: Colombia, Venezuela, Brazil.
1021. Oligotergus Bierig, 1937a: 204

Type species: Philothalpus oculatus Bierig, 1937

Characters/Description: Sharp 1876a: 133 (characters); Navarrete-Heredia et al. 2002: 299 (characters in key);
Newton & Peck 2006: 152 (characters in key); Torres-Rodríguez et al. 2012: 224 (characters), 235 (characters
in key); Aballay et al. 2014: 104 (characters in key); Chatzimanolis & Caron 2016: 132 (characters).
Key: Márquez et al. 2004: 30 (some Mexican species).
Distribution.
South America: Colombia, Venezuela, Guyana, French Guiana, Galapagos Isl., Peru, Bolivia, Brazil, Argentina.
1022. Paraxenopygus Bernhauer, 1911a: 415

Type species: Paraxenopygus tremolerasi Bernhauer, 1911

Characters/Description: Torres-Rodríguez et al. 2012: 225 (characters), 235 (characters in key).
Distribution.
South America: Peru, Brazil, Uruguay, Argentina.
1023. Phanolinopsis Scheerpeltz, 1968: 67

Type species: Phanolinus discedens Sharp, 1884

Characters/Description: Chatzimanolis 2017: 50 (characters).
Key: Chatzimanolis 2017a: 65 (species of the genus).
Distribution.
1024. Phanolinus Sharp, 1884: 363

Type species: Phanolinus flohri Sharp, 1884

Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Torres-Rodríguez et al. 2012: 226
Key: Scheerpeltz 1968: 55 (species of the genus); Navarrete-Heredia 2005: 891 (Mexican species).
Distribution.
South America: Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil.
1025. Plociopterus Kraatz, 1857a: 539

Type species: Staphylinus fetialis Erichson, 1839
Key: Scheerpeltz 1960: 121 (species of the genus).
Distribution.
South America: Colombia, Venezuela, Guyana, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay.
1026. Scaponopselaphus Scheerpeltz, 1972c: 38

Type species: Trigonopselaphus mutator Sharp, 1876

Characters/Description: Chatzimanolis 2015c: 3 (characters).
Distribution.
South America: Colombia, Guyana, Suriname, French Guiana, Peru, Brazil.
1027. Scariphaeus Erichson, 1839a: 342

Type species: Velleius luridipennis Runde, 1835

Characters/Description: Lynch-Arribálzaga 1884: 114 (characters in key), 117 (characters); Scheerpeltz 1936: 531
(characters).
Distribution.
South America: Ecuador, Brazil, Paraguay, Argentina.
1028. Smilax Laporte, 1835: 116

Type species: Smilax americanus Laporte, 1835
Cordylaspis Nordmann, 1837: 17 [synonymized by Blackwelder 1952: 107 but previously noted by
Lacordaire, 1854: 73]
Type species: Cordylaspis tuberculatus Nordmann, 1837

Characters/Description: Erichson 1839a: 339–341 (characters); Navarrete-Heredia et al. 2002: 299 (characters in
key); Torres-Rodríguez et al. 2012: 230 (characters), 235 (characters in key); Chatzimanolis 2016: 285
(characters).
Key: Scheerpeltz 1937a: 335 (species of the Cordylaspis); Chatzimanolis 2016: 300 (species of the genus).
Distribution.
Central America: Mexico?, Nicaragua, Costa Rica, Panama.
Paraguay, Uruguay, Argentina.
1029. Styngetus Sharp, 1884: 361

Type species: Philonthus viduus Erichson, 1840
Key: Navarrete-Heredia 1998: 70 (Mexican species).
Distribution.
South America: Colombia, Venezuela, Peru, Bolivia, Brazil, Paraguay, Argentina.
1030. Terataki Chatzimanolis, 2013: 252 (Fig. 226)

Type species: Creophilus badiipennis Nordmann, 1837
Distribution.
1031. Torobus Herman, 2001a: 29

Type species: Staphylinus purpurascens Nordmann, 1837

Characters/Description: Scheerpeltz 1972c: 37–40 (characters); Chatzimanolis & Ashe 2009: 3–4 (characters);
Chatzimanolis 2017a: 50 (characters); Chatzimanolis 2018: 285 (characters).
Distribution.
1032. Triacrus Nordmann, 1837: 19

Type species: Triacrus dilatatus Nordmann, 1837

Characters/Description: Bernhauer 1931: 84 (characters); Chatzimanolis 2015b: 515 (characters).
Distribution.
1033. Tricholinus Bernhauer, 1922a: 15

Type species: Tricholinus pehlkei Bernhauer, 1922

Distribution.

1034. Trigonopselaphus Gemminger & Harold, 1868: 597 (Fig. 228)
Type species: Trigonophorus myrtillinus Nordmann, 1837
Lypophemus Bernhauer, 1921a: 74 [synonymized by Strand 1921: 180]
Type species: Staphylinus herculeanus Laporte, 1835
Polyphematiana Strand, 1915: 122 [synonymized by Herman 2001a: 28]
Polyphemus Bernhauer in Bernhauer & Schubert, 1914: 397 [synonymized by Scheerpeltz 1933: 1419]
Polyphemus Bernhauer, 1914: 51 [synonymized by Scheerpeltz 1933: 1419]
Type species: Polyphemus banghaasi Bernhauer, 1914
Trigonophorus Nordmann, 1837: 8 [synonymized by Gemminger & Harold 1868: 597]
Type species: Trigonophorus myrtillinus Nordmann, 1837

Characters/Description: Lynch-Arribálzaga 1884: 140 (characters); Bernhauer 1914: 51 (characters); Bernhauer
1921a: 74 (characters); Bernhauer 1931: 84 (characters); Smetana & Davies 2000: 18 (characters);
Chatzimanolis 2015a: 168 (characters).
Key: Lynch-Arribálzaga 1884: 141 (species of Buenos Aires province, Argentina); Bernhauer 1921e: 19 (species
of the genus); Scheerpeltz 1972c: 40 (species of the genus); Chatzimanolis 2015a: 176 (species of the genus).
Distribution.
South America: Ecuador, Peru, Brazil, Paraguay, Argentina.
1035. Xanthopygus Kraatz, 1856b: 539

Type species: Staphylinus xanthopygus Nordmann, 1837
Heteropygus Bernhauer, 1906b: 195 [synonymized by Blackwelder 1943: 449]
Type species: Xanthopygus oliveirae Lynch-Arribálzaga, 1884
Lampropygus Sharp, 1884: 346 [synonymized by Blackwelder 1943: 449]
Type species: Staphylinus xanthopygus Nordmann, 1837

Characters/Description: Moore & Legner 1979: 37 (chracters); Lynch-Arribálzaga 1884: 130 (characters); Hayashi
1997: 488 (characters); Blackwelder 1943: 394 (characters in key), 449 (characters); Navarrete-Heredia et al.
2002: 300 (characters in key); Chatzimanolis 2008: 1769 (characters); Torres-Rodríguez et al. 2012: 231
(characters), 236 (characters in key); Chatzimanolis & Caron 2016: 132 (characters).
Key: Lynch-Arribálzaga 1884: 131 (species of Buenos Aires province, Argentina); Scheerpeltz 1969a: 109
(species of the genus); Navarrete-Heredia 2004: 12 (Mexican species).
Distribution.
West Indies: St. Vincent & the Grenadines, Trinidad-Tobago.
Uruguay, Argentina.
1036. Xenopygus Bernhauer, 1906b: 196 (Figs. 229–230)

Type species: Philonthus analis Erichson, 1840
Leptodiastemus Bernhauer, 1934d: 215 [synonymized by Chatzimanolis & Caron 2016: 132]
Type species: Dysanellus excellens Bernhauer, 1934
Characters/Description: Navarrete-Heredia et al. 2002: 299 (characters in key); Caron et al. 2016: 63 (characters).
Key: Irmler 1979: 31 (species of the genus); Irmler 1982a: 206 (species of the genus); Márquez et al. 2004: 30
(Mexican species); Torres-Rodríguez et al. 2012: 233 (characters), 236 (characters in key); Aballay et al.
2014: 104 (characters in key); Caron et al. 2016: 65 (species of the genus); Chatzimanolis & Caron 2016: 140
(Key to the punctatus species group in Xenopygus).
Distribution.
Other regions: Oceanic.
1037. Zackfalinus Chatzimanolis, 2012: 265 (Fig. 231)

Type species: Zackfalinus kitrinos Chatzimanolis, 2012

Characters/Description: Chatzimanolis 2015c: 17 (characters).
Distribution.
Central America: Mexico, Guatemala. Belize, Nicaragua, Costa Rica, Panama.
South America: Colombia, Venezuela, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay.
Incertae sedis in Tribe Staphylinini Latreille, 1802
1038. Quediomacrus Sharp, 1884: 339

Type species: Quedius puniceipennis Solsky, 1868

Distribution.
1039. Termitoquedius Bernhauer, 1912c: 45

Type species: Termitoquedius iheringi Bernhauer, 1912
Ecitolycus Wasmann, 1925b: 76, 161 [synonymized by Seevers 1965: 329]

Distribution.
1040. Valdiviodes Smetana, 1981: 349

Type species: Valdiviodes ashworthi Smetana, 1981

Characters/Description: Smetana 1981: 354 (characters in key).

