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Ethics Morality and Business The Development of Modern Economic Systems Volume I Ancient Civilizations 1St Edition Edition Dipak Basu Full Chapter
Ethics Morality and Business The Development of Modern Economic Systems Volume I Ancient Civilizations 1St Edition Edition Dipak Basu Full Chapter
Ethics, Morality
and Business: The
Development
of Modern Economic
Systems, Volume I
Ancient Civilizations
Dipak Basu Victoria Miroshnik
Nagasaki University Reitaku University
Nagasaki, Japan Kashiwa, Japan
© The Editor(s) (if applicable) and The Author(s), under exclusive license to Springer
Nature Switzerland AG 2021
This work is subject to copyright. All rights are solely and exclusively licensed by the
Publisher, whether the whole or part of the material is concerned, specifically the rights
of translation, reprinting, reuse of illustrations, recitation, broadcasting, reproduction on
microfilms or in any other physical way, and transmission or information storage and
retrieval, electronic adaptation, computer software, or by similar or dissimilar methodology
now known or hereafter developed.
The use of general descriptive names, registered names, trademarks, service marks, etc.
in this publication does not imply, even in the absence of a specific statement, that such
names are exempt from the relevant protective laws and regulations and therefore free for
general use.
The publisher, the authors and the editors are safe to assume that the advice and informa-
tion in this book are believed to be true and accurate at the date of publication. Neither
the publisher nor the authors or the editors give a warranty, expressed or implied, with
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This Palgrave Macmillan imprint is published by the registered company Springer Nature
Switzerland AG
The registered company address is: Gewerbestrasse 11, 6330 Cham, Switzerland
This book is dedicated to the famous philosopher of Japan
Professor Chikuro Hiroike, founder of the Reitaku University
&
to our uncle Nishith Ranjan Mitra,
the Managing Director of Deb Sahitya Kuthir,
the famous publishing house of India,
who have inspired us always.
Introduction: Civilizations and
Morality
vii
viii INTRODUCTION: CIVILIZATIONS AND MORALITY
As the ethical codes give rise to religions, the unifying force combining
a group of people was always religious. In the early civilizations, religion,
agriculture, and technology were all interrelated. If the religion and ethical
code are not interrelated, then the civilization will be corrupted and may
even die. Religion is a psychosocial character, which combines the mind
of both individuals and the society. Human society continues along with
religion, which guides the society to some specific direction and provides
satisfaction during a crisis.
Religion obviously implies a sense of duty and virtue, and uncondi-
tional respect for moral rules, rights, and needs—“do your duty without
expecting the results,” as Krishna said in Bhagavad Gita. It is also related
to Kant’s “categorical imperative,” on respect for human dignity and
virtue, the rigorous fulfillment of moral principles.
Opposite examples include:
lives; this is descriptive ethics. It may try to tell us what is morally right
and wrong. That’s called normative ethics.
Descriptive ethics raises the question about the role “morality” actually
plays in the actual economy. We may say that business has nothing to do
with morality. However, morality has important roles for the efficiency
of any economic system. In an exchange economy, if there is no trust in
transactions, there cannot be any transaction.
In recent decades, efforts have been made to implant capitalist
economic systems and impose capitalist structural reforms on coun-
tries meant to induce some sort of self-sustaining economic growth.
The efficient functioning of economic institutions is not independent
of its cultural context. If economic theory itself is ignorant of that
context, it will be irrelevant and could not produce any real effects.
The success of an economic system and its resultant institutions are very
much contingent on cultural factors and moral criteria. Moral norms
make economic behavior more predictable and reliable. Thus, they make
economic transactions possible.
Moral norms create an important economic resource, called “social
capital,” denoting intangible resources that can be sustained via social
relationships. Such moral norms include:
These are issues that can undermine these ethical norms. If these norms
may be exclusive to a family organization, there can be nepotism and
favoritism which discriminate against outsiders.
There can be bribery, which discriminates those who play by the rules
and creates an atmosphere of fraud. There can be cartels and price fixing
in which companies form “oligopolies,” restricting competition to create
excessive profits. As a result, consumers and the country have to incur
losses. Prices can go up and there can be mal-distribution of wealth and
assets.
x INTRODUCTION: CIVILIZATIONS AND MORALITY
1. The world sages have taught human society about sacred transcen-
dental beings that permit various values and standpoints with regard
to life to coexist in a peaceful manner.
