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 Matonineae 355–363; Hymenophyllaceae 363–365; Cyatheaceae 365–
375; Polypodiaceae 375–380; Dipteridinae 380–394.

CHAPTER XXII
MARATTIALES (FOSSIL). Pp. 395–411.

Ptychocarpus 397; Danaeites 398; Parapecopteris 398; Asterotheca 398–400;

Hawlea 400; Scolecopteris 401, 402; Discopteris 402–404; Dactylotheca
404–406; Renaultia 406; Zeilleria 407; Urnatopteris 407; Marattiopsis 407–
409; Danaeopsis 409; Nathorstia 410, 411.

CHAPTER XXIII
PSARONIEAE. Pp. 412–426.

CHAPTER XXIV
OPHIOGLOSSALES (FOSSIL). Pp. 427–431.

CHAPTER XXV
COENOPTERIDEAE. Pp. 432–472.

I. Botryoptereae 434–443
II. Zygoptereae 443–470

CHAPTER XXVI
HYDROPTERIDEAE AND SAGENOPTERIS. Pp. 473–483.

Marsiliaceae 473–475; Salviniaceae 475–477; Sagenopteris 477–483.

CHAPTER XXVII
GENERA OF PTERIDOSPERMS, FERNS, AND PLANTAE INCERTAE SEDIS.
Pp. 484–580.

Taeniopteris 485–494; Weichselia 494–496; Glossopteris 496–512;

