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Sonoran Desert

Phenology Calendar

Photo credits: Steve Amesbury.


Phenology: an introduction
What is phenology?
What causes phenology?
Phenology is the study of the timing of cyclic or seasonal plant and animal life
The three key environmental drivers of phenology are
cycle events that are influenced by environmental conditions, including
photoperiod (length of daylight hours), temperature, and the
variations in weather and climate. Derived from the Greek words phaino,
availability of moisture. The combination and interaction of
which means “to appear”, and logos, which means “to study”, the observation
these three drivers impose complex and varying controls on
of phenological events (termed “phenophases”) in nature has been going on
phenology events in different parts of the globe.
for millennia: the earliest humans observed and planned their lives around
· Photoperiod provides plant and animal species with a
plants' growing seasons and the presence of animals for hunting. Written
consistent annual climatic signal. Photoperiod can trigger
records of plant and animal phenology are available from hundreds of years
the growth and death of leaves and animals’ biological
back, including cherry blossom timing in Japan and grape harvesting in
clocks, e.g., reproduction, migration, and molting of feathers
Europe.
or fur.
· Temperature can regulate growth and development
Why is phenology important?
patterns of plant and animal species. Many plants require a
Phenology is nature’s “biological clock”, an integrated measurement of the
chilling period for their seeds to grow; other plants and
interactions between the atmosphere and the Earth. Phenology is valuable in
animals go dormant in either very hot or very cold
many ways:
seasons. In temperate areas plant phenology is often
· Phenology relates to human issues like agricultural production, health (e.g.,
driven by the combination of photoperiod and temperature.
allergies, timing of vector-borne diseases) and recreation and tourism
· Moisture availability (e.g., rain or snow) is important for
· Phenological patterns relate to environmental change due to altered climate
plant and animal species because it affects the ability and
and ecosystems; thus phenology can be utilized in natural resource
rate of cell division, for example in leaves and in eggs of
conservation and management
amphibians. Water limitation is especially important in the
· Phenology can be assessed and monitored at different scales that relate to
deserts of the world, and the combination of precipitation
individual plants or animals, populations, communities and/or ecosystems,
and photoperiod affects phenology in the tropical areas of
offering a potent look into those systems’ health and functioning
the world.

Measurement of phenology
Every living thing experiences and responds to the changing of seasons. Thus, we can observe phenology in individual plants or
animals, populations of the same or different species, and entire communities, as each member responds to changes in light, moisture,
temperature, and even the phenophases of other organisms. Observation of phenological events includes noting, for example, when
individual flowers appear on a rose bush, when different rose bushes flower, when bees begin to show up to pollinate the roses, and what
other plants the bees also pollinate at that time. Tools for observing phenology include the human eye of course, as well as cameras,
satellites, and tools to measure the amount of pollen in the air or bacteria in the soil. Phenology can be measured across minutes, hours,
days, and months, and changes in the timing of phenological events can be measured across seasons or among years to millennia.

Photo credits: National Park Service.


