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PALGRAVE MACMILLAN ASIAN BUSINESS SERIES
Family Business in
China, Volume 2
Challenges and Opportunities
Series Editor
Yingqi Wei, Business School, University of Leeds
Leeds, UK
The Palgrave Macmillan Asian Business Series publishes theoretical and
empirical studies that contribute forward-looking social perspectives on
the study of management issues not just in Asia, but by implication else-
where. The series specifically aims at the development of new frontiers in
the scope, themes and methods of business and management studies in
Asia, a region which is seen as key to studies of modern management,
organization, strategies, human resources and technologies. The series
invites practitioners, policy-makers and academic researchers to join us
at the cutting edge of constructive perspectives on Asian management,
seeking to contribute towards the development of civil societies in Asia
and further a field.
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To submit a book proposal for inclusion in this series please email
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Family Business
in China, Volume 2
Challenges and Opportunities
Ling Chen Jian An Zhu
Zhejiang University Zhejiang University City College
Hangzhou, Zhejiang, China Hangzhou, Zhejiang, China
Hanqing Fang
Business and Information Technology
Missouri University of Science and
Technology
Rolla, MO, USA
© The Editor(s) (if applicable) and The Author(s), under exclusive license to Springer Nature
Switzerland AG 2021
This work is subject to copyright. All rights are solely and exclusively licensed by the Publisher,
whether the whole or part of the material is concerned, specifically the rights of translation, reprinting,
reuse of illustrations, recitation, broadcasting, reproduction on microfilms or in any other physical
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The use of general descriptive names, registered names, trademarks, service marks, etc. in this
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The registered company address is: Gewerbestrasse 11, 6330 Cham, Switzerland
Contents
1 Introduction 1
1.1 Family and Family Business in China 2
1.2 Challenges and Opportunities in Chinese Family
Business 5
1.3 What This Book Is About 6
References 7
2 Family Business in China: Present Status 9
2.1 Defining Family Business in China 10
2.2 Prevalence of Family Business in China 15
2.3 Employment in Chinese Family Business 15
2.4 Economic Scale in Family Businesses 19
2.5 Individual Characteristics of Chinese Family
Entrepreneurs 23
2.6 Differences Between Family and Non-Family
Businesses 26
References 32
v
vi Contents
Postface 169
Index 175
List of Figures
ix
x List of Figures
xi
1
Introduction
The People’s Republic of China1 has been established for over 70 years,
but the history of its private enterprises goes back only a little more
than 40 years. Most of these private enterprises were established after
the launching of reforms and the opening up of the economy in 1978.
Forty years later, the entire society of China has become more dynamic
and creative due to the efforts of a large group of innovative, wealth-
creating entrepreneurs who created new combinations of the production
factors that have made China a powerhouse in the world economy.
In March 2011, The Economist magazine attributed the power that
has driven China’s economic take-off to “Bamboo Capitalism.” It is not
the state-owned enterprises that are changing China, but the private
enterprises whose average return on investment is 10% higher than
that of state-owned enterprises. Thanks to the gradual deregulation
of the economy, entrepreneurial activities have sprung up like after-
the-rain mushrooms. But entrepreneurial activities in China are not
“equally distributed geographically.” In general, the coast areas have more
1 Family business, family firm, and family enterprise are interchangeably used in this book.
© The Author(s), under exclusive license to Springer Nature 1
Switzerland AG 2021
L. Chen et al., Family Business in China, Volume 2,
Palgrave Macmillan Asian Business Series,
https://doi.org/10.1007/978-3-030-51402-0_1
2 L. Chen et al.
2 Shang, Yang (390–338 BC) introduced several social, economic, and legal reforms that funda-
mentally strengthened the power of the State of Qin. These reforms included implementing a
legal system that treated all citizens equally, favoring agriculture over commerce, encouraging
the cultivation of unsettled lands, privatizing land to farmers, assigning free land to soldiers,
and rewarding farmers who exceeded quotas.
