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INTRODUCTION
TO OPTIMUM
DESIGN
FOURTH EDITION
Jasbir Singh Arora
The University of Iowa,
College of Engineering,
Iowa City, Iowa
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This book and the individual contributions contained in it are protected under copyright by the
Publisher (other than as may be noted herein).
Notices
Knowledge and best practice in this field are constantly changing. As new research and experience
broaden our understanding, changes in research methods, professional practices, or medical treatment
may become necessary.
Practitioners and researchers must always rely on their own experience and knowledge in evaluating
and using any information, methods, compounds, or experiments described herein. In using such infor-
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any liability for any injury and/or damage to persons or property as a matter of products liability,
negligence or otherwise, or from any use or operation of any methods, products, instructions, or ideas
contained in the material herein.
ISBN: 978-0-12-800806-5
Introduction to Optimum Design, Fourth Edi- basis for a first course on optimum design.
tion presents an organized approach to opti- More detailed calculations are included in
mizing the design of engineering systems. examples to illustrate procedures. Key ideas
Basic concepts and procedures of optimiza- of various concepts are highlighted for easy
tion are illustrated with examples to show reference.
their applicability to engineering design Practical applications of optimization
problems. The crucial step of formulating techniques are expanding rapidly, and to
a design optimization problem is detailed. illustrate this, engineering design exam-
The numerical process of refining an initial ples are presented throughout the text.
formulation of the optimization problem Chapters: Optimum Design: Numerical
is illustrated and how to correct the initial Solution Process and Excel Solver, Opti-
formulation if it fails to produce a solution mum Design with MATLAB®, and Practical
or does not produce an acceptable solution. Applications of Optimization also address
This as well as other numerical aspects of various aspects of practical applications of
the solution process for optimum design are optimization with example problems. Many
consolidated in chapter: Optimum Design: projects (identified with a *) based on prac-
Numerical Solution Process and Excel Solver. tical applications are included as exercises
The pedagogical aspects of the material at the end of chapters: Optimum Design
are addressed from student as well as teach- Problem Formulation, Graphical Solution
er’s point of view. This is particularly true Method and Basic Optimization Concepts,
for the material of chapters: Introduction Optimum Design: Numerical Solution Pro-
to Design Optimization, Optimum Design cess and Excel Solver, Optimum Design
Problem Formulation, Graphical Solution with MATLAB®, Practical Applications of
Method and Basic Optimization Concepts, Optimization, Discrete Variable Optimum
Optimum Design Concepts: Optimality Con- Design Concepts and Methods, and Nature-
ditions, More on Optimum Design Concepts: Inspired Search Methods.
Optimality Conditions, Optimum Design: Direct search methods that do not use any
Numerical Solution Process and Excel Solver, stochastic ideas in their search are presented
Optimum Design with MATLAB®, Linear in chapter: More on Numerical Methods for
Programming Methods for Optimum Design, Unconstrained Optimum Design and those
More on Linear Programming Methods for that do are presented in chapter: Nature-
Optimum Design, Numerical Methods for Inspired Search Methods. These methods do
Unconstrained Optimum Design, More on not require derivatives of the problem func-
Numerical Methods for Unconstrained Opti- tions, therefore they are broadly applicable
mum Design, and Numerical Methods for for practical applications and they are rela-
Constrained Optimum Design that forms the tively easy to program and use.
xv
xvi Preface to Fourth Edition
The instructor’s manual that accompanies for students. Three courses are suggested,
the text is comprehensive and user-friendly. although several variations are possible.
The original problem statement for each
exercise is included in the solution page for Undergraduate/First-Year Graduate
easy reference. Many MATLAB and Excel
Level Course
files for the problems are also included.
The text is broadly divided into three Topics for an undergraduate and/or first-
parts. Part I presents the basic concepts year graduate course include:
related to optimum design and optimal-
• Formulation of optimization problems
ity conditions in chapters: Introduction to
(see chapters: Introduction to Design
Design Optimization 2 Optimum Design
Optimization and Optimum Design
Problem Formulation, Graphical Solution
Problem Formulation)
Method and Basic Optimization Concepts,
• Optimization concepts using the
Optimum Design Concepts: Optimality Con-
graphical method (see chapter: Graphical
ditions, and More on Optimum Design Con-
Solution Method and Basic Optimization
cepts: Optimality Conditions. Part II treats
Concepts)
numerical methods for continuous variable
• Optimality conditions for unconstrained
smooth optimization problems and their
and constrained problems (see chapter:
practical applications in chapters: Opti-
Optimum Design Concepts: Optimality
mum Design: Numerical Solution Process
Conditions)
and Excel Solver, Optimum Design with
• Use of Excel and MATLAB® illustrating
MATLAB®, Linear Programming Methods
optimum design of practical problems
for Optimum Design, More on Linear Pro-
(see chapters: Optimum Design:
gramming Methods for Optimum Design,
Numerical Solution Process and Excel
Numerical Methods for Unconstrained
Solver and Optimum Design with
Optimum Design, More on Numerical Meth-
MATLAB®)
ods for Unconstrained Optimum Design,
• Linear programming (see chapter: Linear
Numerical Methods for Constrained Opti-
Programming Methods for Optimum
mum Design, More on Numerical Methods
Design)
for Constrained Optimum Design, Practical
• Numerical methods for unconstrained
Applications of Optimization. Part III con-
and constrained problems (see chapters:
tains advanced topics of practical interest on
Numerical Methods for Unconstrained
optimum design, including nature-inspired
Optimum Design and Numerical
metaheuristic methods that do not require
Methods for Constrained Optimum
derivatives of the problem functions in
Design)
chapters: Discrete Variable Optimum Design
Concepts and Methods, Global Optimization The use of Excel and MATLAB should be
Concepts and Methods, Nature-Inspired introduced mid-semester so that students
Search Methods, Multi-objective Optimum have a chance to formulate and solve more
Design Concepts and Methods, and Addi- challenging projects by the semester’s end.
tional Topics on Optimum Design. Note that project exercises and sections with
Introduction to Optimum Design, Fourth advanced material are marked with an aster-
Edition can be used to design several types of isk (*) next to section headings, which means
courses depending on the learning objectives that they may be omitted for this course.