Gangamopteris 512–517; Lesleya 517–519; Neuropteridium 519–523;
 Cardiopteris 523–525; Aphlebia 525–529; Sphenopteris 529–532;
Mariopteris, Diplotmema, Palmatopteris 532–537; Cephalotheca 537;
Thinnfeldia 537–544; Lomatopteris 544–546; Cycadopteris 546; Ptilozamites
546–548; Ctenopteris 548–550; Dichopteris 550–552; Odontopteris 552–556;
Callipteris 557–559; Callipteridium 560; Archaeopteris 552–565; Neuropteris
565–571; Cyclopteris 571, 572; Linopteris 572, 573; Alethopteris 573–576;
Lonchopteris 576; Pecopteris 576–580.
Pp. 609–
INDEX 624
 LIST OF ILLUSTRATIONS
Several of the illustrations are printed from blocks for which I am indebted to
learned societies or to individuals. The sources from which clichés were
obtained are mentioned within square brackets.
Frontispiece. Thamnopteris Schlechtendalii (Eich.). From a photograph given to
me by Dr Kidston and Mr Gwynne-Vaughan. (page 329.)
Fig. Page
112. Sphenophyllostachys 2
113. Sphenophyllostachys Römeri 3
Sphenophyllum trichomatosum
S. majus
114, 115. Sphenophyllostachys fertilis 4, 5
[Council of the Royal Society of London.]
116. Sphenophyllostachys Dawsoni 6
[Mr A. G. Tansley, Editor of the New Phytologist.]
117. Cheirostrobus pettycurensis 8
Pseudobornia ursina
118. Psilotum triquetrum 18
119. Psilotum triquetrum (anatomy) 20
120. Tmesipteris tannensis 22
121. Lycopodium (seven species) 35
122. Lycopodium squarrosum 36
123. Lycopodium cernuum 37
124. Lycopodium obscurum 38
125. Lycopodium (anatomy of stem) 41
126. Lycopodium (anatomy of cones) 45
127–129. Lycopodium cernuum (cone) 47–49
[Council of the Royal Society of Edinburgh.]
130. Selaginella grandis 50
131. Selaginella (anatomy) 52
132. Isoetes echinospora 59
I. lacustris
 Fig. Page
133. Isoetes lacustris (anatomy) 62
134. Pleuromeia Sternbergi 70
135. Selaginellites and Lycopodites 80
136. Lycopodites lanceolatus 81
[Council of the Geological Society of London.]
137. Lycopodites falcatus 83
138. Selaginellites primaevus 86
139. Lycostrobus Scotti 89
140. Picea excelsa 94
141. Lepidodendron Sternbergii 97
142. Sigillaria (leaves) 98
143. Lepidodendron (leaves) 99
144. Lepidodendron Veltheimianum 101
145. Lepidodendron leaf-cushion 102
146. Lepidodendron and Lepidophloios leaf-cushions 104
147. Lepidophloios leaf-cushion 108
148–155. Lepidodendron vasculare 112–122
156. Knorria mirabilis 125
157. Lepidodendron Veltheimianum (Ulodendron) 129
158. Diagrammatic section illustrating the branch-theory 132
of the Ulodendroid scar
[Council of the Manchester Literary and
Philosophical Society.]
159. Pinus clausa 134
160. Lepidophloios scoticus 135
161. Halonia tortuosa 136
162–167. Lepidodendron fuliginosum 143–147
[Council of the Cambridge Philosophical Society.]
168. Lepidodendron vasculare and L. fuliginosum 148
169. Lepidodendron fuliginosum 149
170–172. L. fuliginosum 150–152
173. Lepidodendron obovatum 154
174–176. Lepidodendron aculeatum 155, 156
[Oxford University Press: Annals of Botany.]
177. Stigmaria radiculosa 157
 Fig. Page
178. Stigmarian rootlet 158
179. Lepidodendron Harcourtii and L. fuliginosum 162
180. Lepidodendron Wünschianum 163
181, 182. L. Wünschianum 165, 166
183, 184. L. Wünschianum 168, 169
[Editor of the New Phytologist.]
185. Lepidodendron Veltheimianum 173
186. L. Veltheimianum and L. macrophyllum 176
187. Lepidodendron australe 179
[Dr H. Woodward, Editor of the Geological
Magazine.]
188, 189. Lepidostrobus 183, 184
190. Lepidodendron and Lepidostrobi 186
191. Lepidostrobus 188
192. Spencerites insignis 193
[Oxford University Press: Annals of Botany.]
193. Sigillaria elegans, S. rugosa, S. tessellata, 197
Omphalophloios anglicus
194. Sigillaria McMurtriei 199
195. Sigillaria mammillaris 199
196. Sigillaria Brardi, S. laevigata, and Lepidodendron 200
Wortheni
197. Carica sp. 202
198, 199. Sigillaria 205, 206
200. Sigillaria Brardi 212
201. Sigillariostrobus 216
202. Sigillaria elegans and S. elongata 220
203. Sigillaria Brardi 225
204, 205. Stigmaria ficoides 227, 228
206. Cyperus papyrus 230
207, 208. Stages in the development of Sigillaria 236
209. Stigmariopsis 237
210. Stigmaria 241
211. Bothrodendron punctatum 250
212. Bothrodendron minutifolium, B. punctatum, B. 252
kiltorkense and Lepidostrobus Olryi
 Fig. Page
213. Bothrodendron minutifolium 254
214. Bothrodendron Leslei 258
[Trustees of the British Museum.]
215. Bothrodendron mundum 259
216. Bothrostrobus 263
[Council of the Manchester Literary and
Philosophical Society.]
217. Omphalophloios 265
218. Lepidocarpon Lomaxi 273
219. Miadesmia and Bothrodendron 276
220. Angiopteris evecta and Cycas revoluta 283
221. Osmunda cinnamomea, O. regalis, and Todea 286
barbara
222. Schizaea elegans 287
223. Aneimia rotundifolia 288
224. Aneimia flexuosa, A. phyllitidis, Hymenophyllum, 289
Matonia pectinata, Thyrsopteris elegans,
Gleichenia
225. Gleichenia dicarpa 290
226. Gleichenites Rostafinskii, Gleichenia dicarpa, G. 290
dichotoma
227. Matonia pectinata 292
[Council of the Royal Society.]
228. Matonia pectinata 293
229. Thyrsopteris elegans, Cyathea spinulosa, Dicksonia 294
coniifolia, D. culcita, Davallia concinna, Alsophila
excelsa
230. Dicksonia Bertervana 295
[Trustees of the British Museum.]
231. Dipteris quinquefurcata, D. conjugata, D. Wallichii, 297
and Polypodium quercifolium
232. Davallia aculeata 299
233. Polypodium Billardieri 302
234. Polypodium quercifolium 303
235. Hemitelia capensis 304
236 a, 236 b. Pteris aquilina 305, 306
[Council of the Linnean Society of London.]
 Fig. Page
237. Matonia pectinata, Matonidium, Gleichenia dicarpa, 310
and
Trichomanes reniforme (anatomy)
238. Trichomanes scandens 311
[Editor of the New Phytologist.]
239. Platyzoma microphylla 312
[Editor of the New Phytologist.]
240. Cyathea Imrayana 313
[Editor of the New Phytologist.].
241. Angiopteris evecta and Marattia fraxinea 317
242. Angiopteris evecta and Danaea 318
243, 244. Angiopteris evecta 319
245. Marattia fraxinea, M. Kaulfussii, Kaulfussia, and 320
Marattiopsis Münsteri
246. Ophioglossum vulgatum 322
247. Botrychium virginianum 322
248. Zalesskya gracilis 327
249. Zalesskya diploxylon 328
250. Thamnopteris Schlechtendalii 330
251. Lonchopteris virginiensis 331
252. Osmundites Dunlopi 333
253, 254. Osmundites Kolbei 334, 335
[Editor of the Geological Magazine.]
255. O. Kolbei 336
256. Cladophlebis denticulata, Todites Williamsoni, 340
Discopteris Rallii, Kidstonia heracleensis, and
Todeopsis primaeva
257, 258. Cladophlebis denticulata 342, 345
259. Klukia exilis 348
[Council of the Cambridge Philosophical Society.]
260. Ruffordia Goepperti 349
261. Chrysodium lanzaeanum, Lygodium Kaulfussi, 350
Marattia Hookeri
262. Gleichenites longipennis, G. delicatula, G. 354
Nordenskioldi and G. Zippei
263. Gleichenites hantonensis 356
[Council of the Palaeontographical Society.]
Fig. Page
264. Laccopteris elegans 357
[Council of the Royal Society.]
265. Matonidium Wiesneri, Marattiopsis marantacea, 358
