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Biodiversity Conservation: a Very Short

Introduction David W. Macdonald


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Biodiversity Conservation: A Very Short Introduction
VERY SHORT INTRODUCTIONS are for anyone wanting a stimulating
and accessible way into a new subject. They are written by experts, and
have been translated into more than 45 different languages.
The series began in 1995, and now covers a wide variety of topics in
every d
­ iscipline. The VSI library currently contains over 700 volumes—a
Very Short Introduction to everything from Psychology and Philosophy of
Science to American History and Relativity—and continues to grow in
every subject area.

Very Short Introductions available now:

ABOLITIONISM Richard S. Newman AMERICAN IMMIGRATION


THE ABRAHAMIC RELIGIONS David A. Gerber
Charles L. Cohen AMERICAN INTELLECTUAL
ACCOUNTING Christopher Nobes HISTORY
ADDICTION Keith Humphreys Jennifer Ratner-­Rosenhagen
ADOLESCENCE Peter K. Smith THE AMERICAN JUDICIAL SYSTEM
THEODOR W. ADORNO Charles L. Zelden
Andrew Bowie AMERICAN LEGAL HISTORY
ADVERTISING Winston Fletcher G. Edward White
AERIAL WARFARE Frank Ledwidge AMERICAN MILITARY HISTORY
AESTHETICS Bence Nanay Joseph T. Glatthaar
AFRICAN AMERICAN HISTORY AMERICAN NAVAL HISTORY
Jonathan Scott Holloway Craig L. Symonds
AFRICAN AMERICAN RELIGION AMERICAN POETRY David Caplan
Eddie S. Glaude Jr AMERICAN POLITICAL HISTORY
AFRICAN HISTORY John Parker and Donald Critchlow
Richard Rathbone AMERICAN POLITICAL PARTIES
AFRICAN POLITICS Ian Taylor AND ELECTIONS L. Sandy Maisel
AFRICAN RELIGIONS AMERICAN POLITICS
Jacob K. Olupona Richard M. Valelly
AGEING Nancy A. Pachana THE AMERICAN PRESIDENCY
AGNOSTICISM Robin Le Poidevin Charles O. Jones
AGRICULTURE Paul Brassley and THE AMERICAN REVOLUTION
Richard Soffe Robert J. Allison
ALEXANDER THE GREAT AMERICAN SLAVERY
Hugh Bowden Heather Andrea Williams
ALGEBRA Peter M. Higgins THE AMERICAN SOUTH
AMERICAN BUSINESS HISTORY Charles Reagan Wilson
Walter A. Friedman THE AMERICAN WEST
AMERICAN CULTURAL HISTORY Stephen Aron
Eric Avila AMERICAN WOMEN’S HISTORY
AMERICAN FOREIGN RELATIONS Susan Ware
Andrew Preston AMPHIBIANS T. S. Kemp
AMERICAN HISTORY Paul S. Boyer ANAESTHESIA Aidan O’Donnell
ANALYTIC PHILOSOPHY ASTROPHYSICS James Binney
Michael Beaney ATHEISM Julian Baggini
ANARCHISM Alex Prichard THE ATMOSPHERE Paul I. Palmer
ANCIENT ASSYRIA Karen Radner AUGUSTINE Henry Chadwick
ANCIENT EGYPT Ian Shaw JANE AUSTEN Tom Keymer
ANCIENT EGYPTIAN ART AND AUSTRALIA Kenneth Morgan
ARCHITECTURE Christina Riggs AUTISM Uta Frith
ANCIENT GREECE Paul Cartledge AUTOBIOGRAPHY Laura Marcus
ANCIENT GREEK AND ROMAN THE AVANT GARDE David Cottington
SCIENCE Liba Taub THE AZTECS Davíd Carrasco
THE ANCIENT NEAR EAST BABYLONIA Trevor Bryce
Amanda H. Podany BACTERIA Sebastian G. B. Amyes
ANCIENT PHILOSOPHY Julia Annas BANKING John Goddard and
ANCIENT WARFARE John O. S. Wilson
Harry Sidebottom BARTHES Jonathan Culler
ANGELS David Albert Jones THE BEATS David Sterritt
ANGLICANISM Mark Chapman BEAUTY Roger Scruton
THE ANGLO-SAXON AGE John Blair LUDWIG VAN BEETHOVEN
ANIMAL BEHAVIOUR Mark Evan Bonds
Tristram D. Wyatt BEHAVIOURAL ECONOMICS
THE ANIMAL KINGDOM Michelle Baddeley
Peter Holland BESTSELLERS John Sutherland
ANIMAL RIGHTS David DeGrazia THE BIBLE John Riches
ANSELM Thomas Williams BIBLICAL ARCHAEOLOGY
THE ANTARCTIC Klaus Dodds Eric H. Cline
ANTHROPOCENE Erle C. Ellis BIG DATA Dawn E. Holmes
ANTISEMITISM Steven Beller BIOCHEMISTRY Mark Lorch
ANXIETY Daniel Freeman and BIODIVERSITY CONSERVATION
Jason Freeman David W. Macdonald
THE APOCRYPHAL GOSPELS BIOGEOGRAPHY Mark V. Lomolino
Paul Foster BIOGRAPHY Hermione Lee
APPLIED MATHEMATICS BIOMETRICS Michael Fairhurst
Alain Goriely ELIZABETH BISHOP
THOMAS AQUINAS Fergus Kerr Jonathan F. S. Post
ARBITRATION Thomas Schultz and BLACK HOLES Katherine Blundell
Thomas Grant BLASPHEMY Yvonne Sherwood
ARCHAEOLOGY Paul Bahn BLOOD Chris Cooper
ARCHITECTURE Andrew Ballantyne THE BLUES Elijah Wald
THE ARCTIC Klaus Dodds and THE BODY Chris Shilling
Jamie Woodward THE BOHEMIANS David Weir
HANNAH ARENDT Dana Villa NIELS BOHR J. L. Heilbron
ARISTOCRACY William Doyle THE BOOK OF COMMON PRAYER
ARISTOTLE Jonathan Barnes Brian Cummings
ART HISTORY Dana Arnold THE BOOK OF MORMON
ART THEORY Cynthia Freeland Terryl Givens
ARTIFICIAL INTELLIGENCE BORDERS Alexander C. Diener and
Margaret A. Boden Joshua Hagen
ASIAN AMERICAN HISTORY THE BRAIN Michael O’Shea
Madeline Y. Hsu BRANDING Robert Jones
ASTROBIOLOGY David C. Catling THE BRICS Andrew F. Cooper
BRITISH CINEMA Charles Barr COGNITIVE BEHAVIOURAL
THE BRITISH CONSTITUTION THERAPY Freda McManus
Martin Loughlin COGNITIVE NEUROSCIENCE
THE BRITISH EMPIRE Ashley Jackson Richard Passingham
BRITISH POLITICS Tony Wright THE COLD WAR Robert J. McMahon
BUDDHA Michael Carrithers COLONIAL AMERICA Alan Taylor
BUDDHISM Damien Keown COLONIAL LATIN AMERICAN
BUDDHIST ETHICS Damien Keown LITERATURE Rolena Adorno
BYZANTIUM Peter Sarris COMBINATORICS Robin Wilson
CALVINISM Jon Balserak COMEDY Matthew Bevis
ALBERT CAMUS Oliver Gloag COMMUNISM Leslie Holmes
CANADA Donald Wright COMPARATIVE LITERATURE
CANCER Nicholas James Ben Hutchinson
CAPITALISM James Fulcher COMPETITION AND ANTITRUST
CATHOLICISM Gerald O’Collins LAW Ariel Ezrachi
CAUSATION Stephen Mumford and COMPLEXITY John H. Holland
Rani Lill Anjum THE COMPUTER Darrel Ince
THE CELL Terence Allen and COMPUTER SCIENCE
Graham Cowling Subrata Dasgupta
THE CELTS BarryCunliffe CONCENTRATION CAMPS
CHAOS Leonard Smith Dan Stone
GEOFFREY CHAUCER David Wallace CONDENSED MATTER PHYSICS
CHEMISTRY Peter Atkins Ross H. McKenzie
CHILD PSYCHOLOGY Usha Goswami CONFUCIANISM Daniel K. Gardner
CHILDREN’S LITERATURE THE CONQUISTADORS
Kimberley Reynolds Matthew Restall and
CHINESE LITERATURE Sabina Knight Felipe Fernández-Armesto