Preface to Fourth Edition xvii
Acknowledgments
I would like to thank my colleague Pro- classroom instruction and provided me with
fessor Karim Abdel-Malek, Director of the their input.
Center for Computer-Aided Design and Pro- I acknowledge the input of my current
fessor of Biomedical Engineering at The Uni- and former colleagues at The University of
versity of Iowa, for his enthusiastic support Iowa who have used the book to teach an
for this book and for the exciting research on undergraduate course on optimum design:
optimization-based modeling, simulation Professors Karim Abdel-Malek, Asghar
and biomechanics of human motion. Bhatti, Kyung Choi, Vijay Goel, Ray Han,
I would like to thank the following col- Harry Kane, George Lance, and Emad
leagues for their contribution to some Tanbour.
chapters: Professor Tae Hee Lee – chapter: Thesis works of all of my current and for-
Optimum Design with MATLAB®; Dr Tim mer graduate students on various topics of
Marler – chapter: Multi-objective Optimum optimization have been invaluable in broad-
Design Concepts and Methods; Professor ening of my horizons on the subject.
G. J. Park – chapter: Additional Topics on I would like to thank reviewers of various
Optimum Design; and Dr Marcelo A. da parts of the book for their suggestions.
Silva and Dr. Qian Wang – chapter: Practical I would also like to thank Steve Merken,
Applications of Optimization. Peter Jardim, Kiruthika Govindaraju and
Dr Yujiang Xiang, Dr Rajan Bhatt, their team at Elsevier for their superb han-
Dr Hyun-Joon Chung, John Nicholson and dling of the manuscript and production of
Robert Lucente provided valuable input for the book.
some chapters. I value the support of the Department of
I appreciate the help given by Jun Choi, Civil and Environmental Engineering, Cen-
John Nicholson, Palani Permeswaran, Karlin ter for Computer-Aided Design, College of
Stutzman, Dr Hyun-Jung Kwon, Dr Hyun- Engineering, and The University of Iowa for
Joon Chung, and Dr Mohammad Bataineh this project.
with parts of the instructor’s manual. Finally, I highly appreciate the uncondi-
I would like to thank instructors at vari- tional love and support of my family and
ous universities who have used the book for friends.
xix
Key Symbols and Abbreviations
xxi
C H A P T E R
1
Introduction to Design Optimization
3
4 1. Introduction to Design Optimization
as results without rigorous proofs. However, their implications from an engineering point of
view are discussed.
Any problem in which certain parameters need to be determined to satisfy constraints
can be formulated as an optimization problem. Once this has been done, optimization con-
cepts and methods described in this text can be used to solve it. Optimization methods are
quite general, having a wide range of applicability in diverse fields. It is not possible to
discuss every application of optimization concepts and methods in this introductory text.
However, using simple applications, we discuss concepts, fundamental principles, and basic
techniques that are used in most applications. The student should understand them with-
out getting bogged down with notations, terminologies, and details on particular areas of
application.
The basic concepts are described in this text to aid the engineer in designing systems at
minimum cost.
The design process should be well organized. To discuss it, we consider a system evolu-
tion model, shown in Fig. 1.1, where the process begins with the identification of a need that
may be conceived by engineers or nonengineers. The five steps of the model in the figure are
described in the following paragraphs.
1. The first step in the evolutionary process is to precisely define the specifications for the
system. Considerable interaction between the engineer and the sponsor of the project is
usually necessary to quantify the system specifications.
2. The second step in the process is to develop a preliminary design of the system. Various
system concepts are studied. Since this must be done in a relatively short time, simplified
models are used at this stage. Various subsystems are identified and their preliminary
designs are estimated. Decisions made at this stage generally influence the system’s
final appearance and performance. At the end of the preliminary design phase, a few
promising design concepts that need further analysis are identified.
3. The third step in the process is a detailed design for all subsystems using the iterative
process described earlier. To evaluate various possibilities, this must be done for all
previously identified promising design concepts. The design parameters for the
subsystems must be identified. The system performance requirements must be
identified and formulated. The subsystems must be designed to maximize system
worth or to minimize a measure of the cost. Systematic optimization methods described
in this text aid the designer in accelerating the detailed design process. At the end of
the process, a description of the final design is available in the form of reports and
drawings.
4. The fourth and fifth steps shown in Fig. 1.1 may or may not be necessary for all systems.
They involve fabrication of a prototype system and testing, and are necessary when the
system must be mass-produced or when human lives are involved. These steps may
appear to be the final ones in the design process, but they are not because the system
may not perform according to specifications during the testing phase. Therefore, the
specifications may have to be modified or other concepts may have to be studied. In fact,
this reexamination may be necessary at any point during the design process. It is for this
reason that feedback loops are placed at every stage of the system evolution process, as
shown in Fig. 1.1. This iterative process must be continued until the best system evolves.
Depending on the complexity of the system, this process may take a few days or several
months.
The model described in Fig. 1.1 is a simplified block diagram for system evolution. In
a ctual practice, each block may be broken down into several subblocks to carry out the stud-
ies properly and arrive at rational decisions. The important point is that optimization concepts
and methods are helpful at every stage of the process. Such methods, along with the appropriate
software, can be useful in studying various design possibilities rapidly.
FIGURE 1.2 Comparison of: (a) conventional design method; and (b) optimum design method.