Phenology: examples
Plant Phenology
The most readily visible example of phenology is in the leafing, flowering, and fruiting of
plants. The phenology of trees can be especially interesting to study because specific
individuals can often be monitored for long periods of time. This helps to reduce the
Phenology and humans
effects of individual variation in the timing of phenological events, making trees
Economics. Human activities are in large part affected by the
potentially a more sensitive tool for identifying changes in phenological timing caused by
environmental context within which particular communities live,
global climate change.
and economic activities are not an exception. Phenological
Trees show a variety of phenophases in the development of leaves, flowers, fruits,
events (especially those driven by temperature and
and seeds. Each of these features functions in different ways in the lives of trees --
precipitation) are routinely taken into account when engaging in
leaves are needed for photosynthesis and growth, flowers are the organs of
agriculture, cattle ranching, wildlife management, tourism,
reproduction, and fruits and seeds are involved in plant dispersal – and each responds
regional fruit harvesting, and every other activity that involves a
to different environmental factors. The cycle of leaf development from bud to deciduous
live component and therefore life cycles. Examples wherein
leaf fall is related to rainfall, day length, and temperature. Flower development is often
phenology constitutes an important tool for economic activities
timed to correspond with the availability of pollinators. Fruit and seed production may
include controlling crop plague, determining suitable crops for a
be influenced by the presence of seed dispersal agents like wind and fruit-eating
specific season, determining the best time for the harvesting of
animals. Thus it is possible that the seasonal patterns of each of these phenophases is
game species or regional fruits, etc.
linked to different environmental cues and may be affected in different ways by
Disease. For thousands of years it has been observed that
environmental change.
many human diseases are more likely to occur during specific
times of the year, such as influenza and the common cold.
Animal Phenology
Unlike most phenological phenomena, human diseases with a
Birds, fish, and mammals such as deer and bobcat are all attuned to the seasons of the
phenological signature are typically most active in the winter. In
Earth. Some of the most studied phenological events are the mass migrations of
addition to the common phenological diseases that are
animals between regions, continents and even globally (e.g., whale and sea bird
experienced across the globe, inhabitants of the Sonoran
migrations across entire oceans). Animals such as snowshoe hares, elk and deer
Desert are vulnerable to unique diseases that are only
change the way they look, becoming a different color or growing antlers based on
experienced by a few regions in the world, such as
seasonal changes, and bears and amphibians will hibernate during inhospitable
coccidioidomycosis or "valley fever".
seasons.
Invasive species. Exotic invasive plants often experience
Insect populations are also attuned to the rhythms of climate. Population dynamics
different phenological responses than native plants to annual
are often controlled by climate variables like temperature, which restricts survival and
and seasonal changes in temperature and precipitation,
precipitation, which restricts reproduction for species whose young rely on standing
differentiating them functionally from natives. Frequently
water. Insects are also important members of a complex ecology where species
invasive species are found to respond to precipitation faster
interact on multiple levels. Plants often provide food and, in return, many insects help
than natives and set flower sooner, drawing soil moisture down
plants pollinate. Disruption of the cycle of one species might therefore have significant
and extracting nutrients otherwise available to natives.
consequences on another species. Insects also act as pests by destroying crops or
Understanding the phenology of invasive species is critical to
seriously damaging other vegetation. They can also act as disease vectors for
management and control efforts; chemical and manual
pathogens such as West Nile Virus and Dengue Fever. Thus the phenology of insects is
treatments depend on the status of the soil and the phenology
highly tied to the phenologies of other animals, plants, and human systems.
of the plant.

Photo credits: Amanda Barnard, National Park Service.


Phenology in the Sonoran Desert
Unique climate, unique phenology
The Sonoran Desert, spanning parts of southwestern USA and northwestern Mexico (shown in the bottom right, as defined
by the World Wildlife Fund), has two rainy seasons but is overall hot and dry. These climatic features result in unique
phenological traits of the species living in this biome. Light is rarely limiting and temperatures rarely stay below freezing;
overall there is high sensitivity of both plants and animals to the availability of water and the avoidance of extreme
heat. Many plants can respond to specific rainfall events, flowering numerous times in one growing season; both plants and
animals can become dormant in times of drought. These characteristics provide unique opportunities and challenges in
observing and measuring phenology in the Sonoran Desert.

The above graph summarizes the climate of the Sonoran Desert using
temperature and precipitation data from the Arizona Meteorological
Network (http://ag.arizona.edu/AZMET/). Three points in the Sonoran
Desert of Arizona (Tucson, Yuma and Paloma) for the period 1992-2007
were used to generate daily averages for high (red line) and low (blue line)
temperatures, and the probability of precipitation (green line). In addition,
the daily high (red dot) and low (blue dot) temperatures experienced over
the entire period are plotted to give readers the sense of the range of
temperatures that may be experienced during specific times of the year.
Similar climate figures were made for each month.

Photo credits: Andy Fisher, National Park Service.