1 Introduction 3
span of a private enterprise is only 3.7 years, and for entrepreneurs and
business owners in China, business survival is still the top priority.
Many successful entrepreneurs worry that the title “family business” is
not good for them because family private ownership may be seen as polit-
ically contrary to the economic system of a socialist country. For a long
time in China, family involvement in business has been synonymous
for “obsolete” and “outmoded.” The 40-year history of the development
of Chinese private enterprises has continuously influenced family busi-
nesses, and the sector has increasingly felt the need for traditional family
businesses to change in the direction of being more modern corporate
enterprises.
Although most family businesses cannot yet be called “modern” and
“high-level,” they should not be seen as “low efficiency” and “backward”
(e.g., Chandler, 1990). Indeed, the large numbers of family businesses
and their rapid growth show they must be doing something right. This
combination of family ownership and management, aligned with a long-
term focused orientation, gives family businesses a unique competitive
advantage demonstrating rapid decision-making, high accountability and
strong resilience. At the same time, when corporate leaders, succes-
sors, and important executives come from the same family system, a
closed family human resource pool may limit the development of enter-
prises, and complex family relationships can also hinder the growth of
enterprises.
In China, family businesses differ from other types of organizations
such as state-owned enterprises in terms of their goals and objectives,
strategic planning processes, ownership structures, and firm perfor-
mances. In order to grow their businesses, many entrepreneurs take the
initiative and choose the family governance model where the family
provides the entrepreneur with financial, human, social, and patient
capital. The family holds a high proportion of shares, and many family
members participate in operations to share the benefits of entrepreneur-
ship. Also, the relationship among family members can have a very
synergistic effect, reducing agency costs and improving organizational
performance.
1 Introduction 5
At the same time, faced with the scale discrimination and ownership
discrimination from state-owned banks and other government institu-
tions, family businesses often choose to use a pyramid control structure
in order to hide the true identity of the family. Also it is common for
family businesses to develop political and social relations in order to miti-
gate the strong risk and uncertainty stemming from the discriminations
mentioned before. Controlling families may also encroach on the inter-
ests of small and medium shareholders, which might bring in additional
agency costs to the businesses.
3 Family business in China before 1978 has been discussed in Family business in China, Volume
1: A historical perspective.
1 Introduction 7
References
Chandler, A. D. (1990). Scale and scope: The dynamics of industrial capitalism.
Belknap Press.
Lansberg, I. (1999). Succeeding generations: Realizing the dream of families in
business. Harvard Business Review Press.
2
Family Business in China: Present Status
1 In a broad sense, the term “private enterprise” refers to all economic components except
state-owned and state-controlled enterprises. “Domestic-funded private enterprises” refers to the
broad private economy except the Sino-foreign joint ventures, enterprises with Sino-foreign
cooperation, wholly foreign-owned enterprises, and Hong Kong, Macao, and Taiwan-invested
enterprises. In a narrow sense, the private economy refers to the economies of private enterprises
and privately controlled corporates. We discuss the private economy in a narrow sense.
2 Xi, J. (2018, November 1). Private sector speech at the enterprise forum. Xinhua News Agency.
3 Press conference of the Second Session of the 13th National People’s Congress, March 6,
2019.
and Reform Commission, also noted that “5, 6, 7, 8, 9” does not refer
to strict statistical standards, but rather to the importance of the private
sector to the Chinese economy.
Family business is the most important part of private economy in
China. Nevertheless, there are limited empirical studies on their overall
status and their economic contribution. There are several reasons for this.
First, the history of family business research is relatively short. In fact,
China has only begun to pay attention to family businesses during the
last couple of decades. Concerning the statistical evidence, it is important
for researchers to consider how to define family business, yet the interna-
tional academic community has not yet reached a consensus on how to
clearly distinguish family businesses from non-family businesses. While
there are some family business studies in Germany (Klein, 2000), Sweden
(Morck & Yeung, 2003), and the United States (Astrachan & Shanker,
2003; Heck et al., 2001), the definition and scope of family businesses
and the judgment of what constitutes a family business often diverge.
