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Fundamentals of Digital Marketing Puneet Singh Bhatia Full Chapter
Fundamentals of Digital Marketing Puneet Singh Bhatia Full Chapter
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Preface xv
Acknowledgments xix
About the Author xxi
• UNIT IV (Digital Marketing Execution): The fourth part drills deep into the basic elements
of digital campaigns and their execution across all key Digital Marketing Types (Search,
Display, Social, Partner, Direct, Content, Platform marketing). It completes the ASCOR
Framework stages with coverage of Digital Marketing Measurement and Refinement. This
part concludes with a full chapter on Digital Marketing Execution Elements including
Digital Marketing Revenue, Service Delivery & Payment, Security & Privacy Concerns,
Legal and Ethical Issues.
• UNIT V (Digital Business-Present & Future): This last part sheds light on key present and
future aspects of Digital Marketing starting with the latest trends in digital business to
covering the rise of E-commerce along-with discussing emerging digital business models.
The book ends with a dedicated chapter on emerging career opportunities for digital pro-
fessionals which will help readers of the book decide on career tracks they could follow
being digital marketing professionals.
This book comprises insights gained from my years of corporate work experience and industry
interactions with clients/workmates across multiple sectors, domains and geographies, which I
have aimed to transform into practical and actionable frameworks and models. The title has been
authored keeping in mind the needs of students, faculty, corporates, entrepreneurs and individu-
als who want to master the core concepts of digital marketing. I believe the title will best equip
readers to begin a career in digital marketing and use the concepts as a launchpad to venture into
specific channels, platforms and techniques as they discover them during their digital marketing
careers.
And as is beautifully put in the parting words of the iconic and inspiring Pixar animation
‘Ratatouille’, I hope that with the knowledge gained from this book, you don’t just ‘Inspire
Others’ but regularly ‘Surprise Yourself’ in the true moments of exhilaration this journey presents!
We, at Pearson, wish you all the best for these digital expeditions.
My final acknowledgment to the light of my life, my daughter Aarza, who lovingly (and
unknowingly) gifted a lot of her hours which went into the making of this title for which I am
obligated forever.
I hope that this edition provides you (the reader) the most simple yet practical way of learn-
ing digital marketing concepts from scratch and really helps in developing your knowledge to
emerge as a strong digital marketing professional in future. Request each of you to share feed-
back and suggestions on topics you would like to extend or change in the book and I would try
my best to include the same in forthcoming editions.
Thanks again for providing an opportunity for us to learn this wonderful and emerging area
together.
1
Introduction to
Digital Marketing
LEARNING OBJECTIVES
After studying this chapter, you will be able to:
• Explain the evolution of digital marketing
• Define digital marketing and differentiate it from traditional ‘e-models’ like e-business
and e-commerce
• Outline the underlying technology and frameworks within which digital marketing op-
erates and gain an understanding of digital transformation models
• Describe the structure of digital marketing, opportunities, applications, business modes
and challenges
• Understand key components and application of ASCOR Digital Marketing Framework
across multiple stakeholder scenarios
F lipkart, the poster boy of digital entrepreneurship and flagbearer of e-commerce in India,
has had multiple firsts to its credit which has given it a distinctive advantage as well as a
good positioning in India’s rapidly evolving digital marketing landscape. With the expected
valuation of Indian e-commerce industry estimated to be $100 billion by 2020 (at currently
$17 billion), Flipkart has undoubtedly showed stellar growth from being an exclusively online
bookselling portal to garnering a valuation of $15 billion (figures as per May 2015 last fund-
raising round). According to Forbes.com, the e-commerce platform lists 30 million products,
ships eight million packages a month, and has 45 million registered users (as declared on
