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Trees A Handbook of Forest Botany
Trees A Handbook of Forest Botany
A HANDBOOK OF FOREST-BOTANY
Anthony Walsh
TREES
A HANDBOOK |of| FOREST#BOTANY
TREES
A HANDBOOK |of| FOREST#BOTANY
Anthony Walsh
Trees*!a#Handbook|of|Forest#Botany
^by* Anthony Walsh
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Contents
1@ Forest Plantations 1
2@ Forest**an^Area|of|Density|of|Trees 62
3@ Forest * Botanical Relationships 105
4@ Silviculture*and%Management*in=Forestry 140
5@ Food*and%Plant Products*in=Forestry 162
6@ Forest Structure*and%Patterns 213
7@ Land*and%Farm Resources*in=Forestry 237
1
Forest Plantations
Introduction
Forest plantations covered 187 million hectares*in=2000* |of| w h i c h
Asia accounted{for}62 per cent@*the^forest plantation area represents
!a# significant increase{from]*the^1995 estimate |of| 124 million hectares@
*the^ reported n e w annual planting rate[is]4@5 million hectares globally*
with Asia*and%South A m e r i c a accounting{for}89 per cent@ A b o u t 3
million hectares {are] estimated {to] be successful@ Globally* half *the^
forest plantation estate [is] {for} industrial end#use* one#quarter {for} non#
industrial end#use*and%one#quarter not specified@ Globally**the^m a i n
fast#growing* short#rotation species {are] *in= *the^ genera Eucalyptus and+
Acacia@ Pines *and% other coniferous species {are] *the^ main medium#
rotation utility species* primarily*in=*the^temperate*and%boreal zones@
The potential {for} forest plantations {to] partially m e e t d e m a n d
^from* natural forests{for}w o o d*and%fibre{for}industrial uses[is]increasing@
A l t h o u g h accounting{for}only 5 per cent|of|global forest cover* forest
plantations {were] estimated*in=*the^year 2000{to]supply about 35 per
cent |of| global roundwood@ This figure [is] anticipated {to] increase *to^
44 per cent {by] 2020@ *in= some countries forest plantation production
already contributes *the^ majority |of| industrial wood supply@ There *is$
increasing interest *in= development |of| forest plantations as carbon
2 Trees*!a#Handbook|of|Forest#Botany
Plantation Resources*in=India
Plantation resources {have] been generated *and% maintained *in^
India over *the^ ages@ Emperor Ashoka (273#232 BC) [is] known {to] *have^
^had* trees planted along long stretches |of| main roads *in= his vast
empire@ E m p e r o r Shivaji (1630#1680)[is]reported{to]{have]encouraged
plantation forestry within his empire@ Sher K h a n (1472#1545)* w h o
asserted his independence {from] *the^ Mughal Emperor Humayun and+
built *and% ruled over !a# large empire* [is] known {to] {have] formally
converted*an^old imperial h i g h w a y spanning almost*the^entire north
|of|*the^Indian sub#continent into*the^G r a n d T r u n k R o a d*and%{had]large
stretches|of|*the^roadsides planted with trees@
The earliest plantation |of| *the^ colonial era *in= India [is] reported *to^
be |of| !a# native species* teak (Tectona grandis)* planted *in= 1840 *in^
Nilambur* Kerala@ Regular planting* mainly |of| teak* b e g a n*in=1865
*in& m a n y|of|*the^teak#growing central*and%southern provinces@
In 1910* Eucalyptus spp@ was introduced *in= *the^ Nilgiri Hills |of| *the^
present Tamil Nadu@ Planting |of| other native species was accelerated
after*the^taungya system w a s introduced*in=1911@ These plantations*
however* did not cover*an^extensive area until 1950@
P l a n n e d afforestation {for} soil conservation* industrial wood*
fuelwood*and%fodder started*in=*the^late 1950s@ *the^ total plantation
6 Trees*!a#Handbook|of|Forest#Botany
forms !a# complex regional mosaic |of| communities* determined {by] !a#
combination |of| climatic* soil *and% physiographic factors@ *in= *the^ southern
portion|of|*the^Subatlantic province* Quercus spp@ b e c o m e increasingly
prominent* particularly{at]lower elevations@*in=*the^eastern Pyrenees*
Quercus pubescens [is] found{at]*the^lowest elevations* Fagus sylvatica
{and} Abies alba {at] higher elevations* with Abies alba increasing *in^
importance with altitude@
The Central European province covers*the^former East Germany*
western Poland *and% western Czech Republic *and% [is] characterized *by^
!a# more continental climate than *the^ Atlantic or Subatlantic provinces@
Fagus sylvatica r e a c h e s {its] e a s t e r n m o s t extent here* *and% occurs
extensively north|of|*the^Baltic Sea@{it]{has]m a n y associates* depending
on elevation**and%often{has]!a#more species#rich ground layer than
comparable forests*in=*the^Subatlantic province@ Carpinus betulus#
Quercus petraea communities formerly occupied rich* mesic plains
{and} uplands* but m o s t {have] b e e n converted {to] agriculture@ Betula#
Quercus forests g r o w on nutrient#poor* low p H soils* with Betula
pubescens *and% Quercus robur c o m m o n on wet sites*and%Betula pendula
{and} Quercus petraea on drier sites@ Pinus sylvestris *and% Picea abies *are^
locally intermixed *in= *the^ north *and% east* with Larix decidua common
^to**the^hills |of| southern Poland@
The Sarmatic province extends*in=!a#b a n d eastward{to]nearly 60°
E* {its] north#south boundaries tapering {from] about 50#60° N *in= *the^
west {to] about 53#54° N {at] {its] eastern edge@ {to] *the^ south lies steppe
{and} {to] *the^ north boreal forests@ Generally* *the^ Sarmatic province *is$
characterized {by] !a# progressive west#to#east loss |of| *the^ deciduous
species characteristic|of|*the^three provinces{to]*the^west*and%*an^increase
*in& *the^ importance |of| conifers@ {for} example* there [is] !a# tendency *for^
nutrient#poor sites {to] shift {from] domination {by] Betula *and% Quercus
robur *in= *the^ Subatlantic province {to] domination {by] Pinus *in= *the^
Sarmatic@ Similarly* on more fertile sites* Fagus sylvatica [is] no longer
present* *and% {has] been replaced {by] Tilia* Quercus robur* Carpinus and+
Pinus@ Pinus sylvestris [is] distributed throughout *the^ Sarmatic province
{and}[is]*an^associate or dominant on all but*the^m o s t fertile* moist soils@
O n sandy soils {it] forms nearly pure stands* without deciduous
associates@ Picea abies[is]also common* occurring with Pinus sylvestris*
{and} increasing*in=importance with soil fertility*and%moisture@ Abies
alba occurs*in=*the^eastern part|of|*the^province* with Abies sibirica in
30 Trees*!a#Handbook|of|Forest#Botany
Plantations*and%Carbon Sequestration
In*the^past ten years**the^development |of| forest plantations as
carbon offsets{has]evolved towards!a#market mechanism* although
*an& organized market with carbon prices defined according{to]supply
{and} d e m a n d forces[is]still!a#long w a y off@*the^adoption|of|*the^K y o t o
P r o t o c o l *in= 1 9 9 7 t r i g g e r e d !a# s t r o n g i n c r e a s e *in= i n v e s t m e n t in
34 Trees*!a#Handbook|of|Forest#Botany
studies |of| life history traits *and% species requirements@ Variation *in^
*the^ frequency *and% size |of| natural disturbance events should be
accounted{for}*in=management planning{by]incorporating or mimicking
natural disturbance regimes{at]several spatial*and%temporal scales@
Such planning needs {to] take !a# hierarchical perspective {to] match *the^
mobility *and% requirements |of| biological entities that vary *in= size *from^
small arthropods {to] herds |of| large mammals@
Plantations*and%Wood Energy
Woodfuels {from] plantations or natural or semi#natural forests
^are* particularly important*in=developing countries* providing about
15 per cent |of| their total e n e r g y demand@ Woodfuel provides about
7 per cent |of| energy demand {for} *the^ world as !a# whole *and% *in^
industrialized countries only 2 per cent@ Woodfuel provides more
than 70 per cent|of|energy n e e d s*in=3 4 developing countries*and%more
than 90 per cent*in=13 countries@ Woodfuel m a k e s u p about 80 per
cent |of| total w o o d use*in=developing countries*and%about 89 per cent
*in& Africa
The prediction |of| !a# woodfuel crisis *in= developing countries *in= *the^
1980s was based largely on looking {at] supply *and% demand *from^
forest plantations *and% natural forests@ *the^ reaction {to] *the^ expected
woodfuel crisis w a s{to]plant trees{for}this purpose* often*in=*the^form
|of| traditional plantations@
Many programme failures resulted {from] lack |of| appreciation *for^
*the^ complexities |of| bioenergy supply *and% demand* failure {to] take
into account social aspects*and%people@s n e e d s*and%p o o r p r o g r a m m e
structures@ *the^ importance |of| planted trees o n farmland**in=villages
{and} homesteads *and% along roads *and% waterways as !a# source |of|
woodfuel supply was underestimated@
Rural communities harvest stems* branches* stumps* twigs* leaves
{and} litter{for}woodfuels*in=chronic woodfuel supply areas@*in=these
instances *the^ nutrient recycling process [is] broken* resulting *in^
degradation |of| forest plantation sites@ *in= many rural communities *in^
developing countries* woodfuel [is] considered !a# public free good* *to^
be foraged {from] public natural *and% plantation forests@ Often women
{and} children collect *the^ woodfuel {at] little or no cost@ As !a# result* *the^
growing |of| private forest plantations specifically {for} woodfuel* *in^
which development costs *and% rotation cycles {are] involved* can be
Trees*!a#Handbook|of|Forest#Botany 41
These species account {for} 9@7* 12@0 *and% 6@0 million hectares |of|
plantations* respectively@ All grow *in= *the^ humid tropics@ *in= terms |of|
plantation area* Asia {has] 92 per cent|of|*the^world@s rubber* 86 per
cent |of| *the^ world@s coconut p a l m*and%78 per cent |of| *the^ world@s
African oil palm@ Indonesia*
Thailand*and%Malaysia {have] almost three#quarters|of|*the^rubber
plantations~ Indonesia*and%*the^Philippines {have] about half*the^coconut
resources~*and%Malaysia{has]55 per cent|of|*the^oil p a l m resource@ All
three species {are] g r o w n principally {for} other products rather than
wood* so w h e n o v e r m a t u r e t h e y {are] available {for} fibre#based
industries {at] minimal cost@
Rubberwood [is] harvested when latex productivity declines and+
3
yields 100 m per hectare |of| roundwood* but recovery {for} lumber
*is] only 25{to]4 5 per cent because |of| poor form*and%small size@ M o s t
|of|*the^planted stands*in=Southeast Asia{are]o w n e d {by] smallholders
{and}{are]geographically dispersed* with poor accessibility*and%poor#
quality stems@ Currently*the^major proportion |of| industrially utilized
r u b b e r w o o d comes {from] large#scale plantations@ Quality furniture*
parquet* panelling* reconstituted panels* general utility timber and+
woodfuel* including charcoal* {are] made {from] rubberwood@
However* *the^ rubberwood must be processed within days |of|
harvesting {to] minimize sapstain attack@ *the^ most developed
downstream industries {are] *in= Malaysia*{where}*the^ production |of|
3 3
sawn rubberwood timber rose {from] 88 000 m *in= 1990 {to] 137 000 m
*in&
1997 *and% medium density fibreboard (MDF) production *from^
rubberwood reached 1@16 million cubic metres per annum {by] 1999@
Exports |of| r u b b e r w o o d furniture {have] grown {from] about U S $ 7 4
million*in=1991{to]U S $ 6 8 3 million*in=1998@ R u b b e r w o o d {has] b e c o m e
!a# substitute {for} light tropical forest hardwoods@ {its] acceptance as !a#
sustainable plantation#grown* environmentally friendly timber *has^
given {it] wide appeal@
Coconut palms{are]harvested as*the^copra yields decline (beyond
3
60 years)*and%yield 90 m per hectare |of| coconut wood@ Coconut p a l m
^has* variable properties*and%[is]intrinsically difficult {for} conversion
but can yield !a# relatively low#cost* general#utility timber *for^
construction* panelling* stairs* door jambs* furniture* flooring and+
power poles@ *in= 1993 Indonesia {had] 65 million cubic metres |of|
overmature coconut stems which needed disposal before replanting@
Trees*!a#Handbook|of|Forest#Botany 43
There [is] increasing interest *in= this raw material *in= European and+
North American markets@ {it] [is] unlikely {to] replace conventional timber*
b u t likely{to]find{its]w a y into niche markets@{it]{will]continue{to]b e
used as!a#low#cost construction timber@
Oil p a l m plantations{are]harvested{for}fibre b e y o n d*the^25{to]30
3
year rotations*and%yield about 235 m per hectare@{it][is]estimated that
over 1@6 billion cubic metres |of| fibre {will] b e available*in=*the^years
^to* come {from] established resources *in= Southeast Asia@ {from] 1996 *to^
1999 *the^ area increased {by] 18 per cent@ *in= Malaysia *the^ area *has^
increased {by] 3 million hectares *in= *the^ past 30 years@
Most oil palm plantations (unlike rubber *and% coconut) *in= *the^
main growing countries* Malaysia *and% Indonesia* {are] managed *by^
plantation companies or cooperatives@ Oil palm by#products such as
kernel shells* pressed fibres*and%e m p t y fruit b u n c h e s {are] currently
used*in=heat generation{at]*the^extraction plants@ W a t e r*in=*the^stems
can reach five times*the^weight |of| dry matter@
The high moisture content as well as *the^ high amounts |of|
parenchyma tissue rich *in= sugar *and% starches make conversion into
quality forest products !a# challenge@ *an^ MDF plant *in= Malaysia *is$
currently being planned {to] utilize oil palm stems@
grown *in= *the^ wrong sites* with *the^ wrong species/provenances* *by^
*the^ wrong growers* {for} *the^ wrong reasons@
Examples exist{where}natural forests{have]been cleared{to]establish
forest plantation development or{where}customary owners |of|
traditional lands may {have] been alienated {from] their sources |of| food*
medicine*and%livelihoods@
In s o m e instances p o o r site/species matching *and% inadequate
silviculture {have] resulted*in=p o o r growth* hygiene* v o l u m e yields
{and} e c o n o m i c returns@*in=other instances* changes*in=soil*and%water
status {have] caused problems{for}local communities@ L a n d use conflicts
can occur b e t w e e n forest plantation d e v e l o p m e n t*and%other sectors*
particularly *the^ agricultural sector@
The negative impacts |of| forest plantations can d r a w *the^ focus
a w a y {from] *the^ fact that forest p l a n t a t i o n r e s o u r c e s {are] totally
renewable *and% can be economically* socially* culturally and+
environmentally sustainable with prudent planning* management*
utilization*and%marketing@
• Reducing evaporation{from]*the^soil~
• Reducing transpiration{from]plants~
• Moderating extreme temperatures@
Quite often* protection can be combined with production *by^
choosing tree *and% shrub species that* apart {from] furnishing *the^ desired
sheltering effect* yield needed wood products@
Design |of| Windbreaks *and% Shelterbelts
When considering windbreak or shelterbelt planting* three zones
can be recognized**the^windward zone (from which*the^wind blows)~
*the^ leeward zone (on *the^ side{where}*the^ wind passes)~ *and% *the^
protected zone (that *in= which *the^ effect |of| *the^ windbreak or shelterbelt
is felt)@
Fig@ Functioning|of|!a#Windbreak
The effectiveness|of|*the^windbreak or shelterbelt[is]influenced
^by* {its] permeability@ If {it] [is] dense* like !a# solid wall* *the^ airflow *will^
pass over *the^ top |of| {it] *and% cause turbulence on *the^ leeward side due
^to**the^lower pressure on that side~ this gives!a#comparatively limited
zone |of| effective shelter on*the^leeward side compared{to]*the^zone
that!a#moderately permeable shelter creates@ Optimum permeability
*is] 40{to]50 per cent|of|open space* corresponding{to]!a#density |of| 50
^to* 60 per cent*in=vegetation@ Gaps*in=*the^barriers should be avoided@
Permeability|of|dense shelterbelt can be improved{by]pruning lower
branches{at]0@50#0@8 m{from]*the^soil level@
It[is]generally accepted that!a#windbreak or shelterbelt protects
*an& area over!a#distance up{to]{its]own height on*the^windward side
{and} up {to] 20 times {its] height on *the^ leeward side* depending on *the^
strength |of| *the^ wind@ *in= reducing wind speeds* narrow barriers can
be as effective as wide ones@ Furthermore*!a#narrow shelterbelt has
52 Trees*!a#Handbook|of|Forest#Botany
Choice |of| species should not be dictated {by] *the^ value |of| trees *for^
wood* nor planting expenses weighed against *the^ value |of| *the^ wood
produced@ Costs{for}planting*in=parks{for}recreation should take into
account aesthetic benefits *and% *the^ intangible gains *in= *the^ health and+
well#being |of| *the^ people@
Street Planting
Street plantings{are]often*the^responsibility|of|local municipalities
{and}{are]m a d e{to]beautify*the^cities* provide shade*and%control outdoor
noise *and% traffic pollution@ *in= recent years* many |of| *the^ towns and+
cities *in= *the^ world {have] learned through experience that paving
roadways* streets**and%sidewalks does not complete*the^job@*in=m o s t
communities{where}there {has] been *an^ increase *in= population and+
greater congestion |of| motorized traffic* {it] {has] become important that
*the^ city municipality assumes {its] responsibility |of| providing *the^
amenities which add much pleasure {to] life *in= *the^ city@
For use*in=street planting* trees should*
• Be easy{to]establish* preferably with*the^ability{to]be transplanted
as advanced nursery stock* *and% grow relatively quickly {to] *the^
stage that they provide some amenity value~
• Be healthy*in=*the^environment* relatively long#lived*and%not subject
^to* wind#throw or*the^breakage |of| large limbs~
• Be as maintenance#free as possible@ Trees requiring permanent
pruning*and%removal|of|fallen leaves{will]{have]!a#high maintenance
cost@
• The form*and%height|of|*the^species must be suitable{for}*the^width
|of|*the^street*in=which they{are]{to]be planted~
• Whether {to] use !a# single variety or mix on *the^ same street [is] !a#
matter |of| taste@ !a# variety might be used depending on *the^
geographic section |of| *the^ country *and% *the^ width |of| *the^ street@
Greenbelt Planting