Distribution.
Tribe Xantholinini Erichson, 1839a
1041. Agerodes Motschulsky, 1858b: 208

Type species: Agerodes coeruleus Motschulsky, 1858
Hymeneus Sharp, 1885: 487 [synonymized by Bernhauer & Schubert 1914: 312]
Type species: Hymeneus godmani Sharp, 1885
Characters/Description: Navarrete-Heredia et al. 2002: 296 (characters in key); Janák & Bordoni 2014: 257, 262
(characters); Janák & Bordoni 2015: 18 (characters); Márquez & Asiain 2016: 254 (characters).
Distribution.
Central America: Mexico, Nicaragua, Costa Rica, Panama.
South America: Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil, Paraguay, Argentina.
1042. Agrodes Nordmann, 1837: 161

Type species: Agrodes elegans Nordmann, 1837
Characters/Description: Asiain et al. 2007: 41 (characters).
Key: Asiain et al., 2007: 46 (species of the genus).
Distribution.
South America: Colombia, Ecuador, Peru, Bolivia, Brazil.
1043. Allotrichus Sharp, 1885: 486

Type species: Allotrichus arenarius Sharp, 1885

Distribution.
1044. Eulissus Mannerheim, 1830: 8, 35

Type species: Eulissus chalybaeus Mannerheim, 1830
Dinolinus Casey, 1906: 366, 373 [synonymized by Bernhauer & Schubert 1914: 310]
Type species: Eulissus chalybaeus Mannerheim, 1830
Xanthohypnus Casey, 1906: 367, 374 [synonymized by Bernhauer & Schubert 1914: 310]
Type species: Xantholinus strigiceps Sharp, 1885
Characters/Description: Lynch-Arribálzaga 1884: 178 (characters); Steel 1938: 55 (characters); Navarrete-Heredia
et al. 2002: 296 (characters in key); Janák & Bordoni 2014: 257, 262 (characters); Janák & Bordoni 2015: 18
(characters); Newton 2017: 30 (characters).
Distribution.
South America: Colombia, Venezuela, Suriname, French Guiana, Peru, Bolivia, Brazil, Paraguay, Argentina.
Other regions: Palaearctic [adventive].
1045. Gauropterus Thomson, 1860: 187

Type species: Staphylinus fulgidus Fabricius, 1787

Characters/Description: Coiffait & Sáiz 1964: 511 (characters in key); Smetana 1982: 60 (characters in key), 71
(characters); Bordoni 2002: 62 (characters in key), 184 (characters); Janák & Bordoni 2015: 18 (characters);
Márquez & Asiain 2016: 254 (characters); Bordoni 2016: 416 (characters in key), 508 (characters).
Distribution.
1046. Gyrohypnus Leach in Samouelle, 1819: 172 (Fig. 192)

Type species: Staphylinus fracticornis O. Müller, 1776
Hyponygrus Tottenham, 1940: 49 [synonymized by Coiffait 1962b: 390]
Subgenera:
Gyrohypnus Leach in Samouelle, 1819: 172
Xenoxantholinus Coiffait, 1972: 175, 177, 187
Type species: Xantholinus marginalis Wollaston, 1862
Characters/Description: Lynch-Arribálzaga 1884: 188 (characters); Smetana 1982: 61 (characters in key), 187
(characters); Bordoni 2002: 71 (characters in key), 616 (characters); Bordoni 2016: 416 (characters in key),
579 (characters).
Key: Smetana, 1978: 338 (some species of the genus).
Distribution.
1047. Heterolinus Sharp, 1885: 475 (Fig. 193)

Type species: Xantholinus puncticeps Guérin-Méneville, 1844
Characters/Description: Márquez 2001: 318 (characters); Márquez & Asiain 2016: 254 (characters).
Key: Márquez 2001: 321 (species of the genus).
Distribution.
South America: Colombia, Ecuador, Peru, Bolivia.
1048. Homalolinus Sharp, 1885: 472 (Fig. 195)

Type species: Xantholinus canaliculatus Erichson, 1839
Ehomalolinus Bierig, 1934a: 16 [synonymized by Márquez 2003: 496]
Type species: Ehomalolinus punctipennis Bierig, 1934

Characters/Description: Navarrete-Heredia et al. 2002: 296 (characters in key); Márquez 2003: 496 (characters);
Márquez & Asiain 2016: 254 (characters).
Distribution.
1049. Lepitacnus Smetana, 1982: 103

Type species: Leptacinus pallidulus LeConte, 1880

Distribution.
1050. Leptacinus Erichson, 1839b: 429

Type species: Staphylinus batychrus Gyllenhal, 1827
Leptacinodes Casey, 1906: 369, 401 [synonymized by Bernhauer & Schubert 1914: 292]
Type species: Staphylinus batychrus Gyllenhal, 1827

Characters/Description: Blackwelder 1943: 473 (characters in key), 493 (characters); Coiffait & Sáiz 1964: 510
(characters in key); Smetana 1982: 60 (characters in key), 92 (characters); Bordoni 2002: 71 (characters in
key), 605 (characters); Navarrete-Heredia et al. 2002: 297 (characters in key).
Key: Coiffait & Sáiz 1964: 512 (species of Chile); Blackwelder 1943: 493 (species of the West Indies).
Distribution.
South America: Galapagos Isl., Chile, Argentina.
1051. Linosomus Kraatz, 1857: 647

Type species: Gyrohypnus tenuicornis Nordmann, 1837
Kainolinus Coiffait & Sáiz, 1964: 511, 519 [synonymized by Bordoni 2001: 163]
Type species: Kainolinus approbabilis Coiffait & Sáiz, 1964
Notolinus Casey, 1906: 375 [synonymized by Bordoni 2016: 628]
Type species: Notolinus fumipennis Casey, 1906

Characters/Description: Coiffait & Sáiz 1964: 511 (characters in key as Kainolinus); Bordoni 2016: 628
(characters).
Distribution.
Other regions: Ethiopian, Australian.
1052. Lissohypnus Casey, 1906: 368, 398

Type species: Lissohypnus texanus Casey, 1906
Distribution.
1053. Lithocharodes Sharp, 1876a: 204

Type species: Lithocharodes fuscipennis Sharp, 1876
Lithocharitoides Rye, 1878: 36 [synonymized by Smetana 1982: 120]
Type species: Lithocharodes fuscipennis Sharp, 1876
Nematolinus Casey, 1906: 370, 407 [synonymized by Smetana 1982: 120]
Type species: Leptacinus longicollis LeConte, 1863
Oligolinus Casey, 1906: 372, 420 [synonymized by Smetana 1982: 120]
Type species: Metoponcus floridanus LeConte, 1880
Characters/Description: Lynch-Arribálzaga 1884: 193 (characters); Blackwelder 1943: 473 (characters in key as
Oligolinus), 473 (characters in key), 496 (characters), 504 (characters as Oligolinus); Smetana 1982: 61
Key: Blackwelder 1943: 496 (species of the West Indies), 505 (species of Oligolinus of the West Indies);
Scheerpeltz 1960: 92 (Oligolinus).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Dominican Republic, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
South America: Venezuela, Galapagos Isl., Brazil, Argentina.
1054. Megalinus Mulsant & Rey, 1877: 261

Type species: Staphylinus glabratus Gravenhorst, 1802
Lepidophallus Coiffait, 1956a: 58, 59 [synonymized by Bordoni 2008: 55]
Type species: Xantholinus hesperius Erichson, 1839
Leptophallus Coiffait, 1956a: 59 [synonymized by Coiffait 1983]
Type species: Xantholinus flavocinctus Hochhuth, 1849
Leptophius Coiffait, 1983: 345 [synonymized by Assing 2007a: 43]
Type species: Xantholinus flavocinctus Hochhuth, 1849
Metacyclinus Reitter, 1908: 115 [synonymized by Blackwelder 1943: 473]
Type species: Staphylinus glabratus Gravenhorst, 1802
Characters/Description: Lynch-Arribálzaga 1884: 182 (characters); Coiffait & Sáiz 1964: 512 (characters in key as
Lepidophallus); Sáiz 1970b: 383 (characters as Lepidophallus); Bordoni 2002: 73 (characters in key as
Lepidophallus); 631 (characters as Lepidophallus); Assing 2007a: 43 (characters).
Key: Coiffait & Sáiz 1964: 518 (species of Chile as Lepidophallus).
Distribution.