CHOICE THEORY Michael Allingham CONSCIENCE Paul Strohm
CHRISTIAN ART Beth Williamson CONSCIOUSNESS Susan Blackmore
CHRISTIAN ETHICS D. Stephen Long CONTEMPORARY ART
CHRISTIANITY Linda Woodhead Julian Stallabrass
CIRCADIAN RHYTHMS CONTEMPORARY FICTION
Russell Foster and Leon Kreitzman Robert Eaglestone
CITIZENSHIP Richard Bellamy CONTINENTAL PHILOSOPHY
CITY PLANNING Carl Abbott Simon Critchley
CIVIL ENGINEERING COPERNICUS Owen Gingerich
David Muir Wood CORAL REEFS Charles Sheppard
THE CIVIL RIGHTS MOVEMENT CORPORATE SOCIAL
Thomas C. Holt RESPONSIBILITY Jeremy Moon
CLASSICAL LITERATURE William Allan CORRUPTION Leslie Holmes
CLASSICAL MYTHOLOGY COSMOLOGY Peter Coles
Helen Morales COUNTRY MUSIC Richard Carlin
CLASSICS Mary Beard and CREATIVITY Vlad Glăveanu
John Henderson CRIME FICTION Richard Bradford
CLAUSEWITZ Michael Howard CRIMINAL JUSTICE Julian V. Roberts
CLIMATE Mark Maslin CRIMINOLOGY Tim Newburn
CLIMATE CHANGE Mark Maslin CRITICAL THEORY
CLINICAL PSYCHOLOGY Stephen Eric Bronner
Susan Llewelyn and THE CRUSADES
Katie Aafjes-van Doorn Christopher Tyerman
CRYPTOGRAPHY Fred Piper and EMOTION Dylan Evans
Sean Murphy EMPIRE Stephen Howe
CRYSTALLOGRAPHY A. M. Glazer EMPLOYMENT LAW David Cabrelli
THE CULTURAL REVOLUTION ENERGY SYSTEMS Nick Jenkins
Richard Curt Kraus ENGELS Terrell Carver
DADA AND SURREALISM ENGINEERING David Blockley
David Hopkins THE ENGLISH LANGUAGE
DANTE Peter Hainsworth and Simon Horobin
David Robey ENGLISH LITERATURE Jonathan Bate
DARWIN Jonathan Howard THE ENLIGHTENMENT
THE DEAD SEA SCROLLS John Robertson
Timothy H. Lim ENTREPRENEURSHIP Paul Westhead
DECADENCE David Weir and Mike Wright
DECOLONIZATION Dane Kennedy ENVIRONMENTAL ECONOMICS
DEMENTIA Kathleen Taylor Stephen Smith
DEMOCRACY Naomi Zack ENVIRONMENTAL ETHICS
DEMOGRAPHY Sarah Harper Robin Attfield
DEPRESSION Jan Scott and ENVIRONMENTAL LAW
Mary Jane Tacchi Elizabeth Fisher
DERRIDA Simon Glendinning ENVIRONMENTAL POLITICS
DESCARTES Tom Sorell Andrew Dobson
DESERTS Nick Middleton ENZYMES Paul Engel
DESIGN John Heskett EPICUREANISM Catherine Wilson
DEVELOPMENT Ian Goldin EPIDEMIOLOGY Rodolfo Saracci
DEVELOPMENTAL BIOLOGY ETHICS Simon Blackburn
Lewis Wolpert ETHNOMUSICOLOGY Timothy Rice
THE DEVIL Darren Oldridge THE ETRUSCANS Christopher Smith
DIASPORA Kevin Kenny EUGENICS Philippa Levine
CHARLES DICKENS Jenny Hartley THE EUROPEAN UNION
DICTIONARIES Lynda Mugglestone Simon Usherwood and John Pinder
DINOSAURS David Norman EUROPEAN UNION LAW
DIPLOMATIC HISTORY Anthony Arnull
Joseph M. Siracusa EVANGELICALISM
DOCUMENTARY FILM John G. Stackhouse Jr.
Patricia Aufderheide EVIL Luke Russell
DREAMING J. Allan Hobson EVOLUTION Brian and
DRUGS Les Iversen Deborah Charlesworth
DRUIDS Barry Cunliffe EXISTENTIALISM Thomas Flynn
DYNASTY Jeroen Duindam EXPLORATION Stewart A. Weaver
DYSLEXIA Margaret J. Snowling EXTINCTION Paul B. Wignall
EARLY MUSIC Thomas Forrest Kelly THE EYE Michael Land
THE EARTH Martin Redfern FAIRY TALE Marina Warner
EARTH SYSTEM SCIENCE Tim Lenton FAMILY LAW Jonathan Herring
ECOLOGY Jaboury Ghazoul MICHAEL FARADAY
ECONOMICS Partha Dasgupta Frank A. J. L. James
EDUCATION Gary Thomas FASCISM Kevin Passmore
EGYPTIAN MYTH Geraldine Pinch FASHION Rebecca Arnold
EIGHTEENTH‑CENTURY BRITAIN FEDERALISM Mark J. Rozell and
Paul Langford Clyde Wilcox
THE ELEMENTS Philip Ball FEMINISM Margaret Walters
FILM Michael Wood GLOBAL ECONOMIC HISTORY
FILM MUSIC Kathryn Kalinak Robert C. Allen
FILM NOIR James Naremore GLOBAL ISLAM Nile Green
FIRE Andrew C. Scott GLOBALIZATION Manfred B. Steger
THE FIRST WORLD WAR GOD John Bowker
Michael Howard GÖDEL’S THEOREM A. W. Moore
FLUID MECHANICS Eric Lauga GOETHE Ritchie Robertson
FOLK MUSIC Mark Slobin THE GOTHIC Nick Groom
FOOD John Krebs GOVERNANCE Mark Bevir
FORENSIC PSYCHOLOGY GRAVITY Timothy Clifton
David Canter THE GREAT DEPRESSION AND
FORENSIC SCIENCE Jim Fraser THE NEW DEAL Eric Rauchway
FORESTS Jaboury Ghazoul HABEAS CORPUS Amanda L. Tyler
FOSSILS Keith Thomson HABERMAS James Gordon Finlayson
FOUCAULT Gary Gutting THE HABSBURG EMPIRE
THE FOUNDING FATHERS Martyn Rady
R. B. Bernstein HAPPINESS Daniel M. Haybron
FRACTALS Kenneth Falconer THE HARLEM RENAISSANCE
FREE SPEECH Nigel Warburton Cheryl A. Wall
FREE WILL Thomas Pink THE HEBREW BIBLE AS
FREEMASONRY Andreas Önnerfors LITERATURE Tod Linafelt
FRENCH LITERATURE John D. Lyons HEGEL Peter Singer
FRENCH PHILOSOPHY HEIDEGGER Michael Inwood
Stephen Gaukroger and Knox Peden THE HELLENISTIC AGE
THE FRENCH REVOLUTION Peter Thonemann
William Doyle HEREDITY John Waller
FREUD Anthony Storr HERMENEUTICS Jens Zimmermann
FUNDAMENTALISM Malise Ruthven HERODOTUS Jennifer T. Roberts
FUNGI Nicholas P. Money HIEROGLYPHS Penelope Wilson
THE FUTURE Jennifer M. Gidley HINDUISM Kim Knott
GALAXIES John Gribbin HISTORY John H. Arnold
GALILEO Stillman Drake THE HISTORY OF ASTRONOMY
GAME THEORY Ken Binmore Michael Hoskin
GANDHI Bhikhu Parekh THE HISTORY OF CHEMISTRY
GARDEN HISTORY Gordon Campbell William H. Brock
GENES Jonathan Slack THE HISTORY OF CHILDHOOD
GENIUS Andrew Robinson James Marten
GENOMICS John Archibald THE HISTORY OF CINEMA
GEOGRAPHY John Matthews and Geoffrey Nowell-Smith
David Herbert THE HISTORY OF COMPUTING
GEOLOGY Jan Zalasiewicz Doron Swade
GEOMETRY Maciej Dunajski THE HISTORY OF EMOTIONS
GEOPHYSICS William Lowrie Thomas Dixon
GEOPOLITICS Klaus Dodds THE HISTORY OF LIFE Michael Benton
GERMAN LITERATURE Nicholas Boyle THE HISTORY OF MATHEMATICS
GERMAN PHILOSOPHY Jacqueline Stedall
Andrew Bowie THE HISTORY OF MEDICINE
THE GHETTO Bryan Cheyette William Bynum
GLACIATION David J. A. Evans THE HISTORY OF PHYSICS
GLOBAL CATASTROPHES Bill McGuire J. L. Heilbron
THE HISTORY OF POLITICAL INTERNATIONAL SECURITY
THOUGHT Richard Whatmore Christopher S. Browning
THE HISTORY OF TIME INSECTS Simon Leather
Leofranc Holford‑Strevens IRAN Ali M. Ansari
HIV AND AIDS Alan Whiteside ISLAM Malise Ruthven