0. The optimum design method has block 0, where the problem is formulated as one of
optimization (see chapter: Optimum Design Problem Formulation for detailed discus-
sion). An objective function is defined that measures the merits of different designs.
1. Both methods require data to describe the system in block 1.
2. Both methods require an initial design estimate in block 2.
3. Both methods require analysis of the system in block 3.
4. In block 4, the conventional design method checks to ensure that the performance
criteria are met, whereas the optimum design method checks for satisfaction of all of the
constraints for the problem formulated in block 0.
5. In block 5, stopping criteria for the two methods are checked, and the iteration is stopped
if the specified stopping criteria are met.
6. In block 6, the conventional design method updates the design based on the designer’s
experience and intuition and other information gathered from one or more trial designs;
the optimum design method uses optimization concepts and procedures to update the
current design.
The foregoing distinction between the two design approaches indicates that the conven-
tional design process is less formal. An objective function that measures a design’s merit is
not identified. Trend information is usually not calculated; nor is it used in block 6 to make
design decisions for system improvement. In contrast, the optimization process is more for-
mal, using trend information to make design changes.
A point is an ordered list of numbers. Thus, (x1, x2) is a point consisting of two numbers where-
as (x1, x2, …, xn) is a point consisting of n numbers. Such a point is often called an n-tuple. The
n components x1, x2, …, xn are collected into a column vector as
x1
x2 T
x= = x1 x2 . . . xn (1.1)
xn
where the superscript T denotes the transpose of a vector or a matrix. This is called an n-vector.
Each number xi is called a component of the (point) vector. Thus, x1 is the first component, x2
is the second, and so on.
We also use the following notation to represent a point or a vector in the n-dimensional
space:
x = ( x1 , x2 , . . . , xn ) (1.2)
In 3-dimensional (3D) space, the vector x = [x1 x2 x3]T represents a point P, as shown in
Fig. 1.3. Similarly, when there are n components in a vector, as in Eqs. (1.1) and (1.2), x is
interpreted as a point in the n-dimensional space, denoted as Rn. The space Rn is simply the
collection of all n
-dimensional vectors (points) of real numbers. For example, the real line is
R1, the plane is R2, and so on.
The terms vector and point are used interchangeably, and lowercase letters in roman
boldface are used to denote them. Uppercase letters in roman boldface represent matrices.
1.5.2 Sets
Often we deal with sets of points satisfying certain conditions. For example, we may con-
sider a set S of all points having three components, with the last having a fixed value of 3,
which is written as
{
S = x = ( x1 , x2 , x3 ) x3 = 3 } (1.3)
Information about the set is contained in braces ({ }). Eq. (1.3) reads as “S equals the set of
all points (x1, x2, x3) with x3 = 3.” The vertical bar divides information about the set S into two
parts: To the left of the bar is the dimension of points in the set; to the right are the properties
that distinguish those points from others not in the set (eg, properties a point must possess
to be in the set S).
Members of a set are sometimes called elements. If a point x is an element of the set S, then
we write x ∈ S. The expression x ∈ S is read as “x is an element of (belongs to) S.” Conversely,
the expression “y ∉ S” is read as “y is not an element of (does not belong to) S.”
If all the elements of a set S are also elements of another set T, then S is said to be a subset of
T. Symbolically, we write S ⊂ T, which is read as “S is a subset of T” or “S is contained in T.”
Alternatively, we say “T is a superset of S,” which is written as T ⊃ S.
As an example of a set S, consider a domain of the xl – x2 plane enclosed by a circle of radius
3 with the center at the point (4, 4), as shown in Fig. 1.4. Mathematically, all points within and
on the circle can be expressed as
{
S = x ∈R2 }
( x1 − 4)2 + ( x2 − 4)2 ≤ 9 (1.4)
Thus, the center of the circle (4, 4) is in the set S because it satisfies the inequality in Eq. (1.4).
We write this as (4, 4) ∈ S. The origin of coordinates (0, 0) does not belong to the set because it
does not satisfy the inequality in Eq. (1.4). We write this as (0, 0) ∉ S. It can be verified that the
following points belong to the set: (3, 3), (2, 2), (3, 2), (6, 6). In fact, set S has an infinite number
of points. Many other points are not in the set. It can be verified that the following points are
not in the set: (1, 1), (8, 8), and (−1, 2).
{ }
FIGURE 1.4 Geometrical representation of the set S = x ( x1 − 4 ) + ( x 2 − 4 ) ≤ 9 .
2 2
A constraint of this form is a “less than or equal to type” constraint and is abbreviated as
“≤ type.” Similarly, there are greater than or equal to type constraints, abbreviated as “≥ type.”
Both are called inequality constraints.
This represents the numbers x1, x2, …, xn. Note that “i = 1 to n” represents the range for the
index i and is read, “i goes from 1 to n.” Similarly, a set of k vectors, each having n compo-
nents, is represented by the superscript notation as
x ( j ) ; j = 1 to k (1.7)
This represents the k vectors x(l), x(2), …, x(k). It is important to note that subscript i in Eq. (1.6)
and superscript j in Eq. (1.7) are free indices; that is, they can be replaced by any other variable.
For example, Eq. (1.6) can also be written as xj, j = 1 to n and Eq. (1.7) can be written as x(i), i = 1
to k. Note that the superscript j in Eq. (1.7) does not represent the power of x. It is an index that
represents the jth vector of a set of vectors.
We also use the summation notation quite frequently. For example,
c = x1 y1 + x2 y2 + . . . + xn yn (1.8)
is written as
n
c = ∑ xi yi (1.9)
i=1
There is another way of writing the matrix multiplication of Eq. (1.10). Let m-dimensional
vectors a(i); i = 1 to n represent columns of the matrix A. Then y = Ax is also written as
n
y = ∑ a( j ) x j = a(1) x1 + a( 2 ) x2 + . . . + a( n) xn (1.12)
j=1
The sum on the right side of Eq. (1.12) is said to be a linear combination of columns of matrix
A with xj, j = 1 to n as its multipliers. Or y is given as a linear combination of columns of A
(refer Appendix A for further discussion of the linear combination of vectors).