2. When we take this moral approach and seek to understand the
universe and the divine, we see that the universe reflects the soul
of the divine while humankind takes its place as an element of the
universe, retaining part of that soul.
Buddha said, “Very form is emptiness and emptiness if form”;
we understand that people and things of this world are mutable and
INTRODUCTION: CIVILIZATIONS AND MORALITY xi
May all creatures be happy minded, As a mother at the risk of her life watches
over her own child, so also let everyone cultivate a boundless (friendly) mind
towards all beings ”. (Sutta-nipata, viii, 5–7)
Buddhist ethics finds its foundation not on the changing social customs
but rather on the unchanging laws of nature. Buddhist ethical values are
intrinsically a part of nature and the unchanging law of cause and effect
(karma). The simple fact that Buddhist ethics are rooted in natural law
makes its principles both useful and acceptable to the modern world.
INTRODUCTION: CIVILIZATIONS AND MORALITY xiii
Similarly, Christ in his sermons from the mountain asserted the moral
values of his messages:
Blessed are the poor in spirit: for theirs is the kingdom of heaven.
Blessed are they that mourn: for they shall be comforted.
Blessed are the meek: for they shall inherit the earth.
Blessed are they who do hunger and thirst after righteousness: for they shall
be filled.
Blessed are the merciful: for they shall obtain mercy.
Blessed are the pure in heart: for they shall see God.
Blessed are the peacemakers: for they shall be called the children of God.
Blessed are they who are persecuted for righteousness’ sake: for theirs is the
kingdom of heaven.
(Mathew, 5: 3–12).
In this book, we traced the important philosophers who could not get
proper attention so far. The reason is that normally we ignore eastern
philosophy and social system. Thus, we include analysis of Hindu philos-
ophy and associated history, which was neglected so far in the Western
literature. Given the vastness of the subject, we could not include every
sage we wanted to include.
Reference
Hiroike, C., 1928. Treatise of Moral Science, Vol. 1–9. Kashiwa: Reitaku
University.
Contents
Conclusion 189
Index 191
xvii
List of Diagrams
xix
CHAPTER 1
According to the Puranas, Human history, after the last ice age, possibly
started in about 80,000 years ago (Mazumdar 1917). Manu was
born in about 72,000 years ago. He was called Adima or the first Patri-
arch. He had seven sons who occupied seven continents of the world.
Bhavishya Purana gave clear account of that period. Aryans after their
migrations out of India reached the north of Russia, which was called
Uttara Kuru. Rig Veda mentioned large Aryan settlements in Roosam
(Russia) and Hariyupia (Europe) before 3000 B.C. (Mazumdar 1917).
After the discovery of the Denisovians in some caves in Siberia we
have to push back that time to perhaps 400,000 years ago (Zubova et al.
2017). Denisovians were giants, called Davav in the Puranas, who used
to live in Balochistan before.
Thus, Aryans went everywhere in every continents and all civiliza-
tions came out of the Aryan influence. Aryans, derived from the original
Sanskrit word Arya, means cultivated educated person, according to Sri
Aurobindo, used to have very high moral standard with great regard for
truth, promise, and hospitality. Thus, we have to describe the Aryans if
we want to analyze moral aspects of human civilization. Their sages, or
Rishis, after many years of meditations and learning, derived some great
ideas (Riks) on God and universe.
Ancient Aryans divided the entire world into three land masses: Aswa
Kranta (Eurasia), Ratha Kranta (Africa), Vishnu Kranta (the two Amer-
icas). They divided the world into seven continents: Saka (Europe),
Jambu (Asia), Plaksha (a submerged continent between Asia and Africa),
Pushkara (North America), Kusa (South America), Salmali (Africa), and
Krauncha (Oceania).
Europe was called alternatively Hariyupia, Saka, and Ishurupa.
According to Mahabharata, a son of Krishna, Samva brought a group
of Sakal Brahmins to modern Multan (then called Mitra Sthana or Mula
Sthana). Hue En Shang, the Chinese monk when visited that place, he
has seen a Sun Temple, Sambapura, made of gold.
