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The fruit is a schizocarp, dividing into two mericarps; the plane in
which these separate coincides with that of the union of the carpels,
and the two nut-like mericarps are in most genera kept together for
awhile at the top of a thin, bifid, or undivided stalk (carpophore)
which is in direct continuation with the flower-stalk (Fig. 537). Each
mericarp has most frequently 5 more or less strongly projecting
ridges, the primary ridges (Figs. 530, 532, 534, 535, etc.), of which 3
lie on the back of the mericarp, the dorsal ridges, and 2 on its edge
near the plane of division, the marginal ridges; five of these (10
ridges in all in the entire fruit) are placed opposite the calyx-teeth
and the others between them. In some genera there are in addition 4
secondary ridges to each mericarp between the primary ones (Fig.
528 E: the secondary ridges bear the long bristles). Inside these
secondary ridges, or inside the grooves between the primary ridges,
when the secondary ridges are absent, oil ducts (vittæ,
schizogenous ducts) are found in the pericarp, most frequently one
in each groove; two are also often found on the ventral side of each
mericarp (Figs. 528 E, 530 ol, etc.). The seed is most frequently
united with the pericarp. The embryo is small and lies high up in the
large, most frequently horny endosperm (Fig. 528 D).—The
endosperm does not contain starch, but oil, and presents three
different forms, of important systematic value: (a) those which are
quite flat on the ventral side (i.e. the side turned towards the plane of
splitting) (Figs. 528 E, 530, 531, 534, etc.): the majority of the
genera, Orthospermeæ (e.g. Carum, Pastinaca); (b) those in which
the endosperm on the ventral side is provided with a longitudinal
groove, often deep: Campylospermeæ (e.g. Anthriscus); the
transverse section is nearly a crescent (Fig. 532); (c) those in which
the endosperm is concave on the ventral side (hollow in both
longitudinal and transverse sections): Cœlospermeæ (e.g.
Coriandrum) (Fig. 538).
The genera are distinguished first of all by the endosperm and forms of fruit, the
ridges and oil-ducts; then by the form of the umbel, the calyx and corolla, by the
absence or presence of an involucre, etc.
Fig. 529.—
Hydrocotyle
vulgaris. Transverse
section of fruit.
1. Hydrocotyleæ, Penny-wort Group. Capitula or simple
umbels (all the other groups have compound umbels). No oil-ducts.
Orthospermous.—Hydrocotyle (Penny-wort). The fruit is
considerably compressed laterally (Fig. 529). The calyx-teeth are
small. The leaves are peltate.—Didiscus.—Sanicula (Sannicle). The
umbels are small, capitate, generally collected in a raceme; calyx-
teeth distinct. ♂ -and ♀ -flowers in the same umbel. The fruits are
round, studded with hooked bristles. No carpophore.—Astrantia has
an umbel surrounded by a large, often coloured involucre, with this
exception it is the same as the preceding, but the fruit is slightly
compressed, with 5 equal ridges. Hacquetia (Dondia).—Eryngium
(Sea Holly): leaves often thorny. The flowers are all sessile, the
inflorescence is thus a capitulum; each flower is often subtended by
a bract, which is thorny like the involucre, resembling the burrs of the
Teasel. The sepals are large.—Lagœcia: one of the loculi of the ovary is
suppressed.
Fig. 530.—Fruit of Carum petroselinum: fr endosperm; ol oil-ducts.
Sub-Class 2. Sympetalæ.
The characters which separate this from the first Sub-class, the
Choripetalæ, have been described on page 336. They consist in the
following: the flower is always verticillate, generally with 5 sepals, 5
petals, 5 stamens, and 2 carpels (in the median plane), the calyx is
generally persistent and gamosepalous, the corolla is gamopetalous
and united to the stamens, which are therefore adnate to it, the
ovules have only one thick integument and a small nucellus. (The
exceptions are noted later.)
This Sub-class is no doubt more recent than the Choripetalæ; it is also peculiar
in including fewer trees and shrubby forms than the latter.
The Sympetalæ may be separated into 2 sections:—
A. Pentacyclicæ (five-whorled). The flowers in this section
have 5 whorls equal in number, namely, 2 staminal whorls in addition
to the calyx, corolla, and carpels; in some instances, one of the
staminal whorls is rudimentary or entirely suppressed, but in this
case it is frequently the sepal-stamens which are suppressed, and
the whorl which is present stands opposite the petals. The flowers
are regular. The number of carpels equals that of the sepals, but in