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1 Stlongexam
1 Stlongexam
1 Stlongexam
• Single-celled eukaryotes that are a key component of planktic communities in the ocean;
• Present in the upper water column and in deeper layer;
• Possess splender pseudopodia (called filopodia)
• Ex. Ammonia tepida
Additional characteristics
• Most Radiolians are heterotrophs by two groups (Chlorarachnea and Paulinella) include photosynthetic species
• Radiolaria and Foraminifera harbor algal symbionts to supplement their metabolic needs
Two major groups of Rhizaria include:
1. Phylum Foraminifera
2. Phylum Radiolaria
GROUP: Radiolarians
• Ray animals;
• Body divided into distinct intracapsular & extracapsular zones separated by a perforated membrane or capsule
• Usually of microscopic size when existing as solitary forms, but some species (e.g., some members of Order Spumellaria) form
colonies.
• Example: A radiolarian, Acanthometra elasticum
AMOEBOID PROTOZOANS
Cell Structure:
• simple cell structure (Cell membrane, cytoplasm & nucleus)
• contain specialized organelles e.g. food vacuoles, contractile vacuoles & plastids
Pseudopodia:
• temporary cytoplasmic projections used for movement and feeding
• allowing the organism to change shape and direction
Feeding;
• heterotrophic (phagocytosis or pinocytosis)
Reproduction:
• both asexual (binary fission/multiple fission or budding) & sexual (fusion of gametes to form a zygote)
Ecological Roles:
• consumers, decomposers, & predators
• regulate microbial populations, recycle nutrients & contribute to nutrient cycling
The Amoebozoans
• Free living, does include some parasitic species e.g. Entamoeba histolytica
• Nearly all possess branching tubular mitochondrial cristae sometimes referred to as the “naked ramicristate amoebae”
Representative species:
1. Amoeba proteus
2. Chaos carolinensis
Mastigamoebae
• -possess both locomotory flagella and pseudopodia
• Single-celled
• Heterotrophic, using their flagella or phagocytosis
• Found in various aquatic and terrestrial habitats including freshwater ponds & streams, soil, leaf litter, and marine environments.
Distinct Characteristics
Flagellar structure: Two flagellar types (anterior flagella & posterior flagella)
Flagellar reproduction: may undergo binary fission or form cysts for dispersal
Representative species:
1. Naegleria fowleri
2. Percolomonas
The Rhizaria
• possess splender pseudopodia (filopodia) – may be simple or branching, sometimes highly complex networks (reticulopodia)
• encompasses wide range of protists, including foraminiferans and radiolarians
• both free-living and symbiotic organisms
• diverse feeding strategies (autotrophic & heterotrophic)
• Asexual & sexual reproduction
• believe to have diversified from a common ancestor with other eukaryotic groups such as Alveolata and Stramenopiles
Phylum Foraminifera
• are primarily marine and live on bottom sediments as benthic organisms
• no parasitic species, some are ectocommensal
• secrete multichambered test, typically of calcium carbonate
• reticulopodia – food capture
• repeated extension and shortening – permits slow crawling over the ocean bottom
• Paleontological Significance
Distinct Characteristics
• Test Morphology: Calcium carbonate, agglutinated particles, or organic material
• Pseudopodia
• Chambered Structure: septa (buoyancy control, protection, reproduction)
• Paleotological Importance
• BioIndicator Species
Representative species:
1. Globigerina bulloides
2. Nummulites
Phylum Radiozoa
• members of this phylum radiolarians & acantharians – support their pseudopodia with thin, radiating microtubules that give a spiny,
rayed appearance to many species
• Axopodia
• possess a rigid endoskeleton composed of either silica (in radiolarians) or strontium sulfate (in acantharians)
• prominent in the fossil record
• phototrophic & heterotrophic
• radiolarian body (spherical – intracapsular zone (nucleus is contained) and extracapsular zone (food vacuole formation and digestion
occur)
Distinct Characteristics
• Body is divided into distinct intracapsular and extracapsular zones separated by a perforated membrane or capsule
Representative species:
1. Acanthometra pellucida
2. Aulacantha scolymantha
Flagellates (mastigophorans)
• body covered by pellicle
• locomotion by one or more flagella
• may be solitary or colonial
• cytostome is present in some species
• contractile vacuoles may be present, particularly in freshwater species
Flagellates include:
1. Phytoflagellated protozoans
2. Zooflagellated protozoans
- Free-living forms
- Parasitic forms
Phytoflagellated protozoans
- Photosynthesizing species
- Some contain chlorophyll (e.g. Euglena from Class Euglenida)
- Others use light but require various dissolved organic compounds
- Many species have stigma, a photosensitive organelle
- Some become holozoic if necessary
Zooflagellated protozoans
- animal-like
- assimilate organic material by osmotrophy or phagotrophy
- Most species are free-living in freshwater, saltwater, or soil
- About 25% are parasitic or commensal with plants, invertebrates, and vertebrates
-Trypanosomes, responsible for African sleeping sickness with tsetse fly as vector
Leishmania donovani, sand flies as vectors, causes extreme disfigurement and death
Ex: Flagellate diversity: trypanosomes.