1055. Microlinus Casey, 1906: 372, 419 (Fig. 200)
Type species: Leptolinus pusio LeConte, 1880
2 species are included in this genus from West and South America.
Characters/Description: Blackwelder 1943: 473 (characters in key), 502 (characters); Smetana 1982: 60 (characters
in key), 112 (characters); Navarrete-Heredia et al. 2002: 297 (characters in key); Newton & Peck 2006: 152
(characters in key); Busanello & Caron 2016: 606 (characters).
Key: Busanello & Caron 2016: 610 (species of the genus).
Distribution.
West Indies: Jamaica, Montserrat, Trinidad-Tobago.
South America: Galapagos Isl., Brazil.
1056. Neohypnus Coiffait & Sáiz, 1964: 512, 522

Type species: Neohypnus chilensis Coiffait & Sáiz, 1964
Sungaria Bordoni, 2003: 221 [synonymized by Bordoni 2013: 53]
Type species: Xantholinus mandschuricus Bernhauer, 1923

Characters/Description: Coiffait & Sáiz 1964: 512 (characters in key); Smetana 1982: 62 (characters in key), 198
(characters); Navarrete-Heredia et al. 2002: 297 (characters in key); Aballay et al. 2014: 103 (characters in
key); Newton 2017: 31 (characters).
Key: Coiffait & Sáiz 1964: 522 (species of Chile).
Distribution.
West Indies: Cuba, Jamaica, Haiti, Puerto Rico, US Virgin Isl., Antigua & Barbuda, Saint Kitts & Nevis,
Montserrat, Guadeloupe, Dominica, Saint Vincent & the Grenadines, Barbados, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, French Guiana, Galapagos Isl., Bolivia, Brazil, Paraguay, Uruguay, Chile,
Argentina.
1057. Neoleptacinus Coiffait & Sáiz, 1964: 511, 515

Type species: Leptolinus cribripennis Fairmaire & Germain, 1862

Characters/Description: Coiffait & Sáiz 1964: 511 (characters in key); Sáiz 1968b: 89 (characters).
Key: Sáiz 1968b: 90 (species of the genus).
Distribution.
1058. Neoxantholinus Cameron, 1945b: 783

Type species: Metoponcus rufulus Broun, 1880
Neolinus Coiffait, 1981b: 292 [synonymized by Bordoni 2004: 44]
Type species: Neolinus galapagoensis Coiffait, 1981
20 species (1 in Dominican amber) are included in this genus from America.
Characters/Description: Bordoni 2002: 60 (characters in key), 127 (characters); Navarrete-Heredia et al. 2002: 297
(characters in key); Newton & Peck 2006: 151 (characters in key); Bordoni 2016: 415 (characters in key), 456
(characters).
Distribution.
West Indies: Cuba, Cayman Is., Haiti, Dominican Republic, Montserrat, St. Vincent & Grenadines, Grenada.
South America: Colombia, Venezuela, Ecuador, Galapagos Isl., Brazil.
Other regions: Nearctic, Ethiopian, Oriental, Australian, Oceanic.
1059. Nudobius Thomson, 1860: 188

Type species: Staphylinus lentus Gravenhorst, 1806
Pedinolinus Bernhauer, 1912a: 479 [synonymized by Bordoni 2016: 470]
Type species: Nudobius africanus Bernhauer, 1912

Characters/Description: Smetana 1982: 60 (characters in key), 75 (characters); Bordoni 2002: 61 (characters in
key), 177 (characters); Navarrete-Heredia et al. 2002: 296 (characters in key); Bordoni 2016: 416 (charactes
Key: Smetana 1980: 159 (Mexican species).
Distribution.
Central America: Mexico, Guatemala, Honduras, Costa Rica.
1060. Pachycorynus Motschulsky, 1858a: 204

Type species: Pachycorynus dimidiatus Motschulsky, 1858
Holisomorphus Kraatz, 1859b: 100
Type species: Holisomorphus ceylanensis Kraatz, 1859
Distribution.
Other regions: Ethiopian, Madagascan, Oriental, Oceanic.
1061. Phacophallus Coiffait, 1956a: 50 (Fig. 207)

Type species: Staphylinus parumpunctatus Gyllenhal, 1827
Characters/Description: Smetana 1982: 60 (characters in key), 106 (characters); Bordoni 2002: 68 (characters in
key), 531 (characters); Navarrete-Heredia et al. 2002: 297 (characters in key); Bordoni 2016: 417 (characters
Distribution.
West Indies: Bahamas, Cuba, Jamaica, Puerto Rico, Saint-Kitts & Nevis, Antigua & Barbuda, St. Vincent &
Grenadines, Barbados, Grenada.
1062. Plochionocerus Dejean, 1833: 64 (Figs. 220–222)

Type species: Staphylinus violaceus Olivier, 1795
Araeocnemus Nordmann, 1837: 163 [synonymized by Gemminger & Harold 1868: 599]

Type species: Staphylinus fulgens Fabricius, 1792
Plociocerus Agassiz, 1846: 299 [synonymized by Blackwelder 1952: 315]
Sterculia Laporte, 1835: 118 [synonymized by Blackwelder 1952: 360]

2002: 296 (characters in key); Asiain et al. 2007: 9 (characters); Márquez & Asiain 2016: 254 (characters).
Key: Asiain et al. 2007: 10 (species of the genus).
Distribution.
South America: Colombia, Venezuela, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Argentina.
1063. Renda Blackwelder, 1952: 338

Type species: Sterculia formicaria Laporte, 1835
Plochionocerus Sharp, 1885: 471 [synonymized by Blackwelder 1952: 338]
Type species: Sterculia formicaria Laporte, 1835

Characters/Description: Navarrete-Heredia et al. 2002: 296 (characters in key); Márquez 2010: 7 (characters);
Márquez & Asiain 2016: 254 (characters).
Distribution.
1064. Scytalinus Erichson, 1839a: 305 (Fig. 224)

Type species: Scytalinus serpentinus Erichson, 1839

Characters/Description: Navarrete-Heredia et al. 2002: 296 (characters in key); Márquez & Asiain 2016: 254
(characters).
Distribution.
South America: Venezuela, French Guiana, Peru, Brazil, Uruguay, Argentina.
1065. Somoleptus Sharp, 1885: 494

Type species: Somoleptus aenescens Sharp, 1885
Distribution.
South America: Colombia, Venezuela, Bolivia, Brazil, Paraguay, Argentina.
1066. Stenolinus Bierig, 1937b: 273
Type species: Stenolinus macrothrichus Bierig, 1937
Characters/Description: Blackwelder 1943: 473 (characters in key), 503 (characters).
Distribution.
West Indies: Cuba.
1067. Tesba Sharp, 1876a: 194

Type species: Tesba gigas Sharp, 1876
Distribution.
Central America: Guatemala, Nicaragua.
South America: Peru, Bolivia, Brazil, Paraguay, Argentina.
1068. Thyreocephalus Guérin-Méneville, 1844: 10 (Fig. 227)

Type species: Thyreocephalus jekelii Guérin-Méneville, 1844
Indoscytalinus Heller, 1900: 5 [synonymized by Bernhauer & Schubert 1914: 313]
Type species: Xantholinus albicornis Heller, 1900
Linidius Sharp, 1876a: 196 [synonymized by Steel 1938: 56]
Type species: Linidius recticollis Sharp, 1876
Saurohypnus Sharp, 1885: 501 [synonymized by Smetana 1977: 348]
Type species: Saurohypnus scutellaris Sharp, 1885

Characters/Description: Steel 1938: 55 (characters); Blackwelder 1943: 473 (characters in key), 490 (characters);
Smetana 1982: 60 (characters in key), 66 (characters); Bordoni 2002: 60 (characters in key), 63 (characters in
key), 210 (characters); Navarrete-Heredia et al. 2002: 296 (characters in key); Janák & Bordoni 2014: 257
(characters); Janák & Bordoni 2015: 14, 18 (characters); Márquez & Asiain 2016: 254 (characters); Newton
2017: 30 (characters); Bordoni 2016: 416 (characters in key), 520 (characters).
Key: Smetana 1977: 350 (some Mexican species of Thyreocephalus); Márquez & Asiain 2016: 255 (Mexican
species of Thyreocephalus).
Distribution.
South America: Colombia, Venezuela, Peru, Bolivia, Brazil.
1069. Xantholinus Dejean, 1821: 23

Type species: Staphylinus linearis Olivier, 1795
Afrolinus Coiffait, 1962c: 73
Type species: Xantholinus algericus Coiffait, 1962
Calolinus Coiffait, 1956a: 58, 61
Type species: Xantholinus piochardi Coiffait, 1956
Echinophallus Coiffait, 1956a: 58, 60 [synonymized by Smetana 1978: 339]
Type species: Xantholinus translucidus Scriba, 1870