Gleichenites gracilis, Laccopteris Goepperti, and
L. Muensteri
266. Laccopteris polypodioides 359
[Trustees of the British Museum.]
267. Laccopteris 359
[Trustees of the British Museum.]
268. ? Laccopteris polypodioides 360
[Trustees of the British Museum.]
269. Matonidium Goepperti 362
[Editor of the Encyclopaedia Britannica.]
270. Senftenbergia elegans, Oligocarpia Brongniartii, 364
Trichomanes sp., Hymenophyllum tunbridgense,
Sphenopteris (Hymenophyllites) quadridactylites
271. Coniopteris hymenophylloides 368
[Council of the Manchester Literary and
Philosophical Society.]
272. C. hymenophylloides 369
273. Coniopteris quinqueloba 370
274. Coniopteris arguta 371
275. Coniopteris arguta and C. hymenophylloides 372
276. Oncopteris Nettvalli 373
277. Protopteris punctata 373
278. Laccopteris polypodioides, L. Muensteri, Dicksonia, 374
Onychiopsis Mantelli, Hausmannia Sewardi, H.
Kohlmanni, and Protopteris Witteana
279. Adiantides antiquus and A. lindsayoides 376
280. Onychiopsis Mantelli 379
281. Dictyophyllum exile 381
282. Dictyophyllum Nilssoni, Rhizomopteris Schenki, 382
Camptopteris spiralis, and D. exile
283. Dictyophyllum rugosum 384
[Trustees of the British Museum.]
284. Thaumatopteris Münsteri 386
285. Clathropteris meniscoides 387
Fig. Page
286. Clathropteris egyptiaca 388
[Editor of the Geological Magazine.]
287. Camptopteris spiralis 389
288. Hausmannia dichotoma 391
289. Hausmannia sp. 393
290. Alethopteris lonchitica, Lonchopteris rugosa, 399
Sphenopteris Hoeninghausi, Parapecopteris
neuropteroides,and Pecopteris (Dactylotheca)
plumosa
291. Ptychocarpus unita, Asterotheca Sternbergii, 400
Danaeites sarepontanus, Hawlea Miltoni, H.
pulcherrima, Scolecopteris elegans
292. Dactylotheca plumosa 405
293. D. plumosa 406
294. Nathorstia angustifolia and N. latifolia 410
295. Psaronius 414
296. Psaronius infarctus, P. coalescens, P. musaeformis, 416
and P. asterolithus
297. Pecopteris Sterzeli 419
298. Caulopteris peltigera and Megaphyton insigne 421
299. Ptychopteris 423
300. Dicksonia antarctica 424
301. Rhacopteris sp. 427
302. Noeggerathia foliosa 429
303. Chiropteris Zeilleri 430
[Annals of the South African Museum.]
304. Tubicaulis solenites 435
[Editor of the New Phytologist.]
305. Botryopteris cylindrica 439
306. Botryopteris ramosa 441
307. Botryopteris antiqua 442
308. Clepsydropsis antiqua, Etapteris Scotti, Diplolabis 444
forensis, Zygopteris primaria, Stauropteris
oldhamia
309. Diplolabis forensis, Botryopteris forensis, 445
Corynepteris coralloides, Schizopteris pinnata
 Fig. Page
310. Metaclepsydropsis duplex, Stauropteris oldhamia, 450
Ankyropteris scandens
311. Ankyropteris Grayi 451
312. Thamnopteris Schlechtendalii, Ankyropteris 453
corrugata, A. bibractensis
313. Ankyropteris bibractensis 454
314. Ankyropteris corrugata 457
315. Ankyropteris corrugata 458
[Editor of the New Phytologist.]
316, 317. Ankyropteris corrugata 459, 460
318. Etapteris Scotti 462
[Editor of the New Phytologist.]
319. Etapteris, Botryopteris forensis 463
320. Stauropteris oldhamia 464
[Editor of the New Phytologist.]
321. Stauropteris oldhamia 467
322. Stauropteris oldhamia 468
[Editor of the New Phytologist.]
323. Stauropteris 469
[Editor of the New Phytologist.]
324. Asterochlaena laxa 472
[Editor of the New Phytologist.]
325. Sporocarp-like bodies (? Sagenopteris) 478
326. Regnellidium diphyllum, Sagenopteris rhoifolia 479
327. Sagenopteris Phillipsi 480
[Trustees of the British Museum.]
328. Sagenopteris Phillipsi 481
[Council of the Manchester Literary and
Philosophical Society.]
329. Taeniopteris multinervis, Lesleya Delafondi 487
330. Taeniopteris Carnoti, T. spatulata, T. coriacea 490
331. Taeniopteris Carruthersi 491
[Annals of the South African Museum.]
332. Taeniopteris vittata 493
333. Weichselia Mantelli, W. erratica 495
334. Glossopteris Browniana 499
[Council of the Geological Society of London.]
 Fig. Page
335, 336. Glossopteris Browniana 500, 501
[Trustees of the British Museum.]
337. Vertebraria indica 502
338. Vertebraria indica, Onoclea struthiopteris 503
339. Glossopteris fronds attached to rhizome 504
340, 341. Glossopteris indica, G. angustifolia 506, 507
[Trustees of the British Museum.]
342. Glossopteris angustifolia var. taeniopteroides 508
[Council of the Geological Society.]
343. Blechnoxylon talbragarense 509
344. Glossopteris retifera 511
[Trustees of the British Museum.]
345. Gangamopteris cyclopteroides 515
[Trustees of the British Museum.]
346. Arberia sp. 517
347. Lesleya simplicinervis 518
348. Neuropteridium validum 520
[Trustees of the British Museum.]
349. Neuropteridium intermedium 522
350. Cardiopteris frondosa 524
351. Gunnera manicata 527
352. Sphenopteris obtusiloba, Pecopteris arborescens, 529
Sphenopteris furcata
353. Sphenopteris affinis 531
354. Palmatopteris, Mariopteris, Diplotmema Zeilleri, 535
Neuropteris macrophylla, N. heterophylla, N.
Scheuchzeri, Alloiopteris Essinghii
355. Cephalotheca mirabilis 536
356. Thinnfeldia odontopteroides, Ptilozamites 539
[Council of the Geological Society.]
357. Thinnfeldia odontopteroides 540
[Council of the Geological Society.]
358. Thinnfeldia odontopteroides 541
[Annals of the South African Museum.]
359. Thinnfeldia rhomboidalis 542
360. Lomatopteris jurensis, L. Schimperi, Thinnfeldia 544
rhomboidalis
Fig. Page
361. Ptilozamites Heeri 547
362. Ctenopteris cycadea 549
363. Dichopteris visianica 551
364. Alethopteris lonchitica, Mariopteris muricata, 553
Odontopteris cf. alpina
365. Odontopteris minor 554
366. Odontopteris genuina, Callipteridium gigas, 557
Callipteris Pellati, C. lyratifolia
367. Callipteris conferta 559
368. Archaeopteris hibernica 561
369. Archaeopteris hibernica, A. archetypus, A.fissilis, A. 564
fimbriata
370. Neuropteris with Cyclopteris leaflets 566
[From a block received from Mr Carruthers.]
371. Neuropteris heterophylla 568
372. Neuropteris macrophylla 569
373. Neuropteris Scheuchzeri 570
374. Linopteris neuropteroides 573
375. Alethopteris Serlii 575
376. Pecopteris arborescens 578
 ERRATA IN VOL. I
Page 16, line 4. For “The North American Tulip tree” read The
Tulip tree of North America and China.
„ 66, line 2 from the bottom. For “Browera” read Berowra.
„ 127, line 3 and 4 from bottom. For Achyla and
Palaeachyla read Achlya and Palaeachlya.
„ 145, lines 4 and 5. For “Upper Greensand” read Lower
Eocene.
„ 162, line 3 from bottom. For “Corallina barbata” read
Cymopolia barbata.
„ 170, line 20. For “sporangiaphore” read sporangiophore.
„ 185, line 2. The genera Udotea and Halimeda, members
of the Siphoneae, are incorrectly included under the
Corallinaceae.
„ 191, line 11 from bottom. Omit Chondrus crispus, which is
one of the Florideae and not a Brown Alga.
„ 202, line 13. For “Halmeda” read Halimeda.
„ 250, line 11. For “three” read the.
„ 381, line 10. For “Calamopytus” read Calamopitys.
 CHAPTER XII[1].
SPHENOPHYLLALES (concluded).