(a) Trypanosoma lewisi.
(b) African trypanosome (Trypanosoma brucei brucei)
Representative species
1. Euplectella aspergillum (Venus’ Flower Basket)
• Skeleton contains hexactine siliceous
• Known as "glass sponge" due to its glass-like appearance.
• Surrounded by a trabecular net
• Within the net are finger-like chambers covered in choanocytes, opening into the spongocoel.
• Choanocytes vibrate to move water through the sponge, creating a funnel-like shape.
• At its base, a tuft of elongated spicules attaches it to the ocean bottom.
2. Hyalonema sp. (Glass Rope Sponge)
• Unique sponge species with no organ system or tissues.
• Can grow up to 6ft tall and has a silica-based skeleton.
• Found in deeper waters
• Reproduction: Sexually and Asexually
3. Farrea ananchorata
4. Hexasterophora incertae sedis
5. Oopsacas minuta
Distinct characteristics:
• The entire sponge, including the outer layer, is syncytial.
• Lacks contractile elements.
• No pinacoderm layer.
• The inner, flagellated layer is also syncytial.
• can rapidly conduct electrical impulses across their bodies
CLASS CALCAREA
Ex. Clathrina darwinii
Characteristics
• “Calx”, means limestone or chalk
• Skeletons made entirely of calcium carbonate spicules and lack spongin
• 10 cm less in height and dull in color
• Tree-pointed spires (triactines) is the most common spicule shape
• Only class with having the three canal system (asconoid synconoid and leuconoid)
Taxonomy
• ~ 756 marine species
• 7.5% of sponge diversity
Habitat
• Can be found mainly on the rocky bottoms, and shallow saltwater, particularly in temperate areas.
• Deepest species found at 4400m.
Reproduction
• Can reproduce sexually and asexually
• Asexual reproduction – by budding
• Sexual reproduction – being hermaphroditic
• exception of Genus Leucetta for being viviparous
• Ex. Leucetta chagonensis
Representative organisms:
1. Leucetta chagonensis
2. Leucandra heathi
CLASS HOMOSCLEROMORPHA
General Characteristics
• Absence of a skeleton
• Small size and simple body structure
• Encrusting forms
• Exclusively found in marine environments
• possess a unique type of tetractine spicules (calthrops)
• Basement membrane lining choanoderm and pinacoderm
Family Oscarellidae
• are typically small sponges often ranging from few centimeters in size
• Compared to other sponge group they lack complex canal system or large cavities
• Exhibit a compact, globular or vase-like shape with a single osculum at the apex
• Lack of spicules
• Typically found in shallow marine environments often inhabiting crevices or rocky substrates
• They are filter feeders
Representative organism
1. Oscarella lobularis
~Small globular sponge
~Found in rocky substrates in Mediterranean sea and Northeastern Atlantic Ocean
~Lacks skeleton of spicules instead supported by network of collagen fibers
~They are filter feeder
2. Plakina jani
~found in shallow coastal water
~Has flattened, encrusting or lobate morphology with slightly wrinkled surface
~Lacks spicules or very small, simple spicules compared to other sponges
~Supported by network of collagen fibers
Family Plakinidae
• Exhibits an encrusting or massive growth form
• Flattened body
• Lack of spicules
• Presence of plakins
• Found in marine environments, inhabiting wide range of depths from shallow coastal waters to deeper oceanic regions
• Filter feeders
Representative organism
1. Corticium candelabrum
~Encrusting colonies speading over surfaces of rocks, shells or coral substrates
~Flattened body
~Lack of spicules, no supporting skeletons
~Presence of plakins
~Inhabits shallow coastal waters to moderate depths
~Filter feeders
2. Plakortis halichondrioides
~Sponge species with massive or globular shape
~Typically found in deeper waters
~Exhibit variable coloration, ranging from light brown to orange or reddish-bown
~Appear uneven, with ridges or pores scattered across surface
~Presence of oscula
~Produce bioactive compounds for defense mechanisms
DEMOSPONGIAE
Characteristics
• are asymmetrical
• range from thin encrustations several cm in diameter to huge cake-shaped species 2 m (6.6 ft) in diameter
• skeleton of either 1- or 4-rayed siliceous spicules, spongin fibres, or both; skeleton lacking in a few primitive genera
• ex. Crella incrustans (Crellidae)
