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Bio1LE2 Reviewer
Bio1LE2 Reviewer
4-5% vertebrates)
Evolution of species (Speciation) - how does a species change from one generation to another through time?
- Speciation- process by which one species is split into 2 or more species which may share many of the same characteristics but also produces
tremendous diversity.
- Evolution - change over many generations in the composition of individuals comprising a population of a species
- It is the population that evolves because individuals of the populations are those selected to survive
a. Allopatry (allopatric speciation), Adaptive radiation
i. Physical barrier fragments, isolates populations into subpopulations, resulting in reproductive isolation
1. Series of dispersals leads to a reproductive isolation of populations
2. Porkfish v. panamic porkfish (barrier from south america and north america - adaptive radiation)
ii. Once geographic separation has occurred, the separated gene pools may diverge when different mutations arise and natural selection
and genetic drift may alter allele frequencies in different ways in the separated populations.
iii. With changes occuring between the species, reproductive barriers arise (when populations are subjected to different environmental conditions)
iv. Leads to Adaptive radiation - emergence of new variety of species from a common stock due to allopatric speciation
v. It is biological barriers, not physical separation, that prevents interbreeding
b. Sympatry (sympatric speciation)
i. Occurs in population that lives in a wide/huge common habitat
1. Complete absence of physical barrier
ii. Certain individuals from the original population will become adaptive to certain conditions in certain sections of the habitat (certain unique set of
environmental conditions) and the separated subpopulations will soon be fully adapted to certain unique conditions in certain regions
which would lead to the development of different species through adaptive radiation.
iii. Food resources,environmental conditions
iv. Can occur if gene flow is reduced by factors such as:
1. Polyploidy: plants: origin of species from accident during cell division which results to an extra set of chromosomes
2. Sexual selection: Mate choice based on a characteristic as the main reproductive barrier that normally keeps the gene pools of the two
species separate
3. Habitat differentiation: Subpopulation exploits a habitat or resource not used by the parent population
v. Lead to adaptive radiation
c. Descent with modification (Natural theology, creationist essentialist view)
i. Infinite,supreme being was able to create the universe
ii. Intelligent design- universe was created by an intelligent entity which is why we see many diverse forms that happen today (watchmakers
analogy)
d. Evolution of idea of descent with modification
i. Plato - perfect v. imperfect world (perfect world is populated by perfectly adapted organisms; imperfect world is populated by real imperfect
representations of organisms in the perfect world - organisms are not capable of changing overtime)
ii. Aristotle - scale of increasing complexity (natural scale) - organisms are not capable of changing overtime. All levels of the scale are already
fully occupied
iii. Carolus Linnaeus - father of taxonomy : came up with binomial classification of organisms based on morphological characteristics
a. Still believed that animals were not capable of changing over time
iv. George cuvier - excavation; fossilized record remains become different from one level to another (catastrophism)
v. James hutton - Gradualism : geological features are records of past geological events that happened on earth
vi. Charles Lyell- uniformitarianism : existence of processes that have been happening to shape the same manner and intensity over time.
vii. John Baptist Lamark - fossil record as evidence of continuing evolution of organisms from inanimate material towards greater
complexity and perfection in response to organism’s sentiments interieur (felt needs)
1. Lamarck’s mechanism of evolution (Lamarck’s giraffes)
a. Use and disuse - part being constantly used to satisfy needs will become more prominent over time.
b. Traits acquired during an organism's lifetime is inherited by offspring -prominent traits acquired will be passed onto offspring
c. Flawed - ie. abs cannot be passed on to next generation
d. Upside- Diversity is result of evolution happening overtime
viii. Charles Darwin - Evolution of Species (and John Henslow, Fitzroy, Malthus, Lyell, Wallace)
