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Networks and Geographies of Global Social Policy Diffusion Michael Windzio Full Chapter
Networks and Geographies of Global Social Policy Diffusion Michael Windzio Full Chapter
Global Dynamics
of Social Policy CRC 1342
Global Dynamics of Social Policy
Series Editors
Lorraine Frisina Doetter, University of Bremen, Bremen,
Germany
Delia González de Reufels, University of Bremen, Bremen,
Germany
Kerstin Martens, University of Bremen, Bremen, Germany
Marianne Sandvad Ulriksen, University of Southern
Denmark, Odense, Denmark
This open access series welcomes studies on the waves, ruptures and
transformative periods of welfare state expansion and retrenchment glob-
ally, that is, across nation states and the world as well as across history
since the inception of the modern Western welfare state in the nine-
teenth century. It takes a comprehensive and globalized perspective on
social policy, and the approach will help to locate and explain episodes
of retrenchment, austerity, and tendencies toward de-welfarization in
particular countries, policy areas and/or social risk-groups by reference
to prior, simultaneous or anticipated episodes of expansion or contrac-
tion in other countries, areas, and risks. One of the aims of this series is
to address the different constellations that emerge between political and
economic actors including international and intergovernmental organi-
zations, political actors and bodies, and business enterprises. A better
understanding of these dynamics improves the reader’s grasp of social
policy making, social policy outputs, and ultimately the outcomes of
social policy.
Networks
and Geographies
of Global Social
Policy Diffusion
Culture, Economy, and Colonial
Legacies
Editors
Michael Windzio Ivo Mossig
SOCIUM Institute of Geography
University of Bremen University of Bremen
Bremen, Germany Bremen, Germany
© The Editor(s) (if applicable) and The Author(s) 2022. This book is an open access publication.
Open Access This book is licensed under the terms of the Creative Commons Attribution 4.0
International License (http://creativecommons.org/licenses/by/4.0/), which permits use, sharing, adap-
tation, distribution and reproduction in any medium or format, as long as you give appropriate credit
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The images or other third party material in this book are included in the book’s Creative Commons
license, unless indicated otherwise in a credit line to the material. If material is not included in the
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from the relevant protective laws and regulations and therefore free for general use.
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The registered company address is: Gewerbestrasse 11, 6330 Cham, Switzerland
Acknowledgments
v
About This Book
vii
Contents
ix
x Contents
Index 269
Contributors
xi
xii Contributors
xiii
xiv List of Figures
xv
xvi List of Tables
Introduction1
The global diffusion of social policy is an emerging field in political
science and comparative macro-sociology. Detailed, qualitative studies
can precisely highlight the mechanisms of diffusion at work, e.g.,
learning, emulation, competition, or coercion (Gilardi 2016; Obinger
et al. 2013). Even though this approach can reveal these mechanisms,
it is limited to the respective cases under investigation. At a higher
level of abstraction, researchers can apply statistical models for diffusion
research on a comprehensive set of countries and over a long historical
1This chapter is a product of the research conducted in the Collaborative Research Center
“Global Dynamics of Social Policy” at the University of Bremen. The center is funded
by the Deutsche Forschungsgemeinschaft (DFG, German Research Foundation)—project
number 374666841—SFB 1342.
I. Mossig (B)
Institute of Geography, University of Bremen, Bremen, Germany
e-mail: ivo.mossig@uni-bremen.de; mossig@uni-bremen.de
period. On the one hand, such studies usually abstract from the country-
specific “micro” mechanisms; on the other hand, they provide a “macro”
perspective on the diffusion process in the overall population of countries
around the globe. The empirical studies collected in this volume follow
the second approach.
Our research was conducted in the Collaborative Research Center
1342 (CRC 1342) at the University of Bremen, which is funded by the
German Research Foundation (DFG). The members of the CRC 1342
collected an unprecedented amount of historical data on welfare policies
around the globe to allow for macro-quantitative analyzes of global diffu-
sion in different subfields of social policy covering almost all countries in
the world. This book is a collaborative effort of the quantitative projects
in the CRC 1342 that analyze the diffusion of welfare policies.