• -This sponge can be found in rock pools, under ledges in the intertidal or in rubble banks of boulders
• -possess a leuconoid structure, with a folded choanoderm
Ex. Amphimedon queenslandica
Poecilosclerida
• has a unique feature— chelae, which are meniscoid (crescent-shaped) microscleres with a curved shaft and recurved,
winglike or broadly rounded structures at each end
Haplosclerida
• are frequently cushionshaped; however, encrusting, branching, tubular, vase-, and fan-shaped forms are also quite common;
coloring is not very intense
Habitat and Behavior
• marine environment from the intertidal to the abyssal zone; some species inhabit freshwater
• are immobile animals attached at the base to a substrate, or surface on which they live
• some species successfully compete with corals and other sponges for space by releasing toxic chemicals.
• any member of the sponge family Clionidae (class Demospongiae, phylum Porifera), noted for its ability to dissolve and bore into
calcium-containing substances, such as limestone, coral, and mollusk shells.
Reproduction
• Some are hermaphroditic while others have distinct sexes
• may be viviparous, oviparous, or asexual
• In sexual reproduction, spermatocytes develop from the transformation of choanocytes, and oocytes arise from archeocytes.
• asexual reproduction include both budding and the formation of gemmules
OPISTHOKONTS
• contains the true fungi and their protozoan relatives, and the multicellular animals (including humans) with their protist relatives
• has a posterior ( opistho = G: behind) flagellum ( kont = G:flagellum) in some part of the life cycle.
The Myxozoans
• Chloromyxum, Myxidium
• Molecular data now show that the members of this group, which are all parasitic, mostly in fishes, are highly degenerate metazoans, probably
most closely related to jellyfish and other cnidarians.
• Classification of myxospores is based on the number of shell valves (Bivalvulida, Multivalvulida) and the position of the polar capsules
in the spore.
CHARACTERISTICS
• Hydrozoans are small, relatively common Class under the Phylum Cnidarian.
• Can be found in both marine and freshwater environments.
• exhibit two body forms, either polypoid, which is frequently colonial and a medusoid stage in their life cycles, or both.
Diploblastic
• two embryonic tissue layers that are mostly made up of epitheliomuscular cells/myoepithelial cells
• mesoglea that separates the epidermis and gastrodermis.
• interstitial stem cells or i-cells that produces various somatic cells including the stinging cells called cnidocytes
• Gastrodermis encloses the gastrovascular system which serves both nutritive and transport function.
• zooids (or modules of the colony), all of which are interconnected and share a continuous gastrovascular cavity
• Morphologically distinct polyp types; feeding gastrozooids, reproductive gonozooids, defensive dactylozooids.
• Manubrium - the projection from the middle of the subumbrella surface and contains the mouth at its apex.
• a membranous, shelf-like structure called a velum, which projects in (towards the manubrium) from the inner edge of the ring canal
region.
LIFE CYCLE AND REPRODUCTION
• Reproductive polyps, known as gonozooids (or "gonotheca" in thecate hydrozoans) bud off asexually produced medusae.
• In hydrozoan species with both polyp and medusa generations, the medusa stage is the sexually reproductive phase. Medusae of these
species of Hydrozoa are known as "hydromedusae".
• Alternates between polyp and medusa stage • Fertilization occurs when sperm are carried by water
• Planula develops into a feeding gastrozooid current to a female medusae
• Gastrozooids produced Gonozooids, which produces • Blastula is converted into an oval ciliated planula larva
medusae through invagination
• Medusae mature and break free and swim out of the • Settles down on submerged substratum at one end
opening • Disc appears for attachment and develops a manubrium
• Medusae reporoduce sexually
and tentacles to turn into a simple polyp
• Common freshwater Hydrozoans that hangs underside of a floating plants in clean streams and ponds
• Lacks medusa stage and reproduce asexually(budding) and sexually
Sexual Reproduction of Hydra
• fusion of specialized reproductive cells from two parent hydras.