Several cities *in= arid zones {have] established *in= their municipal area
green belts with!a#number |of| purposes*
• To enhance*the^beauty|of|*the^site~
• To provide!a#recreation area{for}*the^urban dwellers~
• To reduce *the^ harmful effect |of| dry winds *and% dust storm and+
control sand encroachment@
There[is]!a#wide array |of| trees*and%shrubs{for}greenbelt planting@
A n y p r o g r a m m e |of| this sort* however* should be well designed in
60 Trees*!a#Handbook|of|Forest#Botany
Where water [is] available* several species can be selected@ Within *an&
area each row can be given over {to] one or more species@ Mixtures
^are* thus created*and%{are]considered better as they yield different
produce{to]m e e t local needs@
2
Forest**an^Area|of|Density|of|Trees
Introduction
The forest[is]!a#c o m p l e x e c o s y s t e m consisting m a i n l y |of| trees that
buffer*the^earth*and%support!a#m y r i a d |of| life forms@ *the^ trees help
create !a# special environment which* *in= turn* affects *the^ kinds |of|
animals *and% plants that can exist *in= *the^ forest@ Trees {are] *an^ important
c o m p o n e n t |of| *the^ environment@ T h e y clean*the^air* cool{it]on h o t
days* conserve heat{at]night**and%act as excellent sound absorbers@
Plants provide !a# protective canopy that lessens *the^ impact |of|
raindrops on *the^ soil* thereby reducing soil erosion@ *the^ layer |of|
leaves that fall around*the^tree prevents runoff*and%allows*the^water
^to* percolate into*the^soil@ Roots help{to]h o l d*the^soil*in=place@ D e a d
plants decompose {to] form humus* organic matter that holds *the^
water *and% provides nutrients {to] *the^ soil@ Plants provide habitat *to^
different types |of| organisms@ Birds build their nests on *the^ branches
|of| trees* animals*and%birds live*in=*the^hollows* insects*and%other
organisms live*in=various parts |of| *the^ plant@ T h e y produce large
quantities |of| oxygen *and% take *in= carbon dioxide@ Transpiration *from^
*the^ forests affects *the^ relative humidity *and% precipitation *in= !a# place@
The F A O (Food*and%Agriculture Organization){has]defined forest
as land with tree c r o w n cover (or equivalent stocking level) |of| m o r e
Trees*!a#Handbook|of|Forest#Botany 63
of *the^ Forest Resources Assessment 2000 {were] released {by] FAO *in^
2000@ *in= 1990* temperate *and% boreal forests occupied 1@64 billion ha
{and} tropical forests 1@76 billion ha@!a#b r e a k d o w n{by]ecological zone
w a s available only{for}tropical forests@The figures on tropical forests
s h o w that m o s t (88%){are]*in=lowlands~ |of| these* tropical rain forests
accounted{for}4 7 %|of|all lowland tropical forests* followed{by]moist
deciduous forest (38%)*and%dry*and%very dry formations (15%)@
Natural Forests
The forest area estimates described above include undisturbed
forests* forests modified{by]h u m a n s through use*and%m a n a g e m e n t
(or @seminatural@ forests)*and%forests created artificially{by]humankind
(i@e@ forest plantations){by]afforestation or reforestation@ (Afforestation
*is] defined as*the^establishment|of|!a#tree crop on*an^area {from] w h i c h
^it* {has] always* or {for} !a# very long time* been absent@ Reforestation *is$
defined as *the^ establishment |of| !a# tree crop on forest land@) *in= most
industrialized countries* particularly*in=continental Europe* forests
^are* being managed *in= such !a# way that {at] management#unit level !a#
continuum exists {from] low#intensity management* involving natural
regeneration* t h r o u g h m o r e intensive m e t h o d s i n v o l v i n g s o m e
artificial planting{to]highly intensive methods with complete planting
{and} cultivation~ this m a k e s{it]difficult{to]isolate figures{for}natural
forest*and%plantations@*the^distinction b e t w e e n natural or seminatural
forests*and%forest plantations can more easily be m a d e{for}developing
countries*and%some industrialized countries such as N e w Zealand
*in& which forest plantations {have] b e e n established using introduced
species@
Interest *in= natural forests* particularly their role *in= *the^
conservation |of| biological diversity* {has] led {to] efforts {to] compare
forests today with what [is] thought {to] be their original character and+
^to* give complete protection {to] areas |of| forests which {have] {had] no*
or minimal* human interference@ Although there {are] difficulties *in^
identifying *the^ extent |of| natural forest* compounded {by] problems |of|
definition* some information exists that can be used as *an^ indication
|of| broad patterns |of| natural forests*in=various regions@
A n attempt{has]b e e n m a d e{by]*the^W o r l d W i d e F u n d{for}Nature
(WWF) {to] quantify *the^ area |of| forests *in= western Europe that *has^
been relatively undisturbed {by] humans or which {has] retained much
|of|{its]natural character@ Their report distinguishes b e t w e e n @virgin
66 Trees*!a#Handbook|of|Forest#Botany
of w o o d l a n d {to] m a k e pasture{for}sheep*and%goats*and%*the^cutting
|of| ship timber* particularly {for} *the^ construction *in= *the^ 1660@s |of| !a#
French navy consisting |of| 60 ships#of#the#line *and% 40 frigates@ *in^
Britain* as H@ C@ Darby says* *the^ naval wars with *the^ Dutch *in= *the^
seventeenth century*and%with*the^French*in=*the^eighteenth*and%*the^
Napoleonic wars*in=*the^early nineteenth put such!a#strain on *the^
English oak forests that they {have] never recovered@
W h e n $the far#flung navies melt away$ they take*the^forests with
them@ Ship timber[is]no longer one|of|*the^major requirements* although
wood *in= large amount [is] still needed *in= ships *and% shipping@ {it] *is$
estimated that 350*000 board feet |of| lumber {were] used *in= *the^
construction |of| every Liberty ship built*in=*the^United States during
W o r l d W a r II*and%*the^w o o d that goes into boxes* crating*and%d u n n a g e
on!a#m o d e r n general cargo ship runs into 100*000 b o a r d feet or more@
T o d a y *the^ heaviest d e m a n d {for} industrial w o o d products #¬
lumber* pulp* p l y w o o d*and%*the^like ##[is]on*the^midlatitude forests
|of| N o r t h A m e r i c a*and%Europe@ There[is]!a#chance* however* as far as
*the^ pressure |of| this d e m a n d [is] concerned* that i m p r o v e d forestry
m e t h o d s m a y check any further significant reduction |of| *the^ forest
area *in= these regions@ If *the^ area [is] reduced *in= *the^ coming years *the^
reason [is] more likely {to] be *the^ need |of| more agricultural land* or *the^
n e e d |of| space{for}industrial*and%u r b a n expansion@
T h e old n e e d{for}local construction material*and%*the^relatively
new demand {for} industrial products {are] also felt *in= *the^ tropical and+
subtropical forests *and% one |of| *the^ great uncertainties ahead [is] what
^will* happen {to] them if *and% when they become fully exploited *for^
industrial purposes@ If problems |of| tropical silviculture* management
{and} e c o n o m i c s ## i n c l u d i n g a b o r i g i n a l e c o n o m y ## s h o u l d b e
successfully solved* these forests m a y b e c o m e!a#very great source
|of| w o o d products* b u t if*the^p r o b l e m s{are]not solved*the^exploitation
m a y result*in=*an^e n o r m o u s disaster@ O n e|of|*the^complicating factors
*is] that *the^ most potent pressure on *the^ tropical forest comes *from^
*the^ third |of| *the^ ancient demands* namely *the^ need |of| agricultural
land@
Changes*in=Forest Cover*and%Condition
A n analysis |of| changes*in=*the^cover*and%condition|of|*the^world@s
forests requires!a#differentiation between*
Trees*!a#Handbook|of|Forest#Botany 73
former USSR)* wildfires over large areas |of| forest {have] b e c o m e less
frequent*in=*the^industrialized countries@ Further**the^average area
burned {by] wildfire decreased *in= most regions between 1983#90 and+
1991#94@ There {has] been !a# slight reduction *in= total areas burned *in^
*the^ developed world overall as !a# result |of| improved prevention*
detection*and%control systems* although there{has]b e e n*an^increase
|of| 15% *in= *the^ former USSR@ *in= 1990* *the^ average area |of| forest and+
other wooded land affected annually {by] fires *in= Europe* North
A m e r i c a*and%*the^former U S S R together w a s about 4@26 million h a
or 0@22% |of| their total area |of| forest*and%other w o o d e d land@ Pests
{and} diseases remain constant threats* particularly{to]*the^semi#natural
forests*and%plantations*in=*the^industrialized world@ Trees {are] m o r e
likely {to] b e attacked w h e n under stress* w h e t h e r {from] abnormal
weather conditions such as drought or h i g h temperatures* airborne
pollution* uncontrolled movement |of| germplasm or lack |of|
management@ Recent outbreaks |of| pests *and% diseases that may *have^
been related {to] stress include *the^ decline |of| various species |of| oaks
*in& m a n y parts|of|Europe*and%*in=central Russia**the^recurrent infestation
|of| forests*in=Poland{by]*the^n u n moth**the^attack on b e e c h{by]*the^b e e c h
scale*in=western*and%central Europe* b i r c h*and%ash diebacks*in=north¬
eastern USA**the^littleleaf disease |of| shortleaf pine*in=southern USA*
{and}*the^@x#disease@ |of| pines*in=southern California@
significant reduction *in= wooded areas burned [is] likely {to] occur *in= *the^
near future@
Outbreaks |of| pests *and% diseases *in= developing countries *are^
generally reported {for} those plantations *and% planted trees{where}*the^
impact [is] m o s t apparent@ Introduced pests {are] usually extremely
destructive *and% several |of| them {have] {had] very damaging effects *in^
recent years *in= *the^ developing world@ Examples |of| introduced insects
affecting plantations*and%planted trees include*the^Leu#caena psyllid*
Heteropsylla cubana* which{has]spread into Asia*and%*the^Pacific islands
{and}[is]n o w extending across Africa~*the^cypress aphid* Cinara cupressi*
w h i c h[is]established*in=eight eastern*and%southern African countries
{and} [is] causing heavy mortality among !a# number |of| exotic and+
indigenous species but especially *the^ important plantation species
Cupressus lusitanica* threatening {its] future role *in= *the^ plantation
p r o g r a m m e s |of| these countries~*and%*the^European woodwasp* Sirex
noctilio* w h i c h {has] spread into Argentina* U r u g u a y *and% southern
Brazil* affecting principally Pinus taeda but which may become !a#
threat {for} *the^ large Chilean plantation estate |of| Pinus radiata@
Strategies {to] control introduced insects {from] attacking forest
trees include r e g u l a t o r y measures* eradication* integrated pest
management**and%monitoring |of| population levels*and%occurrence
|of| potentially harmful species@
Effective monitoring |of| insect populations {will] b e especially
important*in=v i e w|of|*the^large plantation p r o g r a m m e s being carried
out {to] produce industrial roundwood* which often rely on one or !a#
few exotic species@ Such programmes may be able {to] afford *the^
investment that [is] necessary {for} control measures@ Smaller#scale
p r o g r a m m e s m a y{have]{to]insure against failure due {to] unexpected
attack {by] insects or diseases {by] diversifying into several species
adapted{to]*the^sites*and%end#uses@
There [is] also evidence |of| forest decline due {to] !a# combination |of|
biotic *and% abiotic factors *in= *the^ developing world* with air pollution
likely {to] play *an^ increasing role *in= some cases as industrial and+
transportation infrastructure develops@ Examples |of| such decline
include n e e m (Azadirachta indica)*in=*the^Sahel* framire (Terminalia
ivoren#sis) *in= I v o r y C o a s t *and% Ghana* *and% Eucalyptus globulus
plantations*in=Colombia*and%Peru* while airborne pollution[is]affecting
forests near large cities*and%industrial areas *in= China (e@g@ Pinus
Trees*!a#Handbook|of|Forest#Botany 79
will be fewer but larger diameter trees* although total volume |of|
timber may be somewhat reduced@ This allows forests {to] be managed
*in& different ways {to] yield different assortments |of| products@ How *to^
model these relationships *and% outcomes *in= detail* beyond *the^ purely
empirical* forms*the^bulk |of| this chapter@
Growth Models
The history |of| forest growth m o d e l s[is]not simply characterized
^by**the^development |of| continuously i m p r o v e d m o d e l s replacing
former inferior ones@ Instead* different m o d e l types with diverse
objectives *and% concepts {were] developed simultaneously@ *the^
objectives *and% structure |of| !a# model reflect *the^ state |of| *the^ respective
research area{at]{its]time*and%d o c u m e n t*the^contemporary approach
^to* forest growth prediction@ *the^ history |of| growth modelling thus
also documents *the^ advancement |of| knowledge *in= *the^ science |of|
forest growth@
Beginning with yield tables{for}large regions as!a#basis{for}taxation
{and} planning* model development led {to] regional yield tables and+
site#specific yield tables *and% culminated *in= *the^ construction |of| growth
simulators{for}*the^evaluation |of| stand development under different
m a n a g e m e n t schemes@ Vanclay (1994) provided*an^overview about
growth*and%yield m a n a g e m e n t models*and%their application{to]m i x e d
tropical forests@*the^1980s brought!a#n e w trend with*the^development
|of| ecophysiological models* which give insight into *the^ complex
causal relationships*in=forest growth*and%predict growth processes
under various ecological conditions@*the^emphasis*in=m o d e l research
^has* shifted towards ecophysiological models*and%away {from] m o d e l s
aimed only {at] providing growth*and%yield information {for} forest
management@ These models attempt {to] simulate forest growth on *the^
basis |of| fundamental ecophysiological processes@ *the^ scientific value
|of| ecophysiological models cannot be overrated~ however* they *will^
not be applied *in= forest management {for} *the^ next few years as they
^are* *in= many ways not yet sufficiently validated@ Also* input and+
output variables do not yet meet *the^ demand |of| forest management
practice@
A major change {has] taken place *in= model conception* i@e@ *the^
understanding |of| forest growth on which *the^ model [is] based@ *the^
tables {by] Weise (1880)* Schwappach *and% Wiedemann resulted *from^
!a# purely descriptive analysis |of| sample area data *in= *the^ form |of| total
82 Trees*!a#Handbook|of|Forest#Botany
To date* model research {has] {had] little success *in= substituting *the^
yield tables {for} pure stands {by] *an^ improved information system *for^
pure *and% mixed stands@ This can *in= no way be attributed {to] !a# deficit
*in& methodological principles* data or technical equipment@ Rather*
*the^ causes lie*in=*the^fact that n e w models{are]not properly adapted
^to* practical requirements@ *the^ recent introduction |of| *the^ growth
model SILVA 2@2 {for} forest management use led {to] !a# range |of|
operational requirements *and% outputs demanded {from] *the^
m a n a g e m e n t models that {will] b e used*in=decision#making processes
^at* stand*and%forest enterprise levels@
1@ The natural management |of| forests [is] currently making great
headway@ *in= *the^ long run only those growth models capable |of|
simulating *the^ growth |of| pure *and% mixed stands |of| all age
compositions*and%structural patterns {will] find approval@
2@ Models need{to]be operable{at]stand*and%forest enterprise levels
{and} able{to]simulate growth behaviour under different thinning
regimes *and% different processes |of| artificial *and% n a t u r a l
regeneration@
3@ Flexibility |of| *the^ model [is] essential so as {to] permit simulation |of|
growth reactions {to] site alterations *and% interference factors on !a#
large regional scale@
4@ Apart {from] tree*and%stand characteristics such as volume
production* assortment yield* wood quality*and%financial yield
should also include structural parameters determining *the^
recreational *and% protective functions |of| forests as well as indicators
showing*the^impact |of| hazards or ecological instability@
5@ Forestry practice[is]interested* first*and%foremost**in=calculating
scenarios{at]stand*and%forest enterprise levels@ This can only be
achieved if input*and%output data|of|*the^model consider what
information [is] available*and%which data{are]needed *in= forestry
practice@ Furthermore* achieving this goal also depends on whether
*the^ model forms part|of|!a#comprehensive forestry information
system and* lastly* whether hardware specifications{are]acceptable
*in& practice@
For decades forestry practice {has] b e e n hoping {for} i m p r o v e d
growth models {to] assist with research* planning* operations and+
control *in= forest management@ *the^ general acceptance |of| new models
^by* practitioners calls{for}close cooperation b e t w e e n forest science
{and} forest practice*{from]*the^design*and%development|of|*the^m o d e l
^to*{its]actual introduction*in=forest management@
96 Trees*!a#Handbook|of|Forest#Botany
Trees*in=*the^Urban Environment
Trees*in=*the^urban e n v i r o n m e n t{are]often referred{to]as*the^urban
forest* comprising trees*in=civic woodlands* parks*and%*the^street@
Earlier* urban trees {were] mainly regarded as aesthetic elements*
w h e r e a s today they {are] recognized as h a v i n g!a#positive impact on
*the^ e n v i r o n m e n t as well as providing e c o n o m i c*and%social benefits@
Monetary evaluations reflect *the^ various benefits arising {from] *the^
urban forest* covering such aspects as reduction |of| pollution and+
energy use* environmental amelioration* savings *in= public health
care *and% increase *in= economic investment@ Hence* *the^ value |of| *the^
urban forest [is] being increasingly recognized as !a# vital component
*in&*the^m a i n t e n a n c e |of| !a# sustainable urban e n v i r o n m e n t *in= cities
around*the^world@ Meanwhile**the^population living*in=urban areas
^has* increased rapidly since*the^1950s*and%*the^lack |of| space m a k e s
