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Phytosociology of Pinus Qurecus Forest V
Phytosociology of Pinus Qurecus Forest V
Phytosociology of Pinus Qurecus Forest V
ABSTRACT: Thirteen plant communities were recognized from Nikyal valley on the basis of importance
value viz Myrsine -Pinus- Rhus community, Olea-Punica-Berberis community, Olea-Themeda-Pinus
community, Myrsine-Rhus-Pinus community, Quercus community, Rubus-Quercus-Oxalis community,
Olea-Themeda-Pinus community, Myrsine community, Qurecus-Oxalis- Justica community, Quercus-
Myrsine-Berberis community, Quercus-Rabdopsia- Themeda community, Pinus-Quercus- Indigofera
community, Pinus-Quercus-Oxalis community. The altitude of the area differs from 1535 – 1860m.
The soil texture differs from loam to clay loam type, with slight basic pH. Saturation varies from 42-57,
-1
E.C. from 0.52-0.72, organic matter from 0.57-1.40, Phosphorus from 2.80-6.60 mgkg , Potassium from
-1
60-160 mgkg .
Key word: Vegetation, Nikyal valley, Qurecus dilatata, Pinus roxburghaii community
INTRODUCTION
Nikyal Hills are situated in District Kotli, Azad Jammu and Kashmir at an altitude of 1500-1900m. They are
o
located 30 km away from Kotli towards North. The investigated area lies within longitude 74 04 to 10 East and
o
latitude 33 26 to 29 North. It is surrounded by Kotli on south, on western side by Tatapani, on Northern side by
Mender and on east by Pir-Panjal (Anonymous 2006).
The climate of Nikyal valley is of sub tropical humid type with average annual rainfall of 95.60 mm. The
maximum rainfall occurs during July amounting to 251.52 mm, while least rainfall occurs during November
amounting to 14.44 mm. The hottest months of the year are June and July, with mean daily maximum temperature
o o o o
of 37.69 C and 34.82 C respectively and minimum temperature of 23.61 C and 23.62 C respectively, while the
o o
coldest months of year were December and January mean maximum temperature of 19.99 C and18.09 C
o o
respectively and minimum temperature of 5.49 C and 4.41 C respectively. The average maximum and minimum
relative humidity received by the area is 79.64 and 30.82% respectively (Anonymous 2006).
The study was conducted during monsoon 2012. The area was divided in to thirteen stands on the basis of
altitude and physiognomic characters. Quadrat method was used for vegetation sampling. The size of quadrat was
2 2 2
10 × 10 m for trees, 5 × 5 m for shrubs, 1× 1 m for herbs. In each community five quadrats were laid for trees,
ten for shrubs, and fifteen for herbs. Total quadrats in all the communities were 390. Each species was recorded in
different strata due to its habitat. The herbage cover was determined by the Daubenmire’s cover scale
(Daubenmire, 1959). Trees with diameter at breast height (1.5m) were measured to obtain basal area (Hussain,
1969). Density, frequency and canopy cover thus measured following Curtis & McIntosh (1950) and Hussain
(1969). They were changed to the relative values and summed up to obtain the importance value Index (IVI) in a
particular stand (Curtis and Cottom 1959). Every species was ranked according to their importance values and the
species with the highest importance value in the stand was considered as the dominant species. (Khan et al 2010).
Soil was collected from each community up to the depth of 15cm and analyzed physically and chemically in the soil
and water testing laboratory Rawalpindi
Intl J Agri Crop Sci. Vol., 5 (24), 2952-2960, 2013
Phytosociology is the study of plant communities, their composition and development due to
interrelationship between the species (Allaby, 2004). Some phytosociological studies have been conducted in
some other parts of Azad Kashmir by Amjad et al. (2011), Malik and Malik (2004), Dastagir et al. (1999) , Malik
and Zandial (1996), and Malik et al., (2001; 2007), Malik & Hussain, (1987, 1988, 1990). According to their
observation, the subtropical Olea-Pinus and Olea-Acaia forests reflect highly deteriorated condition due to intensive
human pressure, deforestation and overgrazing. The aim of this study was to reports the existing vegetation
diversity of the area, which might be helpful for future workers involved in the development of this area.