Some of the Aryans after leaving India entered Babylon, Assyria, and
Egypt. They are called Panis in Rig Veda. Most of them settled in Asia
Minor (Historians History of the World, 1902). Rig Veda mentioned
them as great navigators using Satarika Nau (a ship with 100 oars).
Original people of Assyria were not Semitic. Before 4000 B.C.,
Southern Babylon was the original home of the Sumerians from India
and Northern Babylon originally came from central Asia. Modern name
Mesopotamia came from the original Madhya Vedi, according to the
Historians History of the World (vols. 1 and 2, 1902). One of the most
famous King of Babylon was Asur Bani Pal, a pure Sanskrit name. Both
Hittite and Mitranis used to speak Indo-European language. Their gods
were Vedic gods.
HR Hall, curator of the British Museum wrote (Hall 1939), “The
ethnic type of the Sumerians so strongly marked in their statues and relief
was as different from those of the races which surround them as was their
language from those of the Semites; they were decidedly Indian in type. The
face type of the average Indian of today is no doubt much the same as that
of his race ancestors thousands of years ago. And it is by no means improb-
able that the Sumerians were an Indian race. It was in the Indian home,
perhaps the Indus valley; we suppose for them, that their culture developed.
There their writings may have invented and progressed from a purely picto-
rial to a simplified and abbreviated from which afterwards in Babylonia
took on its peculiar cuneiform appearance owing to its being written with a
square ended stylus on soft-clay. There is little doubt that India must have
been one of earliest centres of human civilization and it seems natural to
suppose that the strange un-Semitic people who came from the East to civilise
the West were of Indian origin, especially when we see with our eyes how very
Indian the Sumerians were in type”.
1 ORIGIN OF THE ARYANS 3
Rig Veda (1, 22, 16–21) mentioned Vishnu the leader of the Aryans
and his people used to live in a place called Indra-Laya from 8000 to
5000 B.C., and in 5000 B.C. they started to disperse in different direc-
tions. Manu was the first person, the origin of the Aryans. Priyavrata,
son of Manu, according to Vishnu Purana (II, 1), Garura Purana, Bhaga-
vata (V, 1), Devi-Bhagavata (VIII, 4), divided his kingdom among his
ten sons but three of them refused to be Kings, instead chose to be
hermits. Prince Agnidhara got Jambu Dwipa (Asia), Medhatithi got
Plakssha (submerged continent between Africa and Asia). Vapushman
got Samali (Africa), Dyutiman got Krauncha Dwipa (Australia), Bhavya
got Saka-Dwipa (Europe), Savala got Pushkara-Dwipa (North America),
Jyotishman got Kusa Dwipa (South America). The grandson of Manu,
Agnidhara divided Asia among his nine sons (Vishnu Purana [II, 2],
Garura Purana [I, 54], Brahma Purana [18]).
India was able to assimilate itself to Buddhism by accepting the Aryan
traditions. Gautama Buddha himself was an Aryan, His philosophy is
called the “Aryan Way.” That is the reason it is important to know the
origin of the values and traditions of the Aryan society. This chapter
analyzes the history of the Aryan civilizational values.
Indians and Greeks are the two major Aryan tribes. Their relationship
through the philosophical tradition is very close. Upanishad and Eleatics
are almost similar. Xenophanes taught that God and the Universe are
one, eternal, and unchangeable. Parmenides holds that Universal being is
neither created nor to be destroyed and omnipresent, only reality. These
are similar to the messages of Upanishads. The idea of ancient Greece that
everything came from water is similar to the idea of the Vedas. Ideas of the
Samkhya philosophy also have its reflection in Greece. Samkhya doctrine
said that the material world is produced by Prakiti, the primitive matter,
and ultimately sinks back to it. Anaximander thought that foundation of
all things, a primitive matter, eternal, unfathomable and indefinite, from
which definite substances arise and into which they return. Innumerable
annihilations and reformations of the universe of the Samkhya doctrine
have its reflections in philosophy of Heraclitus.
Samkhya proposed the eternal and endless reality of all products and
eternity and indestructibility of matter. These are reflected in the theories
of Empedocles and Epicurus. Almost all doctrines of Pythagoras used to
exist in India early sixth century B.C., including the Pythagoras theorem
or irrational number in Sulva-sutra (Von Schroeder 1884). The doctrines
of Pythagoras, number is the essence of things, elements of numbers
1 ORIGIN OF THE ARYANS 5
wise man abandons both joy and sorrows by meditation on the self, and
is released from the jaws of death.”