We regard diffusion as a process driven by multiplex ties between
countries in global social networks. In social network research, multi-
plexity means that subjects have network ties in various dimensions. In
our view, global trade, colonial history, similarity in culture, and spatial
proximity link countries to each other. In an epidemic, nowadays an
unfortunately well-known type of diffusion, the share of infected subjects
in the population depends on single events of disease-adoption at the
micro-level; these events, in turn, result from some kind of interaction
between subjects. Hence, networks are the “pipe structure,” or the struc-
tural backbone, of the diffusion process. We will analyze diffusion in
several subfields of social policy, investigating the question of which
network dimensions drive the process. For instance, the introduction of
certain labor regulations might depend more on economic ties, in partic-
ular, global trade, whereas cultural similarity between countries could
.25
1
cumulative logistic density
.8
.2
logistic density
.15
.6
.4
.1
.05
.2
0
0
-1 -.5 0 .5 1 -1 -.5 0 .5 1
values of x values of x
will use logit models, where the hazard rate r (t ) is predicted by j time-
dummies α that indicate e.g., 25-year time intervals, to estimate the
effects of our four networks of trade, colonial history, cultural spheres,
and spatial proximity, and some control variables β’x.
1
r (t) =
1 + exp( −(α1 t1 + ... + α j t j + β1 trade + β2 colony + β3 cultur e + β4 pr oximt y + β x) )
In the column “expo.” in the table to the right of Fig. 1.2, there is
a particular value of exposure for each year in which the two countries
i and j were at risk of adopting (which means that they had not yet
adopted, or T ≥ t ). The event of adoption occurs as a result of a given
exposure in the moment before adoption, so the respective exposure is
lagged by one year. At the bottom of Fig. 1.2, hazard ratios are shown
for the binary explanatory variable WEIRD. Country i is WEIRD and
has 1 event out of 4 time periods at risk and thus a hazard rate of 0.25.
Period t 1 has been dropped because of the lagged exposure, i.e., there
is no data on subjects that adopted at t ≤ 1. Country j (non-WEIRD)
has 1 event out of 2 time periods at risk and thus a hazard rate of 0.5,
so the hazard ratio is (1/4)/(1/2) = 0.5. Computing hazard ratios and
standard errors for continuous variables, such as exposure, is much more
8 I. Mossig et al.
social policy, exposure for its (past) colonial entities increases. However,
this does not hold the other way around. For undirected networks,
exposure would take the same value regardless of direction. Generally,
(unweighted) exposure is included in the logistic hazard models as a
numeric variable ranging from 0, where no alter has adopted the social
policy, to 1, where all alters have adopted the social policy. On a similar
note, the geographical proximity network is time constant, meaning the
tie strength does not change over the duration of analysis, while cultural
spheres, trade, and colonial legacies are time-variant to account for the
declining influence of colonial powers after decolonization, changing
economic partnerships, and evolving cultural characteristics.
We take the four networks as the underlying structures for the diffu-
sion process. As we will see later in the chapter, all networks constitute
different avenues or “pipes” through which communication and infor-
mation about social policies can travel. Taken together, these networks
emphasize different specificities of countries’ interdependencies. By
including different networks, we assume to catch as many instances of
network diffusion as possible through the different mechanisms.
However, social policy diffusion can also depend on domestic factors
such as a country’s level of economic development or financial capa-
bility. The same is true for civil freedom in the political regime (Lindert
2004). Thus, we introduce Gross Domestic Product (GDP) per capita
(Inklaar et al. 2018) and a democratization index as baseline control vari-
ables. The former was linearly interpolated for the whole time frame by
taking the minimum value for every income group based on all obser-
vations before 1800 and filling in any missing values according to the
minimum of the respective income group of the corresponding country
by assuming a logistic growth function. Provided there were no data
available, these were the values to start the interpolation into future
years. This yields a continuous measure of economic development from
1880 to 2010 for almost all countries in our dataset. For the level of
democratization, we use the basic Varieties of Democracy Regime Score
(Lührmann et al. 2018), which in the raw data ranges from 0 to 9
and was linearly interpolated for any missing data points. This method
introduces some noise to the data, as it fills missing data points with non-
natural numbers (decimals). However, filling missing data either with
10 I. Mossig et al.
the introduction of a policy when the country units Slovenia and Croatia
were non-independent observations. This is due to the way we arrange
the dataset for the diffusion analysis: the set of nodes in the network
is constant over time, which implies that Slovenia and Croatia existed
before, during, and after Yugoslavia existed. Our approach to address
this problem is to regard Slovenia and Croatia as “spatial patches,”
remaining well aware of the fact that many countries actually changed
their borders throughout history. From this perspective, Slovenia and
Croatia were spatial patches at risk of adopting a social policy before,
during, and after Yugoslavia existed. Yugoslavia will not be regarded as
a subject in our sample, but Slovenia and Croatia and all other coun-
tries formerly belonging to Yugoslavia are indeed distinct units. These
subjects are not, however, statistically independent from one another!