1. Robert Malthus- Human population increase exponentially
2. Lyell- geological process stay constant in continuously shaping the earth’s landscape
3. Darwin- population size does not increase in proportion to the number of young offsprings produced per generation.
a. Species of finches - all species originate and evolve from a common ancestor
b. Adaptive radiation - When a population become fragmented and are separated over many generations, subpopulations will
increasingly become different from one another as only fit individuals will survive, then become a new species
c. The origin of Species by means of natural selection
ix. Origin of Species by means of natural selection
1. All species originate from a common ancestor through descent with modification (tree of life)
2. Reconciles with Linnaeus’ classification scheme - classification based on the same morphological characteristics.
a. Darwin- same characteristics because they come from the same ancestors
b. Evolution’s mechanism is through Natural Selection (survival of the fittest) -
i. Individuals of a population have unequal chances of surviving
ii. Changes of survival depends on traits inherited from ancestors
iii. Motivating force that drives evolution is the survival of the species
1. Individuals who survive have a survival advantage because of the traits (thus genes) they inherited
iv. Individuals of a population are those fit to survive -> change in composition of individuals of a population ->
origin of a new species.
3. Evidence of Macro-evolution- fossils as a record of progressive evolutionary change (ie. origin of eukaryotic life forms; the origin
of humans; the origin of eukaryotic cells; and extinction of the dinosaurs.)
a. Archaeopteryx to chicken - the feathers of ancestor originated from scales (like the ones on the feet of the chicken) and is not
used for flying but for insulation
b. Divergent evolution - Related species in neighboring habitats appear very similar with each other
i. Darwin’s finches - the beaks of the finches are different and is most likely because of the different resources present in
each their habitats
c. Convergent evolution - Share common characteristics because they share the same lifestyle and the characteristics allow
them to survive in their habitat (penguin, shark, dolphin) - subjected to the same conditions in the same habitat.
d. Parallel evolution - Share same characteristic- developed same characteristic for their survival
i. Lie convergent evolution,but 2 species evolve at the same time or same ecospace (flying squirrel in australia and north
america)
1. Filling similar ecological roles in similar environments, so similar adaptations were selected.
e. Homology- characteristics of a related species as a variation of the same theme but became modified to serve a different
function. (i.e forelimbs of man, bird, tortoise, elephant)
f. Comparative development of vertebrate embryos- Gill slits are characteristics carried over from ancestors
i. Karl Von Baer: Vertebrates appear similar during their stages of early development
ii. Ernst Haeckel : Ontogeny repeats phylogeny (Biogenetic concept: embryo dev. Stages of an animal species follow the
path of its evolutionary history)
1. Development of advanced species passes through stages represented by adult organisms of more primitive species.
Otherwise put, each successive stage in the development of an individual represents one of the adult forms that
appeared in its evolutionary history.
2. Pharyngeal grooves between the pharyngeal arches in the neck of the human embryo not only roughly resembled gill
slits of fish, but directly represented an adult "fishlike" developmental stage, signifying a fishlike ancestor.
g. Biochemical composition of related organisms (genetics)
i. Proteins/peptides are made up of a sequence of amino acids
ii. Composition and sequence of AA of proteins are determined by genes inherited from ancestors
1. Closely related species have almost similar genes, the amino acid sequence of their proteins differ less
(especially when their ancestor is near each other than when their ancestors are distant from each other)
4. Microevolution - evolution at the level of hereditary traits (genes) of a population - deviation from hardy weinberg principle
(Pesticide resistance, herbicide resistance, and antibiotic resistance - focuses on a single population)
a. Hardy Weinberg principle - Explain genetic basis of non-evolving population of organisms.
i. Gene pool of the population remains constant unless acted upon by some factors. Otherwise, it stays in
equilibrium over many generation
ii. Genetic structure of a gene pool - frequency of alleles of a population
iii. Requirements of the HW Principle
1. Large (closed) population size
2. No gene flow/inter-breeding with other related populations (closed population)
3. No net mutations (absence of allele substitutions)
4. Occurrence of random matings in the closed population (no strict preference for mating partner)
5. Absence of natural selection (all offspring has equal chance of survival)
b. Genetic Drift - fragmentation of large population into several isolated/closed sub-populations, resulting in the origin of several
new species (allopatry, sympatry) - has larger effect on smaller populations
i. Bottleneck effect - happens when some catastrophe, like an earthquake or a tsunami, kills off most of a population at
random and leaves only a handful of survivors. The catastrophe has to be something that strikes at random, however,
and kills individuals irrespective of the genes they carry.