^it* tempting {to] use green areas{for}infrastructure*and%buildings@
Collins (1997) e m p h a s i z e s that u r b a n forestry* *the^ planning*
98 Trees*!a#Handbook|of|Forest#Botany
Introduction
Trees represent m a n y different families within *the^ t w o major
divisions |of| *the^ spermatophytes or seed#bearing plants $ *the^
gymnosperms *and% *the^ angiosperms@ Trees |of| *the^ gymnosperm group
^are* more commonly known as conifers or softwoods* while those |of|
*the^ angiosperm group {are] often called broadleaf trees or hardwoods@
T h e s e n a m e s do n o t a p p l y u n i v e r s a l l y $ *the^ w o o d |of| s o m e
gymnosperms*{for}example*[is]considerably harder than that|of|certain
angiosperms@ {for} most practical purposes* however* *the^ forests |of|
*the^ world {are] classified as softwoods* temperate hardwoods and+
tropical hardwoods@
The dominance|of|*the^softwoods*in=*the^northern temperate forests
*is]|of|special e c o n o m i c significance* b e c a u s e m a n{has]found that these
w o o d s {are] particularly useful@ T h u s *the^ s a w timber that [is] u s e d
throughout*the^w o r l d*in=general building*and%construction[is]75 per
cent softwood*and%*an^e v e n larger proportion |of| w o o d pulp c o m e s
^from* this class |of| trees@
Bole or trunk@ T h r o u g h o u t*the^life|of|*the^tree there{are]alternate
periods|of|growth*and%d o r m a n c y*and%*in=general* owing{to]differences
*in&*the^structure|of|*the^w o o d p r o d u c e d{at]*the^b e g i n n i n g*and%{at]the
106 Trees*!a#Handbook|of|Forest#Botany
end |of| !a# growing season* *the^ successive layers appear as !a# series |of|
concentric circles when *the^ tree [is] cut transversely@{where}climatic
conditions{are]such that!a#period |of| growth alternates with!a#period
|of| d o r m a n c y within *the^ year* these w o o d y layers {are] k n o w n as
annual rings@*the^yearly increase*in=w o o d v o l u m e depends primarily
on diameter growth@ {for} *the^ first few years after {its] formation *the^
wood* then known as $sapwood*$ contains some living cells which
store food substances*and%conduct water*and%solutions |of| mineral
nutrients{from]*the^roots{to]*the^c r o w n|of|*the^tree@ Later*the^cells die*
other physiological changes take place*and%*the^wood*{at]this stage
known as $heartwood*$ ceases {to] function except *in= support |of| *the^
crown@ Quite commonly *the^ heartwood [is] darker *in= colour* more
durable*and%{has]better timber properties than*the^sapwood@
The physical structure*and%consequently*the^density*and%technical
properties|of|*the^w o o d laid d o w n{by]*the^cambium* vary considerably
^from* one species{to]another@ *the^ rate |of| diameter growth m a y also
^have* diverse influences on *the^ strength*and%quality |of| *the^ wood*
while rapid growth increases *the^ strength |of| certain hardwoods* *it^
may produce soft* spongy* Jow#grade timber *in= some |of| *the^ softwoods@
The form|of|*the^bole generally varies according{to]*the^strength
required{to]support*the^c r o w n*and%resist w i n d pressures**in=!a#closed
forest*the^bole approaches!a#cylindrical form* whereas*in=e x p o s e d
sites{it][is]m o r e conical owing{to]*the^n e e d{for}mechanical support@
There {are] also considerable differences b e t w e e n species* especially
between *the^ two major groups |of| hardwoods *and% softwoods* *in= *the^
form |of| *the^ crown *and% bole *and% *in= *the^ vigour |of| development and+
persistence |of| side branches@
Usually softwoods produce!a#well#defined bole with relatively
small branches* while *the^ branches |of| many |of| *the^ hardwoods *are^
larger *and% more widely spreading@ Since branches produce knots* *for^
*the^ production |of| clear timber{it][is]desirable{to]r e m o v e *the^ lower
dead*and%dying b r a n c h e s while they{are]fairly small@*in=general* trees
growing*in=!a#closed forest {have] fewer*and%smaller b r a n c h e s than
those growing*in=*the^open@ However**the^rate{at]w h i c h lower branches
^are* killed{by]shade*and%subsequently drop{to]*the^ground varies with
*the^ species@
Important characteristics|of|*the^mature root system also vary
^from* one species{to]another@ S o m e species {have] deeply penetrating
Trees*!a#Handbook|of|Forest#Botany 107
Nutritional*and%Ecological Requirements
For{its]g r o w t h*and%{for}other life processes**the^tree requires food
{and} like other green plants{it]builds u p{its]food materials {from] simple
substances@ *the^ chemical constituents|of|!a#tree {are] chiefly carbon*
h y d r o g e n *and% oxygen* together with s o m e nitrogen*and%certain
minerals such as potash* lime* iron *and% phosphorus@ *the^ carbon *is$
obtained {from] *the^ small amount |of| carbon dioxide gas that [is] always
present*in=*the^atmosphere*and%enters*the^plant through small openings
*in& *the^ leaf surface known as $stomata@$ Oxygen *and% hydrogen *are^
obtained {from] *the^ soil through *the^ root system *in= *the^ form |of| water
carrying nitrogen (nitrates) *and% mineral nutrients *in= solution@ *the^
synthesizing |of| these elements {to] form substances that *the^ tree requires
#in particular *the^ combination |of| carbon* hydrogen *and% oxygen *to^
form carbohydrates $ takes place mainly *in= *the^ leaves *in= *the^ presence
|of|*the^characteristically green matter (chlorophyll)*and%|of|sunlight*
which supplies *the^ necessary energy@ Air* water* chlorophyll and+
light {are] therefore essential {to] *the^ production |of| *the^ carbohydrates#
*in& particular cellulose*and%lignin #which form {by] far *the^ greater
portion|of|*the^tree@ Soil moisture[is]especially important* as!a#deficiency
|of| water inhibits photosynthesis@ Tree species differ markedly *in^
their optimum water requirements *and% also *in= their sensitivity *to^
departures {from] *the^ optimum@
Tree species also differ greatly*in=*the^a m o u n t|of|light they require*
or* alternatively* *in= *the^ degree |of| shade they {will] tolerate@ {it] *is$
customary {to] classify species as shade#enduring or tolerant (of shade)
{and} light#demanding or intolerant (of shade)* b u t{for}m o s t species
*the^ younger *the^ tree *and% *the^ better *the^ soil quality on which {it] *is$
growing* *the^ greater {its] tolerance |of| shade@
108 Trees*!a#Handbook|of|Forest#Botany
be cut through *the^ forest {to] break *the^ continuity |of| tree growth and+
delay or prevent *the^ spread |of| fires@ Clear#cut strips |of| this nature
^are* known as $fireguards@$ Fire itself[is]sometimes used as!a#preventive
measure{where}*the^annual fall |of| litter*and%*the^dying b a c k |of| ground
vegetation produce large accumulations |of| inflammable debris@ *the^
$early burning$ |of| this material when moisture conditions {are] such
as{to]prevent too rapid spread |of| fire {will] reduce*the^danger during
!a# dry period later*in=*the^season@
Recent important advances*in=*the^field |of| fire protection include
*the^ use |of| aircraft* mainly{for}detection*and%|of|meteorological data
^to* appraise more precisely*the^existing degree |of| fire hazard* so that
d a n g e r o u s conditions can b e quickly r e c o g n i z e d *and% adequate
preventive measures taken@
In some European countries forest fires{are]rare*and%such as do
occur cause negligible damage@ *in= Norway* {for} example* forest
properties m a y b e insured against fire{at]!a#low p r e m i u m rate@ There
^are*!a#n u m b e r |of| reasons{for}this* (a)*in=general*the^forests{are]easily
accessible*and%under intensive management* with annual fellings
well distributed*in=!a#large n u m b e r |of| small scattered holdings~ (b)
*the^ forest consciousness |of| *the^ people [is] highly developed~ (c) *the^
climatic conditions tend {to] be favorable* with well#distributed summer
rains*and%*an^absence |of| prolonged hot* dry periods@
Despite*the^damage{it]causes* fire cannot be looked u p o n solely
as*an^e n e m y|of|*the^forest@ Under certain conditions controlled burning
m a y prove !a# useful tool*in=securing *the^ regeneration |of| valuable
forest trees~{it]m a y prepare!a#suitable seedbed* eliminate competing
vegetation* or assist *in= *the^ distribution |of| seed@ Certain conifers* *for^
example* {have] serotinous cones *and% considerable heat [is] required
^to* open them*and%release *the^ seed* these species m a y reproduce
abundantly after fire*and%*the^resulting stand [is] often referred {to] as
!a# $fire type@$ Extensive even#aged coniferous forests|of|*the^northern
t e m p e r a t e r e g i o n o w e their o r i g i n {to] fire* as d o m a n y o t h e r
commercially important forests*in=various parts|of|*the^world@
Insects that d a m a g e*and%destroy trees*and%*the^fungi*and%other
organisms that cause tree diseases* {are] present *in= all forests@ *in^
endemic proportions *and% *in= balance with their environment* they *are^
!a# necessary *and% sometimes innocuous feature |of| *the^ forest community@
Even under these normal conditions* however* some insects and
112 Trees*!a#Handbook|of|Forest#Botany
Reproduction
A tree crop m a y be established{by]artificial or natural means* or
^by* !a# combination |of| *the^ two@ *in= artificial reproduction* seed *is$
sometimes sown* but more commonly young trees that {have] been
g r o w n*in=nurseries {are] planted on*the^site@*in=either case**the^seed
used [is] collected*in=*the^forest* preferably {from] trees |of| good form
{and} vigour@ When *the^ crop [is] established {by] natural means* *the^
young trees may originate vegetatively or {from] seed disseminated
either before felling takes place or {by] trees left standing {for} *the^
purpose {at] *the^ time |of| logging@ !a# new crop |of| young trees that
b e c o m e s established*in=*the^forest before cutting takes place[is]k n o w n
as $advance growth@$
There {are] t w o principal m e t h o d s |of| cutting *the^ mature forest*
each |of| which {has] advantages with respect {to] *the^ regeneration and+
establishment |of| individual tree species*
Clear#Cutting
B y this method* w h i c h results*in=*the^development |of| m o r e or less
even#aged stands**the^forest[is]cut*in=one operation or*in=!a#succession
|of| operations* each |of| w h i c h r e m o v e s either!a#portion |of| *the^ trees
scattered over *the^ whole area or all |of| *the^ trees {from] !a# part |of| *the^
area~ these operations {are] continued until *the^ original stand *is$
completely cleared@ Regeneration may be established {by] natural or
artificial means**the^former being provided {for} either {by] advance
growth* or{by]seed lying*in=*the^surface soil{at]*the^time |of| cutting or
m a d e available{from]*the^crowns |of| felled trees or {from] standing trees
surrounding*the^cutover area@
For species that require overhead protection *in= their youth !a#
$shelterwood cutting$ [is] made* *the^ stand [is] opened up gradually
over !a# period |of| years {by] m a k i n g three or four successive cuts
periodically over*the^whole area until*the^original stand[is]removed@
^for* other species* *the^ seed source may be *the^ critical factor *and% *the^
stand [is] clear#cut except {for} !a# few mature trees* evenly distributed
over*the^area* which{are]preserved{to]provide*the^seed{for}*the^next
crop@ This[is]k n o w n as $seed#tree cutting@$ Quite commonly* portions
|of|!a#stand {are] clear#cut periodically*in=*the^form |of| n a r r o w strips*
wedge#shaped blocks* or patches* until*the^stand{has]b e e n completely
felled@
The Douglas#fir forests on*the^Pacific coast|of|North America* for
114 Trees*!a#Handbook|of|Forest#Botany
Afforestation
Forestry not only seeks{to]ensure*the^reproduction |of| existing
forests* b u t [is] also concerned with*the^p r o b l e m |of| establishing n e w
forests@ Afforestation m a y b e u n d e r t a k e n (a){to]reclaim forest land
cleared{for}other purposes b u t best suited{for}*the^growing |of| forest
crops* (b) {to] establish forests as *an^ e c o n o m i c crop o n land n o t
previously forested* (c){to]stabilize shifting sand*and%eroding soils*
(d) {to] provide protection {for} agricultural*and%other crops against
w i n d*and%s n o w d a m a g e*and%(e){to]i m p r o v e seepage*in=drainage areas
{and} so stabilize stream flow@
Great Britain* Australia* N e w Z e a l a n d*and%South Africa {have] all
undertaken large#scale afforestation with*the^object |of| supplementing
their natural resources@ *the^ trees u s e d {have] b e e n predominantly
Trees*!a#Handbook|of|Forest#Botany 115
Botanical Names
T o n a m e !a# thing [is] *the^ next best thing {to] understanding it@
Indeed* unless y o u n a m e it* y o u {will] not understand it@ Nomina si
nescis* perit et cognitio rerum* wrote*the^father |of| botanical names* if
y o u k n o w not names**the^k n o w l e d g e also |of| things [is] lost@ N a m e s
^are**the^tools with w h i c h our thoughts grasp {at] nature@
The first m a n{to]n a m e plants[is]said{to]{have]b e e n Adam@ T h o u g h
w e {have] n o catalogue |of| *the^ n a m e s w h i c h he invented* w e m a y
suppose that they{were]*the^equivalents**in=his speech*|of|n a m e s given
^by* our o w n ancestors* apple* daisy* oak* cowslip@ H e probably used
n a m e s m u c h as w e ordinarily do**in=!a#rather general way~ fully
r e c o g n i z i n g that {for} certain p u r p o s e s {it] m i g h t b e desirable to
116 Trees*!a#Handbook|of|Forest#Botany
Fig@ Heptica
The science |of| b o t a n y{had]{its]roots*in=*the^n e e d{for}descriptions
|of|*the^plants{by]w h i c h they might b e recognized@ *the^ early herbals
^were* anonymous* compilations |of| c o m m o n beliefs* old wives@ tales*
sometimes with crude*and%inaccurate illustrations@ Later herbalists
b e g a n{to]devote their efforts{to]careful examination*and%description
|of| plants~ their writings began {to] {have] scientific value@ These *were^
*the^ botanists {from] whom our names {for} plants {are] descended@
Naturally they wrote*in=Latin@ Their w o r k w a s often translated
into *the^ vulgar tongues{for}*the^use |of| *the^ unlearned*and%if they
occasionally wrote first*in=*the^vernacular* their writings appeared
also*in=!a#Latin version@ So{have]*the^familiar Latin n a m e s c o m e d o w n
^to* us* Rosa* Lilium* Viola* Delphinium* Cornus* Quercus* Vitis and+
Greek names which *the^ Romans {had] adopted *and% spelled *in= their
o w n way* Hyacinthus* Narcissus* Lotus@
One |of| *the^ first rewards |of| persistent inquiry into nature [is] !a#
distinction between different orders |of| likeness@ All trees {are] alike*
*in&!a#general way@ All oaks{are]m u c h m o r e so@*and%a m o n g oaks w e
can pick out white oaks* red oaks* b l a c k oaks* b u r oaks*and%m a n y
others@ {it] {has] b e e n s h o w n that e v e n primitive m a n m a d e such
Trees*!a#Handbook|of|Forest#Botany 119
Fig@ Linnaea
It [is] sufficient {for} ordinary knowledge {to] name *the^ genus and+
*the^ fact that one can do so should be received with proper acclamation@
Even*the^recognition|of|*the^family represents precise*and%detailed
knowledge@ M a n y amateur gardeners *and% botanists {are] i n d e e d
interested *in= arriving {at] such knowledge |of| their plants *and% *are^
thwarted {by] *the^ professional growers* some |of| whom {are] interested
rather*in=*the^m o r e internecine aspects |of| their art@ Horticultural
names* consequently* b o r d e r on chaos@
The principle |of| hierarchical classification[is]extremely simple
{and}[is]*in=constant use{for}all sorts |of| objects@ A m o n g vehicles w e
readily distinguish*the^horse#drawn*and%*the^mechanically propelled~
a m o n g*the^latter w e can classify some as using gasoline* others fuel
oil* still others coal or alcohol@ Each |of| these groups can*in=turn b e
b r o k e n down@
So we {have] !a# n u m b e r |of| species |of| roses* R o s a carolina* Rosa
setigera* Rosa palustris *and% *the^ rest* which together make up *the^
genus Rosa@ This genus *and% other genera such as Potentilla* Spiraea*
Fragaria make up *the^ rose family* Rosaceae@ This family *and% *the^
saxifrage family *and% *the^ bean family *and% some others compose *the^
rose order* *the^ Rosales@ O n *the^ basis |of| certain structural features
m a n y orders* i n c l u d i n g *the^ Rosales* {are] p l a c e d *in= *the^ class
Dicotyledones@ These *in= turn form !a# part |of| *the^ Angiospermae*
flowering plants~ one|of|*the^major divisions |of| *the^ plant world@
There {are] only!a#few |of| these divisions~*the^one just n a m e d is
126 Trees*!a#Handbook|of|Forest#Botany
by far *the^ largest@ More or less related {to] {it] [is] *the^ division called *the^
Gymnospermae* which includes those plants commonly known as
conifers* pines* spruces* hemlocks* firs~*the^cycads also*and%!a#few
other r e m n a n t s |of| ancient tribes* {are] usually placed here@ !a# third
division (Filices) holds *the^ ferns* ancient*and%modern@ !a# fourth
(Bryophyta) [is] made |of| *the^ mosses *and% their relatives@ *the^ fifth *is$
*the^ Algae* those mostly small but sometimes very large *and% variously
coloured habitants |of| ocean* lakes*and%streams*and%*the^sixth division
*is] c o m p o s e d |of| plants without chlorophyll**the^Fungi@
Clearcutting
In clearcutting* *an^ entire stand |of| trees [is] r e m o v e d *in= one
operation@{from]*the^forester@s point|of|view* clearcutting[is]*the^easiest
w a y{to]m a n a g e!a#forest#and*the^most economical@ Regeneration m a y
come{from]sprouts on stumps* {from] seedlings that survive*the^logging
operation* or {from] seeds that germinate after*the^harvest@ If natural
regeneration [is] delayed longer than desired**the^area[is]planted or
seeded@
Clearcutting systems {are] often used {to] m a n a g e fast#growing
species that require!a#lot |of| light@ Resulting stands {are] even aged
because all*the^trees*in=*an^area{are]cut#and regenerated#at*the^same
time@ Clearcutting{has]b e c o m e controversial*in=recent years because
^it* {has] *the^ potential {to] damage watersheds *and% because {it] tends *to^
eliminate species |of| wildlife dependent on old growth trees@ If clearcuts