Echinophius Coiffait, 1983: 345 [synonymized by Coiffait 1983: 345]
Type species: Xantholinus translucidus Scriba, 1870
Heterolinus Coiffait, 1966: 199 [synonymized by Coiffait 1983: 345]
Type species: Xantholinus khnzoriani Coiffait, 1966
Heterolius Coiffait, 1983: 345
Type species: Xantholinus khnzoriani Coiffait, 1966
Idiolinus Casey, 1906: 368, 375 [synonymized by Bernhauer & Schubert 1914: 300]
Type species: Xantholinus crassicornis Hochhuth, 1851
Neoleptophallus Bordoni, 1985: 84
Type species: Xantholinus minutus Coiffait, 1962
Paracyclinus Bordoni, 1975: 74, 89
Type species: Xantholinus procerus Erichson, 1839
Polydontophallus Bordoni, 1972: 161, 176
Type species: Gyrohypnus meridionalis Nordmann, 1837
Purrolinus Coiffait, 1956a: 58, 65
Type species: Staphylinus tricolor Fabricius, 1787
Stenophallus Bordoni, 1972: 160
Type species: Xantholinus laniger Fauvel, 1899
Tetralinus Bordoni, 1975: 69, 89
Type species: Xantholinus haematodes Kolenati, 1846
Toxophallus Bordoni, 1972: 160
Type species: Xantholinus heinzi Coiffait, 1970
Subgenera:
Helicophallus Coiffait, 1956a: 58, 70
Type species: Xantholinus rufipes P. Lucas, 1846
Typhlolinus Reitter, 1908: 122
Type species: Xantholinus hungaricus Reitter, 1908
Acanthophallus Coiffait, 1956a: 59, 67 [synonymized by Janák 1993: 47]
Type species: Xantholinus raffrayi Sainte-Claire Deville, 1926
Acanthophus Coiffait, 1983: 345 [synonymized by Janák 1993: 47]
Type species: Xantholinus raffrayi Sainte-Claire Deville, 1926
Xantholinus Dejean, 1821: 23
Type species: Staphylinus linearis Olivier, 1795
Meneidophallus Bordoni, 1999: 119 [synonymized by Assing 2008a: 1277]
Type species: Xantholinus roubali Coiffait, 1956
Paraphallus Bordoni, 1972: 162, 197 [synonymized by Janák 1993: 47]
Type species: Xantholinus longiventris Heer, 1839
Phallolinus Ádám, 1987a: 135
Type species: Xantholinus longiventris Heer, 1839

(characters); Smetana 1982: 61 (characters in key), 264 (characters); Bordoni 2002: 71 (characters in key),
589 (characters); Navarrete-Heredia et al. 2002: 297 (characters in key); Janák & Bordoni 2014: 257
(characters); Newton 2017: 31 (characters).
Distribution.
FIGURES 4–15. Habitus; Glypholomatinae (4–5), Microsilphinae (6), Omaliinae (7–11), Proteininae (12–15); 4, after Thayer
1997; 5, after Thayer 1997; 6, after Scheerpeltz 1972d; 7, after Thayer 2003; 8, after Thayer 2003; 9, after Thayer 2003; 10,
after Thayer 2003; 11, after Scheerpeltz 1972d; 12, after Steel 1966; 13, after Cuccodoro 2011; 14, after Bierig 1940; 15, after
Frank & Thomas 1984b; slightly modified. Scale bars: 1 mm.

FIGURES 16–27. Habitus; Micropeplinae (16–18), Neophoninae (19), Pselaphinae (20–27); 16, after Campbell 1968; 17,
after Campbell 1973c; 18, after Tronquet & Degallier 2010; 19 Chile: region of Los Lagos (VMDC); 20, after Reitter 1883b;
21, after Reitter 1889; 22, after Bryant 1915; 23, after Raffray 1887; 24, after Asenjo 2016; 25, after Asenjo et al. 2017; 26,
after O. Park 1952b; 27, after Bruch 1929; slightly modified. Scale bars: 1 mm.
FIGURES 28–39. Habitus; Pselaphinae (28–29), Phloeocharinae (30–31), Tachyporinae (32–38), Habrocerinae (39); 28, after
Bruch 1933c; 29, after Cuccodoro et al. 2012; 30, after Navarrete-Heredia et al. 2002; 31, after Sharp 1887a; 32, after
Klimaszewski et al. 2013; 33, after Sharp 1883; 34, after Sharp 1884; 35, after Scheerpeltz 1972d; 36, after Sharp 1884; 37,
after Campbell 1979; 38, after Bruch 1932; 39, after Klimaszewski et al. 2013; slightly modified. Scale bars: 1 mm.

FIGURES 40–51. Habitus; Habrocerinae (40), Aleocharinae (41–51); 40, after Solier 1849; 41, Brazil: state of Paraná
(MUSM); 42, Chile: region of Antofagasta (MUSM); 43, after Pace 2015e; 44, after Ashe 2002a; 45, after Pace 2014a; 46, after
Pace 2012; 47, after Y.H. Kim & Ahn 2015; 48, after Pace 2014c; 49, Chile: region of Araucanía (MUSM); 50, after Irmler
2011; 51, after Caron & Ribeiro-Costa 2008; slightly modified. Scale bars: 1 mm.
FIGURES 52–63. Habitus; Aleocharinae (52–63); 52, after Bruch 1917; 53, Peru: department of Madre de Dios (CEMT); 54,
Peru: department of Madre de Dios (CEMT); 55, after Pace 2015e; 56, after Pace 2015a; 57, after Pace 2011b; 58, after Pace
2007; 59, after Pace 2008d; 60, after Pace 2015e; 61, after Pace 2014c; 62, after Pace 2011b; 63, after Pace 2011b; slightly
modified. Scale bars: 1 mm.

FIGURES 64–75. Habitus; Aleocharinae (64–75); 64, after Pace 2015a; 65, after Bortoluzzi et al. 2016; 66, after Kanao et al.
2012; 67, after Pace 1999c; 68, Peru: department of Madre de Dios (CEMT); 69, after Pace, 2011b; 70, after Bruch 1933a; 71,
after Pace 2015e; 72, Chile: region of Coquimbo (MUSM); 73, after Pace 1996b; 74, after Pace 1987b; 75, after Kistner 2006;
slightly modified. Scale bars: 1 mm.
FIGURES 76–87. Habitus; Aleocharinae (76–80); Scaphidiinae (81–85); Piestinae (86–87); 76, after Kistner 2006; 77, after
Borgmeier 1950; 78, after Pace 2007; 79, after Kistner 2006; 80, after Pace 2014c; 81, after Matthews 1888; 82, after Fierros-
Lopéz 2005; 83, after Fierros-Lopéz 2005; 84, after Fierros-Lopéz 2006; 85, after Matthews 1888; 86, after Bortoluzzi et al.
2017; 87, Peru: department of Cuzco (CEMT); slightly modified. Scale bars: 1 mm.

FIGURES 88–99. Habitus; Piestinae (88–91); Osoriinae (92–99); 88, after Pérez et al. 2017; 89, Peru: department of Madre
de Dios (MUSM); 90, Peru: department of Cuzco (MUSM); 91, after Brunke et al. 2011; 92, after Sharp 1887a; 93, after Sharp
1887a; 94, after Irmler 1991; 95, after Irmler 2001b; 96, after Bierig 1931; 97, after Sharp 1887a; 98, after Caron et al. 2012;
99, after Bruch 1942; slightly modified. Scale bars: 1 mm.
FIGURES 100–111. Habitus; Osoriinae (100–111); 100, after Sharp 1887a; 101, after Irmler 2007b; 102, after Fauvel 1901;
103, Brazil (CEMT); 104, Brazil: state of São Paulo (CEMT); 105, after Irmler 2003a; 106, after Sharp 1887a; 107, after Irmler
2003c; 108, after Irmler 2003a; 109, after Scheerpeltz 1951; 110, after Sharp 1887a; 111, after Bruch 1942; slightly modified.
Scale bars: 1 mm.