HOBBES Richard Tuck ISLAMIC HISTORY Adam Silverstein
HOLLYWOOD Peter Decherney ISLAMIC LAW Mashood A. Baderin
THE HOLY ROMAN EMPIRE ISOTOPES Rob Ellam
Joachim Whaley ITALIAN LITERATURE
HOME Michael Allen Fox Peter Hainsworth and David Robey
HOMER Barbara Graziosi HENRY JAMES Susan L. Mizruchi
HORMONES Martin Luck JAPANESE LITERATURE Alan Tansman
HORROR Darryl Jones JESUS Richard Bauckham
HUMAN ANATOMY JEWISH HISTORY David N. Myers
Leslie Klenerman JEWISH LITERATURE Ilan Stavans
HUMAN EVOLUTION Bernard Wood JOURNALISM Ian Hargreaves
HUMAN PHYSIOLOGY JAMES JOYCE Colin MacCabe
Jamie A. Davies JUDAISM Norman Solomon
HUMAN RESOURCE JUNG Anthony Stevens
MANAGEMENT Adrian Wilkinson THE JURY Renée Lettow Lerner
HUMAN RIGHTS Andrew Clapham KABBALAH Joseph Dan
HUMANISM Stephen Law KAFKA Ritchie Robertson
HUME James A. Harris KANT Roger Scruton
HUMOUR Noël Carroll KEYNES Robert Skidelsky
IBN SĪNĀ (AVICENNA) Peter Adamson KIERKEGAARD Patrick Gardiner
THE ICE AGE Jamie Woodward KNOWLEDGE Jennifer Nagel
IDENTITY Florian Coulmas THE KORAN Michael Cook
IDEOLOGY Michael Freeden KOREA Michael J. Seth
THE IMMUNE SYSTEM LAKES Warwick F. Vincent
Paul Klenerman LANDSCAPE ARCHITECTURE
INDIAN CINEMA Ian H. Thompson
Ashish Rajadhyaksha LANDSCAPES AND
INDIAN PHILOSOPHY Sue Hamilton GEOMORPHOLOGY
THE INDUSTRIAL REVOLUTION Andrew Goudie and Heather Viles
Robert C. Allen LANGUAGES Stephen R. Anderson
INFECTIOUS DISEASE Marta L. Wayne LATE ANTIQUITY Gillian Clark
and Benjamin M. Bolker LAW Raymond Wacks
INFINITY Ian Stewart THE LAWS OF THERMODYNAMICS
INFORMATION Luciano Floridi Peter Atkins
INNOVATION Mark Dodgson and LEADERSHIP Keith Grint
David Gann LEARNING Mark Haselgrove
INTELLECTUAL PROPERTY LEIBNIZ Maria Rosa Antognazza
Siva Vaidhyanathan C. S. LEWIS James Como
INTELLIGENCE Ian J. Deary LIBERALISM Michael Freeden
INTERNATIONAL LAW LIGHT Ian Walmsley
Vaughan Lowe LINCOLN Allen C. Guelzo
INTERNATIONAL MIGRATION LINGUISTICS Peter Matthews
Khalid Koser LITERARY THEORY Jonathan Culler
INTERNATIONAL RELATIONS LOCKE John Dunn
Christian Reus-Smit LOGIC Graham Priest
LOVE Ronald de Sousa MILITARY STRATEGY
MARTIN LUTHER Scott H. Hendrix Antulio J. Echevarria II
MACHIAVELLI Quentin Skinner JOHN STUART MILL Gregory Claeys
MADNESS Andrew Scull MINERALS David Vaughan
MAGIC Owen Davies MIRACLES Yujin Nagasawa
MAGNA CARTA Nicholas Vincent MODERN ARCHITECTURE
MAGNETISM Stephen Blundell Adam Sharr
MALTHUS Donald Winch MODERN ART David Cottington
MAMMALS T. S. Kemp MODERN BRAZIL Anthony W. Pereira
MANAGEMENT John Hendry MODERN CHINA Rana Mitter
NELSON MANDELA Elleke Boehmer MODERN DRAMA
MAO Delia Davin Kirsten E. Shepherd-Barr
MARINE BIOLOGY Philip V. Mladenov MODERN FRANCE
MARKETING Vanessa R. Schwartz
Kenneth Le Meunier-FitzHugh MODERN INDIA Craig Jeffrey
THE MARQUIS DE SADE John Phillips MODERN IRELAND Senia Pašeta
MARTYRDOM Jolyon Mitchell MODERN ITALY Anna Cento Bull
MARX Peter Singer MODERN JAPAN
MATERIALS Christopher Hall Christopher Goto-Jones
MATHEMATICAL ANALYSIS MODERN LATIN AMERICAN
Richard Earl LITERATURE
MATHEMATICAL FINANCE Roberto González Echevarría
Mark H. A. Davis MODERN WAR Richard English
MATHEMATICS Timothy Gowers MODERNISM Christopher Butler
MATTER Geoff Cottrell MOLECULAR BIOLOGY Aysha Divan
THE MAYA Matthew Restall and and Janice A. Royds
Amara Solari MOLECULES Philip Ball
THE MEANING OF LIFE MONASTICISM Stephen J. Davis
Terry Eagleton THE MONGOLS Morris Rossabi
MEASUREMENT David Hand MONTAIGNE William M. Hamlin
MEDICAL ETHICS Michael Dunn and MOONS David A. Rothery
Tony Hope MORMONISM
MEDICAL LAW Charles Foster Richard Lyman Bushman
MEDIEVAL BRITAIN John Gillingham MOUNTAINS Martin F. Price
and Ralph A. Griffiths MUHAMMAD Jonathan A. C. Brown
MEDIEVAL LITERATURE MULTICULTURALISM Ali Rattansi
Elaine Treharne MULTILINGUALISM John C. Maher
MEDIEVAL PHILOSOPHY MUSIC Nicholas Cook
John Marenbon MUSIC AND TECHNOLOGY
MEMORY Jonathan K. Foster Mark Katz
METAPHYSICS Stephen Mumford MYTH Robert A. Segal
METHODISM William J. Abraham NANOTECHNOLOGY Philip Moriarty
THE MEXICAN REVOLUTION NAPOLEON David A. Bell
Alan Knight THE NAPOLEONIC WARS
MICROBIOLOGY Nicholas P. Money Mike Rapport
MICROBIOMES Angela E. Douglas NATIONALISM Steven Grosby
MICROECONOMICS Avinash Dixit NATIVE AMERICAN LITERATURE
MICROSCOPY Terence Allen Sean Teuton
THE MIDDLE AGES Miri Rubin NAVIGATION Jim Bennett
MILITARY JUSTICE Eugene R. Fidell NAZI GERMANY Jane Caplan
NEGOTIATION Carrie Menkel-Meadow PAUL E. P. Sanders
NEOLIBERALISM Manfred B. Steger IVAN PAVLOV Daniel P. Todes
and Ravi K. Roy PEACE Oliver P. Richmond
NETWORKS Guido Caldarelli and PENTECOSTALISM William K. Kay
Michele Catanzaro PERCEPTION Brian Rogers
THE NEW TESTAMENT THE PERIODIC TABLE Eric R. Scerri
Luke Timothy Johnson PHILOSOPHICAL METHOD
THE NEW TESTAMENT AS Timothy Williamson
LITERATURE Kyle Keefer PHILOSOPHY Edward Craig
NEWTON Robert Iliffe PHILOSOPHY IN THE ISLAMIC
NIETZSCHE Michael Tanner WORLD Peter Adamson
NINETEENTH‑CENTURY BRITAIN PHILOSOPHY OF BIOLOGY
Christopher Harvie and Samir Okasha
H. C. G. Matthew PHILOSOPHY OF LAW
THE NORMAN CONQUEST Raymond Wacks
George Garnett PHILOSOPHY OF MIND
NORTH AMERICAN INDIANS Barbara Gail Montero
Theda Perdue and Michael D. Green PHILOSOPHY OF PHYSICS
NORTHERN IRELAND David Wallace
Marc Mulholland PHILOSOPHY OF SCIENCE
NOTHING Frank Close Samir Okasha
NUCLEAR PHYSICS Frank Close PHILOSOPHY OF RELIGION
NUCLEAR POWER Maxwell Irvine Tim Bayne
NUCLEAR WEAPONS PHOTOGRAPHY Steve Edwards
Joseph M. Siracusa PHYSICAL CHEMISTRY Peter Atkins
NUMBER THEORY Robin Wilson PHYSICS Sidney Perkowitz
NUMBERS Peter M. Higgins PILGRIMAGE Ian Reader
NUTRITION David A. Bender PLAGUE Paul Slack
OBJECTIVITY Stephen Gaukroger PLANETARY SYSTEMS
OBSERVATIONAL ASTRONOMY Raymond T. Pierrehumbert
Geoff Cottrell PLANETS David A. Rothery
OCEANS Dorrik Stow PLANTS Timothy Walker