18 May 2015). Flipkart delivers in over 1,000 Indian cities and towns.
Flipkart began its operations in 2007, with the company’s founders Sachin Bansal and Binny
Bansal (alumni at the prestigious Indian Institute of Technology (IIT), Delhi) pooling in ` 2 lakh
each and deciding to launch Flipkart, out of their two-bedroom apartment in Koramangala,
Bangalore. There was not a single transaction in the first 10 days, with the first order finally
placed by a customer from Andhra Pradesh for the book Leaving Microsoft to Change the World
by John Wood. From then to now, Flipkart has reported a revenue of ` 2,846 crore (FY
2014) with an estimated employee base of 33,000 (2015). In May 2014, Flipkart received
$210 million from DST Global and raised $1 billion in July 2014. Led by existing investors,
Tiger Global and South Africa’s media group Naspers, in May 2015, it raised $550 million.
Till today, the company has privately raised about $2.7 billion through multiple rounds of
funding.
What makes Flipkart so successful in the face of intense competition, difficult funding
climate, and ever-changing preferences of digital customers? The key to their success is widely
attributed to the unique marketing initiatives they had introduced early-on, including cash
on delivery (CoD), on-time deliveries, mobile-first initiatives, strategic acquisitions, in-house
product launches, and so on. Even on the promotional front, Flipkart has been known for
running some unique advertising campaigns like dressing kids as grown-ups enacting key
features of the brand and what it stands for. Exclusive branding through tie-ups with com-
panies like Motorola (for the launch of their Moto-G Series) was also one of their digital
marketing firsts in India. The impact of such initiatives can be judged from the fact that the
entire stock of Xioami Mi3 took just five seconds to sell, when booking was opened for the
second time for 20,000 phones. On the acquisitions side, Flipkart started its string of digital
acquisitions in 2010 obtaining companies like WeRead (social bookdiscovery tool), Mime360
(digital content platform firm), Chakpak.com (bollywood news site), LetsBuy.com (electron-
ics e-tailer), Myntra.com (a US$319 million deal of a competitor portal), Adiquity (global
mobile network), and Appiterate (mobile marketing start-up). The company is also investing
in four firms, including maps provider MapmyIndia, as it seeks to improve its supply chain
efficiencies, an area where it plans to invest over $2.5billion in the coming years.
It also is a great case study on digital seller partnerships, warehousing, and inventory man-
agement. Key initiatives to help suppliers grow their businesses include F1 Stop campaign
(offering registration, training, cataloguing, packaging, and financial assistance), Flipkart
Seller Campus (special four-hour training sessions on inventory management during peak
season), Growth Capital (helping SMEs leverage their business through the e-marketplace
platform), Seller Hub App (supporting sellers reaching out to 45 million Flipkart customers
on-the-go). Through these initiatives, Flipkart hit $1 billion in annual Gross Merchandise
Value (GMV) in 2014 (an year ahead of their target).
Since 2014, in their bid to improve margins, Flipkart has started concentrating on launch-
ing its own set of tablet, mobile phones, and even a phablet. The first among the series of
tablet phones was Digiflip Pro XT 712 Tablet. In July 2014, Flipkart launched its first net-
working router, under its own brand name—DigiFlip WR001 300 Mbps Wireless N Router.
In September 2014, Flipkart launched its in-house home appliances and personal healthcare
brand (Citron) which includes a wide range of cooking utilities and grooming products.
With a lot going their way across their digital journey, there have also been incidents
which have marred the brand’s reputation and posed execution-related challenges to Flipkart.
The key among them was the handling of their ‘Big Billion Day’ online sale on 6 October
2014, which exposed certain flaws like non-availability of products, out-of-stock listings,
backend (technical) issues, post-booking cancellations, leading to customer dissatisfaction
and apologies from the firm to all. A different kind of challenge included Flipkart receiving
mass criticism on the subject of net neutrality after they announced a partnership with Airtel
to use the Airtel Zero platform. On 14 April 2015, Flipkart had to move back to its decision
on using Airtel Zero platform. On the product side, Flipkart faced some setbacks like the
launch of Flyte Digtial Music Store, a legal paid-music download service, which did not get
popular and had to be shut down on 17 June 2013.