This does not pose a problem for the calculation of exposure through the
networks but it does cause a violation of the assumption of independence
of error terms in the maximum likelihood estimation. In the logistic
diffusion model, we address this statistical non-independence by using
cluster-robust standard errors (Zeileis et al. 2020). Our procedure has
the following advantage: it accounts for the statistical non-independence
of observations when they are part of an overarching cluster (spatial
patch) by using the corrected standard errors, but it does not impose
any standard error correction in the hazard model for country-years not
belonging to the respective cluster or to any other cluster.
The analyses in all chapters of this book follow the same rationale:
First, the exposure to already “infected” countries is calculated for each
network, as discussed above. This statistic, i.e., the weighted share of ego’s
network contacts who had already adopted the policy in question at t −1,
is then handed over to a time-discrete hazard model. In this model, the
adoption rate is regressed on exposure, controlling for GDP per capita,
the democracy index, and additional policy field-specific factors. The
resulting robust standard errors correct any statistical non-independence,
potentially affecting standard errors. In most chapters, we present the
coefficients as hazard ratios, representing influence of the predictors on
the risk of policy adoption. The results therefore allow us to determine
which of our networks represent a “pipe structure” for the contagion of
social policies, for example, through exposure to countries that already
12 I. Mossig et al.
Franzese and Hays 2007; Schmitt and Obinger 2013). A simple form of
a spatial weighting matrix is the neighborhood matrix. If two countries
have a shared border, the respective cell of the neighborhood matrix has
a value of 1, and otherwise 0 (Windzio et al. 2019). The neighborhood
matrix thus implies the assumption that only countries with a common
border can influence each other (Obinger et al. 2013). Not least because
of the criticism of this very narrow assumption, the distances between
capital cities were used instead of, or rather in addition to, the neighbor-
hood matrix to define “spatial lags” in the weighting matrices (Schmitt
2019; Simmons and Elkins 2004).
As a justification of the relevance of geographical proximity, it is
often argued that the intensity of communication between countries can
increase due to their proximity. This argument implies the assumption
that the exchange of information between neighboring or geographi-
cally close countries is substantially higher. Even unintended forms of
information exchange occur more easily and thus more frequently. In
addition, policy examples from neighboring or nearby countries are often
regarded as a blueprint for a country’s own national policies, so that a
high degree of mutual influence is assumed due to geographical prox-
imity (Schmitt and Obinger 2013). However, a clear assignment of
geographical proximity to one of the mechanisms from the diffusion
literature—(i) learning, (ii) competition, (iii) imitation, or (iv) coercion
(Obinger et al. 2013; Starke and Tosun 2019)—is difficult. Magetti and
Gilardi (2016) conclude that “Geography is often an important compo-
nent of diffusion, but it cannot be linked straightforwardly to any of
the […] mechanisms. Therefore, it is a catch-all indicator that usually
discriminates between them. It is best used in combination with other
indicators” (Magetti and Gilardi 2016, 93).
Similarly, Simmons and Elkins (2004) note that geographical distance
does not provide a satisfactory explanation for policy diffusion per se.
In line with their view, Beck et al. (2006) point out in a contribu-
tion with the significant title “Space is more than Geography” that,
on the one hand, taking geographical distances into account in spatial
econometrics is a methodological enrichment, but that other measures
for determining interconnectedness between states would produce more
fruitful results. Similarly, Boschma (2005) argues that proximity not
1 Networks of Global Social Policy Diffusion … 15
xi j = 1/ capital distance
Even though there are occasional shifts of the capital in some countries,
for pragmatic reasons the distances are based on the capital cities in 2020.
Therefore, the geographical distances are a time-invariant network.
largely determine the scope of action and also influence the vulnerability
and sensitivity of interstate relations (Glückler and Doreian 2016; Maoz
2011). The significance of economic globalization and the relevance of
intensifying trade linkages for the diffusion of social policy is based on
the assumption that important trading partners influence a country’s
policies more strongly than subordinate trading partners do. As a result
of the globalization process, countries are becoming more closely aligned
with one another, although this does not necessarily mean that social
policy has to converge (Jahn 2016).
The trade networks were defined as follows: The trade data are
collected from the Correlates of War Project (Barbieri and Keshk 2016).