Related links: About this Calendar:
There are countless sources of information about
This calendar was produced as a final project for the class “The Geography
phenology on the internet. Below are a few links that
of Phenology” (GEOG696C) in the Department of Geography and Regional
provide overviews of phenology and its use in monitoring
Development at the University of Arizona. Contributors to the development
the state of the planet; each contains many more links of
of the calendar include Dr. Willem J.D. van Leeuwen (instructor), Steven
interest.
Stewart Amesbury, Armando Barreto Muñoz, Jennifer Davison, Jahan
- Project Budburst Kariyeva, Cory W. Morin, Aaryn Dyami Olsson, Jose Raul Romo Leon and
http://www.windows.ucar.edu/citizen_science/budburst/ James Derek Tamerius.
- Monarch Watch
http://monarchwatch.org Sources:
- USA National Phenology Network: http://www.usanpn.org
-http://www.saguaro-juniper.com/i_and_i/cacti/saguaros/harvest/saguaro_fruit.html
- University of California Santa Barbara Phenology
Stewardship Program: -http://edc.usgs.gov/guides/avhrr.html
http://www.ucsbphenology.christophercosner.com -http://en.wikipedia.org/wiki/SPOT
- Arizona-Sonora Desert Museum Migratory Pollinators -http://geo.arc.nasa.gov/sge/landsat/lpchron.html
Program: http://www.desertmuseum.org/pollination -http://www.itc.nl/research/products/sensordb/Launch_Schedule.aspx
- Intergovernmental Panel on Climate Change: -http://www.sonoraturismo.gob.mx/caceria-en-sonora.htm
http://www.ipcc.ch
-http://www.whatbird.com/
Special Thanks: -http://www.wrcc.dri.edu/
We would like to acknowledge the following - Epple, Anne Orth. 1995. A field guide to the plants of Arizona. Falcon,
parties for the information and pictures that Guilford, CN.
made this project possible: - Kearney, Thomas H., and Robert H. Peebles. 1951. Arizona flora.
T. Beth Kinsey and The Firefly Forest University of California Press, Berkeley.
http://fireflyforest.net/firefly/ - McGinnies, William G. n.d. Flowering period for common desert plants,
- Veronica Cornonado Peraza southwest Arizona. Office of Arid Lands Studies, College of Agriculture,
- The Arizona-Sonora Desert Museum University of Arizona.
http://www.desertmuseum.org/
- Phillips, Steven J., and Patricia Wentworth Comus, eds. 2000. A natural
- The National Parks Service
history of the Sonoran Desert. Arizona-Sonora Desert Museum Press,
http://www.nps.gov/
Tucson.
- USGS Arizona Water Science Center
http://az.water.usgs.gov/ - Turner, Raymond M., Janice E. Bowers, and Tony L. Burgess. 1995.
- Rogelio Lizarraga Sonoran Desert plants, an ecological atlas. University of Arizona Press,
www.bamury.com Tucson.
- Don Knight, Stephen Minter, and Mark - USDA. 2008. Plants database. Natural Resources Conservation Service.
Dimmitt, ASDM Sonoran Desert Digital Library. U.S. Department of Agriculture. http://plants.usda.gov/index.html
January
The coolest period of the year typically begins in
the last week of December and continues through
the first week of January. Temperatures typically
are near 20˚C during the day and often
reach freezing temperatures at night. Winter
storms bring precipitation in the form of rain for
most of the low elevations and snow at higher
elevations. Most vegetation is dormant during this
month, but some desert fruits ripen and trees
begin to sprout new leaves.

Although unusual, occasionally the desert


valleys receive a dusting of snow.

Species such as Freemont cottonwood (Populus


fremontii) may begin to blossom and the fruit of
Christmas cactus (Schlumbergera bridgesii) will
ripen.

Photo credits: (Top right, bottom right, and background) T. Beth Kinsey, the Firefly Forest; (Bottom left) T.R. Van Devender.
Desert mistletoe
(Phoradendron californicum)
cover many trees during this
period. They are a great food
source for birds that are active
such as the curve-billed
thrasher (Toxostoma
curvirostre, right) and the
cactus wren (Campylorhynchus
brunneicapillus, below).