FIGURES 112–123. Habitus; Osoriinae (112–118); Oxytelinae (119–123); 112, after Sharp 1887a; 113, after Bruch 1942; 114,
after Sharp 1887a; 115, after Irmler 2003c; 116, after Sharp 1887a; 117, after Irmler 2003a; 118, after Sharp 1887a; 119, after
Sharp 1887a; 120, after Caron & Ribeiro-Costa 2007; 121, after Caron & Ribeiro-Costa 2007; 122, after Sharp 1887a; 123,
after Makranczy 2014; slightly modified. Scale bars: 1 mm.
FIGURES 124–135. Habitus; Oxytelinae (124–125); Oxyporinae (126–127); Megalopsidiinae (128–131); Scydmaeninae
(132–135); 124, after Palomino & Dale 1989; 125, after Sharp 1887a; 126, after Márquez & Asiain 2006; 127, after Márquez et
al. 2005; 128, after Puthz 2012; 129, after Sharp 1887a; 130, after Puthz 2012; 131, Chile: region of Araucanía (VMDC); 132,
after Jałoszyński 2013a; 133, after Jałoszyński 2013e; 134, after Jałoszyński 2011a; 135, after Lord et al. 2014; slightly

FIGURES 136–147. Habitus; Scydmaeninae (136–139); Steninae (140–141); Euaesthetinae (142–145); Solieriinae (146);
Leptotyphlinae (147); 136, after Jałoszyński 2014b; 137, after Jałoszyński 2012c; 138, after Jałoszyński 2013d; 139, after
Jałoszyński 2014c; 140, after Fauvel 1901; 141, after Fauvel 1901; 142, after Puthz 2008; 143, after Puthz 2006b; 144, after
Puthz 2006b; 145, after Pace 1986d; 146, after Coiffait & Sáiz 1968; 147, after Coiffait & Decou 1972; slightly modified. Scale
bars: 1 mm.
FIGURES 148–159. Habitus; Pseudopsinae (148–150); Paederinae (151–159); 148, after Herman 1975; 149, Ecuador:
province of Napo (CNC); 150, after Asenjo & Ribeiro-Costa 2010; 151, after Bierig 1933; 152, after Sharp 1886; 153, after
Assing 2012a; 154, after Jiménez-Sánchez & Galián 2013; 155, after Bierig 1933; 156, after Bruch 1928; 157, after Assing
2012b; 158, after Borgmeier 1936; 159, after Sharp 1886; slightly modified. Scale bars: 1 mm.

FIGURES 160–171. Habitus; Paederinae (160–171); 160, after Assing 2013; 161, after Bierig 1933; 162, after Coiffait & Sáiz
1968; 163, after Asenjo 2011; 164, after Fauvel 1901; 165, after Herman 2013; 166, after Sharp 1886; 167, after Vieira et al.
2014; 168, after Fauvel 1901; 169, after Campbell & Peck 1989; 170, after Sharp 1885; 171, after Bruch 1930a; slightly
FIGURES 172–183. Habitus; Paederinae (172–175); Staphylininae (176–183); 172, after Fauvel 1901; 173, after Assing 2013;
174, after Herman 1979; 175, after Sharp 1886; 176, after Brunke & Solodovnikov 2014a; 177, Chile: region of Valparaíso
(MUSM); 178, Chile: region Metropolitana de Santiago (MUSM); 179, Chile: region of Araucanía (MUSM); 180, Chile:
region of Valparaíso (MUSM); 181, after Ashe & Timm 1988; 182, Brazil: state of Paraná (MUSM); 183, Chile: region of
Antofagasta (CEMT); slightly modified. Scale bars: 1 mm.

FIGURES 184–195. Habitus; Staphylininae (184–195); 184, Peru: department of Lima (MUSM); 185, Brazil: state of Paraná
(MUSM); 186, after Chatzimanolis 2014a; 187, after Santiago-Jiménez 2008; 188, after Smetana 1982; 189, after
Chatzimanolis 2012; 190, Peru: department of Madre de Dios (CEMT); 191, after Navarrete-Heredia & Márquez 1998; 192,
after Klimaszewski et al. 2013; 193, after Márquez 2001; 194, after Erichson 1840; 195, after Márquez 2003; slightly
FIGURES 196–207. Habitus; Staphylininae (196–207); 196, after Chatzimanolis 2008; 197, Venezuela: state of Merida
(MUSM); 198, Peru: department of Puno (MEKRB); 199, after Chani-Posse & Solodovnikov 2017; 200, after Busanello &
Caron 2016; 201, after Brunke & Solodovnikov 2014b; 202, after Asiain & Márquez 2003; 203, after Asiain & Márquez 2012;
204, after Chani-Posse & Solodovnikov 2017; 205, after Costa-Silva et al. 2018; 206, after Chani-Posse 2014b; 207, after
Klimaszewski et al. 2013; slightly modified. Scale bars: 1 mm.

FIGURES 208–219. Habitus; Staphylininae (208–219); 208, Chile: region Metropolitana de Santiago (MUSM); 209, after
Navarrete-Heredia 2003; 210, after Asenjo & Ribeiro-Costa 2009; 211, Peru: department of Cuzco (MUSM); 212, Argentina:
province of Jujuy (MUSM); 213, Peru: department of Madre de Dios (MUSM); 214, Peru: department of Cuzco (MUSM); 215,
Peru: department of Madre de Dios (MUSM); 216, Peru: department of Lima (MUSM); 217, Peru: department of Cuzco
(MUSM); 218, Brazil: state of Paraná (MUSM); 219, after Smetana 1982; slightly modified. Scale bars: 1 mm.
FIGURES 220–231. Habitus; Staphylininae (220–231); 220, Peru: department of Madre de Dios (MUSM); 221, Peru:
department of Madre de Dios (MUSM); 222, Peru: department of Madre de Dios (MUSM); 223, Chile: region Metropolitana
de Santiago (MUSM); 224, after Erichson 1840; 225, after Solodovnikov 2006; 226, after Chatzimanolis 2013; 227, after
Márquez & Asiain 2016; 228, Peru: department of Loreto (CEMT); 229, Peru: department of Junin (MUSM); 230, Peru:
department of Madre de Dios (CEMT); 231, after Chatzimanolis 2012; slightly modified. Scale bars: 1 mm.

Dominica, St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.
South America: Colombia, Venezuela, French Guiana, Bolivia, Brazil, Paraguay, Uruguay, Argentina.
Incertae sedis in family Staphylinidae
1070. Apticax Schomann & Solodovnikov, 2012: 381

Type species: Apticax volans Schomann & Solodovnikov, 2012
2 fossil species are known from and are restricted to Brazil.

Distribution.
Taxonomic considerations in Subfamily Pselaphinae
Batrisodes Reitter (Pselaphinae)

The species Batrisodes asteriscus (L. W. Schaufuss) distributed in Colombia and Venezuela, does not
belong to Batrisodes or Batrisus where originally was described (Chandler, personal opinion).
Batrisus Aubé (Pselaphinae)

Batrisus crassipes Sharp of Costa Rica, Panama, Colombia; and Batrisus lamellipes Sharp of Panama, are
members of an undescribed genus (Chandler, personal opinion).
Bryaxis Kugelann (Pselaphinae)

Bryaxis atrata L. W. Schaufuss of Colombia, and Bryaxis proportionalis L. W. Schaufuss of Peru do not
belong to Bryaxis (Chandler, personal opinion).
Enoptostomus Schaum (Pselaphinae)

Enoptostomus clandestinus L. W. Schaufuss of Brazil, probably an error for Ctenisis (Chandler, personal
opinion).
Tyrus Aubé (Pselaphinae)