Charvaka Lokyatika are materialistic philosophers. Discarding idealism
and mysticism, the materialists developed the proposition that only this
world exists, there is no supernatural purpose; nature is governed and
directed by itself. Brihaspati was the founder of this school. Four mate-
rial elements, fire, water, air, and, earth, are the basis for everything.
There is no God. This world is that exists (Chattopadhya 1959). Only
direct perception gives man genuine knowledge. There are two kinds of
sense perceptions: internal and external. External perceptions are from
the activity of the senses. Internal perceptions are the result of mental
operations based on data obtained from the senses.
A related system of philosophy is that of Nyaya or logical school.
Gotama is considered as the founder of this school. Greek philoso-
pher Callisthenes was with the army of Alexander, when he invaded
India. He learned Nyaya from India and passed this system to Aris-
totle. Nyaya is the foundation of Aristotle’s logical theory (Brodov 1984).
According to Nyaya, material world exist objectively and existence of the
external objects does not depend on the source of knowledge. Everything,
that is not accessible through sense perception, has no real existence.
Nyaya is concerned with the instruments of cognition. There are four
types of cognitions: perceptions, inference, comparison, and verbal testi-
mony. There are similarity of Nyaya and Charvakas as Brihaspati was the
common originator of the both systems.
life styles and food habits for these three Gunas. Samkhya philosophy
describes the development of this world from finer forms to the dense
forms or material elements. Matter in the general form is the first cause
of the world of objects, but it was not cognizable as it is in its finer
form. Matter subsequently became in its singular form became moving
objects, phenomena, and events developing in space and time. Purusha, or
extremely fine element is beyond any cognition is the origin of conscious-
ness; it is passive, only Prakiti or matter creates everything, when it comes
into contact with the Purusha. At a certain stage of the development of
the world, consciousness arises out of things while quantitative changes
reaching a certain phase of development became qualitative changes
producing new qualities (Ram Mohan Roy 1902/1982). According to
Samkhya doctrine, the world is material. Matter is the ultimate basis of all
that is. It is eternal and one. Matter cannot be either created or destroyed.
Yoga is a method based on Samkhya school of Philosophy to free
oneself to be united with the absolute soul or Atma (soul of the Brahman,
the creator) which would mean the end of all suffering and release from
the cycle of birth and rebirth. Patanjali, the main proponent of the Yuga
school, interpreted the Samkhya philosophy in the spirit of dualism. The
material (Prakiti) and the spiritual (Spiritual) are regarded as opposite
principles that cannot be reduced to each other, but spirit is the supreme.
As Krishna said, “Spirit, linked to molded matter, enter into bond with
qualities by nature, framed and thus married to matter” (Bhagavad Gita,
Ch 13). Man is one of the highest and most complicated material-spiritual
combinations. Birth and death should be considered as changes in the
matter, but not in the soul. There is no death, but long cycle, which
is called reincarnations or Samsara. Soul attains a degree of purity from
matter after going through a number of reincarnations.
Patanjali described man as system composed of several sub-systems,
mineral man, vegetable man, animal man and ultimately human man
when an individual spiritual subsystem, called Purusa (the soul), is intro-
duced into the individual material subsystem (Prakiti). The activity of the
human man then rises to the highest level, as it is endowed with reason
and soul, ready for the liberation of the soul from the material world to
meet the Atma and receives the ultimate release from the Samsara, the
cycle of birth and rebirth.
The final philosophical system is Vedanta (which means the end of
the Vedas or the essence of these) based on Brahma-Sutra of Badarayana
of seventh to fifth century B.C.. It explains the nature of Brahman, its
1 ORIGIN OF THE ARYANS 11
relationship with the individual soul and the world. There are various
explanatory books within this school. Mandukya Karika of Gaurapada
(seventh century B.C.) based on Mandukya Upanishad is one of the
most important. It explains the concept of Maya or Illusion. The world
according to Gaurapada is just as unreal when he is awake, as it is
when he is asleep. The only reality is Brahman. Samkara in the eighth
century explained it further. Brahman is infinite, eternal. The world of
phenomena, of nature and matter are the dreams or illusion of Brahman.