Sphenophyllum.
The account of the Sphenophyllales given in the first volume[2] of
this work must be extended and somewhat modified in the light of
recent work on the fertile shoots of Sphenophyllum.
Sphenophyllostachys Dawsoni (Will.) was described as consisting
of an axis bearing superposed whorls of bracts connate at the base
in the form of a shallow funnel-shaped collar giving off from the
upper surface and close to the axis of the cone two concentric series
of sporangiophores. Occasionally there are three series, as
represented in fig. 112. In another type of strobilus,
Sphenophyllostachys Römeri[3] each sporangiophore terminates in
two pendulous sporangia (fig. 113, A; see also fig. 107, C, vol. i.). It
has already been pointed out that the common occurrence of
detached strobili necessitates their description under distinct specific
names; it is only by a rare accident that we can assign fossil cones
to their vegetative shoots. There are, however, reasons for believing
that Sphenophyllostachys Dawsoni is the strobilus of the plant
originally described by Sternberg[4] from impressions of foliage-
shoots as Rotularia cuneifolia. Another difficulty presented by
petrified material is that of determining, with certainty, whether two
imperfect specimens, differing from one another in features which do
not appear to be of sufficient importance to warrant specific
separation, are forms of one species or portions of specifically
distinct cones. It has been pointed out by Scott[5] that the strobilus
known as Sphenophyllostachys Dawsoni probably includes two
distinct species, one being the cone of Sphenophyllum cuneifolium
Sternb., and the other the cone of S. myriophyllum Crép[6]. The stem
of S. myriophyllum agrees anatomically with the type known as
Sphenophyllum plurifoliatum Will. and Scott[7].