Tyrus compressicollis L. W. Schaufuss of Chile, probably an error for Tyropsis (Chandler, personal opinion).
Acknowledgements
We are grateful to Drs. Mabel Alvarado, James S. Ashe (†), Volker Assing, Julieta Asiain, Mariana R. Chani-Posse,
Stylianos Chatzimanolis, Emilia Constanza-Perez, Vladimir I. Gusarov, Lee H. Herman, Peter Hlaváč, Paweł
Jałoszyński, David H. Kistner, Gerardo Lamas, Dewanand Makhan, György Makranczy, Juan Márquez, José L.
Navarrete-Heredia, Alfred F. Newton, Roberto Pace (†), Nigel Pitman, Volker Puthz, Ernesto Rázuri-Gonzales,
Quiyari Santiago, Michael Schülke, Diana Silva, Margaret K. Thayer, Biodiversity Heritage Library, Gallica and
Sci-hub for providing many of the publications necessary for completion of this work. We also thank György
Makranczy of the Hungarian Natural History Museum (HNHM) for his excellent contributions and comments in
the section of Oxytelinae. Rodrigo Lopes-Ferreira (UFLA) for the providing photographic equipment. The curators
of insect collections: Fernando Z. Vaz-de-Mello (CEMT), Anthony Davies (CNC), Diana Silva (MUSM) and
Gerardo Lamas (MUSM) are thanked for allowing to take photographs of some specimens loaned. The authors,
and editors are acknowledged for allowing us to use the images included in this work. Additionally, we sincerely
appreciated the reviewers Adam J. Brunke and Margaret K. Thayer for their useful comments on manuscript.
Finally, Victor Manuel Diéguez (VMDC) is thanked for the picture of Megalopinus sanguinitriguttatus
(Scheerpeltz) and Vinícius Costa-Silva (UNICAMP) for the picture of Ontholestes brasilianus Bernhauer. This
work was supported by the National Council for Scientific and Technological Development of the Ministry of
Science, Technology, and Innovation of Brazil, CNPq (151524/2018-1 and 300651/2019-8) granted to first author.
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Museum di Vienna (Coleoptera, Staphylinidae) (XXII Contributo alla conoscenza delle Aleocharinae). Annalen des
Naturhistorischen Museums in Wien, 85B, 53–102.
Pace, R. (1983d) Nuove Aleocharinae brasiliane e boliviane del Museo Zoologico dell'Università Humboldt di Berlino
(Coleoptera Staphylinidae) (XXXVIII contributo alla conoscenza delle Aleocharinae). Giornale Italiano di Entomologia,
1, 295–316.
Pace, R. (1983e) Le specie del genere Leptusa del Monte Baldo (Coleoptera, Staphylinidae). Bollettino del Museo Civico di
Storia Naturale di Verona, 9, 121–138. [Note: nominal year of publication 1982]
Pace, R. (1984) Nuove Aleocharinae microftalme Mediterranee e dell'Iran, del Muséum d'Histoire Naturelle di Ginevra
(Coleoptera Staphylinidae). Archives des Sciences (Genève), 37, 211–219.
Pace, R. (1985) Aleocharinae del Venezuela riportate dal Prof. Franz (Coleoptera, Staphylinidae) (LXVI contributo alla
conoscenza delle Aleocharinae). Giornale Italiano di Entomologia, 2, 371–392.
Pace, R. (1986a) Aleocharinae del Perù (Coleoptera, Staphylinidae) (LXXXV Contributo alla conoscenza delle Aleocharinae).
Redia, 69, 417–467.
Pace, R. (1986b) Aleocharinae dell'Africa Orientale (Coleoptera, Staphylinidae). Annales Historico-Naturales Musei
Nationalis Hungarici, 78, 83–143. [Note: nominal year of publication 1985]
Pace, R. (1986c) Due Aleocharinae troglossene del Venezuela (Coleoptera Staphylinidae) (LXIII contributo alla conoscenza
delle Aleocharinae). Bollettino del Museo Civico di Storia Naturale di Verona, 11, 493–503. [Note: nominal year of
publication 1984]
Pace, R. (1986d) Phaenoctavius brasiliensis n. gen., n. sp. del Brasile e Tachinus (Tachinoderus) rougemonti n. sp. della
Thailandia (Coleoptera Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 11, 45–51. [Note:
Pace, R. (1987a) Aleocharinae dell'isola della Guadalupa (Antille) (Coleoptera: Staphylinidae). Folia Entomologica
Hungarica, 48, 187–205.
Pace, R. (1987b) Aleocharinae del Cile (Coleoptera Staphylinidae) (XCVII Contributo alla conoscenza delle Aleocharinae).
Redia, 70, 459–522.
Pace, R. (1987c) Revision delle Aleocharinae dell'Argentina sudorientale descritte da Scheerpeltz nel 1972 (Coleoptera:
Staphylinidae). Folia Entomologica Hungarica, 48, 161–185.
Pace, R. (1988) Aleocharinae del Cile meridionale (Coleoptera, Staphylinidae) (LXXXVI Contributo alla conoscenza delle
Aleocharinae). Lavori della Societa Veneziana di Scienze Naturali, 13, 85–99.
Pace, R. (1989) Monografia del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Memorie del Museo Civico di Storia
Naturale di Verona (A: Biologica), 2 (8), 1–307.
Pace, R. (1990a) Nuovi Falagriini, Hoplandriini ed Aleocharini della Regione Neotropica (Coleoptera, Staphylinidae) (LXXX
contributo alla conoscenza delle Aleocharinae). Giornale Italiano di Entomologia, 5, 157–180.
Pace, R. (1990b) Aleocharinae neotropiche del Museo Ungherese di Storia Naturale (Coleoptera: Staphylinidae). Annales
Historico-Naturales Musei Nationalis Hungarici, 81, 53–107. [Note: nominal year of publication 1989]
Pace, R. (1990c) Aleocharinae nepalesi del Museo di Ginevra. Parte III. Revisione delle specie himalayane del sottogenere
Microdota Mulsant & Rey (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 97 (4), 901–979.
Pace, R. (1991a) Aleocharinae dello Yemen (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di
Verona, 15, 125–150. [Note: nominal year of publication 1988]
Pace, R. (1991b) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera,
Pace, R. (1991c) Aleocharinae nepalesi del Museo di Ginevra, Parte IV: Autaliini ad Athetini (Coleoptera, Staphylinidae).
Revue Suisse de Zoologie, 9 (1) 8, 107–158.
Pace, R. (1992a) Kaweshkarusa daccordii n. gen. e n. sp. del Cile (Coleoptera, Staphylinidae). (CII contributo alla conoscenza
delle Aleocharinae). Bollettino del Museo Civico di Storia Naturale di Verona, 16, 357–364. [Note: nominal year of
publication 1989]
Pace, R. (1992b) Il genere Leptusa Kraatz della sottoregione indocinese (Taiwan e Vietnam). Monografia del genere Leptusa
Kraatz, Supplemento I (Coleoptera: Staphylinidae). Elytron, 5, 111–119. [Note: nominal year of publication 1991]
Pace, R. (1993) Aleocharinae della Cina (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona,
17, 69–125. [Note: nominal year of publication 1990]
Pace, R. (1995) Nuove specie di Leptusa Kr. di Taiwan. Monografia del genere Leptusa Kraatz: Supplemento V (Coleoptera,
Staphylinidae). Bollettino della Società Entomologica Italiana, 126 (3), 243–248. [Note: nominal year of publication
1994]
Pace, R. (1996a) Nuove specie di Leptusa Kraatz di Taiwan. Monografia del genere Leptusa Kraatz, supplemento VI
(Coleoptera Staphylinidae). Bollettino della Società Entomologica Italiana, 128 (1), 29–36.
Pace, R. (1996b) Aleocharinae della Colombia e dell'Ecuador: Parte I (Coleoptera, Staphylinidae) (131º Contributo alla
conoscenza delle Aleocharinae). Revue Suisse de Zoologie, 103 (2), 395–437.
Pace, R. (1996c) Aleocharinae della Colombia e dell'Ecuador: Parte II (Coleoptera, Staphylinidae) (132º Contributo alla
conoscenza delle Aleocharinae). Revue Suisse de Zoologie, 103 (3), 643–676.
Pace, R. (1996d) Amazonopora manausensis, un notevole nuovo genere e nuova specie di Staphylinidae della tribù Diglottini
(Coleoptera) (130º Contributo alla conoscenza delle Aleocharinae). Bollettino della Società Entomologica Italiana, 128
(1), 37–40.
Pace, R. (1997a) Specie del genere Leptusa in Cina. Monografia del genere Leptusa Kraatz: Supplemento VII (Coleoptera,