According to the following regulations, the edge weights were deter-
mined for each year. The volume of trade between each of the two
countries comprises the total trade in goods in one year and is there-
fore undirected. The original trade flows were converted into US$ using
the average exchange rate from 2011 to avoid an inflation-related densifi-
cation of the networks. Due to the extremely different trade volumes, we
logarithmically transformed the trade values. The edge weight of trade
interdependence between two countries i, j is therefore:
log(trade) if trade > 0
xi j =
0 if trade = 0
If a dyad shared any trade volume in any respective year, the log of this
volume was used, otherwise the edge was set to 0 as the dyad did not
share any trade in the respective year. Further decisions regarding the
construction of trading networks concern former countries that have
split up over time, e.g., Austria-Hungary, Czechoslovakia, or Serbia-
Montenegro. In such countries, the trade volume of the shared years was
divided according to the GDP proportion of these countries after these
countries separated from each other. In the case of the Union of Soviet
Socialist Republics (USSR) this refers to the period from 1922 to 1991,
in the case of the Baltic States 1941–1991, and in the case of the former
Yugoslavia the period from 1918 to 1992. “Small” states that once existed
but are not represented in the selected country sample for this anthology
were deleted (e.g., Yemen People’s Republic, Republic of Vietnam, Korea
18 I. Mossig et al.
Germany (1.69), and France (1.57). The following Table 1.1 divides the
country sample into quartiles. 17 countries (10.4% out of 164 countries)
with the highest centrality rating account for 25% of the cumulative
degree centrality. In contrast, the last quartile is occupied by 81 countries
with the lowest centrality values in global trade. Bhutan, for example,
which ranks last, only accounts for 0.05% of global trade. Lesotho
(0.08), the Comoros (0.09), and the Solomon Islands (0.10) also had
a very low-degree centrality in 2010.
the network but also with the increasing tie-strength to these adopters.
The cultural spheres network is time varying. For example, proportions
of dominant religious or ethnic groups as well as dominant language
changed over time.
To gain a better overview of the network and the resulting cultural
spheres, we clustered the network with a Louvain clustering algorithm.
According to the time-variant nature of the network, the result suggests
a five-cluster solution in 1880 but a three-cluster solution in 2010. This
supports the idea that there has been an increasing isomorphism in insti-
tutional structures around the globe (Meyer et al. 1997). Figure 1.4
In our view, the influence of a colonial power does not simply disappear
immediately after the colony becomes officially independent. According
to our assumption, the influence of the colonial power declines much
more gradually over time after official independence is achieved. There
are different variants to compute exposure due to the colonial legacy
based on this function. We apply the function to the standardized expo-
sure as computed by the netdiffuseR package (Vega Yon and Valente
2021), which restricts the range of exposure between zero and one.
Accordingly, the theoretical assumption is that colonial legacy is very
strong and the strength of the colonial power’s influence remains almost
constant after independence. In contrast, if we do not standardize
the exposure, the influence of former colonial powers still exist, but
compared to the standardized computation of exposure, the power
24 I. Mossig et al.
(weighted) trade network than in the cultural spheres network but the
additional tie in the trade network has a particularly high weight in the
computation of exposure. Similarly important are situations when most
alters are already infected and exposure is generally high. Exposure can be
1 (maximum normalized exposure) in a network dimension where ego is
tied to 12 alters, but it can also be 1 in another network dimension where
ego is tied only to 2 alters. Table 1.3 shows the correlations of (weighted)
exposure to alters that adopted compulsory education. Indeed, corre-
lations are considerably higher. Exposure in the network of geographic
proximity is strongly correlated with the network of cultural proximity
(0.919) and also with exposure in the trade network (0.741). Moreover,
trade and cultural proximity are highly correlated as well (0.728).
2 g(g−1)
exp{θ z(x)}
P(X = x) = , where κ(θ) = exp{θ z(x)}
κ(θ)
n=1
temporal ERGM for the period from 1890 to 2010 in 20-year intervals
and eight measurement occasions using bootstrapping methods (Leifeld
et al. 2016). The term “edges” is the intercept of the regression model
and represents the log odds of the network density, given that all covari-
ates are constrained to zero. The positive significant effect of gwesp
(geometrically edgewise shared partners) indicates that transitive closure
much more likely occurs in the empirical network than in a corre-
sponding random network. In contrast, gwdsp (geometrically dyadwise
shared partners) shows a significantly negative effect and points to the
lower probability of open triads (Harris 2014). Aside from these network
structural effects, ties in the trade network depend on spatial proximity
(0.1488* ) and cultural spheres (0.9971* ) but not significantly on colo-
nial legacies. They occur less often if two countries have the same level
of democratization (same regime) and the higher the absolute difference
30 I. Mossig et al.
in GDP per capita between ego and alter is. Unsurprisingly, global trade
is an issue of economically well-performing countries since high levels
of GDP increase the degree (0.0332* ). We also estimated the memory-
term of “tie stability” (Leifeld et al. 2016), which indicates the stability
of ties and non-ties, and thereby accounts for how strongly the state of
the network at t depends on its previous state at t −1 (Table 1.4).