Fig. 112. Sketch of a radial longitudinal section of Sphenophyllostachys.

There are usually two concentric series of sporangia on the
sporophylls, not three as shown in the figure. The upper figure
(after Zeiller) shows the linear bracts in surface-view.
In addition to the two types of cone already mentioned,
Sphenophyllostachys Dawsoni and S. Römeri, others have been
described by Kidston from carbonised impressions. One of these is
the fertile branch of Sphenophyllum majus[8]. The basal portions of
the bracts of each whorl form a narrow collar round the axis of the
cone; the free portion of each bract consists of a lamina divided into
two equal bifid lobes bearing on its upper surface one group, or
possibly two groups, of four sessile sporangia between the narrow
coherent bases of the laminae and the sinus between the terminal
lobes (fig. 113, C). Another characteristic feature is the greater
length of the internodes; this renders the cone less compact and less
sharply differentiated from the vegetative shoots than those of other
species. A specimen in Dr Kidston’s collection illustrates the peculiar
character of the fertile portion of this species; it consists of an axis
bearing a succession of lax sporophylls succeeded above and below
by whorls of sterile leaves. In this species, therefore, we cannot
speak of a compact strobilus at the end of a shoot of limited growth,
but of axes in which sterile and fertile leaves are borne alternately[9],
a condition recalling the alternation of foliage leaves and sporophylls
in Tmesipteris and in Lycopodium Selago.