^are* kept small*and%*the^cutting interval [is] long enough* however*
biological diversity m a y not b e impaired@
Shelterwood
In shelterwood systems* *the^ forest canopy [is] removed over !a#
period |of| years* usually *in= two cuttings@ After *the^ first harvest*
natural regeneration begins*in=*the^understory@{by]*the^time*the^second
harvest[is]made* e n o u g h y o u n g trees {have] g r o w n{to]assure adequate
regeneration@ Shelterwood systems favor species that{are]intermediate
*in& tolerance{to]shade@ Such systems{are]difficult{to]use successfully
{and}{are]*the^least used|of|*the^three silvicultural m e t h o d s described@
Multiple#Use Forestry
Gifford Pinchot**the^first Chief|of|*the^U@S@ Forest Service* w a s also
this country@s first professional forester@ Pinchot advocated*the^use
|of| forest resources#all resources* not just timber#for h u m a n benefit@
Pinchot w a s!a#strong*and%charismatic leader**and%his ideas helped
shape*the^course |of| forestry*in=*the^United States@ Pinchot{had]!a#vocal
opponent *in= J o h n Muir*!a#y o u n g naturalist {from] California w h o
believed that public lands should b e preserved rather than used@
Eventually M u i r*and%Pinchot b e c a m e rivals {for} public approval@
O d d l y enough* there w a s n o
loser*in=this early conservation battle@ Muir@s preservation ethic
became embodied *in= *the^ philosophy |of| *the^ National Parks* and+
Pinchot@s concept |of| wise use became *the^ guiding principle |of| *the^
National Forests@ National Forests {are] still m a n a g e d u n d e r the
Trees*!a#Handbook|of|Forest#Botany 129
concepts |of| multiple use *and% sustained yield@ *the^ dominant uses |of|
National Forests {are] considered {to] be wood* water* wildlife* forage
(for domestic cattle*and%wildlife)**and%recreation@
E x t r a c t i o n |of| m i n e r a l s *and% other v a l u a b l e p r o d u c t s [is] also
considered!a#legitimate use |of| National Forests@ Because Pinchot@s
philosophy left r o o m{for}*the^$highest*and%best use$|of|!a#given area*
*the^ U@S@ National Forests n o w include!a#wilderness system |of| m o r e
than 32 million acres (13 million ha)*in=w h i c h timber harvest [is] not
allowed@ T o d a y {it] [is] generally r e c o g n i z e d that most* if n o t all*
nondestructive uses |of| forest{are]valid@ S o m e areas m a y b e set aside
as parks~ others {for} wildlife habitat or as wilderness@ Still others *will^
be managed {for} timber harvest or multiple benefits@ Today* conflicts
arise primarily over{where}these different uses {will] be dominant@ *in^
*the^ National Forests* such decisions {are] made through !a# land#use
planning process *in= which *the^ public {has] ample opportunities *for^
input *and% involvement@
Forestry Research
At *the^ turn |of| *the^ century* very little was known about *the^
world@s native forests or how {to] manage them@ *in= *the^ United States*
foresters {were] quick {to] recognize *the^ value |of| information about
forests*and%!a#b r a n c h |of| research w a s established*in=*the^Forest Service
*in& 1915@ Early research w a s done primarily*in=support |of| reforestation
efforts* but* as forestry grew *in= size *and% complexity* so did *the^
research@ Today* *the^ USDA Forest Service {has] six regional experiment
stations located*in=important forest regions@ E a c h experiment station
^has* several field laboratories generally with specialized assignments
^for*!a#geographic region or!a#specific subject area**and%n u m e r o u s sites
^for* field research@ *in= addition* *the^ Forest Products Laboratory *in^
Madison* WI* serves as !a# nationwide centre {for} research and+
d e v e l o p m e n t |of| n e w technology relating{to]wood* including tropical
woods@ T w o laboratories{are]dedicated exclusively{to]tropical forest
research**the^International Institute |of| Tropical Forestry *in= Puerto
Rico*and%*the^Institute |of| Pacific Islands Forestry*in=Hawaii@
Research [is] vital {for} modern forest management* which *is$
information intensive@ Today@s foresters require vast quantities |of|
data *and% !a# knowledge |of| ecology* they must understand not only
*the^ parts |of| ecosystems b u t h o w different parts|of|*the^e n v i r o n m e n t
130 Trees*!a#Handbook|of|Forest#Botany
Forest Values
O n e|of|*the^p r o b l e m s *the^ U C S{has]identified [is] that *the^ public
m u s t b e better educated as{to]*the^valuable services that forests
provide* such as biodiversity* carbon storage**and%water purification@
O n e|of|*the^major issues that m u s t b e faced today[is]*the^economics#
w h e n!a#quick sale can b e m a d e o n*the^l u m b e r or other goods taken
^from*!a#forest*{it][is]h a r d{to]convince people{to]turn a w a y*and%choose
^to* protect *an^ environment that helps everyone but [is] intangible@ *the^
market#based versus environmental# based issues create some |of| *the^
biggest challenges*in=dealing with global w a r m i n g today@
Effects |of| Global Warming
N o t all forests{will]{have]*the^s a m e o u t c o m e s under*the^influence
|of| global warming *and% changes *in= precipitation@ Some forests *will^
die back* while others may extend their ranges@ Amounts |of| CO *will^ 2
It [is] expected that temperate forests *in= *the^ United States *will^
migrate northward {from] 62#329 miles (100#530 km) over *the^ course
|of|*the^next century@ If*the^air temperature w a r m s 3@3°F (2°C) over
*the^ same period* models {have] predicted that tree species {will] *have^
^to* migrate 1#3 miles (1@6#4@8 km) per year* which [is] too fast {for} most
temperate species* except{for}those w h o s e seeds{are]carried{by]birds
over greater distances@
It[is]expected that grasslands {will] dominate m a n y |of| these areas@
*the^ wildlife inhabiting *the^ temperate forests {will] also be affected *by^
loss |of| habitat* drought* *and% wildfires@ Diseases {will] also weaken
them@*the^National A s s e s s m e n t Synthesis T e a m (NAST)|of|*the^U@S@
Global Change Research Programme* some forest models predict *an&
overall increase *in= forest productivity with increased temperatures@
T h e y also clarify that other environmental factors* such as severe
weather* could offset any productivity@ N A S T also states that!a#great
deal |of| biodiversity change [is] expected*in=*the^United States@
The maple/beech/birch forests *in= *the^ Appalachian Range *from^
New England {to] West Virginia could completely disappear@ *in= *the^
U p p e r Great Lakes region* aspen* birch**and%red spruce could b e
obliterated@ If global w a r m i n g occurs gradually* m o d e l s suggest that
oak/pine*and%oak/hickory forests{will]replace these threatened areas@
If* however* conditions {are] b a d enough* entire vegetation species
could b e lost@ N A S T also found that w a r m i n g*in=cooler a r e a s $ s u c h
as *the^ northern United States *and% western mountains near *the^
Canadian border$ {will] increase tree richness *and% provide !a# better
habitat{for}reptiles*and%amphibians@ These same m o d e l s also predict
!a# decrease*in=bird*and%m a m m a l richness*in=*the^eastern United States@
N A S T m o d e l s {have] predicted that temperate forests*in=Scandinavia
^will* m o v e n o r t h w a r d*and%replace other forest communities@
Recent Developments
Four e x a m p l e s|of|modern#day forested areas suffering*the^effects
|of| global warming follow@ *in= July 7* 2006* report *in= USA Today* *the^
frequency *and% size |of| large forest fires {have] increased dramatically
*in& *the^ last 20 years* *and% global warming [is] being blamed as part |of|
*the^ cause@ *the^ western region |of| *the^ United States {has] seen *an&
increase*in=wildfire occurrences since 1987~*the^wildfire season[is]n o w
two *and% !a# half months longer than {it] was back then@ Part |of| *the^
problem [is] that spring warm#up [is] happening earlier *and% more quickly
134 Trees*!a#Handbook|of|Forest#Botany
Controversial Findings
A study conducted{by]*the^Carnegie Institution*in=Washington*
D@C@**and%*the^L a w r e n c e Livermore National Laboratory*in=California
led their scientists {to] conclude that forests *in= temperate regions
actually could w o r s e n global warming@ Using c o m p l e x modeling
software {to] simulate changes *in= forest cover*and%their effects on
global climate* *the^ results they obtained {from] their model *were^
surprising@ Ken Caldeira {from] *the^ Carnegie Institution remarked*$We
^were* hoping{to]find that growing forests*in=*the^United States w o u l d
help slow global warming@ But if w e{are]not careful* growing forests
Trees*!a#Handbook|of|Forest#Botany 137
Silviculture and+
Management *in= Forestry
Silviculture*and%Management
A l t h o u g h silviculture*and%m a n a g e m e n t m a y b e looked u p o n as
t w o b r a n c h e s |of| forestry**the^one m a i n l y technical*and%*the^other
e c o n o m i c * *the^ t w o {are] i n t e r d e p e n d e n t *and% m u s t a l w a y s b e
coordinated@ While silviculture may establish *the^ ideal possibility |of|
production within !a# forest* management determines *the^ degree *to^
which this can be realized through regulation *and% within *the^ bounds
|of| good business practice@ Forest management may be defined as *the^
application |of| business methods *and% technical forestry principles *to^
*the^ operation |of| !a# forest property@
C o m p a r e d with other crops*!a#forest requires!a#relatively long
time between {its] establishment *and% {its] harvesting@ Revenue {from] *an&
unmanaged forest may accrue only {at] fairly long intervals@ Managed
forests {are] organized{to]ensure!a#sustained yield|of|*the^forest crop
*in& perpetuity@ *the^ income should include !a# reasonable profit on *the^
investment* annually if possible@
S u s t a i n e d y i e l d |of| t i m b e r d e p e n d s u p o n *the^ s y s t e m a t i c
reproduction |of| !a# crop as{it][is]harvested@ {it] also requires that the
Trees*!a#Handbook|of|Forest#Botany 141
Silviculture
In essence* silviculture [is] applied forest e c o l o g y*and%forest
genetics~{it]m a y b e described as*the^d e v e l o p m e n t*and%use |of| cutting
m e t h o d s*and%cultural treatments|of|*the^forest*
• To ensure adequate regeneration |of| desirable species as soon as
possible after*the^mature stand[is]cut*
• To bring about conditions favorable {to] *the^ optimum yield and+
quality |of| production *in= keeping with *the^ objectives |of| management
{and}*the^condition|of|*the^forest
• To maintain*and%{where}possibleimprove*the^quality or productivity
|of|*the^site@
Engineering |of| Logging
Logging engineering [is] concerned essentially with *the^ felling and+
removal |of| timber {from] !a# forest area@ Even{where}*the^ sole purpose
*is] liquidation|of|*the^resource* logging techniques m a y b e developed
^to*!a#h i g h degree |of| efficiency* as they{have]!a#direct*and%important
bearing o n*the^cost |of| extraction@*in=forests u n d e r sustained#yield
management* logging methods must be developed {to] meet *the^
requirements |of| *the^ management plan as well as {to] maintain
operational efficiency@
L o g g i n g e v e r y w h e r e presents m a n y technical problems* b u t
particularly{where}*the^ forests {are] difficult |of| access* distances *are^
great *and% *the^ topography [is] rugged@ *in= many forest regions logs *are^
transported long distances*and%{at]relatively l o w cost {by] natural
water systems*and%*in=*the^northern forests winter snows often facilitate
*the^ hauling |of| forest products@
F r o m*the^point |of| v i e w |of| forest conservation* p e r h a p s*the^m o s t
important logging objectives are*
Trees*!a#Handbook|of|Forest#Botany 143
Wood Utilization*in=Forest
While {it] [is] not usual {for} those responsible {for} *the^ growing and+
harvesting |of| timber crops {to] be directly concerned with their
conversion*and%manufacture into usable products* there should b e
!a# close liaison between *the^ two operations |of| forest production and+
wood utilization@
Wood {has] inherent qualities that make {it] particularly suitable *for^
many purposes$good strength *in= relation {to] weight* pleasing
appearance* insulation against heat*and%sound* ease |of| fastening
with nails* dowels* screws* or glue*and%case|of|working with relatively
simple tools@ However* w o o d[is]{by]n o m e a n s!a#h o m o g e n e o u s material*
^it* differs {from] one species {to] another *in= structure *and% physical
properties*and%h e n c e*in=*the^use{for}w h i c h{it][is]best suited@
Within environmental limitations{it][is]desirable{to]grow*the^species
that {will] best m e e t*the^requirements |of| local industry*and%available
export markets@
Furthermore*{to]m a k e*the^fullest u s e|of|*the^timber crop {at] all
stages |of| d e v e l o p m e n t*and%{to]m i n i m i z e logging*and%manufacturing
waste*{it][is]important {to] co#ordinate as closely as possible forest
operations*and%*the^wood#using industries*and%also{to]bring about
!a# closer integration|of|*the^wood#using industries themselves $ as*
^for* example**the^sawmill*and%pulp industries@
Forest Policy
In m a n y countries forests{are]|of|vital importance{to]*the^over#all
economy* not only *in= relation {to] industrial development* but also *in^
providing benefits which may be difficult {to] evaluate *in= monetary
terms@ {for} this reason *and% also because |of| *the^ long#term nature |of|
*the^ forest enterprise* most countries seek {to] ensure *the^ perpetuation
|of| their forests*and%{to]safeguard*the^interests |of| present*and%future
generations{by]placing*the^forests under some degree |of| g o v e r n m e n t
control@
144 Trees*!a#Handbook|of|Forest#Botany
making@ Excellent results {have] been obtained {by] application |of| *the^
Dauerwald idea *in= some |of| *the^ European forests* but Dauerwald*
w h i c h requires* so {to] say* Dauerpflege or continuous*and%careful
attention{to]details*and%m u c h h a n d labour* is*{for}e c o n o m i c reasons*
impossible*in=m o s t|of|*the^world*and%certainly impossible on b o t h
small*and%large forest holdings*in=North America@
Every forest operator k n o w s that he cannot always follow even
*the^ best advice|of|*the^silviculturist*{for}he w o u l d go b r o k e if he tried@
Clear#cut patches*in=*the^Pacific Northwest on w h i c h Douglas#fir
seedlings should be coming up {are] *in= many regions taken over *by^
alder or choked with brushy vegetation~ some |of| *the^ young stands
n e e d thinning~*and%m a n y trees should b e p r u n e d{for}best results@
O n l y *the^ larger c o m p a n i e s {are] able {to] perform i m p r o v e m e n t
operations |of| this type*{for}m o n e y spent on pruning* thinning* or
fighting brush might not be returned until 75 or 100 years *have^
elapsed *and% under present circumstances most |of| *the^ operators *are^
unable{to]finance such long#term investments@
The result[is]that m u c h|of|*the^forest[is]growing u p unattended
{and}*the^land [is] not producing so m u c h g o o d timber as{it]might@
According {to] *the^ Annual Report |of| *the^ Forest Experiment Station *at^
Portland *in= 1952* *the^ use |of| pulp chips obtained {from] mill waste *in^
O r e g o n*and%W a s h i n g t o n{has]m e a n t*an^annual salvaging |of| material
*in& recent years equivalent {to] several hundred million board feet |of|
logs* !a# very good saving* but {at] *the^ same time *the^ practice {has] *had^
!a# detrimental effect on *the^ market {for} pulpwood obtained {by] thinning*
m a k i n g g o o d forest m a n a g e m e n t m o r e difficult@ Thus* w h a t [is] g o o d
e c o n o m y{at]*the^mill[is]not always good silviculture*in=*the^forest@
Cultural Promises *and% Blocks
The outlook can b e s u m m a r i z e d as follows* A l t h o u g h shrinking*
*the^ area|of|*the^accessible forest [is] large e n o u g h*and%our technical
skill* even though p r o b l e m s remain*[is]n o w g o o d e n o u g h {to] supply
all m a n k i n d with!a#reasonably adequate a m o u n t |of| w o o d*in=*the^near
future* provided *the^ skill be freely applied over *the^ area *and% *the^
yield |of| *the^ forest be equitably distributed@ But deeply entrenched
cultural factors interfere with both *the^ application |of| *the^ skill and+
*the^ distribution |of| *the^ yield@ This [is] *the^ root |of| *the^ forestry problem
{and}*the^m o s t difficult part|of|it@ W e k n o w h o w{to]change*the^n u m b e r
|of| c h r o m o s o m e s*in=aspen*and%h o w{to]produce pines that{are]resistant
Trees*!a#Handbook|of|Forest#Botany 147
Heske says* $The small woodlot must not be judged {by] {its] yield |of|
timber* {for} {its] importance lies *in= other values* fuelwood* protection
|of| soil*and%water*and%shelter{from]*the^wind@*the^woodlot {will] probably
never b e*an^important source |of| supply{for}*the^timber market$ This
comment also applies {to] North America@ I suspect that many *an&
owner likes his wood lot as {it] [is] *and% does not think |of| {it] as commercial
forest land* but rather as!a#place{for}getting some firewood*and%fence
posts* doing!a#little hunting*and%fishing* or just walking*in=*the^woods@
$There {are] some w h o can live without wild things*$ says A l d o
Leopold* $and some who cannot$ Wood#lot owners wishing *to^
improve timber management on their lands should certainly be
encouraged* b u t w e {have] n o right{to]force*the^farmer into forestry
{and}*the^n u m b e r |of| people w h o like their w o o d s just as they{are]m a y
be larger than w e think@
Impression |of| *the^ Timber Resource
It[is]unrealistic*and%gives!a#false impression|of|*the^timber resource
*in&*the^United States{to]classify some 200 million acres |of| w o o d lots
as commercial forest land* when *in= fact many |of| them {are] not and+
perhaps never {will] be@ {it] might be wiser {to] count them under some
other heading*in=*the^census*and%face*the^timber p r o b l e m on*the^basis
|of| w h a t we k n o w [is] commercial forest land@ Likewise*{it]w o u l d b e