Thus, a given definition might only be applicable to certain economies.
Second, apart from the information disclosure of publicly-listed
companies, few family businesses in China publicize and disclose their
financial and organizational information. Third, while the distinction
between family businesses and non-family businesses has long been
recognized in family business research, it is only in recent inquiries that
scholars have begun to recognize that family businesses are themselves
quite heterogeneous (Chrisman & colleagues, 2012), and family busi-
nesses can have a variety of different goals, resource endowments, and
governance structures. In the next section, we discuss the various defining
standards that were used in this study.
The academic debate about the definition of family business has been
ongoing for years. For a long time, any family-controlled business was
2 Family Business in China: Present Status 11
Below we discuss how we define family businesses and the overall status
of family businesses in China. There are three major points to be made.
First, there are 70 million self-employed businesses in China, and we
include them as a part of the family business population. Although often
classified as individual workers without a legal form of “enterprise” (Table
4 In Southeast Asia, business group often uses a pyramid structure of ownership in which
separate control rights might deviate from cash flow rights.
2 Family Business in China: Present Status 13
5 Although the national private economy sample survey data includes a large and widely
distributed sample of the national private economy, it still cannot be used as an unbiased
estimate of the family business population in a statistical sense. Therefore, according to the
data in this report, in estimating the overall situation of Chinese family businesses, attention
should be paid to possible unpredictable biases.
2 Family Business in China: Present Status 15
Table 2.3 Number and proportion of family business: Third national economic
census
Organizational
Types
(Unit in Patterns of Enterprise Legal
Million) Categories Person Numbers
Economic Enterprise Domestic State-owned and 0.24
Units Legal Entity Enterprise Collective
Enterprise
Incorporated 1.62
Enterprise
Private Enterprise 5.60
Others 0.09
Foreign Hong Kong, 0.19
Enterprise Macao, and
Taiwan
investment,
Foreign
investment
Licensed Self-employed Business Entities 32.79
Economic Units 40.53
Family Business Number 37.94
Proportion 93.6%
By the end of 2017, the number had fallen to 64.7 million, and the
proportion of the total urban employed population dropped from 20.0
to 15.2%. In 2012, the number of urban employees in private enter-
prises was 75.57 million, but that increased to 133.27 million by the
end of 2017. Thus, the proportion of the total urban employed popu-
lation that was employed in private enterprises increased from 20.3 to
31.4%. From the perspective of the proportion of urban employment
population, the market-oriented reforms in China’s economy have been
advancing, and the labor absorption capacity of the private sector has
expanded greatly. Family firms are the most important component of
the private sector, and the total number of jobs absorbed by family firms
has been increasing.
China has an obvious urban–rural dual structure system. There is a
significant difference in the way resources are allocated in urban and
rural areas. The degree of commercialization in rural areas is low, and
the government’s investment in public facilities such as education and
medical care in rural areas is not as good as those in the cities. Rural
household registration does not have unemployment and pensions. Polit-
ical participation is weak, and the percentage of council representatives
is low. The long-term existence of the dual structure is due to the exis-
tence of institutional barriers such as the household registration system6
between urban and rural areas. It is therefore necessary to look at the
contribution of the private economy, considering the comprehensive
employment data of urban and rural areas. At the end of 2012, there
were 56.43 million individual urban employees and 29.86 individual
rural employees.
We can also look at the number of employed individuals from urban
and rural private enterprises. At the end of 2012, there were 75.57
million people employed by urban private enterprises and 37.39 million
employed by rural private enterprises, for a total of 112.96 million
employees in the private sector. At the end of 2017, there were 133.27
million employed by urban private enterprises and 65.54 million by rural
6 The household registration system in China was initially established in the late 1950s. It was
designed to limit population migration especially from rural to urban areas. Right now the
system is still used to control population mobility and to determine the eligibility of social
welfare.