The relationship between Brahman and this world is like the relation-
ship between the ocean and its waves, like an object and its properties.
Our world is the Maya or sleep of the Brahman. Maya is neither real nor
unreal. Sensory and logical cognitions are relative empirical truth, not
the substantive basis of truth. Knowledge about the Brahman is the real
cognition of the truth.
Ramanuja in twelfth century disputed that according to him, Brahman
is manifested in the form of this world, which is real and not an
illusion (Brodov 1984). Sri Chaitannya in fourteenth century is a
follower of Ramanuja’s Bhakti Movement, according to whom the God
is accessible to all, irrespective of caste or even religious differences.
Swami Vivekananda of nineteenth century India and ISKCON (Inter-
national Society of Krishna Consciousness) followed this interpretation
of Ramanuja. The cause of human sufferings does not lie in the illu-
sory world concealing Brahman but we cannot demonstrate enough love
and loyalty to Brahman. For the followers of Ramanuja serving any living
creature is the worship of the Brahman.
Appendix A
Ancient India: possibly 15000 B.C. and after
Vedic Period (Source: Vedas)
Period of Ramayana and Raghu Dynasty (possibly 7500 B.C.)
Period of Mahabharata and Bharat Dynasty (about 4000 B.C.)
Indus Valley Civilization (3500 B.C.)
Birth of Gautam Buddha (563 B.C.)
Invasion of Alexander (326 B.C.) and Chandra Gupta (321 B.C.)
Asoke (265 B.C.) and Expansion of Buddhism
India in South East Asia: Sri Vijaya (Indonesia), Barman (Burma),
Champa (Vietnam), Cambojh (Cambodia and Laos), Suvarnabhumi
(Thailand)
Samudra Gupta and Gupta Dynasty (320 A.D. to 600 A.D.)
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external. Secondary and Tertiary strata——. Genera: Arcomya,
Goniomya, Pleuromya, Machomya.
Fam. 7. Anatinidae.—A fourth (?byssal) pallial orifice, siphons
long, separate or fused; shell thin, sometimes inequivalve, exterior
often granulose, ligament often with ossicle, hinge toothless or with
lamellae. Jurassic——. Genera: Anatina, Plectomya (Secondary),
Periploma, Cochlodesma, Thracia, Tyleria, Alicia, Asthenothaerus.
Fam. 8. Grammysiidae.—Shell equivalve, oval, ligament external,
cardinal margin straight, toothless, pallial line complete. Palaeozoic
——. Principal genus, Grammysia, with many other genera of
toothless hinge, but whose exact affinities are uncertain.
Fam. 9. Praecardiidae.—Shell thin, equivalve or not, radiately
ribbed, margins dentated, sub-umbonal area as in Arca, hinge
toothless. Palaeozoic——. Principal genus, Praecardium.
Fam. 10. Pholadomyidae.—A fourth pallial orifice, siphons very
long, united, foot small; shell thin, equivalve, with radiating ribs,
ligament external, hinge toothless, pallial line sinuate. Jurassic——.
Single genus, Pholadomya.
Fam. 11. Clavagellidae.—Foot rudimentary, siphons long, united,
contained in a long calcareous tube; shell small, one or both valves
soldered in the tube, tube with a centrally fissured disc at the anterior
end, more or less frilled at the posterior end. Cretaceous——.
Genera: Clavagella, Brechites (= Aspergillum).
Order V. Septibranchiata
Mantle edges united at three points, branchiae replaced by a
muscular septum extending from the anterior adductor to the point of
separation of the siphons, septum with symmetrical orifices.
Fam. 1. Poromyidae.—Branchial septum with groups of lamellae
between the orifices, labial palps large, foot long and narrow,
siphons short, papillose, separated, animal hermaphrodite; shell
small, slightly inequivalve, rounded or oblong, nacreous within.
Eocene——. Genera: Poromya, Silenia.
Fam. 2. Cuspidariidae.—Siphons long, united in part, foot short,
animal dioecious; shell small, slightly inequivalve, rostrate, not
nacreous, each valve with ligamentary cartilage spoon-shaped, with
a calcareous ossicle, cardinal and lateral teeth present or absent.