ii. Founder Effect - When a small group separates from a larger population and strikes out on its own, that small group
might be carrying genes that are rare in the original population. These rare genes will now be common among the new
group's descendants. Other genes present in the original population, however, may be absent from the new group
altogether.
c. Gene flow or inter-breeding with another elated population, mutations, or substitution of alleles, non-random matings -
result in the accumulation of genetic variations to destabilize the gene structures of a non-evolving population, resulting in the
origin of one or several new species
d. Occurence of natural selection - individuals have unequal chances of survival.
i. Evolution is a change/alteration in the genetic structure (or allele frequency) of a population which allows only fit
individuals to survive.
ii. Ie.e brown beetles who can camouflage survive and its gene frequency increases because the reen beetle is easily
picked by the predator
Evolution of sex
a. Origin of sex
i. Recombination of genes as a means to repair damaged DNA among individuals of early life forms (asexual reproduction)
1. Asexual reproduction of single cell resulted to clone having very low survival.
b. Sexual Reproduction
i. Recombination of genes/ gene flow during sexual reproduction is a selected trait because genes of offspring derived from 2 mating
partners increase the survival of the offspring and therefore, the species
ii. Variations of the genes of a population are the raw material of natural selection to allow populations to survive, adapt, and evolve.
iii. Special case of natural selection (need for 2 individuals of opposite sex to mate)
1. Consequences
a. Powerful motivation to attract a suitable mate
b. Development of sex-specific anatomical traits and behavior (mate preferences)
2. Mate Preferences
a. Attractive men produce better quality sperm, while more attractive women are more fertile
iv. Evolution is not really about survival but the perpetuation of henes with survival advantage (mates are chosen due to their reproductive
fitness/viability)
v. Product of evolution is shaped by natural selection because it provides survival advantage, motivating organisms to attract a suitable
mating partner who will most likely produce a fit offspring necessary to perpetuate the genes of the species.
Human evolution
- 1st hominids- 2 to 5 million years ago
- Aegyptopithecus- proconsul- Australopithecus - Humans (chimpanzees nearest in primate evolution)
a. Biological evolution (Homo Sapiens)
i. Large skull: Increased intelligence (tool making), abstract thinking (rituals), Rationality (learning experiences)
ii. Bi-pedalism and upright posture: Increased mobility and maneuverability; hands-free mobility
iii. Extended parental care- increased parental bonding with offspring; transmission of accumulated knowledge and culture to succeeding
generations via speech and language
b. Cultural evolution
i. Homo habilis, Homo erectus - Nomads -> hunting-gathering society
ii. Homo sapiens: Agricultural -> industrialized society
iii. Change in knowledge, skills, tech spread from an indiv to another
iv. Language; tools; exploration of the agricultural, fishing resources
v. Allowed humans to survive despite having little resources left from overhunting in the past
c. The Anthropocene Epoch
i. Humans have become the new natural selection agent, which has short-circuited biological evolution
ii. Cultural evolution has outpaced biological evolution; will earth’s other inhabitants be able to cope and survive?
Community Ecology
a. Ecology - Interactions or relationships of biotic and abiotic components of the environment
b. Community: collection of populations interacting with one another and with the non-living components of their common and shared habitat.
Ecology of communities: Why various species are found in a community (composition, abundance, and distribution of a species in a community)
a. Models of communities
● Individualistic model (HA Gleason)- chance collection of populations of various species sharing the same habitat due to similar abiotic
requirements of each population
● Interactive model (RE Clements) - collection of closely linked species bound by mandatory biotic interactions
i. Entire community functions as one whole integrated unit/superorganism
ii. They aid in each others survival
b. Interactions or relationships as natural selection agents
i. Biological interactions in a community (inter and intra species) and abiotic factors act as natural selection agents to initiate traits or
characteristics among individuals of populations in a community
ii. Biological interactions translate into adaptations among various species in a community, resulting in the evolution of populations.