2 Family Business in China: Present Status 19
Note Foreign enterprises here include companies from Hong Kong, Macao,
Taiwan, and other countries
most family businesses in the sample are in the initial and early growth
stages. Similarly, the eastern region of China, where entrepreneurial
activities are very evident, has also become the most concentrated area
for family businesses (56% of the family business sample companies
in the survey are concentrated in the eastern coastal provinces). The
central provinces account for 24% of the survey sample, and the Western
provinces account for 20%. Many startups have family support behind
them, so the family’s control over the equity of these companies is very
obvious.
According to Li’s (2011) analysis of 212 publicly-traded compa-
nies that are listed on the ChiNext (a NASDAQ-style subsidiary for
newly-created high-growth firms), 200 of them are family-controlled
companies. The average shareholding ratio of the largest family share-
holders before an IPO is as high as 54% in some areas, and 51 of them
hold more than 70% of the shares. There are also 64 companies with 50
to 70% of family control, 61 companies with 30 to 50% control, and
only 23 companies with less than 30% control. Among them, the Zhon-
glei Manguanag Electronic Company of the Li Yuezong family held 99%
of the shares before the issuance and 74% after the issuance.
250
200
150
100
50
0
17 21 23 25 27 29 31 33 35 37 39 41 43 45 47 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 81
Table 2.10 Economic, social, and political status of family business owners
Status Economic Status Social Status Political Status
Rank Percentage Percentage Percentage
Number (%) Number (%) Number (%)
1 45 1.4 46 1.4 36 1.1
2 99 3.0 108 3.3 109 3.3
3 405 12.2 402 12.1 315 9.6
4 436 13.2 428 12.9 348 10.6
5 919 27.7 937 28.3 736 22.5
6 634 19.1 577 17.4 510 15.6
7 347 10.5 342 10.3 344 10.5
8 264 8.0 273 8.3 358 10.9
9 120 3.6 144 4.4 279 8.5
10 46 1.4 52 1.6 242 7.4
References
Astrachan, J. H., & Shanker, M. C. (2003). Family businesses’ contribution to
the US economy: A closer look. Family Business Review, 16 (3), 211–219.
Chayanov, A. V. (1986). The theory of peasant economy. In D. Thorner, B.
Kerblay, & R. E. F. Smith (Ed.).
Chrisman, J. J., Chua, J. H., & Steier, L. (2005). Sources and consequences of
distinctive familiness: An introduction. Entrepreneurship Theory and Practice,
29 (3), 237–247.
Chrisman, J. J., Hofer, C. W., & Boulton, W. B. (1988). Toward a system
for classifying business strategies. Academy of Management Review, 13(3),
413–428.
Chrisman, J. J., Chua, J. H., Pearson, A. W., & Barnett, T. (2012). Family
involvement, family influence, and family–centered non–economic goals in
small firms. Entrepreneurship Theory and Practice, 36 (2), 267–293.
Chua, J. H., Chrisman, J. J., & Sharma, P. (1999). Defining the family business
by behavior. Entrepreneurship Theory and Practice, 23(4), 19–39.
De Geus, A. (2002). The living company. Harvard Business Press.
Heck, R. K., Jasper, C. R., Stafford, K., Winter, M., & Owen, A. J. (2001).
Using a household sampling frame to study family businesses: The 1997
national family business survey. In Databases for the study of entrepreneurship.
Emerald Group Publishing Limited.
Klein, S. B. (2000). Family businesses in Germany: Significance and structure.
Family Business Review, 13(3), 157–182.
Li, J. (2011). The essence of the ChiNext issue is that family listed companies
dominate. Economic Information Daily.
Lansberg, I., Perrow, E. L., & Rogolsky, S. (1988). Family business as an
emerging field. Family Business Review, 1(1), 1–8.
Litz, R. A. (2008). Two sides of a one-sided phenomenon: Conceptualizing
the family business and business family as a Möbius strip. Family Business
Review, 21(3), 217–236.