Occasionally, we must use the double summation notation. For example, assuming m = n
and substituting yi from Eq. (1.11) into Eq. (1.9), we obtain the double sum as
n n n n
c = ∑ xi ∑ aij x j = ∑ ∑ aij xi x j (1.13)
i=1 j=1 i=1 j=1
Note that the indices i and j in Eq. (1.13) can be interchanged. This is possible because c is
a scalar quantity, so its value is not affected by whether we sum first on i or on j. Eq. (1.13) can
also be written in the matrix form, as we will see later.
Thus, the dot product is a sum of the product of corresponding elements of the vectors x
and y. Two vectors are said to be orthogonal (normal) if their dot product is 0; that is, x and y
are orthogonal if (x • y) = 0. If the vectors are not orthogonal, the angle between them can be
calculated from the definition of the dot product:
(x • y) = x y cos θ , (1.15)
where u is the angle between vectors x and y, and ‖x‖ represents the length of vector x (also
called the norm of the vector). The length of vector x is defined as the square root of the sum of
squares of the components:
The double sum of Eq. (1.13) can be written in the matrix form as follows:
n n n n
c = ∑ ∑ aij xi x j = ∑ xi ∑ aij x j = x T Ax (1.17)
i=1 j=1 i=1 j=1
Since Ax represents a vector, the triple product of Eq. (1.17) is also written as a dot product:
c = x T Ax = ( x • Ax ) (1.18)
f ( x ) = f ( x1 , x2 , . . . , xn ) (1.19)
We deal with many functions of vector variables. To distinguish between functions, sub-
scripts are used. Thus, the ith function is written as
gi ( x ) = gi ( x1 , x2 , . . . , xn ) (1.20)
If there are m functions gi(x), i = 1 to m, these are represented in the vector form
g1 ( x )
g (x )
g (x) = 2 = g1 ( x ) g2 ( x ) . . . gm ( x )
T
(1.21)
:
gm ( x )
Throughout the text it is assumed that all functions are continuous and at least twice continu-
ously differentiable. A function f(x) of n variables is continuous at a point x* if, for any ε > 0,
there is a d > 0 such that
f ( x ) − f ( x *) < ε (1.22)
whenever ‖x − x*‖ < d. Thus, for all points x in a small neighborhood of point x*, a change in
the function value from x* to x is small when the function is continuous. A continuous func-
tion need not be differentiable. Twice-continuous differentiability of a function implies not only
that it is differentiable two times, but also that its second derivative is continuous.
Fig. 1.5a,b shows continuous and discontinuous functions. The function in Fig. 1.5a is
differentiable everywhere, whereas the function in Fig. 1.5b is not differentiable at points
x1, x2, and x3. Fig. 1.5c is an example in which f is not a function because it has infinite values
at x1. Fig. 1.5d is an example of a discontinuous function. As examples, functions f(x) = x3
FIGURE 1.5 Continuous and discontinuous functions. (a) and (b) Continuous functions; (c) not a function; and
(d) discontinuous function.
and f(x) = sinx are continuous everywhere and are also continuously differentiable. However,
function f(x) = |x| is continuous everywhere but not differentiable at x = 0.
∂ f (x )
; i = 1 to n (1.23)
∂ xi
The n partial derivatives in Eq. (1.23) are usually arranged in a column vector known as the
gradient of the function f(x). The gradient is written as ∂f/∂x or ∇f(x). Therefore,
[∂ f (x )]/∂ x1
∂ f (x ) [∂ f (x )]/∂ x2 (1.24)
∇f (x ) = =
∂x
[∂ f (x )]/∂ x
n
∂2 f ( x )
; i , j = 1 to n (1.25)
∂ xi ∂ x j
We see that there are n2 partial derivatives in Eq. (1.25). These can be arranged in a matrix
known as the Hessian matrix, written as H(x), or simply the matrix of second partial deriva-
tives of f(x), written as ∇2f(x):
∂ 2 f (x )
H(x ) = ∇ 2 f (x ) = (1.26)
∂ xi ∂ x j n× n
Note that if f(x) is continuously differentiable two times, then Hessian matrix H(x) in
Eq. (1.26) is symmetric.
∂g( x)
∇g ( x ) = = ∇g1 ( x ) ∇g2 ( x ) … ∇gm ( x ) n× m (1.27)
∂x
∂gj
A = aij n× m ; aij = ; i = 1 to n; j = 1 to m (1.28)
∂ xi
conversion factors for the most commonly used quantities. For a complete list of conversion
factors, consult the IEEE/ASTM (2010) publication.
Reference
IEEE/ASTM, 2010. American National Standard for Metric Practice. SI 10-2010. The Institute of Electrical and
Electronics Engineers/American Society for Testing of Materials, New York.
2
Optimum Design Problem
Formulation
It is generally accepted that the proper definition and formulation of a problem take more than
50% of the total effort needed to solve it. Therefore, it is critical to follow well-defined proce-
dures for formulating design optimization problems. In this chapter, we describe the process
of transforming the design of a selected system and/or subsystem into an optimum design
problem. Methods for solving the problem will be discussed in subsequent chapters; here we
focus on properly formulating the problem as an optimization problem.
Several simple and moderately complex applications are discussed in this chapter
to illustrate the problem formulation process. More advanced applications are dis-
cussed in Chapters 6 and 7 and 14–19.