Column 2 in Table 1.4 shows the effects on the log odds of ties
in the network of cultural spheres. Again, we find the pattern of high
transitivity (gwesp) and a negative tendency toward open triads (gwdsp).
Having a tie in the network of colonial legacies has a negative effect
on cultural similarity (−0.3214* ), which means that countries colonized
other countries that were culturally rather different. Contrariwise, a tie in
the network of global trade increases the log odds of a tie in the cultural
spheres network—which we also do not interpret in a strict causal sense
because the direction of the influence could also be reversed (0.8275* ).
Our model does not indicate that political regime type in terms of
levels of democratization and economic development corresponds with
culture: if two countries have the same level of democratization, the
log odds of a tie in the cultural spheres network is only insignificantly
increased. The absolute difference in GDP is insignificant as well. Again,
the memory-term indicates a significant effect of the lagged network.
Finally, we analyze the network of colonial legacies, which has quite
a specific topology as shown in Fig. 1.5. This network is rather special
since there are few “hubs” with many ingoing ties, and there is a clear
distinction between node sets of senders and receivers. We find a positive
effect of 2-in-stars. This means that two ingoing ties occur significantly
more often than expected by chance, which is obvious from the visual
representation in Fig. 1.6. In addition, there are positive effects of spatial
proximity and ties in the trade network. While there is no effect of same
regime, effects of GPD per capita are negative on outdegree. Accordingly,
richer countries name other countries as colonizers less often: overall,
richer counties have a considerably lower risk of being colonized.
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Fig. 219.—Young of Cyclopterus spinosus, from the Arctic Ocean, natural size.
Liparis.—Body sub-cylindrical, enveloped in a more or less loose
naked skin; head broad, obtuse. The infraorbital bone is styliform
posteriorly, extending backwards to the margin of the præoperculum.
One dorsal fin, with feeble flexible rays. Villiform teeth in the jaws,
none on the palate.
Small fishes from the northern coasts of the temperate zone,
ranging beyond the arctic circle. Eight species are known, of which
two (L. lineatus and L. montagui) occur on the British coasts.
Second Family—Gobiidæ.
Body elongate, naked or scaly. Teeth generally small, sometimes
with canines. The spinous dorsal fin, or portion of the dorsal fin, is
the less developed, and composed of flexible spines; anal similarly
developed as the soft dorsal. Sometimes the ventrals are united into
a disk. Gill-opening more or less narrow, the gill-membranes being
attached to the isthmus.
Small carnivorous littoral fishes, many of which have become
acclimatised in fresh water. They are very abundant with regard to
species as well as individuals, and found on or near the coasts of all
temperate and tropical regions. Geologically they appear first in the
chalk.
Gobius.—Body scaly. Two dorsal fins, the anterior generally with
six flexible spines. Ventral fins united, forming a disk which is not
attached to the abdomen. Gill-opening vertical, moderately wide.
Fig. 220.—Gobius lentiginosus, from New Zealand.
The “Gobies” are distributed over all temperate and tropical
coasts, and abundant, especially on the latter. Nearly three hundred
species have been described. They live especially on rocky coasts,
attaching themselves firmly with their ventrals to a rock in almost any
position, and thus withstanding the force of the waves. Many of the
species seem to delight in darting from place to place in the rush of
water which breaks upon the shore. Others live in quiet brackish
water, and not a few have become entirely acclimatised in fresh
water, especially lakes. The males of some species construct nests
for the eggs, which they jealously watch, and defend even for some
time after the young are hatched. Several species are found on the
British coast: G. niger, paganellus, auratus, minutus, ruthensparri.
Fossil species of this genus have been found at Monte Bolca.
A very small Goby, Latrunculus pellucidus, common in some
localities of the British Islands and other parts of Europe, is
distinguished by its transparent body, wide mouth, and uniserial
dentition. According to R. Collett it offers some very remarkable
peculiarities. It lives one year only, being the first instance of an
annual vertebrate. It spawns in June and July, the eggs are hatched
in August, and the fishes attain their full growth in the months from
October to December. In this stage the sexes are quite alike, both
having very small teeth and feeble jaws. In April the males lose the
small teeth, which are replaced by very long and strong teeth, the
jaws themselves becoming stronger. The teeth of the females remain
unchanged. In July and August all the adults die off, and in
September only the fry are to be found.