Pace, R. (1997b) Aleocharinae della Colombia e dell'Ecuador: Parte III (conclusione) (Coleoptera, Staphylinidae). Revue
Pace, R. (1998a) Aleocharinae della Cina: parte IV (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 105 (4), 911–982.
Pace, R. (1998b) Aleocharinae delle Filippine al Museo di Bruxelles (Coleoptera, Staphylinidae). Bulletin et Annales de la
Societe Royale Belge d'Entomologie, 134, 67–74.
Pace, R. (1999a) Nuove specie del genere Leptusa Kraatz raccolte da Manfred Kahlen (Monografia del genere Leptusa Kraatz:
Supplemento VIII) (Coleoptera, Staphylinidae). Veröffentlichungen des Tiroler Landesmuseums Ferdinandeum, 79, 207–
214.
Pace, R. (1999b) Aleocharinae di Hong Kong (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 106 (3), 663–689.
Pace, R. (1999c) Aleocharinae del Cile (Coleoptera, Staphylinidae), (114º Contributo alla conoscenza delle Aleocharinae).
Bollettino del Museo Civico di Storia Naturale di Verona, 23, 119–210.
Pace, R. (2000a) Aleocharinae del Cile, nuove o poco note (Coleoptera, Staphylinifdae), (157º Contributo all conoscenza delle
Aleocharinae). Bollettino del Museo regionale di Scienze naturali di Torino, 17 (2), 355–509.
Pace, R. (2000b) Aleocharinae di Papua-Nuova Guinea (Coleoptera, Staphylinidae), (156° Contributo alla conoscenza delle
Aleocharinae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie, 70, 109–163.
Pace, R. (2001a) Nuove specie Cilene del genere Leptusa. Monografia del genere Leptusa Kraatz: Supplemento XII (162º
Contributo alla conoscenza delle Aleocharinae). Lavori della Societa Veneziana di Scienze Naturali, 26, 7–14.
Pace, R. (2001b) Nuove Aleocharinae del Perù e dell'Ecuador (Coleoptera Staphylinidae), (160º Contributo alla conoscenza
delle Aleocharinae). Bollettino del Museo regionale di Scienze naturali di Torino, 18 (2), 417–432.
Pace, R. (2002) Gyrophaenini e Bolitocharini del Sudamerica (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie (n.
s.), 19 (2), 115–139.
Pace, R. (2003a) Homalotini del Borneo (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 110 (2), 255–313.
Pace, R. (2003b) New or little known Aleocharinae from the Australian Region (Coleoptera, Staphylinidae). In: Daccordi, M.
& Giachino, P.M. (Eds.) Results of the Zoological Missions to Australia of the Regional Museum of Natural Sciences of
Turin, Italy. Museo Regionale di Scienze Naturali, Turin, I. Monografie del Museo Regionale di Scienze Naturali, Torino,
35, 111–186.
Pace, R. (2004a) Zur Kenntnis der Aleocharinae aus Vietnam (Coleoptera, Staphylinidae). Veröffentlichungen des
Naturkundemuseums Erfurt, 23, 195–200.
Pace, R. (2004b) Hygronomini e Athetini della Cina con note sinonimiche (Coleoptera, Staphylinidae). Revue Suisse de
Zoologie, 111 (3), 457–523.
Pace, R. (2004c) Aleocharinae della Cina all'Institut royal des Sciences naturelles de Belgique (Coleoptera, Staphylinidae).
Belgian Journal of Entomology, 6, 353–361.
Pace, R. (2007) Nuovi dati faunistici e tassonomici su specie di Aleocharinae del Sudamerica delle tribu Gyrophaenini,
Placusini, Diestotini, Homalotini, Bolitocharini, Paglini e Falagriini (Coleoptera, Staphylinidae). Beiträge zur
Entomologie, 57 (1), 211–239.
Pace, R. (2008a) New records of Aleocharinae from Ecuador and Peru, with the description of new species, new subgenera and
new genera (Coleoptera, Staphylinidae) In: Giachino, P.M. (Ed.), Biodiversity of South America, I. Memoirs on
Biodiversity, 1. World Biodiversity Association Onlus, Verona. pp. 225–398.
Pace, R. (2008b) Nouve specie di Lomechusini mirmecofili e non mirmecofili dell'Amazzonia (Coleoptera, Staphylinidae).
Pace, R. (2008c) Description of Leptonia onorei n. sp. and Orphnebius belloi n. sp. from Ecuador and new synonym of the
genus Leptonia Sharp 1883 (Coleoptera Staphylinidae). Tropical Zoology, 21 (2), 253–258.
Pace, R. (2008d) Nuovi dati faunistici e tassonomici su Aleocharinae del Sudamerica delle tribù Deinopsini, Athetini e
Thamiaraeini (Coleoptera, Staphylinidae). Beiträge zur Entomologie, 58 (2), 399–439.
Pace, R. (2008e) Aleocharinae (Insecta, Coleoptera, Staphylinidae) de Madagascar. Zoosystema, 30 (3), 555–577.
Pace, R. (2009) Aleocharinae dell’Irian Jaya (Nuova Guinea) al Naturkundemuseum di Erfurt (Insecta: Coleoptera:
Staphylinidae). Vernate, 28, 259–329.
Pace, R. (2011a) Due nuovi generi nove nuove species di Aleocharinae raccolte su infiorescenze di palma in Perù, Ecuador e
Brasile (Coleoptera, Staphylinidae). Revue française d’Entomologie (n. s.), 32 (3–4), 123–134. [Note: nominal year of
publication 2010]
Pace, R. (2011b) New and additional records, new genera and new species of Aleocharinae from Venezuela, Ecuador and Peru
(Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 35, (Botanica Zoologia), 43–70.
Pace, R. (2012) New data, new species, a new genus and a new tribe of Aleocharinae (Coleoptera: Staphylinidae) from the
Antilles. Tropical Zoology, 25 (2), 54–61.
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Pace, R. (2014a) Introduzione alla conoscenza della sottofamiglia Aleocharinae della Guyana Francese: Parte I (Coleoptera,
Staphylinidae). Beiträge zur Entomologie, 64 (1), 59–81.
Pace, R. (2014b) Introduzione alla conoscenza della sottofamiglia Aleocharinae della Guyana Francese: Parte II (Coleoptera,
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(Coleoptera, Staphylinidae). Beiträge zur Entomologie, 65 (1), 75–84.
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species (Coleoptera Staphylinidae). In: Memoirs on Biodiversity vol. 3 (Biodiversity of South America II), WBA Project,
Verona, pp. 15–38.
Pace, R. (2015e) Biodiversity of Aleocharinae from Andean and Brazilian sub-regions: new data, new genera and new species
(Coleoptera Staphylinidae). In: Memoirs on Biodiversity vol. 3 (Biodiversity of South America II), WBA Project, Verona,
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Scheerpeltz, O. (1937b) Eine neue Art der Gattung Trogophloeus Mannh. nebst einer Bestimmungstabelle der aus Nord- und
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Scheerpeltz, O. (1937c) Die Gattung Weiserianum Bernh. Mit einer Beschreibung einer neuen Art der Gattung und einer
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Scheerpeltz, O. (1948) Zwei neue Leptusen aus Kärnten (Coleoptera, Staphylinidae). (43. Beitrag zur Kenntnis der
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Scheerpeltz, O. (1952b) Eine neue Aleocharinen-Gattung und -Art aus Kärnten (Col. Staphylinidae). (51. Beitrag zur Kenntnis
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Scheerpeltz, O. (1953) Rhombopisalia, eine neue Untergattung der Grossgattung Leptusa Kr., mit einer Bestimmungstabelle
der bis jetzt aus dieser Untergattung bekannt gewordenen Arten (Col. Staphylinidae). (52. Beitrag zur Kenntnis der
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Scheerpeltz, O. (1955) Das neotropische Subgenus Cyrtoquedius Bernh. Der Gattung Quedius Steph. mit einer
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Scheerpeltz, O. (1957a) Staphylinidae (Col.) von Sumba und Flores (4. Beitrag zur Kenntnis der Staphyliniden der
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Scheerpeltz, O. (1957b) Vorläufige Diagnosen einiger neuen paläarktischen Arten und Formen der Gattungen Paederidus
Muls. Rey, Paederus Fabr. (mit den neuen Untergattungen Eopaederus, Paederus s. str. nov., Heteropaederus,
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Scheerpeltz, O. (1957c) Zwei neue Arten der neotropischen ektoparasitischen en Gattung Edrabius Fauvel (Coleoptera,
Staphylinidae). Revista Chilena de Entomología, 5, 219–224.
Scheerpeltz, O. (1958) Wissenschaftliche Ergebnisse der von Herrn Dr. K. Lindberg, Lund, im Jahre 1956 nach der Türkei und
Armenien unternommenen Reise. Coleoptera - Staphylinidae. (82. Beitrag zur Kenntnis der paläarktischen Staphyliniden).
Entomologisk Tidskrift (supplement), 78 (10), 3–37. [Note: nominal year of publication 1957]
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Scheerpeltz, O. (1959b) Eine neue Art der Gattung Platyola Muls. Rey aus Niederösterreich, nebst einer Bestimmungstabelle
der bis jetzt bekannt gewordenen und etlichen Bemerkungen über einige der anderen bisherigen Arten dieser Gattung (Col.
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Verhandlungen der Naturforschenden Gesellschaft in Basel, 72 (2), 233–264.
Scheerpeltz, O. (1962a) Staphyliniden aus dem Deutschen Entomologischen Institut seinerziet von Bernhauer als neu erkannt,
aber nicht mehr beschrieben. I. Teil: Arten aus der paläarktischen Faunenregion (Coleoptera: Staphylinidae). (113. Beitrag
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Münchner Entomologischen Gesellschaft, 52, 199–218.
Scheerpeltz, O. (1963a) Eine neue Untergattung und eine neue Art dieser Untergattung, sowie eine neue Übersicht der Arten
der Untergattung Trigonodonia Bernh. der Grossgattung Zyras Steph. (Col. Staphylinidae). Entomologische Arbeiten aus
dem Museum G. Frey, 14, 100–109.
Scheerpeltz, O. (1963b) Ergebnisse der von Wilhelm Kühnelt nach Griechenland unternommenen zoologischen Studienreisen.
I. (Coleoptera - Staphylinidae). (124 Beitrag zur Kenntnis der palaarktischen Staphyliniden). Sitzungsberichte der
Österreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse (Abt. I), 172, 413–452.
Scheerpeltz, O. (1965a) Die bis jetzt aus Venezuela bekannt gewordenen Arten der Gattung Paederus Fabricius (12. Beitrag zur
Kenntnis der neotropischen Staphyliniden). Acta Biologica Venezuelica, 4 (8), 213–227.
Scheerpeltz, O. (1965b) Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera
Staphylinidae (except. Megalopsidiinae et Steninae). (5. Beitrag zur Kenntnis der orientalischen Staphyliniden). Arkiv för
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Scheerpeltz, O. (1966a) Ergebnisse der Österreichischen Neukaledonien-Expedition 1965 Staphylinidae, (4. Beitrag zur
Kenntnis der australisch-polynesischen Staphyliniden). Annalen des Naturhistorischen Museums in Wien, 69, 393–418.
Scheerpeltz, O. (1966b) Die neue Systematik der Grossgattung Leptusa Kraatz (Col. Staphylinidae). (133. Beitrag zur Kenntnis
der paläarktischen Staphyliniden). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 105/106, 5–55.
Scheerpeltz, O. (1966c) Eine neue Art der Grossgattung Staphylinus L., neue Subgenera und Bemerkungen über einige bereits
bekannte paläarktische Arten dieser Grossgattung (Col.). (139. Beitrag zur Kenntnis der paläarktischen Staphyliniden).
Nachrichtenblatt der Bayerischen Entomologen, 15 (11/12), 105–117.
Scheerpeltz, O. (1967) Neue Staphyliniden aus frei hängenden Beutelnestern des Stärlings Xanthornus viridis P. L. S. Müller in
Venezuela (Col. Staphylinidae). (13. Beitrag zur Kenntnis der neotropischen Staphyliniden). Deutsche Entomologische
Zeitschrift (n. f.), 14 (3–4), 237–256.
Scheerpeltz, O. (1968) Eine neue Art der Gattung Phanolinus Erichson (Col: Staphylin.) nebst einer Übersicht über die bischer
bekannt gewordenen Arten dieser Gattung. (14. Beitrag zur Kenntnis der neotropischen Staphyliniden). Revista de la
Facultad de Agronomía, Maracay, 4 (4), 51–68.
Scheerpeltz, O. (1969a) Die zentral- und südamerikanischen Arten der Gattung Xanthopygus Kraatz. (Col. Staphylinidae,
Subfam. Staphylininae, Tribus Xanthopygini). (16. Beitrag zur Kenntnis der neotropischen Staphyliniden).
Koleopterologische Rundschau, 46/47, 109–118.
Scheerpeltz, O. (1969b) Die zentral- und südamerikanischen Arten der Gattung Echiaster Erichson. (15. Beitrag zur Kenntnis
der neotropischen Staphyliniden). Koleopterologische Rundschau, 46/47, 85–106.
Scheerpeltz, O. (1970a) Die zentral- und südamerikanischen Arten der Gattung Sciocharis Lynch-Arribalzaga mit einer
Bestimmungstabelle aller bisher bekannt gewordenen Arten der Gattung (Col. Staphylinidae). (17. Beitrag zur Kenntnis
der neotropischen Staphyliniden). Deutsche Entomologische Zeitschrift (n. f.), 17 (1–3), 227–254.
Scheerpeltz, O. (1970b) Studien über die Arten der Gattungen Espeson Schaufuss und Parespeson Bernhauer (Col.
Staphylinidae, Subfam. Piestinae, Tribus Thoracophorini). (18. Beitrag zur Kenntnis der neotropischen Staphyliniden).
Mitteilungen der Münchner Entomologischen Gesellschaft, 59, 115–129. [Note: nominal year of publication 1969]
Scheerpeltz, O. (1970c) Studien über die Arten der Gattung Calocerus Fauvel (Col. Staphylinidae, Subfam. Piestinae, Tribus
Thoracophorini). (19. Beitrag zur Kenntnis der neotropischen Staphyliniden). Mitteilungen der Münchner
Entomologischen Gesellschaft, 59, 130–142. [Note: nominal year of publication 1969]
Scheerpeltz, O. (1971a) Studien an den arten der Neotropischen gattung Brachydirus Nordman (Col. Staphylininae, Subfam.
Staphyliniae, Tribus Xanthopygini). (22. Beitrag zur Kenntnis der neotropischen Staphyliniden). Mitteilungen der
Entomologischen Gesellschaft, Basel (n. f.), 21 (4), 93–109.
Scheerpeltz, O. (1971b) Studien an den Arten der Gattung Hesperus Fauvel (Col. Staphylinidae). Entomologische Arbeiten aus
dem Museum Georg Frey, 22, 150–197.
Scheerpeltz, O. (1972a) Studien an den Arten der neotropischen, ecitophilen Gattung Acanthodonia Bruch (Col. Staph.),
Subfam. Aleocharinae, Tribus Myrmedoniini). (23. Beitrag zur Kenntnis der neotropischen Staphyliniden).
Scheerpeltz, O. (1972b) Studien an den Arten der neotropischen, ecitophilen Gattung Tetradonia Wasmann (Col. Staph.,
Subfam. Aleocharinae Tribus Myrmedoniini). (24. Beitrag zur Kenntnis der neotropischen Staphyliniden).
Scheerpeltz, O. (1972c) Eine neue Art der Gattung Trigonopselaphus Gemminger-Harold, nebst einer Dichotomik der jetzt zu