Jurassic——. Single genus, Cuspidaria, with many sections.
BRACHIOPODA
PART I
RECENT BRACHIOPODA
BY
INTRODUCTION—SHELL—BODY—DIGESTIVE SYSTEM—BODY
CAVITY—CIRCULATORY SYSTEM—EXCRETORY ORGANS—
MUSCLES—NERVOUS SYSTEM—REPRODUCTIVE SYSTEM
—EMBRYOLOGY—HABITS—DISTRIBUTION—
CLASSIFICATION
Introduction
The group Brachiopoda owes its chief interest to the immense
variety and great antiquity of its fossil forms. Whereas at the present
time the number of extant species amounts to but about 120,
Davidson in his admirable monograph[410] on the British Fossil
Brachiopoda has enumerated close upon 1000 fossil species, found
within the limits of the United Kingdom alone.
The amount of interest that the group in question has excited
amongst naturalists is evinced by the invaluable Bibliography of
Brachiopoda, prepared by the same author and his friend W. H.
Dalton.[411] This monument of patient research contains over 160
quarto pages, each with the titles of from eighteen to twenty
separate papers dealing with Brachiopods, published between the
years 1606 and 1885.
Probably the first reference to Brachiopods in zoological literature
is to be found in a work entitled Aquatilium et Terrestrium aliquot
Animalium, published in the year 1606 by Prince Fabio Colonna at
Rome. This work contains the first description of a Brachiopod under
the name of Concha diphya. In a second edition, which is not so rare
in our libraries as the first, the author mentions three more species of
Brachiopods. Towards the end of the same century, Martin Lister of
Oxford, in his Historia sive Synopsis methodica Conchyliorum which
appeared in parts, described and figured a considerable number of
Brachiopods, which, under the name of Anomia, were until the
present century regarded as Molluscs, and placed in the subdivision
Pelecypoda (Lamellibranchiata).
The first satisfactory figure and description of a Terebratula were
published in the year 1766, in Pallas’ Miscellanea Zoologica, still
under the name Anomia. In 1781 O. F. Müller figured a Crania, under
the name Patella anomala, the generic name being subsequently
altered by Cuvier into Orbicula.
Bruguière in the year 1789 was the first to recognise the
relationship between Lingula and the other Brachiopods. He for the
first time saw the stalk of this genus, and compared it with that of the
stalked Barnacles, a class of animals which has been more than
once associated with our group.
Cuvier in his Mémoire sur l’Anatomie de la Lingule, 1797, gave
the first account of the internal anatomy of a Brachiopod. The same
naturalist first described the nephridia, although his mistake in
considering them lateral hearts was not rectified until the middle of
the present century, when Huxley pointed out that these structures
serve as excretory ducts for the genital products.
Duméril in 1807 proposed the somewhat unfortunate name of
Brachiopoda; and although efforts have been made by de Blainville,
who suggested Palliobranchiata, and more recently by Haeckel, who
proposed Spirobranchiata, to arrive at a name which would be both
grammatically and physiologically more correct, the older name has
maintained its position, and is now universally in use.
In 1834 and 1835 Professor Owen published the results of his
researches into the anatomy of the Brachiopoda. He investigated in
these years the structure of Waldheimia flavescens, of a species of
Lingula and of a Discina, called by him Orbicula. He regarded the
group as midway between the Pelecypoda and the Ascidians. The
structure of Lingula was further investigated by Carl Vogt, who in
1851 also supported the view that the Brachiopoda were related to
the Mollusca. But already in 1847 and 1848 Steenstrup had thrown
doubts upon this relationship, and had maintained that the Order
was more closely related to certain members of the Chaetopoda, a
view which afterwards found its ablest supporter in the American
naturalist Morse.
D’Orbigny seems to have been the first observer who drew
attention to the resemblances alleged to exist between the
Brachiopoda and the Polyzoa, and Hancock, in his masterly works
On the Anatomy of the Fresh-water Bryozoa (Polyzoa) and in his
Organisation of the Brachiopoda, dwelt on these resemblances, and
placed the Brachiopoda between the Polyzoa on the one hand and
the Ascidians on the other; a collocation which subsequently resulted
in their inclusion in the now discarded group of Molluscoidea.