There are several other genera, closely allied to Gobius, as
Euctenogobius, Lophiogobius, Doliichthys, Apocryptes, Evorthodus,
Gobiosoma and Gobiodon (with scaleless body) Triænophorichthys.
Sicydium.—Body covered with ctenoid scales of rather small size.
Cleft of the mouth nearly horizontal, with the upper jaw prominent; lips
very thick; the lower lip generally with a series of minute horny teeth.
A series of numerous small teeth in upper jaw, implanted in the gum,
and generally movable; the lower jaw with a series of conical widely-
set teeth. Two dorsal fins, the anterior with six flexible spines. Ventral
fins united, and forming a short disk, more or less adherent to the
abdomen.
Small freshwater fishes inhabiting the rivers and rivulets of the
islands of the tropical Indo-Pacific. About twelve species are known;
one occurs in the West Indies. Lentipes from the Sandwich Islands is
allied to Sicydium.
Periophthalmus.—Body covered with ctenoid scales of small or
moderate size. Cleft of the mouth nearly horizontal, with the upper jaw
somewhat longer. Eyes very close together, immediately below the
upper profile, prominent, but retractile, with a well-developed outer
eyelid. Teeth conical, vertical in both jaws. Two dorsal fins, the anterior
with flexible spines; caudal fin with the lower margin oblique; base of
the pectoral fin free, with strong muscles. Ventral fins more or less
coalesced. Gill-openings narrow.
The fishes of this genus, and the closely-allied Boleophthalmus,
are exceedingly common on the coasts of the tropical Indo-Pacific,
especially on parts covered with mud or fucus. During ebb they leave
the water and hunt for small crustaceans, and other small animals
disporting themselves on the ground which is left uncovered by the
receding water. With the aid of their strong pectoral and ventral fins
and their tail, they hop freely over the ground, and escape danger by
rapid leaps. The peculiar construction of their eyes, which are very
movable, and can be thrust far out of their sockets, enables them to
see in the air as well as in the water; when the eyes are retracted
they are protected by a membranous eyelid. These fishes are absent
in the eastern parts of the Pacific and on the American side of the
Atlantic; but singularly enough one species reappears on the West
African coast. About seven species are known (including
Boleophthalmus), P. koelreuteri being one of the most common
fishes of the Indian Ocean.
First Family—Cepolidæ.
Body very elongate, compressed, covered with very small cycloid
scales; eyes rather large, lateral. Teeth of moderate size. No bony
stay for the angle of the præoperculum. One very long dorsal fin,
which, like the anal, is composed of soft rays. Ventrals thoracic,
composed of one spine and five rays. Gill-opening wide. Caudal
vertebræ exceedingly numerous.
The “Band-fishes” (Cepola) are small marine fishes, belonging
principally to the fauna of the northern temperate zone; in the Indian
Ocean the genus extends southwards to Pinang. The European
species (C. rubescens) is found in isolated examples on the British
coast, but is less scarce in some years than in others. These fishes
are of a nearly uniform red colour.
Second Family—Trichonotidæ.
Body elongate, sub-cylindrical, covered with cycloid scales of
moderate size. Eyes directed upwards. Teeth in villiform bands. No
bony stay for the angle of the præoperculum. One long dorsal fin,
with simple articulated rays, and without a spinous portion; anal long.
Ventrals jugular, with one spine and five rays. Gill-opening very wide.
The number of caudal vertebræ much exceeding that of the
abdominal.
Small marine fishes, belonging to two genera only, Trichonotus
(setigerus) from Indian Seas, with some of the anterior dorsal rays
prolonged into filaments; and Hemerocoetes (acanthorhynchus) from
New Zealand, and sometimes found far out at sea on the surface.
Third Family—Heterolepidotidæ.
Body oblong, compressed, scaly; eyes lateral; cleft of the mouth
lateral; dentition feeble. The angle of the præoperculum connected
by a bony stay with the infraorbital ring. Dorsal long, with the spinous
and soft portions equally developed; anal elongate. Ventrals
thoracic, with one spine and five rays.
Fig. 222.—Scale from the
lateral line of Hemerocœtes
acanthorhynchus, with lacerated
margin.
Sixth Family—Mastacembelidæ.