RESULTS
Thirteen plant communities were established in the Nikyal valley during Monsoon 2012, which are as:
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This stand was present in loamy soil with slightly basic pH; Electrical conductivity 0.72 µs/cm; O.M very low;
-1 -1
saturation 56%; Phosphorus 4.90 mgkg and potassium 160 mgkg (Table 2).
The soil of this community was loamy with slightly basic pH; Electrical conductivity 0.52 µs/cm. Potassium was
-1 -1
60 mgkg with 33 % saturation; Organic matter was high and Phosphorus was 6.20 mgkg (Table 2).
The soil of the community was loamy having a pH of 7.60, Electrical conductivity was 0.62 µs/cm,
-1 -1
saturation was 50%; O.M was low; Phosphorus was 5.20 mgkg and potassium was 80 mgkg (Table 2).
Quercus community
This community was established in Tar-Kundi at an altitude of 1620m, located at 33N 26.235’ and 74E
08.613’ co‐ordinates (Table 3). Qurecus dlatata was dominant species with the importance value of 52.26.
Themeda anathera and Indigofera heterantha were the co-dominant species. Rubus fruticosus, Gaium elegan
and Elaeagnus parvifolia were associated species. 38 species were recorded as rare (Table 1).The dominance of
Quercus was due to low deforestation as it is a non palatable species.
In this community soil was loam with 7.60 pH; Electrical conductivity was 0.57µs/cm; Saturation was 32%;
-1 -1
O.M was low; Phosphorus was 2.80 mgkg and Potassium was 40 mgkg (Table 2).
In this community soil was loamy with pH 7.18; Electrical conductivity was 0.53 µs/cm; Saturation was 54%;
-1 -1
O.M was high; Phosphorus was 3.40 mgkg and Potassium was 80 mgkg (Table 2).
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Myrsine community
At 1670m Myrsine community was established, located at 33N 28.435’ and 74E 04.412’ co‐ordinates
(Table 3) having the importance value of 51.38. Rhus cotinus, Qurecus dlatata and Pinus roxburghii were the co
dominants. The associated components of this stand were Themeda anathera, Berberis lycium and Viola odorata.
34 species had low importance value (Table 1).
Soil of this community was of loam texture with slight basic pH. The Electrical conductivity was 0.71µs/cm;
-1 -1
Saturation was 55%; O.M was 0.65%; Phosphorus was 6.00 mgkg and Potassium was 120 mgkg (Table 2).
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DISCUSSIONS
Vegetation is a unit which possesses distinctive physiognomic structure which differentiates it from other
units (Hussain and Ilahi, 1991). It is dependent on different environmental as well as anthropogenic factors such as
soil, climate, deforestation and overgrazing (Gairola et al., 2008; Timilsina et al., 2007). Structure, composition,
diversity and distribution of vegetation also show variation in response to variables like productivity, Geographical
location, human forest interaction and evolutionary competition (Woodward, 1988; Eriksson, 1996; Criddle et al.,
2003).
Ahmed et al., (2006) characterized the subtropical-tempreate ecotone forest in the Borad leaved forest and
mixed pine forest and Temperate zone in to Temperate Borad leaved forest, temperate mixed forest, and
temperate coniferous forest. The study area has vegetation in fragments. Thirteen different plant communities
were recognized in the present study due to season and time of sampling which linked up to the particular
vegetational units. Local dominance of therophytes alters the shape of the vegetation which may reflect the
seasonal variations (Oosting, 1956).
Most of the shrubs, trees and perennial herbs are similar in all stands. Plant communities which are
recognized in present research work represent the different seral, edaphic remnants or, degraded stages of the
major vegetational units as recorded by Ahmed et al. (2006), Beg (1975) and Champion et al. (1965).