With the target of becoming India’s first $100 billion internet company, there are clear
challenges and opportunities at hand for this market leader. Key challenges include manag-
ing the supplier ecosystem; providing superior service for their ever-growing customer base;
managing an efficient back-end; enticing top talent; improving ease of payments; rural sales
management; and so on. Since Flipkart is eyeing to earn more than 50 per cent of its revenue
from mobiles, it sees this platform as its biggest opportunity. This is because from a mar-
keting and customer data standpoint, mobiles provide a greater in-depth transaction-level
data which would help Flipkart forecast better to provide the right kind of experience. It
has also introduced a new mobile web application called Flipkart Lite which is designed for
those who do not want to install apps for reasons like low storage capacity on their phone.
According to sources, Flipkart plans to phase out commissions and wants sellers to adver-
tise on its platform for a fee, a move aimed at enlisting more vendors and at the same time
earning higher advertising fee. Flipkart is also trying to cut dependence on its largest vendor
(WS Retail), which accounts for majority of the sales on the website.
With the company said to have set the target to reach a GVM of $8 billion, by adding
another 100,000 sellers on its platform, it is well set to target selling 1 billion goods by the end
of 2017. Flipkart appears to be the bellwether for all Indian digital businesses in the future
and with its phenomenal success, goes on to prove day after day that strong product knowl-
edge combined with astute marketing can make digital realities possible.
data to appear on any web page, a request is placed by the client to the server using an HTTP
protocol. For example, a web browser like Netscape could be the client and a music player run-
ning on a computer can be the server.
Throughout this mayhem, the biggest positive takeaway was that digital marketing as it is
known to be today got defined and established. Many interesting business ideas came forth,
which a decade later, would flourish to actually become internet behemoths.
The impact of search technology is pivotal to digital marketing and would be discussed in later
chapters in detail.
Post 2000, there have been multiple other technological advances, apart from search, which
have impacted businesses run on the internet. Key ones of those include impact of file-sharing
technology, integration of communication services, growth of design—user interface (UI) tech-
nologies, and emergence of social collaboration tools. We would touch upon a few of these in
detail to understand their right impact on the growth of digital marketing.
table 1.2 Key Differences between Web 1.0 and Web 2.0
The shift from Web1.0 to Web2.0 and now even extending to Version 3.0, has had a major
impact on how marketing through the internet has evolved and continues to do so, thus opening
newer avenues to reach out to and meet the ever-changing demands of the consumer. We would
cover the impact of these changes in the upcoming chapters.
Firm Infrastructure
Activities
Support
Human R a
Resource Management
gin
ar
Technology
n Development
p
M
Procurement
keting, online marketing, or the most recently coined term, digital marketing. In all these
cases, ‘E’ stands for ‘Electronic Networks’ and describes the application of electronic network
technology—including internet and electronic data interchange (EDI)—to improve and change
business processes.
The reason for application of Porter’s Value Chain model is that it explains in a single con-
solidated view the coverage of each of the e-models and why they are essentially different even
though they are used interchangeably in multiple contexts across business discussions and texts.
That said, there are variations in the way these concepts have been explained across various
standard marketing primers and one would benefit from taking this explanation as an instance
of such attempts.
Understanding E-Models
To understand what each e-model encompasses, we would need to map the key physical process
areas as described in Porter’s model and understand the extent and coverage of technology on
each of them in a holistic manner. To understand, let us take an example of the most ubiquitous
internet term ‘e-business’ whose impact covers each of the four process areas outlined in Fig. 1.2
to appreciate how it impacted business in general and became the pioneer of future e-models.
E-business The term ‘e-business’ was coined by IBM’s marketing and internet teams in 1996
and was looked at as a key differentiated market offering. It involved application of information
and communication technologies (ICT) to support all the physical processes and activities of any
business looking to go virtual.
Here are some examples of the kind of processes that impact across the value chain which
form the basis of any e-business model.