THE OLD TESTAMENT PLATE TECTONICS Peter Molnar
Michael D. Coogan PLATO Julia Annas
THE ORCHESTRA D. Kern Holoman POETRY Bernard O’Donoghue
ORGANIC CHEMISTRY POLITICAL PHILOSOPHY David Miller
Graham Patrick POLITICS Kenneth Minogue
ORGANIZATIONS Mary Jo Hatch POLYGAMY Sarah M. S. Pearsall
ORGANIZED CRIME POPULISM Cas Mudde and
Georgios A. Antonopoulos and Cristóbal Rovira Kaltwasser
Georgios Papanicolaou POSTCOLONIALISM
ORTHODOX CHRISTIANITY Robert J. C. Young
A. Edward Siecienski POSTMODERNISM Christopher Butler
OVID Llewelyn Morgan POSTSTRUCTURALISM
PAGANISM Owen Davies Catherine Belsey
PAKISTAN Pippa Virdee POVERTY Philip N. Jefferson
THE PALESTINIAN-ISRAELI PREHISTORY Chris Gosden
CONFLICT Martin Bunton PRESOCRATIC PHILOSOPHY
PANDEMICS Christian W. McMillen Catherine Osborne
PARTICLE PHYSICS Frank Close PRIVACY Raymond Wacks
PROBABILITY John Haigh ROMANTICISM Michael Ferber
PROGRESSIVISM Walter Nugent ROUSSEAU Robert Wokler
PROHIBITION W. J. Rorabaugh RUSSELL A. C. Grayling
PROJECTS Andrew Davies THE RUSSIAN ECONOMY
PROTESTANTISM Mark A. Noll Richard Connolly
PSEUDOSCIENCE Michael D. Gordin. RUSSIAN HISTORY Geoffrey Hosking
PSYCHIATRY Tom Burns RUSSIAN LITERATURE Catriona Kelly
PSYCHOANALYSIS Daniel Pick THE RUSSIAN REVOLUTION
PSYCHOLOGY Gillian Butler and S. A. Smith
Freda McManus SAINTS Simon Yarrow
PSYCHOLOGY OF MUSIC SAMURAI Michael Wert
Elizabeth Hellmuth Margulis SAVANNAS Peter A. Furley
PSYCHOPATHY Essi Viding SCEPTICISM Duncan Pritchard
PSYCHOTHERAPY Tom Burns and SCHIZOPHRENIA Chris Frith and
Eva Burns-Lundgren Eve Johnstone
PUBLIC ADMINISTRATION SCHOPENHAUER
Stella Z. Theodoulou and Ravi K. Roy Christopher Janaway
PUBLIC HEALTH Virginia Berridge SCIENCE AND RELIGION
PURITANISM Francis J. Bremer Thomas Dixon and Adam R. Shapiro
THE QUAKERS Pink Dandelion SCIENCE FICTION David Seed
QUANTUM THEORY THE SCIENTIFIC REVOLUTION
John Polkinghorne Lawrence M. Principe
RACISM Ali Rattansi SCOTLAND Rab Houston
RADIOACTIVITY Claudio Tuniz SECULARISM Andrew Copson
RASTAFARI Ennis B. Edmonds SEXUAL SELECTION Marlene Zuk and
READING Belinda Jack Leigh W. Simmons
THE REAGAN REVOLUTION Gil Troy SEXUALITY Véronique Mottier
REALITY Jan Westerhoff WILLIAM SHAKESPEARE
RECONSTRUCTION Allen C. Guelzo Stanley Wells
THE REFORMATION Peter Marshall SHAKESPEARE’S COMEDIES
REFUGEES Gil Loescher Bart van Es
RELATIVITY Russell Stannard SHAKESPEARE’S SONNETS AND
RELIGION Thomas A. Tweed POEMS Jonathan F. S. Post
RELIGION IN AMERICA Timothy Beal SHAKESPEARE’S TRAGEDIES
THE RENAISSANCE Jerry Brotton Stanley Wells
RENAISSANCE ART GEORGE BERNARD SHAW
Geraldine A. Johnson Christopher Wixson
RENEWABLE ENERGY Nick Jelley MARY SHELLEY Charlotte Gordon
REPTILES T. S. Kemp THE SHORT STORY Andrew Kahn
REVOLUTIONS Jack A. Goldstone SIKHISM Eleanor Nesbitt
RHETORIC Richard Toye SILENT FILM Donna Kornhaber
RISK Baruch Fischhoff and John Kadvany THE SILK ROAD James A. Millward
RITUAL Barry Stephenson SLANG Jonathon Green
RIVERS Nick Middleton SLEEP Steven W. Lockley and
ROBOTICS Alan Winfield Russell G. Foster
ROCKS Jan Zalasiewicz SMELL Matthew Cobb
ROMAN BRITAIN Peter Salway ADAM SMITH Christopher J. Berry
THE ROMAN EMPIRE Christopher Kelly SOCIAL AND CULTURAL
THE ROMAN REPUBLIC ANTHROPOLOGY
David M. Gwynn John Monaghan and Peter Just
SOCIAL PSYCHOLOGY Richard J. Crisp TRAGEDY Adrian Poole
SOCIAL WORK Sally Holland and TRANSLATION Matthew Reynolds
Jonathan Scourfield THE TREATY OF VERSAILLES
SOCIALISM Michael Newman Michael S. Neiberg
SOCIOLINGUISTICS John Edwards TRIGONOMETRY
SOCIOLOGY Steve Bruce Glen Van Brummelen
SOCRATES C. C. W. Taylor THE TROJAN WAR Eric H. Cline
SOFT MATTER Tom McLeish TRUST Katherine Hawley
SOUND Mike Goldsmith THE TUDORS John Guy
SOUTHEAST ASIA James R. Rush TWENTIETH‑CENTURY BRITAIN
THE SOVIET UNION Stephen Lovell Kenneth O. Morgan
THE SPANISH CIVIL WAR TYPOGRAPHY Paul Luna
Helen Graham THE UNITED NATIONS
SPANISH LITERATURE Jo Labanyi Jussi M. Hanhimäki
THE SPARTANS Andrew J. Bayliss UNIVERSITIES AND COLLEGES
SPINOZA Roger Scruton David Palfreyman and Paul Temple
SPIRITUALITY Philip Sheldrake THE U.S. CIVIL WAR Louis P. Masur
SPORT Mike Cronin THE U.S. CONGRESS Donald A. Ritchie
STARS Andrew King THE U.S. CONSTITUTION
STATISTICS David J. Hand David J. Bodenhamer
STEM CELLS Jonathan Slack THE U.S. SUPREME COURT
STOICISM Brad Inwood Linda Greenhouse
STRUCTURAL ENGINEERING UTILITARIANISM
David Blockley Katarzyna de Lazari-Radek and
STUART BRITAIN John Morrill Peter Singer
SUBURBS Carl Abbott UTOPIANISM Lyman Tower Sargent
THE SUN Philip Judge VATICAN II Shaun Blanchard and
SUPERCONDUCTIVITY Stephen Bullivant
Stephen Blundell VETERINARY SCIENCE James Yeates
SUPERSTITION Stuart Vyse THE VIKINGS Julian D. Richards
SYMMETRY Ian Stewart VIOLENCE Philip Dwyer
SYNAESTHESIA Julia Simner THE VIRGIN MARY
SYNTHETIC BIOLOGY Jamie A. Davies Mary Joan Winn Leith
SYSTEMS BIOLOGY Eberhard O. Voit THE VIRTUES Craig A. Boyd and
TAXATION Stephen Smith Kevin Timpe
TEETH Peter S. Ungar VIRUSES Dorothy H. Crawford
TERRORISM Charles Townshend VOLCANOES Michael J. Branney and
THEATRE Marvin Carlson Jan Zalasiewicz
THEOLOGY David F. Ford VOLTAIRE Nicholas Cronk
THINKING AND REASONING WAR AND RELIGION Jolyon Mitchell
Jonathan St B. T. Evans and Joshua Rey
THOUGHT Tim Bayne WAR AND TECHNOLOGY
TIBETAN BUDDHISM Alex Roland
Matthew T. Kapstein WATER John Finney
TIDES David George Bowers and WAVES Mike Goldsmith
Emyr Martyn Roberts WEATHER Storm Dunlop
TIME Jenann Ismael THE WELFARE STATE David Garland
TOCQUEVILLE Harvey C. Mansfield WITCHCRAFT Malcolm Gaskill
LEO TOLSTOY Liza Knapp WITTGENSTEIN A. C. Grayling
TOPOLOGY Richard Earl WORK Stephen Fineman
WORLD MUSIC Philip Bohlman WRITING AND SCRIPT
WORLD MYTHOLOGY David Leeming Andrew Robinson
THE WORLD TRADE ZIONISM Michael Stanislawski
ORGANIZATION Amrita Narlikar ÉMILE ZOLA
WORLD WAR II Gerhard L. Weinberg Brian Nelson