McKelvey, B. (1982). Organizational systematics: Taxonomy, evolution, classifica-
tion. University of California Press.
Miller, D., Le Breton-Miller, I., Lester, R. H., & Cannella, A. A., Jr. (2007).
Are family firms really superior performers? Journal of Corporate Finance,
13(5), 829–858.
Morck, R., & Yeung, B. (2003). Agency problems in large family business
groups. Entrepreneurship Theory and Practice, 27 (4), 367–382.
2 Family Business in China: Present Status 33
E. Archaeocalamites.
In the Upper Devonian and Culm rocks casts of a well-defined
Calamitean plant are characteristic fossils; stems, leaf-bearing
branches, roots and cones have been described by several authors,
and the genus Archaeocalamites has been instituted for their
reception. Although this genus agrees in certain respects with
Calamites, and as recent work has shown this agreement extends to
internal structure, it has been the custom to regard the Lower
Carboniferous and Devonian plants as genetically distinct. The
surface features of the stem-casts, the form of the leaves, and
apparently the cones, possess certain distinctive characters which
would seem to justify the retention of a separate generic designation.
We may briefly summarise the characteristics of the genus as
follows:—
Pith-casts articulated, with very slightly constricted nodes; the
internodes traversed by longitudinal ribs slightly elevated or almost
flat, separated by shallow grooves. The ribs and grooves are
continuous from one internode to another, and do not usually show
the characteristic alternation of Calamites[834]. Along the nodal line
there are occasionally found short longitudinal depressions, probably
marking the points of origin of outgoing bundles. Branches were
given off from the nodes without any regular order; a pith-cast may
have branch-scars on many of the nodes, or there may be no trace
of branches on casts consisting of several nodes. The leaves[835] are
in whorls; in some cases they occur as free, linear, lanceolate
leaves, or on younger branches they are long, filiform and repeatedly
forked. The structure of the wood agrees with that of some forms of
Arthropitys. The strobili consist of an articulated axis bearing whorls
of sporangiophores, and each sporangiophore has four sporangia.
Our knowledge of the fertile shoots is, however, very imperfect.
Renault[836] has recently described the structure of the wood in
some small silicified stems of Archaeocalamites from Autun. A large
hollow pith is surrounded by a cylinder of wood consisting of wedge-
shaped groups of xylem tracheids associated with secondary
medullary rays; at the apex of each primary xylem group there is a
carinal canal. The primary medullary rays appear to have been
bridged across by bands of xylem at an early stage of secondary
thickening, as in the Calamite of fig. 83, D.
Our knowledge of the cones of Archaeocalamites is far from
satisfactory. Renault[837] has recently described a small fertile branch
bearing a succession of verticils of sporangiophores; each
sporangiophore stands at right angles to the axis of the cone and
bears four sporangia, as in Calamostachys. It is not clear how far
there is better evidence than that afforded by the association of the
specimen with pith-casts of stems, for referring this cone to
Archaeocalamites, but the association of vegetative and fertile
shoots certainly suggests an organic connection. The cone
described by the French author agrees with Equisetum in the
absence of sterile bracts between the whorls of sporangiophores. It
is an interesting fact that such a distinctly Equisetaceous strobilus is
known to have existed in Lower
Carboniferous rocks.
Stur[838] has also described
Archaeocalamites at considerable length;
he gives several good figures of stem-casts
and foliage-shoots bearing long and often
forked narrow leaves. The same writer
describes specimens of imperfectly
preserved cones in which portions of whorls
of forked filiform leaves are given off from
the base of the strobilus[839]. Kidston[840]
published an important memoir on the
cones of Archaeocalamites in 1883, in
which he advanced good evidence in
support of the view that certain strobili,
which were originally described as
Monocotyledonous inflorescences, under
the generic name Pothocites[841], are the
fertile shoots of this Calamarian genus.