(a) Impact on procurement function: management of physical procurement through IT, for
example, Supply Chain System
(b) Impact on production function: improvements in internal production or resourcing, for
example, production/knowledge portal
(c) Impact on delivery function: technology for internal logistics, for example, intermediaries
collaboration portal
(d) Impact on consumption function: integration with internal customer management tools,
for example, CRM platforms
These examples illustrate the key objective of ‘e-business,’ which was set up to help transition of
the physical processes of any business to make them more technology enabled and efficient. The
key point to note here is that ‘e-commerce’ as a concept has a more internal focus to transform
internal processes and tools and enable them for better external collaborations.
The Concept of E-commerce Follows E-business E-commerce is considered to be the sales
aspect of e-business and involves a wide variety of internet-based business models. Typically,
an e-commerce strategy incorporates various elements of the marketing mix to drive users to a
website or an online marketplace (like Ebay) for the purpose of purchasing a product or service.
Comparison of E-models
Systems for
Improvements in Technology for Integration with
physical
E-
production facilities
commerce platforms processes
Marketing
Business moving from production orientation to customer, delivery and search orientation
Baourakis defines it as: ‘The trading of goods and information through the internet’ (Baourakis,
Kourgiantakis, and Migdalas, 2002).
The major difference between e-commerce and e-business is that while e-business is aimed
at improvements in efficiency and productivity, e-commerce is a more revenue-oriented concept
which focusses on bridging internal and external systems through improved collaboration across
all commerce-oriented functions, from procurement to delivery.
As in the case of ‘E-business,’ let us go through some examples of e-commerce implementa-
tion across the value chain which would clarify the differences more clearly.
(e) Impact on procurement function: cross-business integration with portals/systems of other
suppliers, for example, portal-based procurement
(f) Impact on production function: integration of common production facilities, for example,
value added production chains
(g) Impact on delivery function: collaboration with external delivery platforms for example,
web-based marketplaces
(h) Impact on consumption function: improvement in transaction/sales/payment-focused
processes, for example, customer payment platform
As can be seen above, e-commerce has a stronger orientation towards technology investments
for commerce and better collaboration with the internal–external tools to carry out the sales
process.
As can be seen in Table 1.3, e-commerce in India had started with the launch of IndiaMART
in 1996, almost during the same time as the rise of the dotcom era. The recent growth in
e-commerce had started with the rise of Flipkart in 2007 by two ex-Amazon employees which
gave Indian customers the first taste of how orders, placed through a website, can be fulfilled effi-
ciently, in time, and of the expected quality. (A detailed case on Flipkart and its unique growth
characteristics is covered in the beginning of this chapter.)
1998: Electronic postal stamps can be purchased and downloaded for printing from the web.
1999: Alibaba Group is established in China. Business.com was sold for US $7.5 million to
e-companies, which was purchased in 1997 for US $149,000. The peer-to-peer file sharing
software Napster was launched
2000: The dotcom bust
2001: Alibaba.com achieved profitability in December 2001.
2002: eBay acquires PayPal for $1.5 billion. Niche retail companies like Wayfair and Netshops are
founded with the concept of selling products through several targeted domains, rather than
a central portal.
2003: Amazon.com posts its first yearly profit.
2004: DHgate.com, China’s first online B2B transaction platform is established, forcing other B2B
sites to move away from the ‘yellow pages’ model.
2007: Business.com was acquired by R.H. Donnelley for $345 million.
2009: Z
appos.com was acquired by Amazon.com for $928 million. Retail Convergence, an operator
of private sale website RueLaLa.com, was acquired by GSI Commerce for $180 million, plus
up to $170 million in earn-out payments based on the performance through 2012.
2010: Groupon reportedly rejects a $6 billion offer from Google. Instead, the group buying
websites went ahead with an IPO on 4 November 2011. It was the largest IPO since Google.
2011: Quidsi.com, the parent company of Diapers.com, was acquired by Amazon.com for
$500 million in cash plus $45 million in debt and other obligations. GSI Commerce, a
company specializing in creating, developing and running online shopping sites for brick
and mortar businesses, was acquired by eBay for $2.4 billion.