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David W. Macdonald

BIODIVERSITY
CONSERVATION
A Very Short Introduction
Great Clarendon Street, Oxford, OX2 6DP,
United Kingdom
Oxford University Press is a department of the University of Oxford.
It furthers the University’s objective of excellence in research, scholarship,
and education by publishing worldwide. Oxford is a registered trade mark of
Oxford University Press in the UK and in certain other countries
© David W. Macdonald 2023
The moral rights of the author have been asserted
All rights reserved. No part of this publication may be reproduced, stored in
a retrieval system, or transmitted, in any form or by any means, without the
prior permission in writing of Oxford University Press, or as expressly permitted
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above should be sent to the Rights Department, Oxford University Press, at the
address above
You must not circulate this work in any other form
and you must impose this same condition on any acquirer
Published in the United States of America by Oxford University Press
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Library of Congress Control Number: 2023930670
ISBN 978–0–19–959227–2
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Links to third party websites are provided by Oxford in good faith and
for information only. Oxford disclaims any responsibility for the materials
contained in any third party website referenced in this work.
For Dawn
‘It seems to me that the natural world is the greatest source of excitement;
The greatest source of visual beauty; the greatest source of intellectual interest.
It is the greatest source of so much in life that makes life worth living.’
Sir David Attenborough
Contents

Acknowledgements xix

List of illustrations xxiii

Part 1: Setting the scene

1 What is biodiversity, and why does it matter? 3

2 What’s the problem? 19

3 What is the purpose of biodiversity conservation? 39

Part 2: The Big Five

4 Invasive species 59

5 The trade in wildlife 77

6 Wildlife disease 92

7 Human–wildlife conflict 106

8 Climate change 121

Part 3: The way ahead

9 Who pays, and how? 137

10 What next? 157


Species list 179

References 185

Further reading 198

Index 201
Biodiversity Conservation

xviii
Acknowledgements

Almost everything worth saying seems to have been said already


by Mark Twain: ‘I didn’t have time to write a short letter, so
I wrote a long one instead.’ Having been tasked to write a very
short book about a very big topic, I can confirm it takes time.
Further, as readers will learn within, biodiversity is so diverse that
an egalitarian book, especially a very short one, could devote less
than a punctuation mark to each species, so inevitably I have
fallen prey to bias: my examples draw disproportionately on
vertebrates, especially mammals, for which I don’t much
apologize—­they illustrate well the problems and principles of
biodiversity conservation, which is what matters. I also draw more
than randomly on the work of the Wildlife Conservation Research
Unit (WildCRU) at the University of Oxford for the good reasons
that, as its founding Director, I know it best, and its mission ‘to
achieve practical solutions to conservation problems through
original scientific research’ goes to the heart of the topic of this
book. Even the examples I have mentioned generate a
bibliography too vast to include: for those hungry for the full
story the references are given at www.wildcru.org/VSI_
Biodiversity_Conservation/.

Above all I am grateful to my wife Dr Dawn Burnham—­who has


read, shaken, and stirred every word—­for her professional insights
and personal support. I also thank particularly my friend and
colleague Dr Christopher O’Kane for his unfailing effectiveness
(made possible by Peter and Gyongyver Kadas). In addition to
Dawn and Christopher, the whole book was fruitfully criticized by
risen conservation stars Drs Darragh Hare and Laura Perry, and
by rising representatives of the next generation, Chrishen Gomez
and Claire Marr, to whom I’m limitlessly grateful, along with John
Salmon who also read the whole manuscript as my representative
normal reader (if that’s an apt description of a top surgeon). To
these stalwarts, I add thanks to specialist critics of individual
chapters, Dr Luca Chiaverini, Dr Alayne Cotterill, Professor Chris
Dye, Dr Kim Jacobsen, Dr Kerry Kilshaw, Professor Andrew
Loveridge, Dr Ewan Macdonald, Dr Silvio Marchini, Dr Axel
Moehrenschlager, Dr Chris Newman, Dr Ugyen Penjor, Professor
Alex Teytelboym, Dr Peter Tyrrell, and others of the extraordinary
WildCRU diaspora (all of whom, but none more than me, have
Biodiversity Conservation

been enabled by the remarkable support of Tom and Dafna


Kaplan).

I am so pleased to be years late in delivering this book. Had I been


punctual I’d have pre-­empted biodiversity conservation’s
formative growth spurt that has left the subject almost
unrecognizably metamorphosed from its persona just a few years
ago (although I haven’t forgotten the help of Drs Ros Shaw and
Ruth Feber with an earlier instar). My relief at not writing the
book while the topic was still larval is tempered only by my apology
for the frustration I caused to OUP’s Latha Menon, the series’
mother. I am particularly grateful to her for providing me with
comfort that the era of editors that edit is not entirely past.

Readers will see, in these pages, just how radical have been the
accelerating impacts on biodiversity and biodiversity conservation,
within scarcely a professional lifetime. I began my career in
Borneo, and so too did my son, Ewan, but between our
generations a third of the forest had gone. A vocation once
rewarded by reading a pawprint in the mud, or by staring deeply
into the eyes of another species, or even getting sufficiently into its
xx
skin to know what it would do next, and to understand why,
arrives now at the shuddering intersection of teragrams of carbon,
global pollutants, viral genomes, market forces, property rights,
and geopolitics, with its meta-­analyses and models. There is both
thrill and chill in this new reality. Twenty years ago the present
would have been barely imaginable, so writing now, on her first
birthday, I can only wonder how things will seem when my
granddaughter, Hannah, is old enough to read this book.
Wherever biodiversity conservation’s journey from groundedness
to geopolitics may go next, it is perilously urgent.

Acknowledgements

xxi
List of illustrations

1 Popa langur 4 Clements, G. R., and Hearn, A. J.,


©Thaung Win 2020. Predicting biodiversity richness
in rapidly changing landscapes:
climate, low human pressure or
2 The major biomes of protection as salvation? Biodiversity
the world 12 and Conservation, 29(14),
From <http://evans9j.blogspot.com/ pp. 4035–57, with permission
2015/02/biomes-­our-­first-­activity-­
will-­be-­to.html>, with permission 5 African lion density across (a)
recent historical (1960–1970s)
3 Biodiversity hotspots situated lion distribution; and (b)
within high (black) biodiversity extant range showing lion
areas 15 population densities 21
From Myers, N., Mittermeier, From Loveridge, A. J., Sousa, L. L.,
R. A., Mittermeier, C. G., Da Fonseca, Cushman, S., Kaszta, Ż., and
G. A. and Kent, J., 2000. Biodiversity Macdonald, D. W. (2022). Where
hotspots for conservation priorities. have all the lions gone? Establishing
Nature, 403(6772), pp. 853–8, with realistic baselines to assess decline
permission and recovery of African lions.
Diversity and Distributions, 28,
4 (a) Predictive map of species 2388–2402. https://doi.org/10.1111/
ddi.13637, with permission
richness in mainland
South-­East Asia, Borneo, and
6 Cecil the lion 22
Sumatra; (b) the locations of
Andrew Loveridge
the same hotspots depicted
relative to the locations of
7 Main threats to globally
current protected areas 17
threatened birds 25
Adapted from Macdonald, D. W.,
Chiaverini, L., Bothwell, H. M., From Birdlife International 2016.
Kaszta, Ż., Ash, E., Bolongon, G., Can, http://datazone.birdlife.org/sowb/
Ö. E., Campos-­Arceiz, A., Channa, P., casestudy/a-­range-­of-­threats-­drives-­
declines-­in-­bird-­populations, with communications, 7(1), pp. 1–9,
permission with permission

8 Lord Howe Island stick 14 Regions with contiguous


insect or tree lobster 52 countries where an invasive
species spreads from the
9 Orchid Dracula country of first establishment,
mendozae 52 where it has no impact, into
countries of subsequent
10 Geographical distribution of invasion, where it has an
the costs due to alien species impact 75
for 1970–2017 62 From Faulkner, K. T., Robertson,
From Diagne, C., Leroy, B., Vaissière, M. P., and Wilson, J. R., 2020.
A. C., Gozlan, R. E., Roiz, D., Jarić, I., Stronger regional biosecurity is
Salles, J. M., Bradshaw, C. J., and essential to prevent hundreds of
Courchamp, F., 2021. High and rising harmful biological invasions. Global
economic costs of biological invasions Change Biology, 26(4), pp. 2449–62,
worldwide. Nature, 592(7855), with permission
pp. 571–6, with permission
15 The unacceptable face of
Biodiversity Conservation