sounder {to] think |of| *the^ areas given over {to] shifting cultivation *in= *the^
tropical regions as farm land* which they are* instead |of| calling them
potential timberland@ We {have] as yet no adequate practical way |of|
changing either *the^ system |of| shifting agriculture *in= *the^ tropics or
*the^ w o o d lots*in=*the^midlatitudes@ B o t h {are] firmly rooted cultural
institutions@
S o m e|of|*the^problems{are]perhaps n o m o r e than vexatious* b u t
nevertheless tend{to]hinder good management@*in=*the^United States*
^for* example* most |of| *the^ land [is] m a p p e d according{to]*the^well#
k n o w n township*and%range system* w h i c h[is]!a#g o o d system*in=level
country* b u t{its]suitability*in=*the^rough terrain|of|*the^western forest
lands can be questioned@*the^sections lie over*the^country like squares
on!a#checkerboard*and%property lines run straight across mountains*
ridges* valleys*and%rivers without regard{to]*the^lay|of|*the^land@
O w n e r s h i p s {are] chaotically m i x e d u p*and%*the^chaos [is] worst
precisely{where}*the^best timber grows##that is**in=*the^r e d w o o d belt
|of| California*and%*the^Douglas#fir region |of| O r e g o n*and%Washington@
150 Trees*!a#Handbook|of|Forest#Botany
for *the^ Future that *in= trying {to] control insects *and% diseases we *are^
seriously handicapped {by] !a# lack |of| fundamental knowledge@
Furthermore* w e do not k n o w {to] w h a t extent these epidemics
^are**the^result |of| our opening|of|*the^forests@{it][is]not inconceivable
that opening *the^ virgin forests might be tantamount {to] inviting !a#
more intensive attack upon them {by] insects *and% diseases@ *the^ truth
|of| *the^ matter [is] that modern silviculture *and% forest management *are^
still *in= *the^ experimental stage *and% {it] seems unwise {to] experiment
with our last reserves |of| virgin timber* particularly since we *have^
*an& experimental field closer {at] hand@
If{it][is]true* as {has] often b e e n claimed* that good m a n a g e m e n t
can double *the^ yield |of| *the^ forest *and% if we think we {are] skilled *in^
managing timberland* {it] would seem only common sense not {to] go
into*the^virgin forests until w e {have] p r o v e d our skill{by]first putting
*in& order s o m e 75 or 100 million acres |of| already accessible but
unproductive*and%poorly stocked forest lands that lie strewn over
*the^ United States*in=!a#cut#over* burned#over*and%generally wretched
condition as reminders|of|*the^fact that!a#short while ago w e did not
m a n a g e well@
$ W e s h o u l d r e c o g n i z e *the^ i m p o r t a n c e |of| p l a c i n g h e a v y
investments on d e n u d e d lands or poor quality stands that occupy
*the^ best quality lands@ This[is]m o r e important {for} long#run future
production than rushing into our few remaining virgin stands$ O n e
w o u l d expect that this sensible*and%important suggestion should
^have* stimulated !a# lively discussion among *the^ letter writers* or *for^
that matter on *the^ editorial page* but {to] my knowledge *the^ response
^has* been !a# dead silence@ *the^ prevalent mode |of| thinking seems *to^
be that we must open all *the^ timberlands as soon as possible and+
then* presumably* w e shall put*the^whole h o u s e |of| forestry*in=order@
The real drive behind *the^ demand {for} opening *the^ reserves* *is$
*the^ old yearning {for} virgin timber* but *the^ tradition {has] been
rationalized*in=recent times{by]saying that{it][is]not good conservation
^to* leave *the^ virgin forests alone@ *the^ argument [is] that wood *is$
$wasted$ *in= *the^ mature forests because old trees {are] dying@ {it] [is] !a#
specious argument* b a s e d on*the^old mistaken idea that*the^standing
timber constitutes*the^resource* w h e r e a s*the^real resource[is]*the^land
that produces*the^timber@{it][is]true that trees{are]dying*in=*the^mature
forests* b u t trees{are]also growing up* as they {have] b e e n doing for
Trees*!a#Handbook|of|Forest#Botany 153
Forestry*and%Silviculture*in=*the^Modern Sense
Forestry [is] *in= one sense !a# very ancient practice@ {it] [is] related *in^
Genesis that Abraham planted !a# grove {at] Beersheba~ according *to^
Plato* *the^ Greeks |of| *the^ fifth century B@C@ raised forest trees~ Cato
says that R o m a n farmers b e g a n{at]*an^early time{to]cultivate plantations
|of| trees {to] provide timber {for} local needs~ *and% Pliny describes *at^
length *the^ system |of| coppicing woods *in= regular rotation {to] obtain
firewood* vineyard stakes*and%*the^like@
But forestry*and%silviculture*in=*the^m o d e r n sense {are] not very
old* dating {from] *the^ work |of| Heinrich Cotta *and% G@ L@ Hartig *in^
Germany *in= *the^ 1780@s *and% {from] that |of| Bernard Lorentz *and% Adolphe
Parade*in=France shortly thereafter@ *the^ age |of| m o d e r n silviculture*
^it* might be said* [is] about *the^ same as three or four generations |of|
forest trees *and% *in= terms |of| certain species as only two or three
generations@ !a# plant b r e e d e r w o r k i n g with annuals w o u l d hardly
feel confident about*the^outcome |of| his e x p e r i m e n t after only t w o
or three generations|of|*the^plant he [is] studying@
H e can observe m a n y successive generations*and%[is]able{to]correct
!a# mistake after only!a#few years {have] elapsed* but*the^forester* as
R@ S@ T r o u p puts it* rarely lives long e n o u g h{to]see*the^result |of| his
mistake@{it]w o u l d b e captious n o t{to]recognize*the^great contribution
|of| silviculture*and%*the^remarkable i m p r o v e m e n t s *in= forestry that
^have* b e e n achieved*in=recent years* but w e m u s t keep*in=v i e w the
154 Trees*!a#Handbook|of|Forest#Botany
Food*and%Plant Products*in=Forestry
Introduction
Pursuant{to]*the^authority granted{by]*the^Federal Plant Pest Act
(FPPA) |of| M a y 23* 1957* as a m e n d e d (7 U@S@C@ 150 aa t h r o u g h 150
jj)**and%*the^Plant Quarantine Act (PQA) |of| August 20* 1912* as amended
(7 U@S@C@ 151 t h r o u g h 164* 166**and%167)@ U S D A {has] regulatory
authority over*the^m o v e m e n t into or within*and%t h r o u g h*the^United
States |of| plants* plant products* plant pests**and%any product or
article w h i c h m a y contain!a#plant pest{at]*the^time |of| movement@
These articles {are] regulated *in= order {to] prevent *the^ introduction*
spread* or establishment|of|plant pests n e w{to]or not widely prevalent
*in&*the^United States@ *the^ regulations i m p l e m e n t i n g this statutory
authority {are] found*in=7 C F R Parts 3 0 0 t h r o u g h 399@
$Plant Pest*$ as defined{by]statute* m e a n s any living stage |of| any
insects* mites* nematodes* slugs* snails* protozoa* or other intertebrate
animals* bacteria* fungi* or parasitic plants or reproductive parts
thereof* viruses* or any organisms similar {to] or allied with any |of| *the^
foregoing* or any infectious substances* which can directly or indirectly
injure or cause disease or d a m a g e*in=any plants or parts thereof* or
a n y p r o c e s s e d * m a n u f a c t u r e d * o r o t h e r p r o d u c t s |of| plants)@
$Movement*$ as defined {by] statute* m e a n s {to] ship* deposit for
Trees*!a#Handbook|of|Forest#Botany 163
transmission *in= *the^ mail* otherwise offer {for} shipment* offer *for^
entry* import* receive {for} transportation* carry* or otherwise transport
or move* or allow{to]b e moved*{by]mail or otherwise)@
The current permit system requirements{for}*the^m o v e m e n t into
or within*and%through*the^United States |of| plants* plant products*
plant pests* *and% other articles regulated {by] FPPA *and% PQA *were^
fully described *in= *the^ December 31* 1984 Notice {at] 49 FR 50900#01@
*the^ procedures{for}issuing permits{for}*the^m o v e m e n t |of| plant pests
^were* discussed separately {from] plants* plant products*and%other
articles w h i c h m a y contain plant pests {at] 4 9 F R 50901#02@ U S D A
regulates*the^importation |of| noxious w e e d s through!a#permit system
similar{to]that established{for}plant pests@ *the^ existing regulations
*in& 7 CFR Part 360 which designate plants as noxious weeds and+
establish procedures {for} obtaining *an^ import permit {were] described
at 4 9 F R 50902@
Regulation|of|*the^Introduction|of|Organisms*and%Products Altered
or P r o d u c e d T h r o u g h Genetic Engineering W h i c h {are] or W h i c h
There [is] R e a s o n{to]Believe {are] Plant Pests
The F P P A*and%P Q A{are]applicable{to]*the^m o v e m e n t |of| plants*
plant products**and%other articles*and%plant pests developed through
genetic engineering if such plants* plant products* other articles* or
plant pests present !a# risk |of| plant pest introduction* spread* or
establishment@
U n d e r*the^authority granted{by]*the^F P P A*and%PQA* U S D A is
proposing new regulations which would impose restrictions on *the^
introduction |of| organisms *and% products altered or produced through
genetic engineering w h i c h{are]plant pests or w h i c h there [is] reason
^to* believe {are] plant pests@ *in= accordance with *the^ provisions |of| *the^
FPPA *and% PQA@ USDA must determine *the^ plant pest status |of|
plants* plant products or articles {to] be moved into or within or
through*the^United States@ *the^ evaluation process{for}determining
what safeguards* if any* can be imposed which would allow *the^
movement |of| *the^ plant pest without risk that *the^ plant pest would
be disseminated {were] described *in= *the^ December 31* 1984 Notice *at^
49 FR 50901#02@ {for} genetically engineered material {from] dissimilar
source organisms (inter#generic combinations)**the^determination
m a y b e complex@ Information about genetically engineered organisms
produced through*the^use|of|donor* vector/vector agent*and%recipient
164 Trees*!a#Handbook|of|Forest#Botany
Principles|of|Plant Selection
Selection species[is]important{for}*the^success |of| social forestry@
*the^ selection should b e b a s e d on*the^following criteria*
1@ trees should be fast growing* early maturing*and%yielding*
168 Trees*!a#Handbook|of|Forest#Botany
Chilling Injury
Injury can occur{to]w a r m season plants or products e x p o s e d to
Trees*!a#Handbook|of|Forest#Botany 175
cuts off oxygen {to] *the^ root system *and% causes root death@ Plants *in^
flooded locations can die quickly {from] such injury@ Too much water
can b e as serious!a#p r o b l e m as not e n o u g h water@ {for} this reason*
!a# greenhouse manager must carefully monitor *the^ water level *in^
containers {to] provide *the^ proper amount |of| water {to] meet *the^ crop@s
needs@*in=field situations* drainage tiles or pipes{are]often installed
^to* allow excess water {to] be removed@ Water [is] lost {from] soils *by^
evaporation {from] *the^ soil surface@ Evaporation |of| water {from] soil
combined with loss |of| water through *the^ plant (transpiration) *is$
referred {to] {by] *the^ combined word evapotranspiration* or loss |of|
water {by] plant *and% soil combined@
Applying Water {to] Plants
Horticultural production{has]developed{where}water[is]abundant@
Applying water {to] supplement rainfall [is] !a# standard practice *for^
many horticultural crops@ Irrigation should replace water removed
^by* plants@ Considerations*in=irrigation amounts*and%frequency depend
on *the^ needs |of| *the^ crop* *the^ depth |of| roots *and% *the^ ability |of| *the^
soil {to] absorb water@ !a# certain portion |of| *the^ water [is] lost *to^
evaporation before {it] reaches *the^ soil@ On !a# windy day !a# sprinkler
system putting out!a#fine mist m a y lose 25#30%|of|*the^water applied
before {it] reaches *the^ soil@ Drip or trickle irrigation allows small
a m o u n t s |of| water{to]b e applied frequently{to]!a#portion|of|*the^root
zone@ This reduces*the^a m o u n t |of| water n e e d e d {to] g r o w!a#crop@
Effects |of| Limited Water Availability
For m a n y horticultural plants there[is]!a#period |of| critical water
n e e d w h e n water m a y not b e available@{for}most plants this[is]during
*the^ period |of| flowering*and%fruit development@!a#s y m p t o m |of| water
need [is] wilting |of| *the^ foliage@ Plants {are] affected {by] stress before *the^
external symptom |of| wilting occurs@ Emphasis on water levels
necessary {to] support plant growth *and% scheduling irrigation *to^
minimize water use [is] important@ Water needs can be predicted *from^
climatic factors such as temperature* humidity* w i n d *and% solar
radiation combined with particular characteristics |of| *the^ crop and+
soil type@ Microcomputers assemble various weather factors and+
develop predictions*and%schedules{for}adding supplemental water@
Fluctuations*in=water available can cause sudden splitting|of|cherries*
tomatoes *and% cabbage@ Uniform water supplies {are] necessary *to^
prevent these problems@
Trees*!a#Handbook|of|Forest#Botany 179
Light
Light[is]essential{for}photosynthesis@ Green plants{are]*the^original
solar collectors@ T h e y transfer energy{from]*the^sun{to]!a#usable form
|of| energy{for}*the^plant@ Light varies*in=intensity* quality*and%duration
or photoperiod@
Intensity
The intensity or amount |of| light available{has]!a#direct influence
on *the^ amount |of| photo#synthesis that can occur@ Some plants *have^
adapted {to] growing *in= areas |of| low light availability such as !a# jungle
floor@ This[is]w h y tropical foliage plants g r o w well*in=limited indoor
light@ Grasses{are]native{to]*the^open plains*and%require high light
intensity{for}growth@
When light intensity [is] lowered* *the^ rate |of| photosynthesis *is$
reduced@ When *the^ level |of| photosynthesis equals respiration* {it] *is$
referred {to] as*the^compensation point@ B e l o w compensation point
growth ceases@ S u d d e n exposure{to]high light intensity can cause
leaf scorch or sunburning@ *the^ s y m p t o m s {are] large* brown* dead
areas on leaves@ Plants growing *in= low light {have] !a# thinner layer |of|
wax on leaf surfaces@ This helps *the^ leaf {to] capture more light@
However* m o v i n g{it]suddenly into bright light results*in=rapid water
loss@ H i g h light intensity can also cause fading |of| flowers* especially
*in& hot s u m m e r weather@*an^area |of| special interest{to]*the^greenhouse
industry[is]*the^use|of|high intensity lights{to]supplement natural light
{and}{to]extend*the^photoperiod@ Crops|of|high value m a k e*the^expense
|of| installing *and% o p e r a t i n g h i g h intensity s u p p l e m e n t a l lights
economical@
Light Quality
Light quality refers {to] *the^ $colour$ |of| *the^ light or *the^ portion |of|
*the^ light spectrum@ *the^ biochemical constituents |of| chlorophyll* *the^
compound responsible {for} photosynthesis* absorb only light *from^
particular portions |of| *the^ light spectrum@ Artificial lights supply light
|of| particular portions |of| *the^ spectrum@ Light absorbed {by] plants *is$
^from* *the^ blue#violet *and% orange#red parts |of| *the^ spectrum@ Specialized
l a m p s {have] b e e n developed {to] provide light rich *in= wavelengths
^from* this part |of| *the^ spectrum@ These l a m p s {have] *the^ additional
benefit |of| creating dark green foliage*and%!a#deep* intense flower
colour@ Excellent plant g r o w t h can b e a c h i e v e d w i t h o r d i n a r y
fluorescent bulbs@ Other portions|of|*the^spectrum such as ultraviolet
180 Trees*!a#Handbook|of|Forest#Botany
light {are] important *in= *the^ coloration |of| some fruits *and% *the^
development |of| autumn colour *in= leaves@
Relative Humidity*and%Wind
Relative humidity [is] *the^ amount |of| water vapour *in= *the^ air@ *it^
*is] closely related {to] temperature@ As *the^ temperature goes down*
relative humidity goes up@ Relative humidity[is]measured as!a#percent@
W h e n{it]reaches 1 0 0 % w e {have] rain* snow* fog* or sleet@ This[is]w h y
^it* rains as!a#cold front goes through@
A s *the^ relative h u m i d i t y drops* evaporation increases@ Plants
leaves give off m o r e water{at]4 0 % than 8 0 % relative humidity@ This
water loss increases if*the^w i n d [is] blowing@ Tropical foliage plants
^are* native|of|humid* jungle e n v i r o n m e n t s*and%often suffer*in=*the^dry*
indoor air*in=winter@ G r o u p i n g plants together*and%placing plants on
!a# layer|of|pebbles*in=standing water helps increase relative humidity@
L o w e r i n g*the^interior temperature |of| h o m e s[is]beneficial*in=reducing
some|of|*the^l o w h u m i d i t y p r o b l e m s with plants@
H u m i d i t y [is] often related{to]*the^d e v e l o p m e n t |of| certain plant
diseases@ Warm* d a m p conditions{are]ideal{to]encourage*the^g r o w t h
|of| fungi@ M a n y fungal spores m u s t {have] water {to] b e g i n growing@
Damping off [is] !a# common problem {for} seedlings@ {it] results *from^
fungi rotting small plants {at] *the^ soil line@ Increasing air circulation
^to* lower humidity *and% reducing watering can generally stop *the^
problem@
Wind [is] !a# problem with plant production *in= some areas@ *in^
addition {to] blowing away valuable soil* wind can also cause injury
^to* tender plant stems{by]sandblasting*the^stems{at]*the^soil line@ N e w
transplants*and%n e w l y planted trees{are]often shaded as!a#protection
^from* hot sun*and%wind@*in=m a n y areas|of|*the^Great Plains|of|*the^U@S@*
*the^ planting |of| rows |of| trees *in= !a# windbreak protects homes and+
animals@ Strong winds *in= storms damage many trees *and% shrubs each
year@ W i n d d a m a g e [is] especially severe w h e n{it][is]*in=combination
with ice@ *the^ plant species[is]also important@ T r e e s such as silver
maple* Siberian e l m*and%w i l l o w{are]m o r e frequently injured{by]w i n d
than oak* w a l n u t*and%sugar maple@
Uses* *in= Ayurveda *the^ leaves* seeds *and% *the^ roots {are] used *for^
treatment |of| cough* asthma@
Chemical C o m p o u n d s Obtained* Vasicine[is]obtained {from] this
plant
Adulasa
Botanical Name* Justicia adhatoda* A d h a t o d a Vasica
English Name* M a l a b a r nut
Hindi Name* Adulasa* A d o s a
Sanskrit Name* Atrush