Body elongate, eel-like, covered with very small scales. Mandible
long, but little moveable. Dorsal fin very long, the anterior portion
composed of numerous short isolated spines; anal fin with spines
anteriorly. Ventrals none. The humeral arch is not suspended from
the skull. Gill-openings reduced to a slit at the lower part of the side
of the head.
Freshwater-fishes characteristic of and almost confined to the
Indian region. The structure of the mouth and of the branchial
apparatus, the separation of the humeral arch from the skull, the
absence of ventral fins, the anatomy of the abdominal organs,
affords ample proof that these fishes are Acanthopterygian eels.
Their upper jaw terminates in a pointed moveable appendage, which
is concave and transversely striated inferiorly in Rhynchobdella, and
without transverse striæ in Mastacembelus: the only two genera of
this family. Thirteen species are known, of which Rh. aculeata, M.
pancalus and M. armatus are extremely common, the latter attaining
to a length of two feet. Outlying species are M. aleppensis from
Mesopotamia and Syria, and M. cryptacanthus, M. marchei, and M.
niger, from West Africa.
First Family—Sphyrænidæ.
Body elongate, sub-cylindrical, covered with small cycloid scales;
lateral line continuous. Cleft of the mouth wide, armed with strong
teeth. Eye lateral, of moderate size. Vertebræ twenty-four.
This family consists of one genus only, Sphyræna, generally
called “Barracudas,” large voracious fishes from the tropical and sub-
tropical seas, which prefer the vicinity of the coast to the open sea.
They attain to a length of eight feet, and a weight of forty pounds;
individuals of this large size are dangerous to bathers. They are
generally used as food, but sometimes (especially in the West
Indies) their flesh assumes poisonous qualities, from having fed on
smaller poisonous fishes. Seventeen species.
The Barracudas existed in the tertiary epoch, their remains being
frequently found at Monte Bolca. Some other fossil genera have
been associated with them, but as they are known from jaws and
teeth or vertebræ only, their position in the system cannot be exactly
determined; thus Sphyrænodus and Hypsodon from the chalk of
Lewes, and the London clay of Sheppey. The American Portheus is
allied to Hypsodon. Another remarkable genus from the chalk,
Saurocephalus, has been also referred to this family.[44]
Second Family—Atherinidæ.
Body more or less elongate, sub-cylindrical, covered with scales
of moderate size; lateral line indistinct. Cleft of the mouth of
moderate width, with the dentition feeble. Eye lateral, large or of
moderate size. Gill-openings wide. Vertebræ very numerous.
Small carnivorous fishes inhabiting the seas of the temperate and
tropical zones; many enter fresh water, and some have been entirely
acclimatised in it. This family seems to have been represented in the
Monte Bolca formation by Mesogaster.
Atherina.—Teeth very small; scales cycloid. The first dorsal is
short and entirely separated from the second. Snout obtuse, with the
cleft of the mouth straight, oblique, extending to or beyond the anterior
margin of the eye.
The Atherines are littoral fishes, living in large shoals, which habit
has been retained by the species acclimatised in fresh water. They
rarely exceed a length of six inches, but are nevertheless esteemed
as food. From their general resemblance to the real Smelt they are
often thus misnamed, but may always be readily recognised by their
small first spinous dorsal fin. The young, for some time after they are
hatched, cling together in dense masses, and in numbers almost
incredible. The inhabitants of the Mediterranean coast of France call
these newly hatched Atherines “Nonnat” (unborn). Some thirty
species are known, of which A. presbyter and A. boyeri occur on the
British coast.
Atherinichthys, distinguished from Atherina in having the snout
more or less produced; and the cleft of the mouth generally does not
extend to the orbit.
These Atherines are especially abundant on the coasts and in the
fresh waters of Australia and South America. Of the twenty species
known, several attain a length of eighteen inches and a weight of
more than a pound. All are highly esteemed as food; but the most
celebrated is the “Pesce Rey” of Chile (A. laticlavia).
Tetragonurus.—Body rather elongate, covered with strongly
keeled and striated scales. The first dorsal fin is composed of
numerous feeble spines, and continued on to the second. Lower jaw
elevated, with convex dental margin, and armed with compressed,
triangular, rather small teeth, in a single series.
This very remarkable fish is more frequently met with in the
Mediterranean than in the Atlantic, but generally scarce. Nothing is
known of its habits; when young it is one of the fishes which
accompany Medusæ, and, therefore, it must be regarded as a
pelagic form. Probably, at a later period of its life, it descends to
greater depths, coming to the surface at night only. It grows to a
length of eighteen inches.
Third Family—Mugilidæ.