2012: US ecommerce and the online retail sales was projected to reach $226 billion, an increase
of 12 per cent over 2011.
2014: Overstock.com processes over $1 million in bitcoin sales. India’s e-commerce industry is
estimated to have grown more than 30 per cent from a year earlier to $12.6 billion in 2013.
procurement to sales, e-business can also include enhancement of internal management pro-
cesses like employee services, training, internal information-sharing, video-conferencing, and
recruitment which help improve internal productivity.
Moving ahead from business and commerce, the most commonly used marketing term to denote any form
of internet operations is e-marketing. By definition, e-marketing involves utilization of electronic
medium to perform marketing activities and achieve the desired marketing objectives for an
organization. This term is typically misused for e-business and e-commerce and, as per proper definition,
should also be used to denote and represent marketing-related activities.
The focus of e-marketing is to facilitate the transition of physical marketing and sales-based
activities to a more automated, technology supported system which can drive the product discov-
ery process. Two examples which make the concept of e-marketing clearer include:
(a) Delivery of products/services through collaborations with external partners, for example,
logistics platform implementation
(b) Improvement of internal sales and marketing systems for stronger customer connect, for
example, loyalty automation platforms
Before we move on to explain ‘Online Marketing,’ we should understand how it differs from
e-marketing. Typically (as shown in Fig.1.2), e-marketing has a broader coverage which includes
not only online/internet marketing aspects but also more enterprise-specific marketing areas like
e-CRM, digital customer data, which gives it a broader scope than both of them. This is also the
reason why in the e-models comparison diagram, e-marketing has been shown in the traditional
bucket along with e-business and e-commerce.
The Concept of Online/Internet Marketing Online/Internet marketing started with refer-
ence to the internet and e-mail-based aspects of physical marketing and went on to grow and
represent activities aimed at transitioning all marketing activities to web and online platforms—
most primary of them being web-portals, e-mails, and blogs.
Along with the transition to web, the other most crucial characteristic of online market-
ing was the collaborations which started developing with other top platforms/websites holding
inventory for commerce including Amazon/E-bay. These collaborations formed the basis of the
growth of online marketing to its present avatar called digital marketing whose mainstay is deep
technology and multi-platform marketing engagement.
Key examples which bring forth the power of digital marketing in the context of the Porter
model include:
(a) Delivery function: Collaboration with advanced technology intermediaries, for example,
Marketing Automation Tools for Lead Generation/Reverse Auction Platforms
(b) Consumption function: Implementation of digital marketing platforms, channels, and con-
cepts, for example, Search Marketing, Display Advertising, Social Media Marketing, etc.
While in the above section we have made introductions to key e-concepts and the basics of
online and digital marketing, before we delve deeper into defining digital marketing, its types,
models, and applications, it is imperative to also spend some time to understand the basis behind
digital becoming so important to companies globally and how it has become a broad subject of
study.
Digital marketing typically corresponds to consumption function, wherein the main scope of
this book lies. The importance of explaining digital transformation elements (before we cover
digital marketing in detail) is to help visualize it as a part of the overall digital transformation
strategy. Before we move further that since the scope of digital marketing itself has expanded
tremendously, this textbook aims to cover digital marketing strategies in depth, while touching
upon other digital transformation elements, as and when they seem important in relation to
digital marketing itself.
Internal Teams
Producrement Production Function Automating
Automation Consumption
Customers
Suppliers
Transformation Analytics
Types
Cloud
Automation
Data Technologies
Digitization
(b) Consumer-initiated contact (Pull marketing): Pull marketing involves a consumer plac-
ing his intent and specific interest for a particular type of information and being offered
that information along with relevant marketing messages suited to his intent, query, or
profile-based interests. In the same example of a newspaper, as above, consider that the
company decides to provide information in a way that each article of the newspaper is
tagged and categorized based on multiple criteria to be searched upon a digital platform.