• Sexual reproduction introduces genetic diversity into the population and allows for the creation of offspring with unique combinations
of traits.
Order Siphonophora
• Large oceanic Hydrozoans belong to this order
• Association of polypoid and medusoid individuals
• Include the Portuguese man-of-war; free-floating hydrozoan colonies in which medusoid and polypoid morphs are present
simultaneously in a number of different incarnations.
CLASS ANTHOZOA
CLASSIFICATION
• The class Anthozoa, contains approximately 7, 500 extant species
• Divided into two subclasses: Hexacorallia and Octocorallia
• Hexacorallia
-is comprised of six orders, including the Actiniaria, Antipatharia, Ceriantharia, Corallimorphia, Scleractinia, and Zoantharia
• Octocorallia
-is comprised of three orders, namely Alcyonacea, Helioporacea, and Pennatulacea
DISTINCT CHARACTERISTICS
- absence of a medusa stage
- absence of operculum and cnidocil
- mitochondrial DNA is circular (as in most eukaryotes) rather than linear
- presence of a ciliated groove (siphonoglyph) in the pharyngeal wall leading from the mouth
- coelenteron partitioned by distinct sheets of tissue (mesenteries/septa)
ANATOMY
-Anthozoa are considered ‘simple’ as they contain only two layers of cells, an inner layer called endoderm and an outer layer called ectoderm
- All anthozoans are marine and all exploit the polyp body form and lifestyle exclusively;
• Polyp – is a tubelike sac (gastrovascular cavity) with a single opening at the top of the sac that serve as both the mouth and anus and
thus called the mouth-anus
• Anthozoan mouth opens into a tubular pharynx rather than directly into the gastrovascular cavity, and one or two discrete, ciliated
grooves, called siphonoglyphs
• gastrovascular cavity is partitioned by numerous sheets of tissue called mesenteries or septa
• sac (gastrovascular cavity) with a single opening at the top of the sac that serve as both the mouth and anus and thus called the
mouth-anus
REPRODUCTION
• Can occur either vegetatively (“asexually”) or sexually
• Asexual Reproduction- occurs “vegetatively” involving the complete detachment of body segments from animal to form genetically
identical individuals or genets
• Sexual Reproduction- involves the fusion of gametes- sperm and egg- to form a distinct fertilized embryo which develops into a free-
swimming larva.
Representative Organism
1. Agaricia tenuifolia
• Encrusting or plate-like colonies
• Polyps have thin, elongated tentacles
• Typically brownish or yellowish
• Colonies can reach sizes of up to 30 cm
• Typically found at depths of 5 to 20 meters
2. Pocillopora damicornis
• Branching or bushy colonies
• Polyps have small, cylindrical tentacles
• Variable, including brown, pink, green, or purple
• Colonies can reach sizes of up to 1 meter
• Found in shallow, clear tropical waters
Subclass Ocotocorallia (Alcyonaria)
• Polyps bear eight tentacles and are subdivided by eight complete mesenteries
• The tentacles of octocorallians are pinnate; that is, they bear numerous lateral outfoldings called pinnules
• The polyps of octocorals may be embedded in a thick matrix of mesoglea; these are the soft corals.
Representative Organism
1. Clavularia
• Colonial, with polyps connected by stolons or creeping bases
• Each polyp has a cylindrical or club-like shape, with tentacles surrounding the oral disc
• Individual polyps are typically small, around a few millimeters in size
• Variable, including shades of brown, green, pink, or purple
2. Pennatula phosphorea
• Soft, flexible cylindrical body with a central axis called a rachis
• Numerous polyps arranged along lateral branches called pinnules, giving a feather-like appearance
• Can grow up to several decimeters in height
• Typically pale yellow or light brown, may exhibit bioluminescence
1. Aurelia aurita
• Translucent and whitish in color, often shaded with blue, pink, or blue.
• Shallow dome-shaped bell and range in diameter from 10 to 35 cm.
• Numerous short, fine, fringe-like tentacles around the margin.
• The four white, horseshoe-shaped gonads.
• Four oral arms hang down from the center of the bottom surface of the bell.