The climate of Nikyal valley is of mountane temperate type (Champion et al, 1965, Beg, 1975). However
due to the obvious differences in physiographic, edaphic, and local climatic conditions in different slopes at different
elevations, they support different plant communities (Ahmad, 1986). At the base of hill Pinus roxburghii, Olea
ferruginea and Punica granatum was dominant tree species. Ilyas et al. (2012) and Ahmed et al. (2009; 2006) also
reported these species as dominant at same altitude in Sawat and lower Dir. Myrsine africana, Rhus cotinus and
Berberis lycium were the dominant shrubby species and Themeda anathera was dominant grass. Ahmed et al.
(2006) reported such type of forest under lower pine forest of subtropical zone, however some previous worker
such as Champion et al. (1965) consider these forest as a Himalayan Pine forest. Hussain (1969) and Beg (1975)
consider these forests as dry temperate and Himalayan dry Chir-Pine respectively. Similarly Naqvi (1976) describe
these forests under mountain zone. Deforestation and the influence of the local occupant lead to cutting down of
other tree for their need viz fuel, fodder and construction purpose due to which the population of woody plants
decreases in the area.
With the increase in altitude Qurecus dlatata become the major component along with the Rubus
fruticosus, Myrsine africana, Oxalis corniculata, Justica paploroides while the Pinus roxburghii become the co-
dominant. With slight deviation the results are in line with Ilyas et al. (2012) and Khan et al., 2010. Qurecus dilatata
is non palatable species that is why it dominates. According to Champion et al., (1965) Quercus dilatata found in
higher and relatively narrow altitude (1828m 2286m ) on southern aspects and prefer colder conditions. Moreover
he reported it as one of the natural climax subtype of moist temperate forest that has almost been eliminated due to
anthropogenic activities. So our findings in this regard are also in line with them.
However at the top of hills Quercus- Rabdopsia- Themeda community (1760m), Pinus- Quercus –
Indigofera community (1820m) and Pinus- Quercus -Oxalis community (1870m) were recognized. The co
dominant components in these communities were Plectranthus rugosus, Fragaria nubicola and Galium elegan,
while the grasses such as Chrysopogon aucheri, Themeda anathera and Sorghum halepense were the associated
components. With slight deviation the presents results are in line with those of Hussain & Ilahi (1991), Beg (1975),
Ahmad (1986, 2006), Shaheen et al., (2011) and Ilyas et al. (2012). The area lies in moist temperate zone where
rain fall is high and temperature is low. So these communities might be the remnants of original temperate forest
(Malik 2005; Hussain et al., 1992; Hussain and Ilahi, 1991). Similar degraded vegetation is also reported by
Hussain et al. (1992) Haridasan et al. (1993) Sakya and Bania (1998). So our findings are also in line with them.
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The composition and distribution of plant communities in the investigated area appears to be controlled by
multifarious environmental factors including soil, climate, topography, and biotic pressure. These factors undergo
changes of different degrees due to interactions with each other and may result in micro-gradients (Hanson and
Churchill, 1965). On smaller scale Soil characteristics also influence plant distribution i.e. more local scale
(Bakkenes et al., 2002). Topography also play important role in structural characteristics of vegetation. It affects the
vegetation by altering other environmental factors. Different altitudes, aspects and slopes support different
communities in Nikyal hills. The co- dominant species in particular community is also prominent in other community
and much overlap seen in species composition in the investigated area. Ahmed et al. (2009) recorded the similar
findings while working on Olea ferruginea forests of lower Dir.
Table 1. Importance value of different plant species recorded from Kotli hills during Monsoon, 2010.