Vasaka or Malabar nut [is] *an^ herb native {to] India@ {it] [is] found *in^
almost every part |of| India@
Vasaka[is]!a#m e m b e r |of| Acanthaceae plant family@ This[is]!a#m a n y
b r a n c h e d shrub@ *the^ leaves {are] opposite* elliptic lanceolet* acute
with n a r r o w base@ *the^ flowers{are]*in=axillary spikes@ *the^ white
coloured flowers with rose coloured streaks *in= *the^ throat* bloom *in^
August {to] November@
Uses* *the^ leaves {are] very efficacious as !a# remedy {for} all sorts |of|
coughs *and% Cold* bronchitis* asthma@ *the^ leaves *and% extracts |of|
leaves{are]useful*in=Ayurvedic Medicines@
Trigonella Foenum # Gracum
Botanical Name * Trigonella F o e n u m # G r a c u m
Family Name * Fabaceae
Common Name * Fenugreek* Greek Hayes* Alholva* Bird@s Foot*
Boyotu* Chinagreye* Fenegriek* F o e n u m G r a e c u m
Part used* Leaves*and%Seeds
Habitat* Wild *in= Kashmir* Punjab *and% upper gangetic plains and+
also cultivated *in= other parts |of| India as !a# pot#herb@
Uses* Fenugreek seed{has]b e e n u s e d{for}stomach upset* swelling
(inflammation) |of| *the^ upper air passages or throat* appetite* *for^
lowering blood sugar *and% {for} softening *the^ stool@ {it] also {has] been
used as!a#gargle{to]relieve sore throat*and%as*an^external dressing
^for* swelling (local inflammation)@
The Seeds{are]very nourishing*and%{are]given {to] convalescents
{and}{to]encourage weight gain* especially*in=anorexia nervosa@ Research
^has* shown that*the^Seeds can inhibit cancer|of|*the^liver* lower blood
cholesterol levels *and% also {have] *an^ antidiabetic effect@ *the^ Seeds and+
leaves {are] anticholesterolemic* anti#inflammatory* antitumor*
Trees*!a#Handbook|of|Forest#Botany 187
Annona Squamosa
Botanical Name * A n n o n a S q u a m o s a
Family Name * Annonaceae
Common Name * C u s t a r d Apple* S u g a r Apple* S w e e t s o p *
Kaner@apra* P o m m e Canelle* Tapotapo* F u n Li Chi* Anon* A n o n a
Blanca* Ati* Ates
Part Used* A n n o n a seeds* A n n o n a leaves* A n n o n a bark
Habitat* Throughout India
Uses*There {are] medicinal application|of|*the^custard apple tree@
*the^ bark*and%leaves contain annonaine**an^alkaloid@!a#bark decoction
*is] used {to] stop diarrhea* while *the^ root [is] used *in= *the^ treatment |of|
dysentery@ !a# decoction |of| *the^ leaves [is] used as !a# cold remedy and+
^to* clarify urine@ A n n o n a squamosa Linn@* family Annonaceae*[is]said
^to* show varied medicinal effects* including insecticide* antiovulatory
{and} abortifacient@The fruits |of| A n n o n a {are] Haematinic* cooling*
sedative* stimulant* expectorant* maturant* tonic@ They {are] useful *in^
anaemia* burning sensation@ *the^ seeds {are] abortifacient and+
insecticidal*and%{are]usefull*in=destroying lice*in=*the^hair@
Argyreia Nervosa
Botanical name * Argyreia Nervosa
Family Name * Convolvulaceae
Common Name * H a w a i i a n B a b y W o o d r o s e Seeds* E l e p h a n t
Creeper Seeds@ Original Argyreia Seeds
Part Used* Seeds* w o o d
Habitat* Hills* 900 m*and%above*in=India
Uses* *the^ roots {are] acrid* bitter* astringent* sweet* emolliet@ {it] *is$
used *in= vitiated conditions kapha *and% vita* emaciation* wounds*
ulcers* anorexia* dyspepsia* latulence*@ {it] [is] widely used *for^
Psychotropic *and% ornamental purposes@
The seeds |of| Argyreia nervosa contain about 0@3per cent ergot
alkaloids* including chanoclavine* lysergol* ergotmetrine*and%ergine@
These {are] all derivatives |of| Lysergic acid*and%m o s t{are]collectively
n a m e d lysergic acid amides (LSA)@ L S D (Lysergic Acid Diiethylamide)
*is] also!a#lysergic acid amide*and%although resembling *the^ natural
lysergic acid amides b o t h *in= structure and *and% pharmacological
activity*{it]{has]not b e e n found*in=nature@ M a n y|of|*the^ergot alkaloids
can cause constriction |of| bloodvessels* w h i c h m a y result*in=necrosis
|of|*the^tissue*and%gangrene*in=h u m a n s*and%animals
190 Trees*!a#Handbook|of|Forest#Botany
Azadirachta Indica
Botanical Name * Azadirachta Indica
Family Name * Meliaceae
Common Name * Lilac* M a r g o s a Tree* Neem* N e e m Chal
Part Used* Leaves* Flower* Oil* Seed
Habitat*{it][is]evergreen*and%grows throughout India@
Uses* Vermifuge* Insecticide* Astringent* Tonic*and%Antispetic@
^it* posseses anti diabitic* anti bacterial *and% anti viral properties and+
used sucessfully *in= cases |of| stomach* worms *and% ulcers@ Root bark
posseses anstringent* tonic*and%antiperiodic properties@{it][is]also
useful*in=Malarial fever@*the^oil[is]used*in=m a k i n g N e e m b a s e d soaps*
s h a m p o o s*and%toothpaste@
Bacopa Monniera
Botanical Name * B a c o p a Monniera
Family Name * Scrophulariaceae
Common Name * Herpestis Monnieria* T h y m e L e a v e d Gratiola*
Brahmi
Part Used* W h o l e Plant
Habitat* G r o w s*in=M a r s h y places*and%cultivated South India@
Uses* {it] [is] Diuretic* Cardiac* Nervine *and% Tonic@ {it] [is] reported *to^
improve intellect* treatment |of| asthma* hoarseness* insanity* epilepsy@
^it*[is]!a#potent nervous tonic*and%[is]anti anxiety agent@{it][is]considered
good fort heart@
Boerhaavia Diffusa
Botanical Name * Boerhaavia Diffusa
Family Name * Nyctaginaceae
Common Name * H o g Weed* Pig Weed* Horse Purslane* Tar Vine
Part Used* Root* L e a v e s*and%Seeds@
Habitat* G r o w s as c o m m o n w e e d
Uses* A c c o r d i n g {to] A y u r v e d a Boerhaavia [is] bitter* anaemia*
cooling* heart diseases* astringent{to]bowels* useful*in=biliousness*
b l o o d impurities* leucorrhoea* asthma* alternatives etc@ *the^ leaves
^are* u s e f u l *in= d y s p e p s i a * t u m o u r s * a b d o m i n a l pains* s p l e e n
enlargement@ According {to] Unani system |of| medicine* *the^ leaves *are^
appetizer* alexiteric* useful *in= opthalmia *and% *in= joint pains@ Seeds *are^
tonic expectorant* carminative* useful*in=lumbago* scabies@*the^seeds
^are* considered as promising b l o o d purifier@
Parts Offerd* Roots
Trees*!a#Handbook|of|Forest#Botany 191
Buckwheat Hulls
Botanical Name * F a g o p y r u m Esculentum
Family Name * Polygonaceae
Common Name * B u c k w h e a t
Parts used* Buckwheat* B u c w h e a t hulls
Habitat* Native {to] Central*and%Northern Asia
Uses*{it][is]u s e d{for}wide range |of| cirulatory problems* b u c k w h e a t
*is] best taken as Tea or tablet@
T h e natural filling {has] m a n y advantages over conventional
fiberfill* foam* or feather fillings@Buck wheat{has]*the^m o s t important
quality{for}*the^relief |of| pain[is]that*the^material conforms{to]*the^exact
contours|of|*the^h e a d*and%neck providing!a#compliant passive support@
Advantages|of|*the^b u c k w h e a t hull filling{are]*the^cool*in=*the^summer*
w a r m*in=*the^winter insulating properties**the^ability{to]adjust*the^size
^to* your individual needs {by] removing or adding back filler so *the^
pillow fits you just right@ *the^ filling also lasts {for} many years making
*the^ pillow*the^best value cervical pillow on*the^market@ L o n g after
other pillows {have] collapsed*and%w o r n out!a#b u c k w h e a t hull pillow
^will* still b e fully functional@ B u c k w h e a t hull p i l l o w s {are] also
environmentally friendly@
Calamus
Botanical Name* * A c o r u s C a l a m u s
Family Name* Araceae
Common Name* Calamus* Sweet#flag* Sweet Root* Bach
Part Used* Dried R h i z o m e
Habitat* North Temperate*and%subtropical regions upto 2 2 0 0 m
altitude*in=Himalayas@
Uses*{it][is]Nauseant* Stomachic* Anthelmentic* Stimulants* Emetic*
Expectorant* Carminative* Antispasmodic*and%Nervine Sedative@
R h i z o m e [is] u s e d*in=*the^cure |of| host |of| diseases such as epilepsy*
mental~ ailments* chronic diarrhoea *and% dysentary@ {it] [is] used *in^
incense sticks *and% dhup *and% [is] widely used as insecticide {for} lice*
bedbugs* w o r m s etc@
Cassia Angustifolia
Botanical Name * Cassia Augustifolia
Family Name* Caesalpinaceae
Common Name* Senna* Indian Senna* Tinnervelly Senna* Cassia
Senna
192 Trees*!a#Handbook|of|Forest#Botany
Food Quality
Food quality [is] !a# multi#faceted parameter *and% [is] generally *the^
predominant driver behind economics |of| *the^ food industry@ Within
*the^ Plant Products*and%F o o d Quality p r o g r a m m e *the^ research foci
^are**the^major c o m p o n e n t s m a k i n g u p food quality* taste* texture*
nutritional value*and%safety@
Plant Products
There[is]*an^increasing i m p e t u s{to]ensure that*the^resources w e
use{for}b o t h food*and%non#food u s e s{are]m a n a g e d*in=!a#sustainable
manner@ *the^ utilisation |of| crops{for}applications other than simple
nutrition[is]increasingly b e i n g seen as*the^w a y forward with respect
^to* !a# sustainable economy@ *the^ Plant Products sub#programme *will^
focus on *the^ non#food aspects |of| crops *and% plants such as bioactivity@
Forestry*and%Forest Products
In*the^country|of|Portugal*!a#total|of|*an^estimated 3 8 %|of|*the^total
land area |of| *the^ country [is] devoted {to] forest covers *and% most |of| *the^
major forest products like paper *and% wood pulp* cork etc@ *are^
significant contributors {to] *the^ progressing Portuguese economy and+
*in& recent years*!a#considerable increase*in=production|of|these products
^have* allowed greater quantity |of| exports thus resulting*in=profits@
Some |of| *the^ most important forest products obtained *from^
Portugal {are] cork* wood pulp* paper pulp *and% paper board*
eucalyptus* cellulose *and% wood {for} furniture etc@ {it] [is] known that *in^
Portugal *the^ amount |of| land area devoted {to] forest cover [is] much
m o r e than that dedicated{to]agriculture*and%about 3 1 %|of|*the^forest
area[is]occupied{by]*the^wild pines w h i c h refer{to]*an^estimated area
|of| 976*000 hectares while *the^ cork plantations cover 2 3 % area i@e@
713*000 hectares with eucalyptus accounting{for}2 3 %|of|*the^forest
area@ *the^ abundance |of| forest resources {has] therefore resulted *in= *the^
growth |of| various industries dedicated {to] *the^ seasoning *and% packaging
{and} modification |of| these products {to] prepare them {for} sale *in= *the^
foreign markets@ *the^ most important industry that {has] evolved *is$
that |of| *the^ paper mills *and% *in= Portugal *the^ pulp *and% paper industry
*is] one |of| *the^ p r o m i n e n t players*in=*the^international market with
Portugal occupying*the^fifth position a m o n g*the^various E u r o p e a n
countries*in=terms |of| production@
well@ {it] [is] derived that *an^ estimated 720 hectares |of| forest land *is$
devoted {to] *the^ production |of| cork oak trees@
The Mediterranean type climate|of|Portugal immensely facilitates
*the^ production |of| cork oak trees*and%*the^bark|of|*the^cork oak tree
*is] k n o w n as cork@
The cork oak tree can survive {for} over 150 years *and% when {it] *is$
about 25 years old* *the^ rough *and% rugged *and% thick bark |of| *the^ cork
oak tree [is] r e m o v e d {to] b e harvested {for} over!a#decade*and%after
harvesting{it][is]considered{to]b e suitable{for}use@{it][is]quite natural
^to* raise *the^ question {at] this point that *in= which way can cork aid *in^
forest conservation@ There {are] various reasons behind that@
First *and% foremost* {it] [is] {to] be remembered that as soon as *the^
bark |of| *the^ cork oak tree or *the^ cork [is] removed* {it] [is] replaced *by^
!a# new layer which begins growing once more@ Therefore* *the^ cork
*is]!a#renewable resource@ Cork[is]!a#vegetable tissue*and%{at]*the^time
|of|{its]harvesting n o chemical or artificial process[is]involved*and%as
!a# result {it] always retains {its] naturalness*and%therefore *the^ forest
e n v i r o n m e n t r e m a i n s unhampered@ Moreover* apart {from] b e i n g
ecological* corks{are]also biodegradable substances@*in=*the^present
era* when *the^ world [is] endangered {by] *the^ growing effects |of|
environmental pollution aggravated {by] *the^ growing amount |of|
garbage*and%chemical wastes that h a r m*the^natural surroundings*
cork [is] natural *and% {it] can be recycled {for} reuse later@ Thus cork *is$
extremely helpful *in= maintaining *the^ ecological balance *and% aiding
forest conservation@
Pine Nuts*and%Forest Conservation
Portugal [is] located *in= *the^ south western part |of| *the^ continent |of|
Europe *and% {it] [is] filled with picturesque locations *and% beautiful
landscapes displaying exquisite scenic beauty@{it][is]interesting{to]note
that*in=Portugal* about 3 8 %|of|*the^total land area[is]occupied{by]forest
cover w h i c h[is]even m o r e than*the^a m o u n t |of| land surface dedicated
^to* agriculture *in= *the^ country@ According {to] official estimates* *the^
major section |of| *the^ forest area [is] dominated {by] *the^ wild pine trees
202 Trees*!a#Handbook|of|Forest#Botany
which occupy 31% |of| *the^ total forest area which [is] equivalent *to^
around 976*000 hectares *and% [is] followed {by] corks *and% eucalyptus@
Of these wild pines* there exists also another species k n o w n as
*the^ stony pines which {are] cultivated *in= Portugal@ *the^ stony pines *are^
also popularly referred {to] as umbrella pines *in= *the^ country *and% *the^
statistics*and%figures revealed{by]*the^Portuguese National Forest
Inventory reveal that*the^stony pines occupy!a#total area |of| about
83*900 h a |of| forest areas@ These trees g r o w extensively*in=Portugal
{and} one|of|*the^major causes facilitating{its]cultivation [is] definitely
due {to] *the^ favorable weather conditions provided {by] *the^
Mediterranean climate |of| *the^ region@
pulp@ *in= Portugal* *the^ various forest resources {have] encouraged *the^
birth |of| industries which {are] engaged *in= *the^ seasoning |of| these
products *and% one |of| *the^ premier industries so developed [is] that |of|
*the^ paper pulp@
In Portugal* paper mills {are] present *in= different parts |of| *the^
country *and% *the^ most prominent manufacturers {are] *the^ Portucel
Soporcel group@
Paper*and%Paper Board
In most areas{where}forest covers exceed *the^ area devoted *for^
agriculture* *the^ forest resources obtained play !a# major role *in= *the^
export market@ This [is] especially true *in= case |of| Portugal which *is$
located *in= *the^ south western part |of| *the^ European continent *where^
*the^ forest resources comprise about 10% |of| *the^ exports@ One |of| *the^
foremost products obtained{from]*the^Portuguese forests {are] paper
{and} paperboard@*the^forest cover*in=Portugal occupies about 3 8 % of
204 Trees*!a#Handbook|of|Forest#Botany
the total land area *and% !a# major part [is] comprised |of| timber and+
eucalyptus cultivations which {are] *the^ primary raw materials {for} *the^
paper*and%paperboard industries@
Wood Pulp
In Portugal!a#total |of| about 3 8 % area[is]devoted{to]forest cover
compared {to] 31% which [is] reserved {for} agricultural purposes@ *the^
forest resources *in= Portugal {are] extremely important {for} *the^ economic
growth |of| *the^ country as !a# considerable amount |of| *the^ produce *is$
exported *in= huge quantities {to] mostly various countries |of| *the^
Europeans u n i o n itself@{it]can b e estimated that about 3 1 %|of|*the^total
forestry area *in= Portugal [is] occupied {by] pines *and% around 21% *is$
acquired {by] eucalyptus@
Trees*!a#Handbook|of|Forest#Botany 205
wild pine trees which occupy around 31% |of| *the^ forest area *and% *is$
followed {by] *the^ eucalyptus *and% cork oak plantations@ *the^ holm oak
trees{are]also prevalent*in=*the^Mediterranean regions*and%*in=Portugal
w h i c h[is]dominated{by]*the^Mediterranean type |of| climate* h o l m oak
trees g r o w*in=abundance occupying approximately*an^area|of|464*000
ha@
In fact *the^ forest cover |of| Portugal [is] divided into t w o major
parts@ *the^ first comprises |of| *the^ plantation area w h i c h includes
eucalyptus*and%*the^maritime pines while*the^other[is]*the^comprised
^by**the^indigenous species w h i c h includes cork oak*and%h o l m oak
trees@
The h o l m oak or Quercus ilex{are]prevalent*in=*the^Mediterranean
regions*and%*in=*the^areas adjoining*the^Atlantic Ocean* they{are]mostly
^to* be found@ {from] *the^ sea level they can grow up {to] *an^ altitude |of|
about 1400 meters *and% *the^ warm temperature which [is] characteristic
|of|*the^Mediterranean region [is] conducive{to]{its]growth@
species planted *and% cultivated *in= Portugal since antiquity {are] *the^
cork oak trees@ *the^ cork oaks *and% *the^ holm oak trees occupy extensive
areas|of|*the^forest@
There {are] many different kinds |of| oak forests *and% *in= Portugal *the^
cork *and% holm oak forests {are] extremely predominant *and% *its^
proximity {to] *the^ Mediterranean basin along with *the^ climatic
characteristics aids {its] growth@ There {are] various reasons behind *the^
widespread cultivation |of| *the^ cork oaks *and% holm oaks@ *the^ cork
oaks*and%h o l m oaks w h i c h g r o w extensively*in=*the^forests |of| Portugal
can be cultivated *in= almost any soil type *and% there {are] no hard and+
fast rules with regard {to] {its] growth though *in= order {to] occupy huge
tracts |of| lands as forests@ Especially*the^cork oak trees{are]extremely
helpful*in=preserving*the^ecological balance|of|*the^forest environment@
This [is] possible as *the^ premier product which [is] obtained *from^
*the^ cork oak trees i@e@ cork [is] !a# vegetable tissue *and% !a# renewable
resource*and%recyclable*and%therefore environment friendly@*in=fact
oak trees*in=*the^forest covers|of|Portugal facilitate forest conservation
immensely@ *in= Portugal* *the^ f o r e s t d e p a r t m e n t [is] e x t r e m e l y
instrumental*in=*the^preservation |of| oak trees*and%therefore* cutting
d o w n|of|these trees{are]considered illegal*and%[is]!a#punishable offence@
Paper Mill$Portucel Soporcel Group
Most |of| *the^ forest resources *and% products obtained {from] *the^
forests |of| Portugal {are] extremely important because |of| their enriched
value*in=*the^export market@ *the^ major products include cork* paper
pulp *and% wood pulp *and% paperboard etc@ *and% these products *are^