11 The 10 costliest taxa for bushmeat 78


cumulative damage and
management costs (2017 16 Causes and effects of illegal or
prices) 1970–2017 63 unsustainable wildlife trade
From Diagne, C., Leroy, B., Vaissière, on species, ecosystems,
A. C., Gozlan, R. E., Roiz, D., Jarić, I., and society 79
Salles, J. M., Bradshaw, C. J., and
From Cardoso, P., Amponsah-­
Courchamp, F., 2021. High and rising
Mensah, K., Barreiros, J. P., Bouhuys,
economic costs of biological invasions
J., Cheung, H., Davies, A., Kumschick,
worldwide. Nature, 592(7855),
S., Longhorn, S. J., Martínez-­Munoz,
pp. 571–6, with permission
C. A., Morcatty, T. Q., and Peters, G.,
2021. Scientists’ warning to
12 Parakeets peek out of holes in humanity on illegal or unsustainable
a tree in Richmond Park 64 wildlife trade. Biological
Conservation, 263, p. 109341,
Bruno Guerreiro/Getty
with permission

13 Global invasion threat for


17 The spread of African Swine
the 21st century 73
Fever threatens South-­East
From Early, R., Bradley, B. A., Dukes,
Asia’s 11 wild pig species 97
J. S., Lawler, J. J., Olden, J. D.,
Blumenthal, D. M., Gonzalez, P., From Luskin, M. S., Meijaard, E.,
Grosholz, E. D., Ibañez, I., Miller, Surya, S., Walzer, C., and Linkie, M.,
L. P., and Sorte, C. J., 2016. Global 2021. African Swine Fever threatens
threats from invasive alien species in Southeast Asia’s 11 endemic wild pig
the twenty-­first century and national species. Conservation Letters, 14(3),
response capacities. Nature p.e12784, with permission

xxiv
18 Incidence rate of bovine 22 Conservation problems, and
tuberculosis in cattle within their solutions, can occur
and outside 30 badger cull anywhere along a continuum
areas of the High Risk Area of of species recovery 117
England, during badger cull From Macdonald, D. W. and
years (September to August) Sillero-­Zubiri, C., 2004.
Conservation: from theory to
2013/14–2018/19 101
practice, without bluster. In
From Langton, T. E., Jones, M. W., Macdonald, D. W. and Sillero-­Zubiri,
and McGill, I., 2022. Analysis of the C., eds., The Biology and
impact of badger culling on bovine Conservation of Wild Canids. Oxford
tuberculosis in cattle in the high-­risk University Press, pp. 353–72, with
area of England, 2009–2020. permission
Veterinary Record, 190(6), p.e1384,
with permission
23 Global average sea level has
risen by about 16 cm
19 Integrated wildlife monitoring
(6 inches) since the late 19th
as the combination of
century, and faster
population monitoring,
recently 124
passive (scanning), and active
From Shum, C. K. and Kuo, C. Y.,
(targeted) disease
2010. Observation and geophysical
surveillance 104

List of illustrations
causes of present-­day sea-­level rise.
From Cardoso, B., García-­Bocanegra, In Climate Change and Food Security
I., Acevedo, P., Cáceres, G., Alves, in South Asia. Springer, pp. 85–104,
P. C,, and Gortázar, C., 2021. Stepping with permission
up from wildlife disease surveillance
to integrated wildlife monitoring in
24 The doughnut of social and
Europe. Research in Veterinary
Science, 144, pp. 149–56, with planetary boundaries 140
permission From Raworth, K., 2017. Why it’s
time for Doughnut Economics. IPPR
Progressive Review, 24(3),
20 The ground inside (treated)
pp. 216–22, with permission
and outside (untreated) the
bomas, and the resulting
25 Under anthropocentric
difference in number of maize
economics the intended uses
cobs and cob length 112
of (and impacts on) natural
capital are constrained to
21 Common approaches used to
preclude overexploitation that
mitigate human–wildlife
diminishes human well-­being,
conflict and promote
and uses of human capital are
human–wildlife
constrained to preclude unfair
coexistence 116
or undignified treatment of
From Nyhus, P. J., 2016. Human–
other humans 142
wildlife conflict and coexistence.
Annual Review of Environment and From Vucetich, J. A., Damania, R.,
Resources, 41, pp. 143–71, with Cushman, S. A., Macdonald, E. A.,
permission Burnham, D., Offer-­Westort, T.,

xxv
Bruskotter, J. T., Feltz, A., Eeden, From TNC (<https://www.nature.
L. V., and Macdonald, D. W., 2021. org/en-­us/newsroom/blue-­bonds-­
A minimally nonanthropocentric belize-­conserve-­thirty-­percent-­of-­
economics: what is it, is it necessary, ocean-­through-­debt-­conversion/>)
and can it avert the biodiversity with permission
crisis? BioScience, 71(8), pp. 861–73,
with permission
29 Success/failure of
conservation translocation
26 ‘Living with Tigers’ Project
according to major taxa 160
conceptualized pathway for
From Soorae, P. S. ed., 2021. Global
interventions, intended Conservation Translocation
outcomes, and impacts 149 Perspectives, 2021: Case Studies from
From Fitzmaurice, Amy, Liedekerke, Around the Globe. IUCN SSC
V., Carter, Neil, Trout, E., Parker, B., Conservation Translocation Specialist
Manandhar, Prajwol, Dickson, G., Group, Environment Agency, with
Senn, H., Alibhai, S., Chaudhary, T., permission
Chapagain, P., Poudel, Prabin, Thapa,
Shyam, Zimmermann, A., 30 Global distribution of
Macdonald, D., Subedi, Bishnu,
critically endangered
Paudel, Sakuntala, Thapa, Sima, and
Chaudhary, Gautam (2022). Impact megafauna 163
Biodiversity Conservation

evaluation of the Living with Tigers From Farhadinia, M. S., Johnson,


Project: Do human–felid coexistence P. J., Zimmermann, A., McGowan,
interventions benefit both wildlife P. J., Meijaard, E., Stanley-­Price, M.,
and local people? Final Report 2021, and Macdonald, D. W., 2020. Ex situ
with permission management as insurance against
extinction of mammalian megafauna
in an uncertain world. Conservation
27 Cost per head of livestock
Biology, 34(4), pp. 988–96, with
incurred by attempted permission
protection against predators
by lethal and non-­lethal 31 Wild boar sow with
means 153 piglets 168
Based on McManus, J. S., Dickman, Philip Mugridge/Alamy
A. J., Gaynor, D., Smuts, B. H., and
Macdonald, D. W., 2015. Dead or
alive? Comparing costs and 32 Remaining ranges of Persian,
benefits of lethal and non-­lethal Arabian, Indochinese, and
human–wildlife conflict mitigation Amur leopard subspecies, and
on livestock farms. Oryx, 49(4),
the locations of
pp. 687–95, with permission
borderlands 172
From Farhadinia, M. S., Rostro-­García,
28 Flow diagram of The Nature
S., Feng, L., Kamler, J. F., Spalton, A.,
Conservancy (TNC) ‘Blue Shevtsova, E., Khorozyan, I.,
Bonds for Ocean Al-­Duais, M., Ge, J., and Macdonald,
Conservation’ D. W., 2021. Big cats in borderlands:
challenges and implications for
programme 156
transboundary conservation of

xxvi
Asian leopards. Oryx, 55(3), 34 Nature-­based Solutions as an
pp. 452–60, with permission
umbrella term for ecosystem-­
related approaches 176
33 The Conservation Quartet, From Cohen-­Shacham, E., Walters,
which, in 1986, I designed to G., Janzen, C., and Maginnis, S., 2016.
conceptualize the four Nature-­based solutions to address
interacting components of the global societal challenges. IUCN:
Gland, Switzerland, 97, pp. 2016–36,
mission of the WildCRU 174 with permission.

List of illustrations

xxvii
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orange-blossoms, and covered at the instant it is dished, with
strawberry, apple, or any other clear jelly.
A COMMON RICE PUDDING.

Throw six ounces of rice into plenty of cold water, and boil it gently
from eight to ten minutes; drain it well in a sieve or strainer, and put it
into a clean saucepan with a quart of milk; let it stew until tender,
sweeten it with three ounces of sugar, stir to it, gradually, three large,
or four small eggs, beaten and strained; add grated nutmeg, lemon
rind, or cinnamon to give it flavour, and bake it one hour in a gentle
oven.
Rice, 6 oz.: in water, 8 to 10 minutes. Milk, 1 quart: 3/4 to 1 hour.
Sugar, 3 oz.; eggs, 3 large, or 4 small; flavouring of nutmeg lemon-
rind, or cinnamon: bake 1 hour, gentle oven.
QUITE CHEAP RICE PUDDING.

Boil the rice in water, as for a currie, and while it is still warm, mix
with it a pint and a half of milk, and three fresh or four or five French
eggs (at many seasons of the year these last, which are always
cheap, are very good, and answer excellently for puddings.)
Sweeten it with pale brown sugar, grate nutmeg on the top, and bake
it slowly until it is firm in every part.
RICHER RICE PUDDING.