Kidston’s conclusions are based on the
occurrence on the Pothocites cones, of
leaves like those of Archaeocalamites, on
the non-alternation of the sporangiophores
of successive whorls, and on the close
resemblance between his specimens and
those described by Stur. Good specimens
of the cones, formerly known as Pothocites,
may be seen in the Botanical Museum in
the Royal Gardens, Edinburgh; as they are Fig. 103.
in the form of casts without internal Archaeocalamites
scrobiculatus
structure it is difficult to form a clear (Schloth.).
conception as to their morphological From a specimen in the
features. Woodwardian Museum,
Cambridge. From the
The fossils included under Carboniferous
Archaeocalamites have been referred by limestone of
different authors to various genera, and
considerable confusion has arisen in both Northumberland. ½ nat.
generic and specific nomenclature. The size.
following synonomy of the best known
species, A. scrobiculatus (Schloth.) illustrates the unfortunate use of
several terms for the same plant.
Affinities of Archaeocalamites.
This genus agrees very closely with Calamites both in the
anatomical structure of the stem and in the verticillate disposition of
the leaves. The strobili appear to be Equisetaceous in character, and
there is no satisfactory evidence of the existence of whorls of sterile
bracts in the cone, such as occur in Calamostachys and in other
Calamitean strobili. The continuous course of the vascular bundles
of the stem from one internode to the next is the most striking feature
in the ordinary specimens of the genus; but it sometimes happens
that the grooves on a pith-cast shew the same alternation at the
node as in Calamites. This is the case in a specimen in the Göppert
collection in the Breslau Museum, and Feistmantel[855] has called
attention to such an alternation in specimens from Rothwaltersdorf.
In the true Calamites, on the other hand, the usual nodal alternation
of the vascular strands is by no means a constant character[856].
Stur[857], Rothpletz[858], and other authors have pointed out the
resemblance of Archaeocalamites to Sphenophyllum. The deeply
divided leaves of some Sphenophyllums and those of
Archaeocalamites are very similar in form; and the course of the
vascular strands in Sphenophyllum may be compared with that in
Archaeocalamites. But the striking difference in the structure of the
stele forms a wide gap between the two genera. We have evidence
that the Calamites and Sphenophyllums were probably descended
from a common ancestral stock, and it may be that in
Archaeocalamites, some of the Sphenophyllum characters have
been retained; but there is no close affinity between the two plants.
On the whole, considering the age of Archaeocalamites and the
few characters with which we are acquainted, it is probable that this
genus is very closely related to the typical Calamites, and may be
regarded as a type which is in the direct line of development of the
more modern Calamite and the living Equisetum. Weiss[859] includes
Archaeocalamites as one of his subgenera with Calamitina and
others, and it is quite possible that the genus has not more claim to
stand alone than other forms at present included in the
comprehensive genus Calamites.
The student will find detailed descriptions of this genus in the
works which have been referred to in the preceding pages.
CHAPTER XI.
II. SPHENOPHYLLALES.
I. SPHENOPHYLLUM.
The genus Sphenophyllum is placed in a special class, as
representing a type which cannot be legitimately included in any of
the existing groups of Vascular Cryptogams. Although this
Palaeozoic genus possesses points of contact with various living
plants, it is generally admitted by palaeobotanists that it constitutes a
somewhat isolated type among the Pteridophytes of the Coal-
Measures. Our knowledge of the anatomy of both vegetative shoots
and strobili is now fairly complete, and the facts that we possess are
in favour of excluding the genus from any of the three main divisions
of the Pteridophyta.
In Scheuchzer’s Herbarium Diluvianum there is a careful drawing
of some fragments of slender twigs, from an English locality, bearing
verticils of cuneiform leaves, which the author compares with the
common Galium[860]. As regards superficial external resemblance,
the Galium of our hedgerows agrees very closely with what must
have been the appearance of fresh green shoots of Sphenophyllum.