available as !a# result |of| *the^ swift supply |of| *the^ required raw materials
like timber* eucalyptus {from] *the^ forest cover |of| Portugal@
The leading organization *in= Portugal which trades *in= *the^
production *and% export |of| pulp *and% paper [is] *the^ Portucel Soporcel
Group which was formed {by] *the^ merging |of| *the^ Portucel *and% *the^
Soporcel@ {it] [is] {to] be remembered {at] this point that *the^ paper *and% pulp
industry [is] one |of| *the^ premier among *the^ industries |of| Portugal and+
*the^ extensive *and% efficient operations |of| *the^ Portucel Soporcel group
208 Trees*!a#Handbook|of|Forest#Botany
It [is] {to] be noted that *in= Portugal one |of| *the^ dominant causes *for^
*the^ loss |of| forest resources {are] *the^ wildfires that {are] quite common
{and}{it]{has]b e e n seen*in=!a#particular year* there{has]b e e n*an^alarming
a m o u n t |of| 30*000 forest fires w h i c h [is] frightening@
While this [is] !a# major cause {for} destruction |of| forest area* *the^
absence |of| proper measures *and% safeguards against this natural
disaster further aggravates this problem@
However* *in= recent times* {it] must b e pointed out* extensive
measures{are]being devised{by]forming public opinion*and%spreading
awareness*and%thus preserve*the^natural resources@*in=recent times
afforestation [is] !a# prominent measure introduced {to] preserve *the^
existing forest area *and% increase {it] as well *and% *the^ policies |of| *the^
E u r o p e a n U n i o n Regulations* {it] can b e estimated that *the^ forest
cover*in=Portugal[is]*in=*the^process |of| expansion annually@
Forest Fires*in=Portugal
In Portugal* *the^ entire forest cover engages around 38% |of| *its^
total land surface *and% *the^ various forest products {are] extremely
important contributors{to]*the^export market thus aiding e c o n o m i c
growth*and%prosperity@*the^various forest products like w o o d pulp*
paper pulp*and%paper board* cork* cellulose* eucalyptus {have] recorded
!a# significant increase *in= production *in= *the^ recent times *and% this *has^
positively affected *the^ economy as *in= *the^ year 2000 itself* there *has^
b e e n noticed*an^estimated rise*in=exports{by]about 2 5 % c o m p a r e d
^to* that*in=*the^year 1999@
however* despite all this* one |of| *the^ major threats lurking *in= *the^
corner {for} Portuguese forest covers {are] *the^ widespread forest fires
that {are] extremely c o m m o n*in=*the^country*and%cause widespread
210 Trees*!a#Handbook|of|Forest#Botany
devastation *and% havoc thus ruining huge tracts |of| forest land and+
destroying plantations@ *in= fact* *in= !a# widespread survey conducted
amidst Portuguese nationals *in= *the^ recent past {has] shown that !a#
whopping section |of| 97@7% |of| *the^ population [is] |of| *the^ opinion that
one |of| *the^ major threats {to] *the^ forest cover *in= Portugal {are] due *to^
forest fires@
What {has] further aggravated *the^ problem [is] *the^ absence |of|
adequate measures@ Forest wildfires *in= Portugal {have] devastative
effects on *the^ environment like causing soil erosion *and% affecting *the^
ecological balance |of| *the^ forest environment along with *the^ massive
loss |of| human lives@ Moreover* *the^ number |of| technicians and+
firefighters who {are] appointed {for} *the^ protection |of| *the^ forest resources
^are* often incompetent *and% inefficient *and% this added {to] *the^ lack |of|
proper forest management {has] added {to] *the^ problem@
Therefore* what [is] required [is] {to] draw up !a# full proof plan *in^
order {to] ensure that *the^ amount |of| forest area lost *in= recent times
be replaced {by] expanding *the^ current forest cover with *the^
implementation |of| various methods like conceiving !a# proper forest
m a n a g e m e n t plan* increase vegetation {by] planting n e w saplings*
prevent intrusion into forest area{for}*the^sake |of| urbanization*
prevention |of| forest fires *and% natural hazards etc@ This process *is$
theoretically referred {to] as forest transition *and% {it] {has] been
experimented*in=various countries|of|*the^world |of| w h i c h Portugal
*is] one too@
In purely technical terms* forest transition [is] !a# geographical
process w h i c h talks|of|*the^reversal|of|*the^factors governing usage
|of| land{by]m e a n s |of| recovery|of|*the^net forest area lost*in=!a#particular
period{to]*the^a m o u n t |of| forest area acquired*in=!a#period@*in=Portugal
forest transition could be observed after *the^ commencement |of|
industrialization@
Rules |of| *the^ Forestry Department *in= Portugal
Portugal located *in= *the^ south western part |of| Europe [is] *the^
premier producer |of| cork *in= *the^ world *and% {it] accounts {for} about 50%
|of| *the^ world@s total cork production *and% about 720*000 hectares |of|
land *in= Portugal [is] dedicated {to] *the^ cultivation |of| Forestry *in= Portugal
trees {from]{where}cork [is] obtained@ *the^ steadily rising graph |of| *the^
Portuguese economy {has] immense contributions {from] *the^ Portuguese
cork industry with their cork constituting 70% |of| export trade *in^
recent times@
The Portuguese cork industry [is] highly developed *and% *the^
versatility |of| cork as !a# substance together {its] ecological properties
{and}*the^ability{to]recycle*the^material{has]facilitated{its]growth@ Cork
*is] used*in=performing multifarious functions like{for}building and
212 Trees*!a#Handbook|of|Forest#Botany
architectural purposes *and% *in= floors* *in= *the^ automobile* military and+
space industries* *in= footwear *and% leather goods* as chemical
c o m p o n e n t s*in=pharmaceutical industries*and%m o s t importantly as
cork stoppers u s e d as closures{for}wine bottles@*in=Portugal* w h i c h
*is] k n o w n{for}{its]wines*and%vineyards*{it][is]doubtless that cork stoppers
^are* extremely useful {for} domestic purposes@ Cork w o o l s {are] also
used*in=mattresses*and%cushions@
Forest Structure*and%Patterns
Introduction
The interactions b e t w e e n disturbances*and%forest trees generate
!a# c o m p l e x range |of| forest structures*and%patterns@ Perhaps*the^best
route through this complexity starts with t w o contrasting stand
structures (for descriptive purposes!a#stand[is]taken{to]b e!a#compact
patch |of| w o o d l a n d |of| about 2 ha)@
1@ Even#aged* stands {are] even#aged if*the^canopy trees all started
growth *in= approximately *the^ same year@ !a# limited age spread |of|
perhaps 10 years [is] permissible within *the^ definition@ Such stands
originate after!a#stand#destroying disturbance or*an^abrupt change
*in& land use@ Colonizing trees may take longer than 10 years{to]fill
*the^ available space* but as*the^stand develops{its]age range narrows*
latecomers{will]{have]smaller* lower crowns than those that started
growth immediately*and%this disadvantage ensures that their
mortality rate[is]greater@
2@ Mixed#age* stands comprising!a#wide range|of|age#classes*{from]old
trees{to]saplings* intimately intermixed@ Such stands tend{to]form
!a# mosaic |of| even#aged groups{at]*the^scale |of| one or two canopy
trees@ They function{by]!a#process |of| relatively constant gap
formation**in=which gaps formed{by]*the^loss |of| canopy trees or
major branches {are] filled {by] groups |of| saplings@ These groups
compete amongst themselves as they grow* so that eventually only
214 Trees*!a#Handbook|of|Forest#Botany
The distribution |of| dead wood [is] generally irregular* {for} example !a#
gap created {by] *the^ fall |of| !a# single tree {has] *an^ accumulation |of| crown
b r a n c h e s*and%debris on one margin* leaving*the^rest|of|*the^gap with
!a# clean floor@ *in= many forest types *the^ underwood |of| shrubs and+
groups |of| advance regeneration survives gap formation* {its] original
irregularities {will] b e increased{by]s o m e flattening as*the^canopy tree
falls@
In larger gaps {it] [is] common {for} one or two subcanopy trees *to^
survive* thereby retaining !a# light shelterwood within *the^ gap@
Irrespective |of| size* gaps{have]*an^internal geometry w h i c h ensures
that*in=*the^Northern H e m i s p h e r e *the^ northern side receives more
direct sunlight than *the^ southern side*and%that *the^ eastern side
b e c o m e s drier*in=afternoon sun than*the^western side* w h i c h receives
direct sunlight during*the^more#humid mornings@ Furthermore* light
intensity[is]greater{at]*the^centre |of| gaps than{at]*the^edges@*in=general*
larger gaps permit!a#larger representation |of| shade#intolerant species@
Internal heterogeneity created {by] fallen trunks *and% b r a n c h w o o d
generates irregularities*in=browsing intensity*and%small#scale shading
patterns**and%thus also contributes{to]*the^variety |of| regenerants@
Both gaps *and% *the^ representation |of| stages {have] been used *to^
quantify canopy turnover@ Gaps {are] formed* or canopy space *is$
vacated*{at]about 1% per annum@ *the^ rate varies {from] year{to]year
{and} on!a#scale|of|decades but{where}it{has]b e e n m e a s u r e d*in=different
forests* gaps {have] formed{at]0@5#2% annually@ This implies!a#residence
time |of| about 100 years {for} trees that reach *the^ canopy@{where}*the^
extent *and% duration |of| growth stages {have] been quantified* {it] *has^
been possible {to] compute *the^ duration |of| *the^ growth cycle {at] some
300 years*and%{to]determine*the^degree{to]w h i c h*the^age distribution
|of| patches departs{from]*the^theoretical steady state@
In practice* individual trees live far longer than 100 years@ Firstly*
trees take several decades {to] reach*the^canopy* particularly those
shade#tolerant species that remain virtually static when they *are^
heavily shaded@ Even fast#growing light#demanding species take 30
years or so{to]reach canopy height@ Secondly* s o m e canopy species
^are* inherently short#lived* leaving space{for}other species{to]reside
longer*in=*the^canopy@ Thirdly* gap formation [is] n o t absolutely
correlated with age |of| trees* so that s o m e individuals live far longer
than *the^ majority@ *in= temperate forests* individuals |of| oak* tulip*
Trees*!a#Handbook|of|Forest#Botany 217
Forest Structure
Rainforests {are] divided into different strata* or layers* with
vegetation organized into!a#vertical pattern{from]*the^top|of|*the^soil
^to**the^canopy Each layer[is]!a#unique biotic c o m m u n i t y containing
different plants*and%animals adapted{for}life*in=that particular strata@
Only *the^ emergent layer [is] unique {to] tropical rainforests* while *the^
others {are] also found *in= temperate rainforests@
218 Trees*!a#Handbook|of|Forest#Botany
Forest Floor
The forest floor* *the^ bottom#most layer* receives only 2% |of| *the^
sunlight@ Only plants adapted {to] low light can grow *in= this region@
A w a y {from] riverbanks* s w a m p s*and%c l e a r i n g s * { w h e r e } d e n s e
u n d e r g r o w t h[is]found**the^forest floor[is]relatively clear|of|vegetation
because |of| *the^ l o w sunlight penetration@ This m o r e open quality
permits*the^easy m o v e m e n t |of| larger animals such as* ungulates like
*the^ okapi (Okapia johnstoni)* tapir (Tapirus sp@)* Sumatran rhinoceros
(Dicerorhinus sumatrensis)* *and% apes like*the^western lowland gorilla
(Gorilla gorilla)* as well as many species |of| reptiles* amphibians* and+
insects@ *the^ understory also contains decaying plant *and% animal
matter* w h i c h d i s a p p e a r s quickly* b e c a u s e *the^ warm* h u m i d
conditions promote rapid decay@ M a n y forms |of| fungi growing here
help decay*the^animal*and%plant waste@
Understory Layer
The understory layer lies b e t w e e n *the^ canopy*and%*the^forest
floor@*the^understory[is]h o m e{to]!a#n u m b e r |of| birds* small mammals*
insects* reptiles**and%predators@ E x a m p l e s include leopard(Panthera
pardus)* poison dart frogs (Dendrobates sp@)* ring#tailed coati (Nasua
nasua)* boa constrictor (Boa constrictor)* *and% many species |of|
Coleoptera@ *the^ vegetation {at] this layer generally consists |of| shade#
tolerant shrubs* herbs* small trees**and%large w o o d y vines w h i c h
climb into*the^trees{to]capture sunlight@ O n l y about 5 % |of| sunlight
b r e a c h e s *the^ c a n o p y {to] arrive {at] *the^ u n d e r s t o r y c a u s i n g true
understory plants{to]seldom grow{to]3 m (10 feet)@ A s*an^adaptation
^to* these l o w light levels* understory plants {have] often evolved m u c h
larger leaves@ Many seedlings that {will] grow {to] *the^ canopy level *are^
*in& *the^ understory@
Canopy Layer
The canopy[is]*the^primary layer|of|*the^forest forming!a#roof over
*the^ two remaining layers@{it]contains*the^majority|of|*the^largest trees*
typically 30#45 m *in= height@ Tall* broad#leaved evergreen trees *are^
*the^ dominant plants@ *the^ densest areas |of| biodiversity {are] found *in^
*the^ forest canopy* as {it] often supports !a# rich flora |of| epiphytes*
including orchids* bromeliads* m o s s e s*and%lichens@ These epiphytic
plants attach{to]trunks*and%branches*and%obtain water*and%minerals
^from* rain*and%debris that collects on*the^supporting plants@*the^fauna
*is] similar{to]that found*in=*the^emergent layer* b u t more diverse@ It
Trees*!a#Handbook|of|Forest#Botany 219
are classified {by] *the^ amount |of| rainfall received each year* which *has^
allowed ecologists {to] define differences *in= these forests that look so
similar*in=structure@ According{to]Holdridge@s classification|of|tropical
ecosystems* true tropical rainforests{have]*an^annual rainfall greater
than 800 c m*and%annual temperature greater than 24 degrees Celsius@
However* most lowland tropical rainforests can be classified as tropical
moist or w e t forests* w h i c h differ *in= regards {to] rainfall@ Tropical
rainforest ecology# dynamics* composition**and%function#{are]sensitive
^to* changes *in= climate especially changes *in= rainfall@ *the^ climate |of|
these forests [is] controlled {by] !a# band |of| clouds called *the^ Intertropical
Convergence Zone located near *the^ equator *and% created {by] *the^
convergence |of| *the^ trade winds {from] *the^ northern *and% southern
hemispheres@ *the^ position |of| *the^ b a n d varies seasonally* m o v i n g
north *in= *the^ northern summer *and% south *in= *the^ northern winter* and+
ultimately controlling *the^ wet *and% dry seasons *in= *the^ tropics@ These
regions {have] experienced strong w a r m i n g{at]!a#m e a n rate |of| 0@26
degrees Celsius per decade which coincides with !a# global rise *in^
temperature resulting {from] *the^ anthropogenic inputs |of| greenhouse
gases into*the^atmosphere@ Studies {have] also found that precipitation
^has* declined*and%tropical Asia {has] experienced*an^increase*in=dry
season intensity whereas A m a z o n i a{has]n o significant pattern change
*in& precipitation or dry season@ Additionally* El N i n o # S o u t h e r n
Oscillation events drive *the^ interannual climatic variability *in^
temperature *and% precipitation *and% result *in= drought *and% increased
intensity |of| *the^ dry season@ As anthropogenic warming increases *the^
intensity *and% frequency |of| ENSO {will] increase* leaving tropical
rainforest regions susceptible {to] stress *and% increased mortality |of|
trees@
Soil Types
Soil types {are] highly variable *in= *the^ tropics *and% {are] *the^ result |of|
!a# combination |of| several variables such as climate* vegetation*
topographic position* parent material**and%soil age M o s t tropical
soils {are] characterized {by] significant leaching*and%poor nutrients~
h o w e v e r there {are] s o m e a r e a s t h a t c o n t a i n fertile soils@ Soils
throughout*the^tropical rainforests fall into t w o classifications w h i c h
include*the^ultisols*and%oxisols@ Ultisols{are]k n o w n as well weathered*
acidic red clay soils* deficient *in= major nutrients such as calcium and+
potassium@ Similarly* oxisols {are] acidic* old* typically reddish* highly
Trees*!a#Handbook|of|Forest#Botany 221
Nutrient Recycling
This high rate|of|decomposition[is]*the^result|of|phosphorus levels
*in&*the^soils* precipitation* h i g h temperatures*and%*the^extensive
m i c r o o r g a n i s m communities@*in=addition{to]*the^bacteria*and%other
microorganisms* there{are]*an^abundance |of| other decomposers such
as fungi*and%termites that aid*in=*the^process as well@ Nutrient recycling
*is] important because below ground resource availability controls *the^
above ground biomass *and% community structure |of| tropical
rainforests@ These soils{are]typically phosphorus limited* which inhibits
222 Trees*!a#Handbook|of|Forest#Botany
more sophisticated images@ But they tend {to] reflect only *the^
peculiarities |of| *the^ plot *and% provide little quantification |of| *the^ vertical
organization|of|*the^whole@
Leaf distribution # jess W h o l e canopy L z (leaf#height density)
can b e estimated{by]assembling structural measurements |of| individual
crowns m a d e {from] observations{from]*the^g r o w n or harvested stems@
Lots|of|m e a s u r e m e n t s*in=crop plants* very few{for}w h o l e forests@ This
s h o w s vertical stratification |of| leaf distribution {for} *an^ e a s t e r n
deciduous forest* gathered with crane data@ Fish eye Optical point
measurements more common@ Hemispheric fisheye photography uses
potential light environment{at]!a#point {to] assess canopy structure@
Several devices n o w estimate L A I{from]in#canopy light measurements*
relying on Beer@s l a w*and%*the^absorbance |of| light@
R e m o t e sensing{has]done!a#lot@ Use*the^spectral quality |of| canopy
light**the^ratio |of| red/far red light transmitted through*the^canopy@
Several measures calculated {from] combining different reflectance
bands relate {to] *the^ amount |of| green canopy biomass* such as *the^
$normalized difference vegetation index (NDVI) or $greenness$ index@
Index takes advantage|of|*the^strong reflectance*in=*the^near infrared
b u t w e a k reflectance*in=*the^red wavelengths |of| green canopies@ Other
remote sensing techniques use !a# similar approach* with sensors |of|