Wash very clean four ounces of whole rice, pour on it a pint and a
half of new milk, and stew it slowly till quite tender; before it is taken
from the fire, stir in two ounces of good butter, and three of sugar;
and when it has cooled a little, add four well-whisked eggs, and the
grated rind of half a lemon. Bake the pudding in a gentle oven from
thirty to forty minutes. As rice requires long boiling to render it soft in
milk, it may be partially stewed in water, the quantity of milk
diminished to a pint, and a little thick sweet cream mixed with it,
before the other ingredients are added.
Rice, 4 oz.; new milk, 1-1/2 pint; butter, 2 oz.; sugar, 3 oz.; eggs, 4;
rind of 1/2 lemon: 30 to 40 minutes, slow oven.
RICE PUDDING MERINGUÉ.

Swell gently four ounces of Carolina rice in a pint and a quarter of


milk or of thin cream; let it cool a little, and stir to it an ounce and a
half of butter, three of pounded sugar, a grain or two of salt, the
grated rind of a small lemon, and the yolks of four large, or of five
small eggs. Pour the mixture into a well-buttered dish, and lay lightly
and equally over the top the whites of four eggs beaten as for
sponge cakes, and mixed at the instant with from four to five heaped
tablespoonsful of sifted sugar. Bake the pudding half an hour in a
moderate oven, but do not allow the meringue to be too deeply
coloured; it should be of a clear brown, and very crisp. Serve it
directly it is taken from the oven.
Rice, 4 oz.; milk, or cream, 1-1/4 pint; butter, 1-1/2 oz.; sugar, 3
oz.; rind, 1 lemon; yolks of eggs, 4 or 5; the whites beaten to snow,
and mixed with as many tablespoonsful of sifted sugar: baked 1/2
hour, moderate oven.
Obs.—A couple of ounces of Jordan almonds, with two or three
bitter ones, pounded quite to a paste, will improve this dish, whether
mixed with the pudding itself, or with the meringué.
A GOOD GROUND RICE PUDDING.

Mix very smoothly five ounces of flour of rice (or of ground rice, if
preferred), with half a pint of milk, and pour it into a pint and a half
more which is boiling fast; keep it stirred constantly over a gentle fire
from ten to twelve minutes, and be particularly careful not to let it
burn to the pan; add to it before it is taken from the fire, a quarter of
a pound of good butter, from five to six ounces of sugar, roughly
powdered, and a few grains of salt; turn it into a pan, and stir it for a
few minutes, to prevent its hardening at the top; then mix with it, by
degrees but quickly, the yolks of eight eggs, and the whites of two,
the grated or rasped rind of a fine lemon, and a glass of brandy. Lay
a border of rich paste round a buttered dish, pour in the pudding,
strain a little clarified butter over the top, moisten the paste with a
brush, or small bunch of feathers dipped in cold water, and sift plenty
of sugar on it, but less over the pudding itself. Send it to a very
gentle oven to be baked for three-quarters of an hour.
Rice-flour (or ground rice), 5 oz.; new milk, 1 quart: 10 to 12
minutes. Butter, 4 oz.; sugar, 5 to 6 oz.; salt, 1/2 saltspoonful; yolks,
8 eggs; whites, 2; rind, 1 large lemon; brandy, large wineglassful: 3/4
hour, slow oven.
Obs.—These proportions are sufficient for a pudding of larger size
than those served usually at elegant tables; they will make two small
ones; or two-thirds of the quantity may be taken for one of moderate
size. Lemon-brandy or ratifia, or a portion of each, may be used to
give it flavour, with good effect; and it may be enriched, if this be
desired, by adding to the other ingredients from three to four ounces
of Jordan almonds, finely pounded, and by substituting cream for
half of the milk.
COMMON GROUND RICE PUDDING.

One pint and a half of milk, three ounces and a half of rice, three
of Lisbon sugar, one and a half of butter, some nutmeg, or lemon-
grate, and four eggs, baked slowly for half an hour, or more, if not
quite firm.
GREEN GOOSEBERRY PUDDING.

Boil together, from ten to twelve minutes, a pound of green


gooseberries, five ounces of sugar, and rather more than a quarter of
a pint of water: then beat the fruit to a mash, and stir to it an ounce
and a half of fresh butter; when nearly, or quite cold, add two ounces
and a half of very fine bread-crumbs, and four well whisked eggs.
Bake the pudding gently from half to three-quarters of an hour. To
make a finer one of the kind, work the fruit through a sieve, mix it
with four or five crushed Naples biscuits, and use double the quantity
of butter.
Green gooseberries, 1 lb.; sugar, 5 oz.; water, full 1/4 pint: 10 to 12
minutes. Bread-crumbs, 2-1/2 oz.; eggs, 4: 1/2 to 3/4 hour.
POTATO PUDDING.

With a pound and a quarter of fine mealy potatoes, boiled very dry,
and mashed perfectly smooth while hot, mix three ounces of butter,
five or six of sugar, five eggs, a few grains of salt, and the grated rind
of a small lemon. Pour the mixture into a well-buttered dish, and
bake it in a moderate oven for nearly three-quarters of an hour. It
should be turned out and sent to table with fine sugar sifted over it;
or for variety, red currant jelly, or any other preserve, may be spread
on it as soon as it is dished.
Potatoes, 1-1/4 lb.; butter, 3 oz.; sugar, 5 or 6 oz.; eggs, 5 or 6;
lemon-rind, 1; salt, few grains: 40 to 45 minutes.
Obs.—When cold, this pudding eats like cake, and may be served
as such, omitting, of course, the sugar or preserve when it is dished.
A RICHER POTATO PUDDING.

Beat well together fourteen ounces of mashed potatoes, four


ounces of butter, four of fine sugar, five eggs, the grated rind of a
small lemon, and a slight pinch of salt; add half a glass of brandy,
and pour the pudding into a thickly-buttered dish or mould,
ornamented with slices of candied orange or; pour a little clarified
butter on the top, and then sift plenty of white sugar over it.
Potatoes, 14 oz.; butter, 4 oz.; sugar, 4 oz.; eggs, 5; lemon-rind, 1;
little salt; brandy, 1/2 glassful; candied peel, 1-1/2 to 2 oz.: 40
minutes.
Obs.—The potatoes for these receipts should be lightly and
carefully mashed, but never pounded in a mortar, as that will convert
them into a heavy paste. The better plan is to prepare them by
Captain Kater’s receipt (Chapter XVII.), when they will fall to powder
almost of themselves; or they may be grated while hot through a wire
sieve. From a quarter to a half pint of cream is, by many cooks,
added always to potato puddings.
A GOOD SPONGE CAKE PUDDING.

Slice into a well-buttered tart-dish three penny sponge biscuits,


and place on them a couple of ounces of candied orange or lemon
rind cut in strips. Whisk thoroughly six eggs, and stir to them boiling
a pint and a quarter of new milk, in which three ounces of sugar have
been dissolved; grate in the rind of a small lemon, and when they are
somewhat cooled, add half a wineglassful of brandy, while still just
warm, pour the mixture to the cakes, and let it remain an hour; then
strain an ounce and a half of clarified butter over the top, or strew
pounded sugar rather thickly on it, and bake the pudding three
quarters of an hour or longer in a gentle oven.
Sponge cakes, 3; candied peel, 2 oz.; eggs, 6; new milk, 1-1/4
pint; sugar, 3 oz.; lemon-rind, 1; brandy, 1/2 glass; butter, 1 oz.;
sifted sugar, 1-1/2 oz.: 3/4 hour.
CAKE AND CUSTARD, AND VARIOUS OTHER INEXPENSIVE
PUDDINGS.

Even when very dry, the remains of a sponge or a Savoy cake will
serve excellently for a pudding, if lightly broken up, or crumbled, and
intermixed or not, with a few ratifias or macaroons, which should also
be broken up. A custard composed of four eggs to the pint of milk if
small, and three if very large and fresh, and not very highly
sweetened, should be poured over the cake half an hour at least
before it is placed in the oven (which should be slow); and any
flavour given to it which may be liked. An economical and clever
cook will seldom be at a loss for compounding an inexpensive and
good pudding in this way. More or less of the cake can be used as
may be convenient. Part of a mould of sweet rice or the remains of a
dish of Arocē Docē (see Chapter XXIII.), and various other
preparations may be turned to account in a similar manner; but the
custard should be perfectly and equally mingled with whatever other
ingredients are used. Macaroni boiled tender in milk, or in milk and
water, will make an excellent pudding; and sago stewed very thick,
will supply another; the custard may be mixed with this last while it is
still just warm. Two ounces well washed, and slowly heated in a pint
of liquid, will be tender in from fifteen to twenty minutes. All these
puddings will require a gentle oven, and will be ready to serve when
they are firm in the centre, and do not stick to a knife when plunged
into it.
BAKED APPLE PUDDING, OR CUSTARD.