In this case, when a consumer makes a specific search for a piece of content and chooses
matching information/news, promotions are sent based on his profile preferences, real-
time location, and the topic of the article he found interested in reading. This would then
become be an example of pull marketing.
Based on key characteristic of these two types of marketing, the development of new digital
platforms can be explained and understood. Digital marketing includes all those techniques and concepts
that utilize the intent (pull-based) action of the consumer to market products and services (in a push-based
manner) which would be most needed, relevant, and of interest to him/her.
Let us understand a few key types of digital marketing areas to see how digital technologies
have been built around the combinations of pull–push marketing concepts:
(a) Search marketing: Search was one of the most pioneering pull–push marketing con-
cepts which involved providing a technology platform to consumers to help them express
their intent towards finding a particular piece of information and building a push mar-
keting-based business around that search. (We would cover this particular topic and its
technologies in detail in subsequent chapters and sections).
(b) Display advertising: Similar to print advertisements, display advertising was built on pull-
ing data from consumers’ readership interests on any particular website wherein display
ads were pushed next to the content being read for consumers to interact, click, and buy
on landing pages created for this action.
(c) Social media marketing: With the power of social networks coming to internet-based
platforms, a marketer could not only use the information pulled from customer intent but
also from the intent of his social networks which would be marketed back to him in the
form of sponsored and native advertisements.
BRANCH A. PHANEROCEPHALA.
Sub-Order 1. Nereidiformia.[367]
Fam. 1. Syllidae.—These are small worms, the majority being less than an
inch long, so that they are not easily observed. The body consists of a fair
number of segments.[368] In many genera a dorsal bundle of unjointed,
natatory chaetae makes its appearance at maturity. The palps, which are
grooved, are in some cases so united with one another and with the
prostomium as to be scarcely recognisable. (For head see p. 262, and for feet
see p. 264.) The pharynx is armed with one or more teeth. There is a special
gizzard, following the pharynx, and provided with thick, muscular walls of
peculiar structure. Following the gizzard, the oesophagus receives in many
genera a pair of T-shaped diverticula, that are used for storing water, which is
swallowed with food. These diverticula are absent in Autolytus and other free-
swimming forms. The reproduction of the members of this family is
interesting, and has already been described (p. 278).
Fig. 160.—Syllis armillaris Müll. × 2. (From Johnston.) The head is towards the
right.
Syllis.—The tentacles and cirri are moniliform; the palps large; there is a
single dorsal tooth, which is provided with a poison gland, the duct of which
opens near its apex; it is used rather for stabbing its prey than for grasping
and tearing. S. krohnii Ehlers, is abundant under stones, and forms tubes of
sand; it is nearly an inch long, and consists of some eighty-five to ninety-five
segments marked with yellow bands. It may readily be identified by longer
dorsal cirri, terminally dilated, alternating with shorter ones. S. cornuta
Rathke, has a translucent green body, about half an inch long; no alternation
of cirri. Mediterranean, Atlantic, on the Norwegian coast, off Spitzbergen, and
on the Madeira coast. S. armillaris Müll. is very common at low water; it is
pale yellowish-brown, with a couple of dusky marks on each segment; and
measures 2 inches. The dorsal cirri are quite short, consisting of only eight to
ten joints. In Pionosyllis the tentacles and cirri are not moniliform; a single
dorsal tooth. P. malmgreni M‘I. under stones. Sphaerosyllis.—The dorsal cirri
are swollen at the base, and are not moniliform; the long palps are fused
along nearly their whole extent. S. hystrix Clap. is only about one-eighth of an
inch in length. Exogone Oerst. Grubea Qfg.
Autolytus.—The small palps are entirely fused with the prostomium; the
pharynx, which is bent upon itself, is armed with a circle of denticles. Dorsal
cirri somewhat foliaceous. There are no ventral cirri. The male and female
differ from one another and from the asexual "stock" (see p. 279). A. pictus
Ehl. is abundant under stones. It measures about two-thirds of an inch in
length, is darkly coloured with a median lighter band; the anterior dorsal cirri
are long. A. prolifer Müll. is common.