Species Importance value No of stands in
which species is
Maximum Minimum Average
present
Achillea millefolium L. 1 2.71 1.86 2
Achyranthes aspera Wall 2.7 4.32 3.51 2
Adiantum venustum D.Don 1.5 7.8 2.95 10
Agrostis canina auct 4.87 4.87 1
Ajuga bracteosa Wallich 0.57 6.12 2.16 7
Anaphalis margaritacea (L.) Bth. 0.59 1.09 0.84 2
Androsace rotundifolia Hardw 2.46 2.46 1
Andropogon gerardii Vitman 3.97 4.97 4.47 2
Arisaema jacquemontii Blume 3.09 11.15 5.66 4
Aristida adscensionis L. 0.55 5.73 2.37 7
Bergenia ligulata (Str) Hot 1.03 1.28 1.16 2
Berberis lycium Royle 2.42 31.32 13.90 13
Biden biterata (Lour.) Merr and Sherf 0.86 8.29 4.18 7
Brachiaria eruciformis (J.E Smith)
2.58 22.89 8.02 7
Griseb
Brachiaria repans
2.49 7.09 4.79 2
(L.) Gardner and Hubbard
Bryophyllum pinnatm Kurz 1.81 3.56 2.73 3
Bupleurum falacatum L. 1.2 7.42 3.27 5
Chrysopogon aucheri (Boiss) Stapf 5.91 22.68 11.32 4
Clematis grata Wallich 2.13 7.78 5.85 3
Colebrookia oppositifolia Sm. 1.54 2.32 4.91 4
Commelina benghaliensis
1.28 9.44 3.47 7
Linn.,sp. Pl.
Contoneaster acuminatus ILinley 0.84 6.69 3.96 5
Conyza ambigua Hook. Arn 2.99 2.99 1
Conyza canadensis L. 1.49 1.94 1.72 2
Cymbopogon jwarancusa
3.75 4.89 4.32 2
(Jones) Schult.
Cynodon dactylon (L.) Pers 10.43 12.77 11.60 2
Cynoglossum lanceolatum Forssk 2.7 8.86 4.76 8
Cyprus difformis L. 4.13 4.13 1
Cyperus niveus Retz 2.62 5.75 3.69 6
Cyperus rotundus L. 1.17 8.06 3.73 6
Cyperus globosus Forssk. 1.15 1.15 1
Debregeasia salicifolia (D.Don)
1.74 4.57 2.65 4
Rendle
Dichanthium annulatum (Forssk).Stapf 4.25 8.16 5.73 3
Dicliptera roxburghiana
1.01 11.63 7.21 9
Nees
Dodonaea viscosa L. Jack 4.78 4.78 1
Dryopteris stewartii Fress 1.33 2.23 1.89 3
Elaeagnus parvifolia Wall. ex Royle 1.54 11.18 6.27 4
Eragrostis japonica Thunb. Trin. 5.14 8 6.57 2
Eriophorum comosum Wall 2.17 8.15 3.91 4
Euphorbia helioscopia L. 1.95 4.92 3.48 4
Euphorbia prostrata Ait 1.06 2.95 2.51 2
Euphorbia wallichii HK.f 1.09 5.61 3.86 4
Ficus palmata Forrsk 0.65 7.59 2.20 11
Fragaria nubicola Landle ex Lacaita 1.72 19.47 15.98 4
Galium elegan Wall. 3.24 21.02 11.41 13
Gerbera gossypina Royle 1.61 11.8 6.18 8
Geranium rotundifolium L. 2.15 6.45 4.72 9
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Table 2. Soil analysis result from different communities of Kotli Hills during Monsoon 2012.
S.No Name of Height E.C Saturation O.M P K PH Texture
community (m) µs/cm % % mgkg-1 mgkg-1
KEY
MPR = Myrsine -Pinus- Rhus community M = Myrsine community
0PB = Olea-Punica- Berberis community QOJ = Qurecus-Oxalis- Justica community
OTP = Olea - Themeda -Pinus community QMB = Quercus- Myrsine- Berberis community
MRP = Myrsine - Rhus-Pinus community QRT = Quercus- Rabdopsia- Themeda community
Q = Quercus community PQI = Pinus- Quercus – Indigofera community
RQO = Rubus- Quercus-Oxalis community PQO= Pinus- Quercus -Oxalis community
OTP = Olea –Themeda-Pinus community
CONCLUSION
Nikyal valley is under severe deforestation pressure for fuel and timber value. Alternative sources could be
provided by the government and valley should be protected for 30 years to promote natural vegetation (especially
trees and shrubs cover). There is an urgent need to promote the ethics among the people that improvement and
conservation of natural resources are critical for land and soil management
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