Body more or less oblong and compressed, covered with cycloid
scales of moderate size; lateral line none. Cleft of the mouth narrow
or of moderate width, without or with feeble teeth. Eye lateral, of
moderate size. Gill-opening wide. The anterior dorsal fin composed
of four stiff spines. Vertebræ twenty-four.
The “Grey Mullets” inhabit in numerous species and in great
numbers the coasts of the temperate and tropical zones. They
frequent brackish waters, in which they find an abundance of food
which consists chiefly of the organic substances mixed with mud or
sand; in order to prevent larger bodies from passing into the
stomach, or substances from passing through the gill-openings,
these fishes have the organs of the pharynx modified into a filtering
apparatus. They take in a quantity of sand or mud, and, after having
worked it for some time between the pharyngeal bones, they eject
the roughest and indigestible portion of it. The upper pharyngeals
have a rather irregular form; they are slightly arched, the convexity
being directed towards the pharyngeal cavity, tapering anteriorly and
broad posteriorly. They are coated with a thick soft membrane, which
reaches far beyond the margin of the bone, at least on its interior
posterior portion; this membrane is studded all over with minute
horny cilia. The pharyngeal bone rests upon a large fatty mass,
giving it a considerable degree of elasticity. There is a very large
venous sinus between the anterior portion of the pharyngeal and the
basal portion of the branchial arches. Another mass of fat, of
elliptical form, occupies the middle of the roof of the pharynx,
between the two pharyngeal bones. Each branchial arch is provided
on each side, in its whole length, with a series of closely-set gill-
rakers, which are laterally bent downwards, each series closely
fitting into the series of the adjoining arch; they constitute together a
sieve, admirably adapted to permit a transit for the water, retaining at
the same time every other substance in the cavity of the pharynx.
The lower pharyngeal bones are elongate, crescent-shaped, and
broader posteriorly than anteriorly. Their inner surface is concave,
corresponding to the convexity of the upper pharyngeals, and
provided with a single series of lamellæ, similar to those of the
branchial arches, but reaching across the bone from one margin to
the other.
The intestinal tract shows no less peculiarities. The lower portion
of the œsophagus is provided with numerous long thread-like
papillæ, and continued into the oblong-ovoid membranaceous cœcal
portion of the stomach, the mucosa of which forms several
longitudinal folds. The second portion of the stomach reminds one of
the stomach of birds; it communicates laterally with the other portion,
is globular, and surrounded by an exceedingly strong muscle. This
muscle is not divided into two as in birds, but of great thickness in
the whole circumference of the stomach, all the muscular fasciculi
being circularly arranged. The internal cavity of this stomach is rather
small, and coated with a tough epithelium, longitudinal folds running
from the entrance opening to the pyloric, which is situated opposite
to the other. A low circular valve forms a pylorus. There are five
rather short pyloric appendages. The intestines make a great
number of circumvolutions, and are seven feet long in a specimen
thirteen inches in length.
Fig. 229.—Mugil proboscideus.
Some seventy species of Grey Mullets are known, the majority of
which attain to a weight of about four pounds, but there are many
which grow to ten and twelve pounds. All are eaten, and some even
esteemed, especially when taken out of fresh water. If attention were
paid to their cultivation, great profits could be made by fry being
transferred into suitable backwaters on the shore, in which they
rapidly grow to a marketable size. Several species are more or less
abundant on the British coasts, as Mugil octo-radiatus (Fig. 105, p.
254), M. capito, M. auratus (Fig. 106, p. 254), and M. septentrionalis
(Fig. 107, p. 254), which, with the aid of the accompanying figures,
and by counting the rays of the anal fin, may be readily distinguished
—M. octo-radiatus having eight, and M. capito and M. auratus nine
soft rays. A species inhabiting fresh waters of Central America (M.
proboscideus) has the snout pointed and fleshy, thus approaching
certain other freshwater and littoral Mullets, which, on account of a
modification of the structure of the mouth, have been formed into a
distinct genus, Agonostoma. Myxus comprises Mullets with teeth
more distinct than in the typical species.
This genus existed in the tertiary epoch, remains of a species
having been found in the gypsum of Aix, in Provence.
First Family—Gastrosteidæ.
Body elongate, compressed. Cleft of the mouth oblique; villiform
teeth in the jaws. Opercular bones not armed; infraorbitals covering
the cheek; parts of the skeleton forming incomplete external mails.
Scales none, but generally large scutes along the side. Isolated
spines in front of the soft dorsal fin. Ventral fins abdominal, joined to
the pubic bone, composed of a spine and a small ray.
Branchiostegals three.