Myrianida fasciata Milne Edwards, with its foliaceous cirri, occurs off our
coasts (see Fig. 149, p. 280). Atlantic, Mediterranean.
Psamathe Johnston, has many segments; head with two tentacles and a pair
of three-jointed palps. P. fusca Jnstn. occurs amongst coralline Algae, to
which it bears some resemblance, which is heightened by the moniliform cirri.
It is a small worm, less than an inch in length. Mediterranean. Castalia
punctata Müll. is dirty green or brownish, with a narrow purplish band on each
side. It occurs in deeper water than the preceding. In Ophiodromus the head
has three tentacles; the palps are two-jointed; there are six pairs of
peristomial cirri; the parapodia are biramous. O. vittatus Sars is dredged in
numbers off the Scotch coast, and is found also at low tides. It measures 2
inches in length. A closely allied species lives in the ambulacral grooves of
the starfish Astropecten.
A. With twelve pairs of elytra.—In P. squamata Linn. the elytra entirely cover
the body and conceal the head, each elytron overlapping the next posterior
one, and those of the two sides overlapping. General colour sandy-brown,
speckled, lighter or darker. The fringed elytra are very firmly fixed to the body.
The notopodial chaetae scarcely project from below the elytra. The worm is
common between tide-marks and in the coralline region, is about one to one
and a half inches in length, and about one-third of an inch in width. Atlantic. P.
clava Montagu, may attain a larger size, though it is generally smaller. The
elytra are dark, usually grey, mottled with white or light grey, unfringed, and do
not overlap to so great an extent as in P. squamata, so that the middle of the
back and the hinder part of the body is more or less exposed. This is never
the case in the preceding species, but even here it is subject to variation in
extent, depending on the amount of food contained by the worm or on the
ripeness of the genital products. It occurs in the Mediterranean.[371]
In Hermione the "felt" is absent, so that the elytra are exposed. H. hystrix Sav.
occurs in ten to thirty fathoms of water all over the British area and
Mediterranean. It resembles in its general appearance a fat Polynoid, with
strong chaetae. Laetmonice filicornis Kinb. also occurs on our north-west
coasts, and L. producta Gr. has been dredged in 500 fathoms off the west
coast of Ireland; it has been recorded also from Kerguelen and from Japan,
so that it has a very wide distribution.[373]
In Eulalia an additional (fifth) tentacle arises from the middle of the back of the
prostomium. E. viridis Müll. is a dark green worm smaller than the preceding;
common between tide-marks, hiding in cavities and tunnels in limestone
rocks, which have been bored by the mollusc Saxicava; it is rare where such
rocks are absent. It might have been thought that its vivid colour would
harmonise with its surroundings, but it is most abundant in regions where
Fucus abounds and Ulva is absent. It is evident then that the colour is not
protective; it may perhaps be of warning significance, for the mucus secreted
in quantities by glands on the cirri of the Phyllodocids is probably
objectionable to their enemies. Phalacrophorus Grf. and Pontodora Grf. may
be mentioned as pelagic genera.
N. dumerilii Aud. and Edw. is rather smaller and narrower than the two
preceding species; it is reddish-violet in colour, marked with darker transverse
lines in each segment. It is readily recognised by the two dark brown spots on
the upper surface of the base of the notopodium in most of the segments, and
by the great length of the peristomial cirri, the longest of which reaches the
fifteenth segment. It is sometimes found enclosed in a cocoon-like tube of
hardened grey mucus, more or less covered with foreign particles, such as
sand grains. Atlantic, Mediterranean, Japan.
Fig. 168.—Nereis cultrifera Gr. × 6. Head with buccal region everted, to show
the arrangement of the jaws. (From Ehlers.) Cf. N. diversicolor, Fig. 125, p.
248.