different wavebands on several satellite platforms@ (LIDAR) Also
^from* !a# profiling airborne laser # {to] sense *the^ elevation |of| both *the^
ground *and% canopy* yielding *the^ contour |of| canopy height over !a#
long transect@ So advances*in=quantifying*and%visualizing canopy
structure [is] growing*in=sophistication@
O l d growth m e s s y*the^term $structural complexity$ w a s coined
as *an^ integrative concept {to] represent *the^ complex 3#D structure |of|
old#growth forests@ However* many |of| *the^ measures *and% indices
used{to]characterize structural complexity such as species composition*
tree#size distribution *and% abundance |of| snags *and% woody debris *are^
derived {from] 2#D* ground#based measurements@ Ishii young old
forest O n e |of| *the^ objectives{for}developing indices |of| structural
complexity [is] {to] quantitatively distinguish old#growth forests *from^
younger stands *and% {to] establish criteria {for} enhancing old#growth
forest structure*in=m a n a g e d stands{for}conservation purposes* such
as creating wildlife habitat@ *in= order {to] m e e t such m a n a g e m e n t
objectives* w e must integrate various aspects |of| forest structure
Trees*!a#Handbook|of|Forest#Botany 227
Horizontal Variation
Light gap Forests{are]not spatially uniform* b u t{are]horizontally
heterogeneous{at]various scales@ M o s t w o r k {has] dealt with foliage#
228 Trees*!a#Handbook|of|Forest#Botany
Temporal Changes
Light e n v over year C a n o p y structure changes seasonally*in=all
forest* b u t[is]m o s t dramatic*in=deciduous stands@ But even evergreen
forests* *the^ quantity |of| leaf area varies over *the^ year@ G e n stand
development M o r e substantial changes occur on!a#succession time
scale@ T o t a l a m o u n t stabilizes* b u t {its] v e r t i c a l *and% h o r i z o n t a l
distribution change slowly@ B o b v p stands This difference {has] b e e n
shown through comparative studies |of| 2#D stand structure@ {at] *the^
individual tree level* can be seen@ Jess images |of| stand ages But *at^
*the^ stand level* can also be quantified@ Species composition *and% tree
size distributions b e c o m e m o r e diverse with increasing stand age
{and} specific structural elements such as large* old trees*and%snags
characterize older stands
Deterministic Processes that Drive Development |of| Canopy Structure
In early stages |of| stand development* deterministic processes
such as timing|of|establishment following disturbance* height#growth
rate**and%c r o w n interactions determine canopy structure@*in=mixed#
species natural forests* differences among species *in= timing |of|
establishment *and% initial height#growth rates result *in= vertical
Trees*!a#Handbook|of|Forest#Botany 229
Forest Conditions*and%Area
The area covered{by]forests[is]very likely{to]change u n d e r climate
change* with shifts occurring b e t w e e n forest types due {to] changing
temperature*and%precipitation regimes* while*in=some regions* forest
area [is] expected {to] expand (e@g@ temperate regions) *and% *in= others *to^
contract (e@g@ boreal* tropical *and% mountain forests)@ Such changes
^have* been occurring *in= *the^ past following *the^ natural changes *in^
temperature *and% precipitation that accompanied *the^ different ice
ages@ Currently* however*{it][is]very difficult{to]separate forest area
change due{to]climate change {from] area changes due{to]other factors@
Globally* planted forests*and%natural regeneration {have] increased
*the^ forest areas*in=*the^United States* Europe* China**and%some countries
*in& Latin America *and% *the^ Caribbean e@g@ Chile* Uruguay* Cuba and+
Costa Rica@ On *the^ other hand* some countries *in= Africa* Asia and+
*the^ Pacific *and% *the^ tropical countries |of| Latin America continue *to^
be subject {to] deforestation* mainly due {to] conversion {to] small# and+
large#scale agriculture*and%livestock while deforestation*in=*the^boreal
forests |of| Siberia[is]mainly due{to]forest fires@ A l t h o u g h*the^boreal
forests {are] expected {to] move northward* temperate forests *are^
expected {to] increase their area northward {to] !a# greater extent than
*the^ boreal forests* thus reducing *the^ total area |of| boreal forests@
In *the^ future* {it] [is] expected that *the^ combination |of| climate
change* land u s e conversion*and%un#sustainable land use practices
234 Trees*!a#Handbook|of|Forest#Botany
favour *the^ growth |of| insect populations that [is] detrimental {to] *the^
health |of| forests@
This [is] more likely{to]occur*in=forests dominated {by] few tree
species or{where}specific temperatures or moisture levels control
insect populations@{for}example**the^spread |of| *the^ mountain pine
beetle* Dendroctonus ponderosae**in=boreal forests*{has]b e e n largely
attributed {to] *the^ absence |of| consistently low temperatures over !a#
long period |of| time* which allowed *an^ existing outbreak {to] spread
across montane areas*and%into*the^colder boreal forests@
Trees*!a#Handbook|of|Forest#Botany 235
Land*and%Farm Resources*in=Forestry
Introduction
L a n d [is] a m o n g *the^ m o s t i m p o r t a n t resources u s e d {by] U@S@
agriculture$and {its] chief finite resource@ Decisions on land use *are^
made daily {by] operators |of| *the^ Nation@s 2@1 million farms* contingent
on market conditions* public policies**and%farm/ranch characteristics@
This chapter examines land use* farm real estate values* *and% *the^
relationship between land ownership* farm size* *and% other
characteristics@ A m o n g *the^ findings*
• Land continues {to] shift between agriculture*and%other uses@
Cropland {has] declined over recent decades* but losses do not
threaten*the^Nation@s capacity{to]produce food*and%fibre@
• Real estate[is]!a#major asset{for}farm households@ Values{have]been
rising steadily since 1987* but *the^ inflation#adjusted (real) value |of|
U@S@ farm real estate [is] still below {its] 1982 peak@
• Small family farms account{for}most land owned{by]farms@
Land Use* Environment *and% *the^ Sustainability
Land#use changes can affect *the^ environment *and% *the^
sustainability |of| production@ Because impacts on *the^ environment$
including erosion* water quality**and%wildlife h a b i t a t $ a r e typically
not reflected*in=private profit calculations* land#use choices that are
238 Trees*!a#Handbook|of|Forest#Botany
E R S identifies five groups |of| small family farms (sales less than
$250*000)* limited#resource* retirement* residential/lifestyle* farming#
occupation/low#sales* *and% farming#occupation/high#sales@ *the^
typology also includes large family farms* very large family farms*
{and} nonfamily farms@*in=addition* very small farms (sales less than
$10*000) m a k e u p m o r e than half |of| all farms@ V e r y small farms
account{for}!a#particularly large share |of| farms*in=*the^limited#resource
(72 percent)* retirement (76 percent)**and%residential/lifestyle (76
percent) groups@ Production* however*[is]concentrated a m o n g larger
farms~ small farms account {for} only 27 percent |of| *the^ total value |of|
production@
The smallness |of| m o s t farms {has] implications{for}conservation
{and}*the^environment@ *an^ E R S study found that smaller corn farms
^are* less likely {to] use conservation tillage than {are] larger farms@ *the^
practice [is] more practical {for} larger farms because they {have] more
acres over which{to]spread*the^cost |of| n e w or retrofitted equipment
necessary{to]adopt conservation tillage@ Small farms w h o s e operators
^are* retired or farm part#time{are]also less likely{to]adopt conservation
tillage* possibly because |of| hesitancy{to]change familiar production
practices@ Small farms* however* participate widely *in= *the^
Conservation Reserve Programme (CRP) *and% *the^ Wetlands Reserve
P r o g r a m m e (WRP)@
Distribution |of| Conservation Programme Payments {by] Type |of| Farm
These farms harvest most |of| *the^ land planted {to] p r o g r a m m e
c o m m o d i t i e s*and%therefore receive three#quarters |of| c o m m o d i t y
p r o g r a m m e payments@ However* C R P*and%W R P $ t h e t w o major
conservation p r o g r a m m e s $ a r e targeted{at]particular types |of| land*
not commodities@ Since small farms o w n 70 percent|of|*the^land held
^by* farms* they play!a#large role*in=natural resource*and%environmental
policy@ Retirement* residential/lifestyle**and%low#sales farms account
^for* nearly two#thirds |of| conservation p a y m e n t s*and%!a#similar share
|of|*the^land farmers enrolled*in=*the^C R P*and%WRP@ Participating
farmers*in=e a c h|of|*the^three groups tend{to]enroll large shares |of| their
land *in= these programmes* 46 percent |of| *the^ land operated on
retirement farms* 28 percent on residential/lifestyle farms**and%23
percent on low#sales farms@ *in= contrast* enrollment ranges {from] 5 *to^
9 percent {for} participating high#sales* large* *and% very large farms@
Because their m a i n j o b[is]off#farm* residential/lifestyle operators
242 Trees*!a#Handbook|of|Forest#Botany
a whole@{it][is]{by]far*the^best*and%perhaps*the^only w a y{to]put*an^end
^to* speculation*in=land*and%{to]capture subsequent increases*in=land
values@ These surpluses*{where}realised{by]*the^public authorities*
should benefit*the^community*in=more ways than one@$ Not surprising
*the^ role m o d e l s |of| Indian T o w n Planners $ Delhi M a s t e r Plan*
Chandigarh* Gandhinagar*and%Navi M u m b a i {were] all based on public
ownership |of| land@ Whether public ownership *in= fact achieved *the^
land policy objectives *in= such cases may be !a# matter |of| debate@ But
!a# verdict on Delhi experience was~
1@ It{has]not been possible{for}DDA{to]provide land{at]affordable
prices{to]low income beneficiaries resulting*in=large scale jhuggi
jhopadi colonies@
2@ In*the^absence|of|price signals land{has]been sub optimally used*
resulting*in=over provision{to]powerful groups* and
3@ DDA@s policy {to] auction very few plots {at] !a# time *and% treating *the^
maximum price quoted *in= such biding as *the^ real market price *has^
*in& fact meant artificially increasing *the^ land price through deliberate
scarcity@$
However* securing large#scale public ownership |of| land implied
c o m p u l s o r y acquisition |of| land@ There w a s considerable discontent
a m o n g s t *the^ original l a n d o w n e r s a b o u t *the^ m a n n e r *in= w h i c h
compensation w a s determined*and%paid@*the^L a n d Acquisition A c t
1894 initially provided*the^date |of| declaration|of|intention{to]acquire
*the^ land as*the^reference date {for} determining *the^ market value@
However no time limit was laid down {for} actual payment |of|
compensation@ 1984 amendments introduced *the^ time limit |of| three
years *and% also provided {for} payment |of| interest {from] *the^ date |of|
award {to] actual payment or possession |of| land *and% solatium |of| 30%
|of| market value@ However *the^ market value [is] {to] be reckoned *at^
current use value {at] *the^ exclusion |of| expected rise *in= value on account
|of| future use@ *the^ proposed changes *in= *the^ LA Act *and% *the^ R and+
R Policy attempt {to] remove many |of| these lacunae@ But planned
urban development [is] not being recognised as !a# public purpose *for^
which powers |of| eminent domain could be used *and% *in= practical
terms *the^ proposed method |of| deciding compensation and+
rehabilitation package would make recourse {to] compulsory
acquisition |of| land expensive{for}lands that also require substantial
investment *in= trunk infrastructure@ This would compel search *for^
new paradigm *in= respect |of| urban land@ *the^ thinkers *in= *the^ first
Trees*!a#Handbook|of|Forest#Botany 245
The most recent options {for} rule changes under consideration *are^
summarized *in= !a# $Non#Paper$ *the^ co#chairs |of| *the^ contact group on
LULUCF@
The extent**and%type |of| land use directly affects wildlife habitat
{and} thereby impacts local*and%global biodiversity@ H u m a n alteration
|of| landscapes {from] natural vegetation {to] any other use typically
results*in=habitat loss* degradation**and%fragmentation* all |of| w h i c h
can {have] devastating effects on biodiversity@ Land conversion [is] *the^
single greatest cause |of| extinction |of| terrestrial species@ *an^ example
|of| land conversion b e i n g!a#chief cause|of|*the^critically e n d a n g e r e d
status|of|!a#carnivore[is]*the^reduction*in=habitat{for}*the^African Wild
Dog* Lycaon pictus@ |of| particular concern [is] deforestation* *where^
logging or burning {are] followed {by] *the^ conversion |of| *the^ land *to^
agriculture or other land uses@ Even if some forests {are] left standing*
*the^ resulting fragmented landscape typically fails{to]support m a n y
species@
Lake Winnipesaukee Ice#Oui
Lake W i n n i p e s a u k e e Ice#Out occurs w h e n all*the^ice on Lake
Winnipesaukee**the^largest lake*in=N e w Hampshire*{has]b r o k e n u p
after winter@ O v e r*the^years this{has]b e e n decided u p o n{by]!a#variety
|of| means~ as |of| 2010* D a v e E m e r s o n m a k e s*the^call@ E m e r s o n flies
two {to] three times !a# day over Lake Winnipesaukee {to] check on *the^
ice@ Ice#Out [is] declared when *the^ MS Mount Washington can make
^it*{to]every one|of|{its]ports* Centre Harbor* Wolfeboro* Alton* Weirs
Beach *and% Meredith@ {it] [is] also considered *the^ unofficial start {to] *the^
boating season as well as *the^ end |of| winter *in= New Hampshire@ *the^
earliest Ice#Out recorded was *in= 2010 on March 24* *and% *the^ latest *it^
occurred was *in= 1888 on May 12@ Because *the^ Ice#Out designation
*is] decided u p o n{by]one man*{it][is]highly unscientific@ *the^ call does
not m e a n that *the^ lake [is] entirely devoid |of| ice* nor does*the^M S
M o u n t W a s h i n g t o n actually go {to] each |of| {its] ports@ {it] [is] simply
believed that*the^ship could@ Ice#Out records {have] b e e n kept since
1887 as!a#w a y{to]keep track |of| w h e n b o t h commercial*and%passenger
transportation lanes b e c a m e usable*in=*the^lake@*in=1974* Dr@ William
K@ W i d g e r t compiled data {from] k n o w n records{to]create!a#list |of| past
dates@
Originally *the^ Ice#Out designation w a s decided {by] people on
shore@ Later*{for}roughly 50 years**the^call w a s m a d e{by]B o b Aldrich*
250 Trees*!a#Handbook|of|Forest#Botany
^by* *the^ world@s land vegetation@ Most |of| this vegetation [is] *in= *the^
Northern hemisphere* since this [is]{where}most |of| *the^ land [is] located@
*the^ level decreases {from] northern spring o n w a r d s as n e w plant
g r o w t h t a k e s c a r b o n d i o x i d e o u t |of| *the^ a t m o s p h e r e through
photosynthesis *and% rises again *in= *the^ northern fall as plants and+
leaves die off *and% decay {to] release *the^ gas back into *the^ atmosphere@
D u e*in=part{to]*the^significance |of| Keeling@s findings**the^N O A A
b e g a n monitoring C O levels w o r l d w i d e*in=*the^1970s@ Today* C O
2 2
levels {are] monitored {at] about 100 sites around *the^ globe@ Keeling
died*in=2005@ Supervision|of|*the^m e a s u r i n g project w a s taken over
^by* his son* R a l p h Keeling*!a#climate science professor{at]*the^Scripps
Institution@
Land Alienation*and%Pauperization
The colonial state i m p o s e d*the^land laws based on this worldview
252 Trees*!a#Handbook|of|Forest#Botany
facts |of| this association are* first* that large farms $ including large
family#type* larger#than#family**and%industrial farms*in=Rodefeld@s
terms $ are* m u c h as*in=previous decades* exhibiting!a#demonstrable
increase*in=their shares|of|aggregate production* sales* assets* profits*
use |of| hired labour**and%so on@
Second* however* {it] [is] argued that small farms {are] increasing *in^
numbers *and% holding their own *in= terms |of| acreage *and% aggregate
production@ This bolstered status |of| small farms[is]considered{to]be
m a d e possible largely{by]access |of| their operators*and%other family
m e m b e r s {to] off#farm w a g e income@ U@S@ small farmers {have] b e e n
noted{to]have* on average* total family i n c o m e s*in=excess |of| those
|of| medium#sized farmers*and%*the^U@S@ median family income because
|of| *the^ higher than average educational attainments |of| members |of|
small farm households *and% their consequent ability {to] acquire highly
remunerative off#farm work@ While outside|of|*the^South small farmers
^have* tended{to]{have]higher educational levels than larger* full#time
operators{for}several decades**the^educational disparities b e t w e e n
small* part#time *and% larger* full#time operators {are] suggested {to] *have^
increased@ This {has] often been explained *in= terms |of| *the^ 1970s
$population turnaround$ *in= which well#educated persons* many |of|
urban origin* {were] attracted {to] farm residential situations {for} lifestyle
reasons@
Finally*{it][is]asserted that there{has]b e e n!a#$disappearing middle$
$ that medium#size (traditional family#type) farms {have] decreased
*in& numbers *and% *in= shares |of| aggregate output* assets* profits* and+
*the^ like@ {it] {has] been suggested that these medium#size farms enjoy
neither *the^ advantages |of| bigness or |of| smallness@ *in= particular*
$middle farmers*$ because they {have] operations too large {to] b e
farmed on!a#part#time basis* tend{to]lack*the^labour resources (and
often*the^educational qualifications){to]participate effectively*in=off#
farm labour markets@ T h u s $the middle$[is]tending {to] $disappear*$
albeit slowly@ Approximately 5* 70* *and% 25 percent |of| U@S@ farms *are^
seen {to] lie* respectively* *in= *the^ $large*$ $small*$ *and% $medium$
categories@
These stylized facts appear {to] b e largely accurate* although
Ehrensaft et al@**and%Barlett {have] raised empirical*and%theoretical
objections{to]*the^disappearing middle thesis@ S a l a m o n*and%Davis#
Brown@s findings suggest that*the^$disappearing middle$ m a y have
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