Weigh a pound of good boiling apples after they are pared and
cored, and stew them to a perfectly smooth marmalade, with six
ounces of sugar, and a spoonful or two of wine; stir them often that
they may not stick to the pan. Mix with them while they are still quite
hot, three ounces of butter, the grated rind and the strained juice of a
lemon, and lastly, stir in by degrees the well-beaten yolks of five
eggs, and a dessertspoonful of flour, or in lieu of the last, three or
four Naples’ biscuits, or macaroons crushed small. Bake the pudding
for a full half hour in a moderate oven, or longer should it be not
quite firm in the middle. A little clarified butter poured on the top, with
sugar sifted over, improves all baked puddings.
Apples 1 lb.; sugar, 6 oz.; wine 1 glassful; butter, 3 oz.; juice and
rind, 1 lemon; 5 eggs: 1/2 hour, or more.
Obs.—Many cooks press the apples through a sieve after they are
boiled, but this is not needful when they are of a good kind, and
stewed, and beaten smooth.
DUTCH CUSTARD, OR BAKED RASPBERRY PUDDING.

Lay into a tart-dish a border of puff-paste, and a pint and a half of


freshly-gathered raspberries, well mixed with three ounces of sugar.
Whisk thoroughly six large eggs with three ounces more of sugar,
and pour it over the fruit: bake the pudding from twenty-five to thirty
minutes in a moderate oven.
Break the eggs one at a time into a cup, and with the point of a
small three-pronged fork take off the specks or germs, before they
are beaten, as we have directed in page 424.
Raspberries, 1-1/2 pint; sugar, 6 oz.; eggs, 6: 25 to 30 minutes.
GABRIELLE’S PUDDING, OR SWEET CASSEROLE OF RICE.

Wash half a pound of the best Carolina rice, drain it on a hair-


sieve, put it into a very clean stewpan or saucepan, and pour on it a
quart of cold new milk. Stir them well together, and place them near
the fire that the rice may swell very gradually; then let it simmer as
gently as possible for about half an hour, or until it begins to be quite
tender; mix with it then, two ounces of fresh butter and two and a half
of pounded sugar, and let it continue to simmer softly until it is dry
and sufficiently tender,[151] to be easily crushed to a smooth paste
with a strong wooden spoon. Work it to this point, and then let it cool.
Before it is taken from the fire, scrape into it the outside of some
sugar which has been rubbed upon the rind of a fresh lemon. Have
ready a tin mould of pretty form, well buttered in every part; press the
rice into it while it is still warm, smooth the surface, and let it remain
until cold. Should the mould be one which opens at the ends, like
that shown in the plate at page 344, the pudding will come out easily;
but if it should be in a plain common one, just dip it into hot water to
loosen it; turn out the rice, and then again reverse it on to a tin or
dish, and with the point of a knife mark round the top a rim of about
an inch wide; then brush some clarified butter over the whole
pudding, and set it into a brisk oven. When it is of an equal light
golden brown draw it out, raise the cover carefully where it is
marked, scoop out the rice from the inside, leaving only a crust of
about an inch thick in every part, and pour into it some preserved
fruit warmed in its own syrup, or fill it with a compôte of plums or
peaches (see Chapter XXIII.); or with some good apples boiled with
fine sugar to a smooth rich marmalade. This is a very good as well
as an elegant dish: it may be enriched with more butter, and by
substituting cream for the milk in part or entirely but it is excellent
without either.
151. Unless the rice be boiled slowly, and very dry, it will not answer for the
casserole.
Rice, 1/2 lb.; new milk, 1 quart: 1/2 hour. Fresh butter, 2 oz.;
pounded sugar, 2-1/2 oz.; rasped rind, 1 lemon: 1/2 hour or more.
Obs.—The precise time of baking the pudding cannot well be
specified: it only requires colour.
VERMICELLI PUDDING WITH APPLES OR WITHOUT, AND
PUDDINGS OF SOUJEE AND SEMOLA.

Drop gradually into an exact quart of boiling milk four ounces of


very fresh vermicelli, crushing it slightly with one hand and letting it
fall gently from the fingers, and stirring the milk with a spoon held in
the other hand, to prevent the vermicelli from gathering into lumps.
Boil it softly until it is quite tender and very thick, which it will be
usually in about twenty minutes, during which time it must be very
frequently stirred; then work in two ounces of fresh butter and four of
pounded sugar; turn the mixture into a bowl or pan, and stir it
occasionally until it has cooled down. Whisk five good eggs until they
are very light, beat them gradually and quickly to the other
ingredients, add the finely grated rind of a lemon or a little lemon-
brandy or ratifia, and pour the pudding when nearly cold into a
buttered dish, and just cover the surface with apples pared, cored,
and quartered; press them into the pudding-mixture, to the top of
which they will immediately rise again, and place the dish in a very
gentle oven for three-quarters of an hour, or longer if needed to
render the fruit quite tender. The apples should be of the best quality
for cooking. This is an exceedingly nice pudding if well made and
well baked. The butter can be omitted to simplify it.
Milk, 1 quart; vermicelli, 4 oz.: boiled about 20 minutes. Butter 2
oz.; (when used) pounded sugar, 1/4 lb.; eggs, 5: baked slowly 3/4
hour or more.
For a plain common vermicelli pudding omit the apples and one
egg: for a very good one use six eggs, and the butter; and flavour it
delicately with orange-flower water, vanilla, or aught else that may be
preferred. We have often had an ounce or two of candied citron
sliced very thin mingled with it.
Puddings of soujee and semola are made in precisely the same
manner, with four ounces to the quart of milk, and ten minutes
boiling.
RICE À LA VATHEK, OR RICE PUDDING À LA VATHEK.

(Extremely Good.)
Blanch, and then pound
carefully to the smoothest
possible paste four ounces of
fine Jordan almonds and half a
dozen bitter ones, moistening
them with a few drops of water
to prevent their oiling. Stir to
them by slow degrees a quart of boiling milk, which should be new,
wring it again closely from them through a thin cloth, which will
absorb it less than a tammy, and set it aside to cool. Wash
thoroughly, and afterwards soak for about ten minutes seven ounces
of Carolina rice, drain it well from the water, pour the almond-milk
upon it, bring it very slowly to boil, and simmer it softly until it is
tolerably tender, taking the precaution to stir it often at first that it
may not gather into lumps nor stick to the pan. Add to it two ounces
of fresh butter and four of pounded sugar, and when it is perfectly
tender and dry, proceed with it exactly as for Gabrielle’s pudding, but
in moulding the rice press it closely and evenly in, and hollow it in the
centre, leaving the edge an inch thick in every part, that it may not
break in the oven. The top must be slightly brushed with butter
before it is baked, to prevent its becoming too dry, but a morsel of
white blotting paper will take up any portion that may remain in it.
When it is ready to serve, pour into it a large jarful of apricot jam, and
send it immediately to table. If well made it will be delicious. It may
be served cold (though this is less usual), and decorated with small
thin leaves of citron-rind, cut with a minute paste-cutter. The same
preparation may be used also for Gabrielle’s pudding, and filled with
hot preserved fruit, the rice scooped from the inside being mixed with
the syrup.
GOOD YORKSHIRE PUDDING.

To make a very good and light Yorkshire pudding, take an equal


number of eggs and of heaped tablespoonsful of flour, with a
teaspoonful of salt to six of these. Whisk the eggs well, strain, and
mix them gradually with the flour, then pour in by degrees as much
new milk as will reduce the batter to the consistence of rather thin
cream. The tin which is to receive the pudding must have been
placed for some time previously under a joint that has been put down
to roast one of beef is usually preferred. Beat the batter briskly and
lightly the instant before it is poured into the pan, watch it carefully
that it may not burn, and let the edges have an equal share of the
fire. When the pudding is quite firm in every part, and well-coloured
on the surface, turn it to brown the under side. This is best
accomplished by first dividing it into quarters. In Yorkshire it is made
much thinner than in the south, roasted generally at an enormous
fire, and not turned at all: currants there are sometimes added to it.
Eggs, 6; flour, 6 heaped tablespoonsful, or from 7 to 8 oz.; milk,
nearly or quite 1 pint; salt, 1 teaspoonful: 2 hours.
Obs.—This pudding should be quite an inch thick when it is
browned on both sides, but only half the depth when roasted in the
Yorkshire mode. The cook must exercise her discretion a little in
mixing the batter, as from the variation of weight in flour, and in the
size of eggs, a little more or less of milk may be required: the whole
should be rather more liquid than for a boiled pudding.
COMMON YORKSHIRE PUDDING.

Half a pound of flour, three eggs (we would recommend a fourth),


rather more than a pint of milk, and a teaspoonful of salt.

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