The importance of properly formulating a design optimization problem must be stressed be-
cause the optimum solution will be only as good as the formulation. For example, if we for-
get to include a critical constraint in the formulation, the optimum solution will most likely
violate it. Also, if we have too many constraints, or if they are inconsistent, there may be no
solution for the problem. However, once the problem is properly formulated, good software
is usually available to solve it.
It is important to note that the process of developing a proper formulation for optimum de-
sign of practical problems is iterative in itself. Several iterations usually are needed to revise
Introduction to Optimum Design. http://dx.doi.org/10.1016/B978-0-12-800806-5.00002-0
Copyright © 2017 Elsevier Inc. All rights reserved.
19
20 2. Optimum Design Problem Formulation
the formulation before an acceptable one is finalized. This iterative process is further dis-
cussed in chapter: Optimum Design: Numerical Solution Process and Excel Solver.
For most design optimization problems, we will use the following five-step procedure to
formulate the problem:
Step 1: Project/problem description
Step 2: Data and information collection
Step 3: Definition of design variables
Step 4: Optimization criterion
Step 5: Formulation of constraints
Formulation of an optimum design problem implies translating a descriptive state-
ment of the problem into a well-defined mathematical statement.
We will describe the tasks to be performed in each of the foregoing five steps to develop
a mathematical formulation for the design optimization problem. These steps are illustrated
with some examples in this section and in later sections.
3 ≤ thickness ≤ 15 mm (b)
1 1 1 4 1
I= w × w 3 − ( w − 2 t ) × ( w − 2 t )3 = w − (w − 2t)4, mm 4 (d)
12 12 12 12
1 w 1 (w − 2t) 1 3 1
Q = w 2 × − ( w − 2 t )2 × = w − (w − 2t)3, mm 3 (e)
2 4 2 4 8 8
V = P, N (g)
Mw
σ= , N/mm 2 (h)
2l
VQ
τ= , N/mm 2 (i)
2 It
PL3
q= , mm (j)
3EI
Note that the design variables are defined precisely including the units to be used for them.
It is also noted here that an alternate set of design variables can be selected: wo = outer width of the
section, and wi = inner width of the section. The problem can be formulated using these design vari-
ables. However, note that all the expressions given in Eqs. (c)–(j) will have to be re-derived in terms
of wo and wi. Thus the two formulations will look quite different from each other for the same design
problem. However, these two formulations should yield same final solution.
THIRD SUB-CLASS—CYCLOSTOMATA.
Skeleton cartilaginous and notochordal, without ribs and without
real jaws. Skull not separate from the vertebral column. No limbs.
Gills in the form of fixed sacs, without branchial arches, six or seven
in number on each side. One nasal aperture only. Heart without
bulbus arteriosus. Mouth anterior, surrounded by a circular or
subcircular lip, suctorial. Alimentary canal straight, simple, without
coecal appendages, pancreas or spleen. Generative outlet
peritoneal. Vertical fins rayed.
The Cyclostomes are most probably a very ancient type.
Unfortunately the organs of these creatures are too soft to be
preserved, with the exception of the horny denticles with which the
mouth of some of them is armed. And, indeed, dental plates, which
are very similar to those of Myxine, are not uncommon in certain
strata of Devonian and Silurian age (see p. 193). The fishes
belonging to this sub-class may be divided into two families—
First Family—Petromyzontidæ.
Body eel-shaped, naked. Subject to a metamorphosis; in the
perfect stage with a suctorial mouth armed with teeth, simple or
multicuspid, horny, sitting on a soft papilla. Maxillary, mandibulary,
lingual, and suctorial teeth may be distinguished. Eyes present (in
mature animals). External nasal aperture in the middle of the upper
side of the head. The nasal duct terminates without perforating the
palate. Seven branchial sacs and apertures on each side behind the
head; the inner branchial ducts terminate in a separate common
tube. Intestine with a spiral valve. Eggs small. The larvæ without
teeth, and with a single continuous vertical fin.
“Lampreys” are found in the rivers and on the coasts of the
temperate regions of the northern and southern hemispheres. Their
habits are but incompletely known, but so much is certain that at
least some of them ascend rivers periodically, for the purpose of
spawning, and that the young pass several years in rivers, whilst
they undergo a metamorphosis (see p. 170). They feed on other
fishes, to which they suck themselves fast, scraping off the flesh with
their teeth. Whilst thus engaged they are carried about by their
victim; Salmon have been captured in the middle course of the Rhine
with the Marine Lamprey attached to them.
FOURTH SUB-CLASS—LEPTOCARDII.
Skeleton membrano-cartilaginous and notochordal, ribless. No
brain. Pulsating sinuses in place of a heart. Blood colourless.
Respiratory cavity confluent with the abdominal cavity; branchial
clefts in great number, the water being expelled by an opening in
front of the vent. Jaws none.
This sub-class is represented by a single family (Cirrostomi) and
by a single genus (Branchiostoma);[48] it is the lowest in the scale of
fishes, and lacks so many characteristics, not only of this class, but
of the vertebrata generally, that Hæckel, with good reason,
separates it into a separate class, that of Acrania. The various parts
of its organisation have been duly noticed in the first part of this
work.
The “Lancelet” (Branchiostoma lanceolatum, see Fig. 28, p. 63),
seems to be almost cosmopolitan within the temperate and tropical
zones. Its small size, its transparency, and the rapidity with which it
is able to bury itself in the sand, are the causes why it escapes so
readily observation, even at localities where it is known to be
common. Shallow, sandy parts of the coasts seem to be the places
on which it may be looked for. It has been found on many localities of
the British, and generally European coasts, in North America, the
West Indies, Brazil, Peru, Tasmania, Australia, and Borneo. It rarely
exceeds a length of three inches. A smaller species, in which the
dorsal fringe is distinctly higher and rayed, and in which the caudal
fringe is absent, has been described under the name of
Epigionichthys pulchellus; it was found in Moreton Bay.
