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Fig. 128.—Sclerotinia ciborioides: a sclerotium with three apothecia slightly
magnified; b ascus with eight ascospores; c germinating ascospore.
Fig. 129.—Dasyscypha willkommii: a
portion of bark of Larix decidua with sessile,
cup-shaped apothecia (nat. size); b two
paraphyses on either side of an ascus with
eight ascospores.
Order 2. Mollisiaceæ. Mollisia cinerea, principally on decaying wood.
Family 6. Helvellales.
These Fungi have the appearance of clubs, bells, or mushrooms,
consisting of an upright stalk bearing a large and fleshy head, on the
exterior surface of which the hymenium is spread. The ascocarps
are probably gymnocarpic from the beginning, and on this account
these plants are placed in a separate family. The development of the
ascocarps is unknown. The Morchella (Morell) grows on the ground;
some species are edible. 1 order.
Order. Helvellaceæ. Spathulea is yellow and club-shaped, and forms “fairy
rings” in woods.—Geoglossum (Earth-tongue) projects above the ground as a
black tongue, or as a club-shaped body. Several species are found in meadows
and on heaths.—Helvella has a stalk, bearing an irregularly folded head, on the
external surface of which is the hypothecium.—Morchella (Morell, Fig. 130), the
stalk bears on its summit the conical or spherical head, the external surface of
which is reticulate and bears the asci.—Mitrula. Verpa.
Fig. 130.—Morchella esculenta: a an entire
specimen, about one half natural size; b
longitudinal section through the head.
Fig. 141.—A A portion of the thallus of Parmelia parietina with ascocarps (a)
and pycnidia (b). B A portion of the thallus of Cetraria islandica with pycnidia at the
end of small lobes. C A lobe with pycnidia and ejected microconidia. (Magnified).
Geographical Distribution. The Lichens are the most hardy
plants, and are the first to appear on hitherto bare rocks which they
gradually disintegrate, and hence prepare the way for the growth of
other plants. They are to be found from the Polar regions to the
Equator; from the highest snow-free mountain-peaks down to the
level of the sea; on the stems of trees; on rocks, soil, some even on
inundated places; on stones in woodland streams, and on beaches;
but they are never found upon rotten organic remains. Some grow
gregariously in enormous masses, and form wide-stretching carpets,
e.g. Reindeer Moss (Cladonia rangiferina), species of Cetraria and
other fruticose Lichens.
Uses. On account of the cell-wall being composed of
Lichenstarch (Lichenin), the Iceland-Lichen and Manna-Lichen
(Lecanora esculenta) are used as food; the latter grows on stones, in
the deserts of Asia and North Africa, and is often torn loose in large
masses and carried away by the wind. The Reindeer-Lichen is not
only the principal food of the reindeer, but it is also used in the
manufacture of Danish brandy. Cetraria islandica (Lichen islandicus)
is officinal. Colouring materials (lacmus, orseille, persio) are made
from several species, especially from Roccella tinctoria (from the
rocky coasts of the Mediterranean). Parmelia saxatilis and
particularly Lecanora tartarea are used for colouring purposes in the
Northern countries.
About 2,000 species of Lichens have been described. If we
disregard the Basidiolichenes, which will be considered on page 176,
the remaining Lichens (Ascolichenes) may be divided into the two
following orders according to the structure of the fruit-bodies:—
Order 1. Pyrenolichenes. The ascocarps (apothecia) are
spherical or flask-shaped, as in the Pyrenomycetes, more rarely
linear (Graphis).
According to the nature of the thallus, these Lichens may be divided into:—
a. Thallus homoiomerous, but not gelatinous, branching according to the mode
of growth of the Algæ: Ephebe (Fig. 133), with Algæ of the genus Stigonema.
b. Thallus homoiomerous, gelatinous: Lichina.
c. Thallus heteromerous, crustaceous: Verrucaria, Pyrenula; Graphis (Fig. 139),
which may be considered as Hysteriaceæ with gonidia; several species of Graphis
are common on bark.
d. Thallus heteromerous, foliaceous: Endocarpon.
e. Thallus heteromerous, fruticose: Sphærophorus.
Order 2. Discolichenes. These, as in the Discomycetes, have
open apothecia, which, as a rule, are cupular, more rarely
hemispherical (Cladonia).
According to the nature of the thallus, these Lichens may be divided into:—
a. Thallus homoiomerous, but not gelatinous, branching according to the mode
of growth of the Algæ: Cœnogonium.
b. Thallus homoiomerous, gelatinous: Collema (Fig. 132), with Algæ of the
genus Nostoc; Leptogium.
c. Thallus heteromerous, crustaceous: Pertusaria (Fig. 140), Lecidea, with
apothecia open from the beginning; Lecanora, with apothecia, which in the
beginning are closed, later on open, but with a rim formed by the thallus (Fig. 138);
Bæomyces, whose apothecia are borne on a stem formed by the thallus.
d. Thallus heteromerous, foliaceous: Parmelia (P. saxatilis; P. parietina, Wall-
Lichen, Fig. 141, is yellow, very frequent on tree-stems, stone-walls, tiles); Physcia
(P. ciliaris, frequent on tree-stems); Sticta (S. pulmonacea, Lung-Lichen, on tree-
stems); Peltigera, especially on the Moss among trees; Umbilicaria, on rocks.
e. Thallus heteromerous, fruticose: Cetraria (C. islandica), “Iceland Moss,” with
an olive-brown, flat, furrowed, fringed thallus, on heaths; C. nivalis, white, in the
Polar regions; Evernia, Ramalina, Usnea (U. barbata, Beard-Lichen, Fig. 143);
Roccella, Stereocaulon, Cladonia, of which the genus C. rangiferina, Reindeer-
Moss (Fig. 142) is important; Cladonia has two kinds of thallus, one scaly and leaf-
like, the other erect, which bears the apothecia and may be fruticose (Fig. 142), or
cupular (Fig. 144); they grow in soil in forests and on heaths.
Fig. 142.—Cladonia rangiferina: s ascocarp.