A twig of the same species of Sphenophyllum is figured by
Schlotheim[861] in the first part of his work on fossil plants; he regards
it as probably a fragment of some species of Palm. Sternberg[862]
was the first to institute a generic name for this genus of plants, and
specimens were described by him in 1825 as a species of the genus
Rotularia. The name Sphenophyllites was proposed by Brongniart[863]
in 1822 as a substitute for Schlotheim’s genus, and in a later work[864]
the French author instituted the genus Sphenophyllum. Dawson[865]
was the first to make any reference to the anatomy of this genus; but
it is from the examination of the much more perfect material from St.
Étienne, Autun, and other continental localities, the North of England
and Pettycur in Scotland, by Renault, Williamson, Zeiller and Scott,
that our more complete knowledge has been acquired.
The affinity of Sphenophyllum has always been a matter of
speculation; it has been compared with Dicotyledons, Palms,
Conifers (Ginkgo and Phyllocladus), and various Pteridophytes, such
as Ophioglossum, Tmesipteris, Marsilia, Salvinia, Equisetum and the
Lycopodiaceae[866].
DEFINITION.
a. Stems.
i. Primary structure.
In a transverse section of a young Sphenophyllum stem such as
that diagrammatically sketched in fig. 105, A, we find in the centre
the xylem portion of a single stele with a characteristic triangular
form. The primary xylem consists mainly of fairly large tracheae with
numerous pits on their walls; towards the end of each arm the
tracheids become scalariform, and at the apex there is a group of
narrower spiral protoxylem elements. In the British species there is a
single protoxylem group at the apex of each arm, but Renault has
described some French stems in which the stele appears to be
hexarch, having two protoxylem groups at the end of each of the
three rays of the stele. The primary xylem strand of Sphenophyllum
has therefore a root-like structure, the tracheids having been
developed centripetally from the three initial protoxylem groups. This
type of structure is typical of roots, but it also occurs in the stems of
some recent Vascular Cryptogams.
Fig. 104. Diagrammatic longitudinal section of Sphenophyllum.
c, outer cortex; b, space next the stele, originally occupied by phloem
etc.; a, xylem strand. (After Renault[871].) × 7.
As a rule the tissue next the xylem has not been petrified, but in
exceptionally well-preserved examples it is seen to consist of a band
of thin-walled elements, of which those in contact with the xylem may
be spoken of as phloem, and those beyond as the pericycle.
Succeeding this band of delicate tissue there is a broader band of
thicker-walled and somewhat elongated elements, constituting the
cortex. The specimen drawn in fig. 105, A, shows very prominent
grooves in the cortex opposite the middle of each bay of the primary
wood. It is these grooves that give to the ordinary casts of
Sphenophyllum branches the appearance of longitudinal lines
traversing each internode. In a longitudinal section of a stem, the
cortical tissue (fig. 104, c) is found to be broader in the nodal
regions, thus giving rise to the tumid nodes referred to in the
diagnosis. The increased breadth at the nodes does not mean that
the xylem is broader in these regions, as it is in Calamite stems.
Small strands of vascular tissue are given off from the three edges of
the triangular stele (fig. 105 A) at each node; these branch in
passing through the cortex on their way to the verticils of leaves. The
space b in the diagrammatic section of fig. 104 was originally
occupied by the phloem and inner cortex. In some species of
Sphenophyllum the apex of each arm of the xylem strand, as seen in
transverse section, is occupied by a longitudinal canal surrounded by
spiral tracheids, as in the primary xylem of the old stem shown in fig.
105, C.
Fig. 105. Sphenophyllum.
A. Transverse section of young stem.
B. Transverse section of the wood of a young stem; px, protoxylem; x,
secondary xylem. (A and B. Sphenophyllum plurifoliatum.) × 20.
C. Transverse section of an old stem; (S. insigne); a, phloem; b,
periderm; c, fascicular secondary xylem; d, interfascicular
secondary xylem. × 9. (No. 914 in the Williamson Collection.)
D. Longitudinal section of the reticulate tracheae and medullary rays; r, r,
r, of S. plurifoliatum. × 36.
E. Similar section of S. insigne. × 75. (D and E after Williamson and
Scott.)