APPENDIX.
DIRECTIONS FOR COLLECTING AND PRESERVING FISHES.
Whenever practicable fishes ought to be preserved in spirits.
To insure success in preserving specimens the best and
strongest spirits should be procured, which, if necessary, can be
reduced to the strength required during the journey with water or
weaker spirit. Travellers frequently have great difficulties in procuring
spirits during their journey, and therefore it is advisable, especially
during sea voyages, that the traveller should take a sufficient
quantity with him. Pure spirits of wine is best. Methylated spirits may
be recommended on account of their cheapness; however,
specimens do not keep equally well in this fluid, and very valuable
objects, or such as are destined for minute anatomical examination,
should always be kept in pure spirits of wine. If the collector has
exhausted his supply of spirits he may use arrack, cognac, or rum,
provided that the fluids contain a sufficient quantity of alcohol.
Generally speaking, spirits which, without being previously heated,
can be ignited by a match or taper, may be used for the purposes of
conservation. The best method to test the strength of the spirits is
the use of a hydrometer. It is immersed in the fluid to be measured,
and the deeper it sinks the stronger is the spirit. On its scale the
number 0 signifies what is called proof spirit, the lowest degree of
strength which can be used for the conservation of fish for any length
of time. Spirits, in which specimens are packed permanently, should
be from 40 to 60 above proof. If the hydrometers are made of glass
they are easily broken, and therefore the traveller had better provide
himself with three or four of them, their cost being very trifling.
Further, the collector will find a small distilling apparatus very useful.
By its means he is able not only to distil weak and deteriorated spirits
or any other fluid containing alcohol, but also, in case of necessity, to
prepare a small quantity of drinkable spirits.
Of collecting vessels we mention first those which the collector
requires for daily use. Most convenient are four-sided boxes made of
zinc, 18 in. high, 12 in. broad, and 5 in. wide. They have a round
opening at the top of 4 in. diameter, which can be closed by a strong
cover of zinc of 5 in. diameter, the cover being screwed into a raised
rim round the opening. In order to render the cover air-tight, an
indiarubber ring is fixed below its margin. Each of these zinc boxes
fits into a wooden case, the lid of which is provided with hinges and
fastenings, and which on each side has a handle of leather or rope,
so that the box can be easily shifted from one place to another.
These boxes are in fact made from the pattern of the ammunition
cases used in the British army, and extremely convenient, because a
pair can be easily carried strapped over the shoulders of a man or
across the back of a mule. The collector requires at least two, still
better four or six, of these boxes. All those specimens which are
received during the day are deposited in them, in order to allow them
to be thoroughly penetrated by the spirit, which must be renewed
from time to time. They remain there for some time under the
supervision of the collector, and are left in these boxes until they are
hardened and fit for final packing. Of course, other more simple
vessels can be used and substituted for the collecting boxes. For
instance, common earthenware vessels, closed by a cork or an
indiarubber covering, provided they have a wide mouth at the top,
which can be closed so that the spirit does not evaporate, and which
permits of the specimens being inspected at any moment without
trouble. Vessels in which the objects are permanently packed for the
home journey are zinc boxes of various sizes, closely fitting into
wooden cases. Too large a size should be avoided, because the
objects themselves may suffer from the superimposed weight, and
the risk of injury to the case increases with its size. It should hold no
more than 18 cubic feet at most, and what, in accordance with the
size of the specimens, has to be added in length should be deducted
in depth or breadth. The most convenient cases, but not sufficient for
all specimens, are boxes 2 feet in length, 1½ foot broad, and 1 foot
deep. The traveller may provide himself with such cases ready
made, packing in them other articles which he wants during his
journey; or he may find it more convenient to take with him only the
zinc plates cut to the several sizes, and join them into boxes when
they are actually required. The requisite wooden cases can be
procured without much difficulty almost everywhere. No collector
should be without the apparatus and materials for soldering, and he
should be well acquainted with their use. Also a pair of scissors to
cut the zinc plates are useful.
Wooden casks are not suitable for the packing of specimens
preserved in spirits, at least not in tropical climates. They should be
used in cases of necessity only, or for packing of the largest
examples, or for objects preserved in salt or brine.
Very small and delicate specimens should never be packed
together with larger ones, but separately, in small bottles.
Mode of preserving.—All fishes, with the exception of very large
ones (broad kinds exceeding 3–4 feet in length; eel-like kinds more
than 6 feet long), should be preserved in spirits. A deep cut should
be made in the abdomen between the pectoral fins, another in front
of the vent, and one or two more, according to the length of the fish,
along the middle line of the abdomen. These cuts are made partly to
remove the fluid and easily decomposing contents of the intestinal
tract, partly to allow the spirit quickly to penetrate into the interior. In
large fleshy fishes several deep incisions should be made with the
scalpel into the thickest parts of the dorsal and caudal muscles, to
give ready entrance to the spirits. The specimens are then placed in
one of the provisional boxes, in order to extract, by means of the
spirit, the water of which fishes contain a large quantity. After a few
days (in hot climates after 24 or 48 hours) the specimens are
transferred into a second box with stronger spirits, and left therein for
several days. A similar third and, in hot climates sometimes a fourth,
transfer is necessary. This depends entirely on the condition of the
specimens. If, after ten or fourteen days of such treatment the
specimens are firm and in good condition, they may be left in the
spirits last used until they are finally packed. But if they should be
soft, very flexible, and discharge a discoloured bloody mucus, they
must be put back in spirits at least 20° over proof. Specimens
showing distinct signs of decomposition should be thrown away, as
they imperil all other specimens in the same vessel. Neither should
any specimen in which decomposition has commenced when found,
be received for the collecting boxes, unless it be of a very rare
species, when the attempt may be made to preserve it separately in
the strongest spirits available. The fresher the specimens to be
preserved are, the better is the chance of keeping them in a perfect
condition. Specimens which have lost their scales, or are otherwise
much injured, should not be kept. Herring-like fishes, and others with
deciduous scales, are better wrapped in thin paper or linen before
being placed in spirits.