N. (Nereilepas) fucata Sav. lives in the topmost whorls of empty whelk shells
and in those occupied by hermit crabs. The ground colour is tile-red, with two
milk-white bands along the dorsal surface. The dorsal lobe of the foot is
slightly foliaceous, glandular, and vascular.
Eurythoe borealis Oerst. measuring 6 inches, occurs all round the British
area, from the Shetlands, where it occurs in deep water, to the Channel Isles,
where it lives on shore, under stones, etc. (For parapodium of Amphinome
see p. 264.)
Euphrosyne.—The body is short, oval, and flattened. The parapodia are not
distinct processes, but the chaetae extend from each side of each segment
nearly to the middle dorsal line, and are absent ventrally (Fig. 137, C, p. 265).
The dorsal and ventral cirri are more or less filiform, and there is an
intermediate similar process on the back (? = lip of chaetigerous sac).
Amongst the chaetae are a number of curious branched processes—usually
called "gills."[380] The presence of these and of the chaetae give the upper
surface of the body a fluffy appearance. E. foliosa Aud. and Edw. is fairly
common under stones on our southern shores. It is about an inch in length
and is of a cinnamon-red colour.
Eunice has five tentacles, two great palps, and a pair of nuchal cirri; the gills
are pectinate, and there are four anal cirri. E. harassii Aud. and Edw. is about
8 inches long. It is reddish-brown, with white spots down the back, one to
each segment, and others at the sides. The gills begin at the sixth segment,
and when fully developed have eleven branches. The dorsal cirrus is not
longer than the gill. E. philocorallia Buch.[381] forms its tube amongst the
branches of Lophohelia prolifera, in 200 fathoms, off the west coast of Ireland.
Onuphis Oerst. has a head like the preceding, from which it differs in having
pectinated gills and two nuchal cirri like Eunice. In making its tube it employs
small pebbles, bits of shell, and even echinid spines, which it glues together
with mucus, so that it bears a general resemblance to its surroundings. O.
conchylega Sars, has a flattened, scabbard-like tube, which can be carried
about by its owner. Atlantic.
Fig. 171.—Glycera meckelii Aud. and Edw. with pharynx everted, × 1. (Règne
Animal.)
Goniada is distinguished from the preceding by the fact that the parapodia
suddenly change in size and character at about one-third the length of the
body. The pharynx has numerous paragnaths. G. maculata Oerst. occurs off
our coasts.
Fam. 11. Sphaerodoridae.—The dorsal and ventral cirri of each segment are
spherical. The chaetae are usually jointed, and there is an aciculum to each
parapodium. Ephesia Rthke. (E. gracilis R. = Sphaerodorum peripatus Jnstn.)
is exceptional in having unjointed chaetae. North Sea, Arctic Ocean, and the
Channel. The family, which is much modified, is allied in some respects to the
Syllidae.
Sub-Order 2. Spioniformia.
Spio seticornis Fabr. is a small worm less than an inch in length, colourless
except for the red blood in its vessels. It builds long and flexible tubes of sand
in the clefts of rocks and under stones in the upper part of the littoral zone.
The prostomium is notched at the anterior margin. The gills commence on the
twelfth segment, and do not extend to the end of the body. A membrane-like
cirrus exists also on the second chaetigerous segment. The podal membrane
is adnate to the gill throughout its extent. Four short anal cirri occur.
Greenland and Scandinavia.
P. ciliata Jnstn. inhabits soft mud tubes near low water; it also makes U-
shaped galleries in stones and shells, and the tube projects from each mouth.
The worm is about ½ inch long, consists of some forty segments, and is
yellowish or flesh-coloured. The prostomium resembles that of Spio; the
peristomium is raised into a slight collar at each side. The anus is surrounded
by an incomplete funnel. The species has almost a world-wide distribution,
having been recorded from Iceland, Australia, the Philippine Islands, as well
as from the European seas. P. coeca Oerst. often lives commensally with a
sponge, having a protective odour.