In 1854 Huxley[412] published what is, with the possible exception
of Hancock’s monograph, mentioned above, the most important work
upon the anatomy of the Brachiopoda with which we are acquainted.
He corrected numerous errors of his predecessors and added many
new facts to our knowledge of the group. He was the first to describe
the true nature of the lateral hearts of Cuvier, and to describe the
true heart, afterwards so carefully figured by Hancock.
A further step was made in 1860 and 1861 by the discovery and
description of the larvae of Brachiopoda, by F. Müller and Lacaze-
Duthiers. Since that time we owe what little advance has been made
in the embryology of the group to the researches of Morse and of
Kowalevsky.
Modern methods of research—section cutting, etc.—were first
applied to the group by the Dutch naturalist, van Bemmelen,[413]
from whose admirable historical account of our knowledge of the
group many of the above facts have been gathered. These methods
have thrown considerable light upon the histology of the group, but
have not added very much to our knowledge of the structure or the
affinities of the Brachiopoda. The modern views as to the latter point
may be best discussed after some account of the anatomy of the
various genera has been given.
The Shell
The body of a Brachiopod is enclosed within a bivalve shell, but
the two halves are not, as they are in the Pelecypoda, one on each
side of the body, but occupy a different position with regard to the
main axes of the body. What this position is, has formed the subject
of a good deal of discussion. For our purpose, however, it will suffice
to distinguish the two valves by the most commonly accepted terms
of dorsal and ventral. The former is, as a rule, the smaller of the two,
and usually lies on the lower surface of the animal in life. Adopting
the orientation indicated above, the stalk by means of which the
Brachiopoda are attached to the rocks and stones, etc., upon which
they live, becomes posterior, and the broader edge of the two shells,
which are capable of being opened to some extent, is anterior.
The Body
The shell of a Brachiopod is secreted partly by the general
surface of the body which is situated at the hinder end of the shell,
and partly by the two leaf-like extensions of the body, which are
termed the dorsal and ventral mantles. These are, in fact, folds of the
body wall, and into them the body cavity and certain of its contents,
such as the liver and generative glands, etc., extend. The space
between the two folds of the mantle, which is limited behind by the
anterior wall of the body, is termed the pallial or mantle cavity. On
each side of the middle line the anterior wall of the body is produced
into two “arms,” which occupy as a rule a considerable part of the
mantle cavity. These arms may be but flattened portions of the
general body wall, which occupies a large part of what in other
genera is the mantle of the dorsal valve, as in Cistella and Argiope;
[417] or they may be outgrowths of the body wall in the form of long
processes, which are coiled and twisted in a very characteristic
manner in the various genera. In any case the cross section of the
arm shows a groove, one side of which forms a continuous lip, and
the other takes the form of a single row of tentacles, which are richly
ciliated and capable of considerable movement. The whole arm in
Rhynchonella can be protruded from the shell, as was noted years
ago by O. F. Müller, and although his statement to this effect has
often been doubted, its truth was confirmed by Professor Morse,[418]
who writes: “In the year 1872, while studying living Rhynchonella in
the St. Lawrence, I observed a specimen protrude its arms to a
distance of 4 c.m. beyond the anterior borders of the shell, a
distance nearly equalling twice the length of the shell.” The same
observer also mentions that Lingula has the power of partially
protruding its arms. In most genera the cirrhi or tentacles can alone
be protruded.
Fig. 314.—View of the left half of Cistella
(Argiope) neapolitana, which has been cut
in two by a median longitudinal incision, to
show the disposition of the organs. Partly
diagrammatic. The inorganic part of the
shell only is shown. The tubular
extensions of the mantle and the organic
outer layer are not indicated, and hence
the pores appear open.
1. The ventral valve.
2. The dorsal valve.
3. The stalk.
4. The mouth.
5. Lip which overhangs the mouth and
runs all round the tentacular arms.
6. Tentacles.
7. Ovary in dorsal valve.
8. Liver diverticula.
9. Occlusor muscle; its double origin is
shown.
10. Internal opening of left nephridium.
11. External opening of left nephridium.
12. Ventral adjuster. The line from 10
crosses the dorsal adjuster.
13. Divaricator muscle.