The spirits used during this all-important process of preservation
loses, of course, gradually in strength. As long as it keeps 10° under
proof it may still be used for the first stage of preservation, but
weaker spirits should be re-distilled; or, if the collector cannot do this,
it should be at least filtered through powdered charcoal before it is
mixed with stronger spirits. Many collectors are satisfied with
removing the thick sediment collected at the bottom of the vessel,
and use their spirits over and over again without removing from it by
filtration the decomposing matter with which it has been
impregnated, and which entirely neutralises the preserving property
of the spirits. The result is generally the loss of the collection on its
journey home. The collector can easily detect the vitiated character
of his spirits by its bad smell. He must frequently examine his
specimens; and attention to the rules given, with a little practice and
perseverance, after the possible failure of the first trial, will soon
insure to him the safety of his collected treasures. The trouble of
collecting specimens in spirits is infinitely less than that of preserving
skins or dry specimens of any kind.
When a sufficient number of well-preserved examples have been
brought together, they should be sent home by the earliest
opportunity. Each specimen should be wrapped separately in a piece
of linen, or at least soft paper; the specimens are then packed as
close as herrings in the zinc case, so that no free space is left either
at the top or on the sides. When the case is full, the lid is soldered
on, with a round hole about half an inch in diameter near one of the
corners. This hole is left in order to pour the spirit through it into the
case. Care is taken to drive out the air which may remain between
the specimens, and to surround them completely with spirits, until
the case is quite full. Finally, the hole is closed by a small square lid
of tin being soldered over it. In order to see whether the case keeps
in the spirit perfectly, it is turned upside down and left over night.
When all is found to be securely fastened, the zinc case is placed
into the wooden box and ready for transport.
Now and then it happens in tropical climates that collectors are
unable to keep fishes from decomposition even in the strongest
spirits without being able to detect the cause. In such cases a
remedy will be found in mixing a small quantity of arsenic or
sublimate with the spirits; but the collector ought to inform his
correspondent, or the recipient of the collection, of this admixture
having been made.
In former times fishes of every kind, even those of small size,
were preserved dry as flat skins or stuffed. Specimens thus prepared
admit of a very superficial examination only, and therefore this
method of conservation has been abandoned in all larger museums,
and should be employed exceptionally only, for instance on long
voyages overland, during which, owing to the difficulty of transport,
neither spirits nor vessels can be carried. To make up as much as
possible for the imperfection of such specimens, the collector ought
to sketch the fish before it is skinned, and to colour the sketch if the
species is ornamented with colours likely to disappear in the dry
example. Collectors who have the requisite time and skill, ought to
accompany their collections with drawings coloured from the living
fishes; but at the same time it must be remembered that, valuable as
such drawings are if accompanied by the originals from which they
were made, they can never replace the latter, and possess a
subordinate scientific value only.
Very large fishes can be preserved as skins only; and collectors
are strongly recommended to prepare in this manner the largest
examples obtainable, although it will entail some trouble and
expense. So very few large examples are exhibited in museums, the
majority of the species being known from the young stage only, that
the collector will find himself amply recompensed by attending to
these desiderata.
Scaly fishes are skinned thus: with a strong pair of scissors an
incision is made along the median line of the abdomen from the
foremost part of the throat, passing on one side of the base of the
ventral and anal fins, to the root of the caudal fin, the cut being
continued upwards to the back of the tail close to the base of the
caudal. The skin of one side of the fish is then severed with the
scalpel from the underlying muscles to the median line of the back;
the bones which support the dorsal and caudal are cut through, so
that these fins remain attached to the skin. The removal of the skin
of the opposite side is easy. More difficult is the preparation of the
head and scapulary region; the two halves of the scapular arch
which have been severed from each other by the first incision are
pressed towards the right and left, and the spine is severed behind
the head, so that now only the head and shoulder bones remain
attached to the skin. These parts have to be cleaned from the inside,
all soft parts, the branchial and hyoid apparatus, and all smaller
bones, being cut away with the scissors or scraped off with the
scalpel. In many fishes, which are provided with a characteristic
dental apparatus in the pharynx (Labroids, Cyprinoids), the
pharyngeal bones ought to be preserved, and tied with a thread to
the specimen. The skin being now prepared so far, its entire inner
surface as well as the inner side of the head are rubbed with
arsenical soap; cotton-wool, or some other soft material is inserted
into any cavities or hollows, and finally a thin layer of the same
material is placed between the two flaps of the skin. The specimen is
then dried under a slight weight to keep it from shrinking.
The scales of some fishes, as for instance of many kinds of
herrings, are so delicate and deciduous that the mere handling
causes them to rub off easily. Such fishes may be covered with thin
paper (tissue-paper is the best), which is allowed to dry on them
before skinning. There is no need for removing the paper before the
specimen has reached its destination.
Scaleless Fishes, as Siluroids and Sturgeons, are skinned in the
same manner, but the skin can be rolled up over the head; such
skins can also be preserved in spirits, in which case the traveller
may save to himself the trouble of cleaning the head.