- I.e. in a population where most bunnies are not disease resistant, when they are hit by a virus, the only surviving ones will be the
disease resistant rabbit. Hence, the non drug resistant bunnies will be extinct while the disease resistant rabbit will be able to
reproduce.This will result in the future generation of bunnies to all be disease resistant.
c. Biological interactions
i. Predation: predator-prey interaction
1. Parasitism and herbivory parasitism
2. Coping with predator-prey interactions
a. Result in the co-evolution of adaptive traits (Predator adaptations and Anti-predator adaptation)
i. Predator adaptation: adjust characteristics to allow them to catch more prey
ii. Anti predator adaptations: Cryptic coloration (blend in with its environment or have color), mimicry (look like the predator),
aposematic coloration (make themself look prominent and dangerous), organisms living with a lot of members of the
same species (protect each other by number)
3. Dynamics of predator-prey interactions
a. Increase population of prey = increase population of predator
b. Consequence of predation as biological pest control- hazardous for farmer
ii. Competition: Intra- and inter-species interactions
1. May result in:
a. Direct physical contact (Aggressive, ritualistic, territorial, hierarchical behavior)
b. Lower population size (high death rate, low birth rate)
2. Competitive exclusion principle: 2/more species competing for common resources cannot co-exist (especially if complete
competition)
a. Natural selection favors individuals with traits/adaptations which are able to reduce or eliminate competition
i. Evolution of population is shaped and guided by competition
3. Character displacement
a. Species competing long enough can result in character displacement (composition of one competing population from another
population = competition as natural selection agent)
i. Present day characteristics are the result of past species interactions
ii. Allopatric - eventually develop similar characteristics because they do not live together, thus do not compete for
resources they both need/prefer
iii. Sympatric - eventually develop different sizes of beaks/ characteristics because they do not stay in one island due to
competition
4. Resource Partitioning- Resource partitioning is the division of limited resources by species to help avoid competition in an
ecological niche. In any environment, organisms compete for limited resources, so organisms and different species have to
find ways to coexist with one another.
a. Based on physical location - lizards may live in same habitat but different place (one on the ground, one on the trees)
b. Based on food - chemical characteristic, same food but different part, or different eating pattern
c. By partitioning out resources, species can have long-term coexistence with one another in the same habitat (reduce inter
and intra species competition). This allows both species to survive and thrive rather than one species causing the other to go
extinct. Only the individuals with the appropriate traits to reduce competition (the stronger species) will survive.
iii. Symbiosis:
1. Commensalism - one benefits while one does not experience any effects
2. Mutual Symbiosis - each organism benefits and cannot exist without each other
- I.e. Bifidobacteria - lives in the intestines of people, each party cannot live without the other.
iv. Evolution of population is shaped by biological interactions/relationships (comp, pred, symb)
1. Biological interactions may translate into traits or long term adaptations having a survival advantage among various species
in the community, resulting in the evolution of populations.
Ecology of Population
Ecosystem
- Sum of all living communities and their interactions with biotic and abiotic components of the environments consisting of chemical elements
and energy
- Living and nonliving components consist of chem elements that contain chemical energy and are bonded together by chemical energy.
- Purpose of interaction among biotic and abiotic factors in the environment: need to acquire energy to power life processes that organisms need
to grow, breed, and SURVIVE.
- Energy comes from biosphere (living and on living)
Vital processes of ecosystem
- Flow of energy
- Recycling of chemical elements found in living and nonliving matter
d.
2. Phosphorus Cycle - (phosphate in soil from decomposers and geological uplifting used in leaching and sedimentation for the formation
of new rocks) - organic fertilizer
a.
3. Carbon Cycle - CO2 released by organisms (cellular respiration) and burning of wood and fossil fuels are used by plants and converted
to detritus to make non-renewable fossils fuels (oil, natural gas)
a. Carbon does not exist as carbon element but forms chemical compounds where carbon is sourced by the environment (mostly
CO2)
b.