Fig. 113.
A. Sphenophyllostachys Römeri. (Solms-Laubach.)
B. Sphenophyllum trichomatosum Stur.
C. Sphenophyllum majus. Bronn. (A–C. After Kidston.)

Another form of cone, also from the Middle Coal Measures, is

referred by Kidston to Sphenophyllum trichomatosum Stur[10] (fig.
113, B): this is characterised by the more horizontal position of the
bracts, which “do not appear to be so much or so suddenly bent
upwards in their distal portion as in some other species of
Sphenophyllum,” and by sessile sporangia borne singly on the upper
face of each bract.
 Fig. 114. Sphenophyllostachys fertilis (Scott). (After Scott.) Diagram of a
node in longitudinal section, showing one sporophyll and the
base of the opposite one. v.l. ventral lobe of sporophyll; v.s. one
of the segments into which it divides; v.s′. stump of another
segment; d.l. dorsal lobe; d.s., d.s′. segments of dorsal lobe.
A more recent addition to our knowledge of the fertile shoots of
Sphenophyllum is due to Scott who has described a new type of
cone under the name Sphenophyllum fertile[11]. The petrified
specimen on which the species was founded was discovered by Mr
James Lomax in the Lower Coal Measures of Lancashire; it
represents a portion of a cone 6 cm. long and approximately 12 mm.
broad. The axis contains a single vascular cylinder agreeing in
essentials with the type of stem structure known as Sphenophyllum
plurifoliatum. The nodal regions, which exhibit the slight swelling
characteristic of the genus, bear several (probably twelve)
appendages connate at the base and forming a narrow flange
encircling the axis. Each bract, the base of which forms part of the
narrow collar surrounding the axis, consists of two lobes, ventral and
dorsal, divided palmately into several (sometimes four) segments or
sporangiophores (fig. 115). Each sporangiophore terminates distally
in an oblong or oval lamina bearing two sporangia on its adaxial face
(fig. 114). The space between the axis and the periphery of the cone
is thus occupied by crowded peltate laminae, each with its pair of
sporangia. A single vascular bundle supplies each sporangiophore
and bifurcates in the distal lamina into two branches which extend to
the bases of the sporangia. The sporangia agree in structure with
those of other species of Sphenophyllum: the spores are of one size
and elliptical, characterised by the presence of several sharp ridges