dieser Gattung gehörigen Arten, Bemerkungen über die aus dieser Gattung auszucheidenden Arten und neue, zum Teil auf
diesen ausgeschiedenen Arten gegründete Gattungen. (Col. Staphylinidae, Subfam. Staphylininae, Tribus Xanthopygini).
(11. Beitrag zur Kenntnis der neotropischen Staphyliniden). Mitteilungen der Münchner Entomologischen Gesellschaft,
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Zootaxa 4621 (1): 001–406 ISSN 1175-5326 (print edition)

Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis,

ANGÉLICO ASENJO1, ULRICH IRMLER2, JAN KLIMASZEWSKI 3, DONALD S. CHANDLER4,

Accepted by Z.-W. Yin: 31 Dec. 2018; published: 21 Jun. 2019

FIRST PUBLISHED IN 2019 BY

© 2019 Magnolia Press

ISSN 1175-5326 (Print edition)

2 · Zootaxa 4621 (1) © 2019 Magnolia Press ASENJO ET AL.

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4 · Zootaxa 4621 (1) © 2019 Magnolia Press ASENJO ET AL.

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Key words: diversity, distribution, species, Latin America, Staphylinidae

6 · Zootaxa 4621 (1) © 2019 Magnolia Press ASENJO ET AL.

Material and methods

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8 · Zootaxa 4621 (1) © 2019 Magnolia Press ASENJO ET AL.

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10 · Zootaxa 4621 (1) © 2019 Magnolia Press ASENJO ET AL.

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FIGURE 2. Number of species described by authors.

12 · Zootaxa 4621 (1) © 2019 Magnolia Press ASENJO ET AL.

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Subfamily GLYPHOLOMATINAE Jeannel, 1962a

0001. Glypholoma Jeannel, 1962a: 482 (Fig. 4)

Type species: Glypholoma pustuliferum Jeannel, 1962

6 species are included in this genus from South America.

0002. Proglypholoma Thayer, 1997: 167 (Fig. 5)

Only one species is known from and is restricted to South America.

Subfamily MICROSILPHINAE Crowson, 1950

0003. Microsilpha Broun, 1886: 889 (Fig. 6)

3 species are included in this genus from South America.

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Tribe Anthophagini Thomson, 1859

0004. Amphichroum Kraatz, 1857b: 947

Only one species is known in this genus from Central America.

0005. Anthobium Leach in Samouelle, 1819: 175 (Fig. 7)

2 species are included in this genus from Central America.

Tribe Corneolabiini Steel, 1950a

0006. Metacorneolabium Steel, 1950b: 62

10 species are included in this genus from South America.

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Tribe Omaliini W.S. MacLeay, 1825

0007. Acrolocha Thomson, 1858: 38

Only one species is known in this genus from Central America.

0008. Crymus Fauvel, 1904a: 92

0009. Dropephylla Mulsant & Rey, 1880: 242 (Fig. 8)

Only one species is known from Central America.

0010. Ischnoderus Fauvel, 1867: 46

16 · Zootaxa 4621 (1) © 2019 Magnolia Press ASENJO ET AL.

2 species are included in this genus from South America.

0011. Omaliopsis Jeannel, 1940: 118

0012. Omalium Gravenhorst, 1802: 111 (Fig. 9)

10 species are included in this genus from Latin America.

0013. Phloeonomus Heer, 1839: 184 (Fig. 10)

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0014. Phyllodrepa Thomson, 1859: 52 (Fig. 11)

5 species are included in this genus from South America.