Some Sharks are known to attain to a length of 30 feet, and
some Rays to a width of 20 feet. The preservation of such gigantic
specimens is much to be recommended, and although the difficulties
of preserving fishes increase with their size, the operation is
facilitated, because the skins of all Sharks and Rays can easily be
preserved in salt and strong brine. Sharks are skinned much in the
same way as ordinary fishes. In Rays an incision is made not only
from the snout to the end of the fleshy part of the tail, but also a
second across the widest part of the body. When the skin is removed
from the fish, it is placed into a cask with strong brine mixed with
alum, the head occupying the upper part of the cask; this is
necessary, because this part is most likely to show signs of
decomposition, and therefore most requires supervision. When the
preserving fluid has become decidedly weaker from the extracted
blood and water, it is thrown away and replaced by fresh brine. After
a week’s or fortnight’s soaking the skin is taken out of the cask to
allow the fluid to drain off; its inner side is covered with a thin layer of
salt, and after being rolled up (the head being inside) it is packed in a
cask, the bottom of which is covered with salt; all the interstices and
the top are likewise filled with salt. The cask must be perfectly water-
tight.
Of all larger examples of which the skin is prepared, the
measurements should be taken before skinning so as to guide the
taxidermist in stuffing and mounting the specimens.
Skeletons of large osseous fishes are as valuable as their skins.
To preserve them it is only necessary to remove the soft parts of the
abdominal cavity and the larger masses of muscle, the bones being
left in their natural continuity. The remaining flesh is allowed to dry
on the bones, and can be removed by proper maceration at home.
The fins ought to be as carefully attended to as in a skin, and of
scaly fishes so much of the external skin ought to be preserved as is
necessary for the determination of the species, as otherwise it is
generally impossible to determine more than the genus.
A few remarks may be added as regards those Faunæ, which
promise most results to the explorer, with some hints as to desirable
information on the life and economic value of fishes.
It is surprising to find how small the number is of the freshwater
faunæ which may be regarded as well explored; the rivers of Central
Europe, the Lower Nile, the lower and middle course of the Ganges,
and the lower part of the Amazons are almost the only fresh waters
in which collections made without discrimination would not reward
the naturalist. The oceanic areas are much better known; yet almost
everywhere novel forms can be discovered and new observations
made. Most promising and partly quite unknown are the following
districts:—the Arctic Ocean, all coasts south of 38° lat. S., the Cape
of Good Hope, the Persian Gulf, the coasts of Australia (with the
exception of Tasmania, New South Wales, and New Zealand), many
of the little-visited groups of Pacific islands, the coasts of north-
eastern Asia north of 35° lat. N., and the western coasts of North and
South America.
No opportunity should be lost to obtain pelagic forms, especially
the young larva-like stages of development abounding on the
surface of the open ocean. They can be obtained without difficulty by
means of a small narrow meshed net dragged behind the ship. The
sac of the net is about 3 feet deep, and fastened to a strong brass-
ring 2 or 2½ feet in diameter. The net is suspended by three lines
passing into the strong main line. It can only be used when the
vessel moves very slowly, its speed not exceeding three knots an
hour, or when a current passes the ship whilst at anchor. To keep the
net in a vertical position the ring can be weighted at one point of its
circumference; and by using heavier weights two or three drag-nets
can be used simultaneously at different depths. This kind of fishing
should be tried at night as well as day, as many fishes come to the
surface only after sunset. The net must not be left long in the water,
from 5 to 20 minutes only, as delicate objects would be sure to be
destroyed by the force of the water passing through the meshes.
Objects found floating on the surface, as wood, baskets,
seaweed, etc., deserve the attention of the travellers, as they are
generally surrounded by small fishes or other marine animals.
It is of the greatest importance to note the longitude and latitude
at which the objects were collected in the open ocean.
Fishing in great depths by means of the dredge, can be practised
only from vessels specially fitted out for the purpose; and the
success which attended the “Challenger,” and North American Deep-
sea explorations, has developed Deep-sea fishing into such a
speciality that the requisite information can be gathered better by
consulting the reports of those expeditions than from a general
account, such as could be given in the present work.
Fishes offer an extraordinary variety with regard to their habits,
growth, etc., so that it is impossible to enumerate in detail the points
of interest to which the travellers should pay particular attention.
However, the following hints may be useful.
Above all, detailed accounts are desirable of all fishes forming
important articles of trade, or capable of becoming more generally
useful than they are at present. Therefore, deserving of special
attention are the Sturgeons, Gadoids, Thyrsites and Chilodactylus,
Salmonoids, Clupeoids. Wherever these fishes are found in sufficient
abundance, new sources may be opened to trade.
Exact observations should be made on the fishes the flesh of
which is poisonous either constantly or at certain times and certain
localities; the cause of the poisonous qualities as well as the nature
of the poison should be ascertained. Likewise the poison of fishes
provided with special poison-organs requires to be experimentally
examined, especially with regard to its effects on other fishes and
animals generally.
All observations directed to sex, mode of propagation, and
development, will have special interest: thus those relating to
secondary sexual characters, hermaphroditism, numeric proportion
of the sexes, time of spawning and migration, mode of spawning,
construction of nests, care of progeny, change of form during growth,
etc.
If the collector is unable to preserve the largest individuals of a
species that may come under his observation he should note at least
their measurements. There are but few species of fishes of which the
limit of growth is known.
The history of Parasitic Fishes is almost unknown, and any
observations with regard to their relation to their host as well as to
their early life will prove to be valuable; nothing is known of the
propagation of fishes even so common as Echeneis and Fierasfer,
much less of the parasitic Freshwater Siluroids.
The temperature of the blood of the larger freshwater and marine
species should be exactly measured.
Many pelagic and deep-sea fishes are provided with peculiar
small round organs of a mother-of-pearl colour, distributed in series
along the side of the body, especially along the abdomen. Some
zoologists consider these organs as accessory eyes, others (and it
appears to us with better reason) as luminous organs. They deserve
an accurate microscopic examination made on fresh specimens; and
their function should be ascertained from observation of the living
fishes, especially also with regard to the question, whether or not the
luminosity (if such be their function) is subject to the will of the fish.