January
Sun Mon Tue Wed Thu Fri Sat
1 2 3

4 5 6 7 8 9 10

11 12 13 14 15 16 17

January is breeding season for the mule 18 19 20 21 22 23 24

deer (Odocoileus hemionus) . Males put


much of their energy into competing for
25 26 27 28 29 30 31
mates, often making them more
susceptible to attack by cougars (Puma
concolor), who are also very active Record Low Record High
Max Monthly Precipitation (1993): 5.58 inches
during this month. (1913): 6F (1971): 90F

Photo credits: (Top right) Rhonda Spencer, Arizona-Sonora Desert Museum; (Top middle) Warren Lynn, Arizona-Sonora
Desert Museum; (Top left) Jeanne Broom, Arizona-Sonora Desert Museum; (Bottom left) Paul Berquist, Arizona-Sonora
Desert Museum; (Background) T. Beth Kinsey, the Firefly Forest.
February
On the cusp of springtime in Arizona, February can
provide balmy days and snowy storms with seemingly
equal probability. Shifts in bird phenology increase in
the Sonoran Desert in February. Cinnamon teal (Anas
cyanoptera), Western grebes (Aechmophorus
occidentalis) and other early north-bound migrants
begin to arrive. Wintering ducks show brighter
plumage, preparing for breeding season. Wintering
raptors such as ferruginous hawks (Buteo regalis) and
red-tailed hawks (Buteo jamaicensis, right) remain
abundant.

Photo credits: (Top right, bottom left) Beth Kinsey, the Firefly Forest; (Background) Jennifer Davison.
February
Sun Mon Tue Wed Thu Fri Sat
Nesting is underway for Costa’s hummingbirds 1 2 3 4 5 6 7

(Calypte costae, male shown above) in


February. Some individuals are year-round
residents, while others migrate to the Sonoran 8 9 10 11 12 13 14
Desert in late winter or early spring, continuing
on to the highlands of Arizona and California
when temperatures get too hot. With nesting 15 16 17 18 19 20 21
sites in desert scrub (right), chaparral and
tropical deciduous forest, Costa's
hummingbirds rely not only on the flowers 22 23 24 25 26 27 28
becoming abundant at this time of year but the * 1986,
SPOT-1
insects that frequent those flowers.
Record Low Max Monthly Precipitation (1905): 4.15 inches Record High
(1899): 17F (1957): 93F

Photo credits: (Left) T. Beth Kinsey, the Firefly Forest; (Right)


(Right) Willem J. D. van Leeuwen;
Leeuwen; (Background) Jennifer Davison.
* Satellite launch dates.
March

A great diversity of plant and animal species live in


streamside riparian habitats in the Sonoran Desert. Two
tree species dominate these habitats, the Freemont
cottonwood (Populus fremontii; above left) and the
Goodding willow (Salix gooddingii; above right). Flowering
in both these species generally begins in early spring.

Photo credits: Steve Amesbury.


Spring wildflowers begin to bloom in February
and peak in March following wet winters in the
Sonoran Desert. The Mexican gold poppy
(Escholtzia mexicana, right) may bloom in such
profusion that entire hillsides turn orange for
several weeks. Many shrubs and trees flower
at this time, too, including the invader, salt
cedar (Tamarix ramossissima, below). Salt-
cedar grows in rpiarian areas, where it
competes with native riparin trees. Although it
is not native to this region, some native
species of birds have adopted it for nesting.

March
Sun Mon Tue Wed Thu Fri Sat
1 2 3 4 5 6 7

8 9 10 11 12 13 14

15 16 17 18 19 20 21
Vernal
Equinox

22 23 24 25 26 27 28

29 30 31

Record Low Record High


Max Monthly Precipitation (1905): 3.88 inches
(1897): 22F (1988): 98F

Photo credits: (Bottom left and background) Steve Amesbury; (Top right) Beth Kinsey, the Firefly Forest
April
The trees of the Sonoran Desert flower later than the
wildflowers, including the blue palo verde (Parkinsonia florida;
below left) and the foothills palo verde (P. microphylla; below
right). Both bloom during April, but the flowers of the blue
palo verde appear one to two weeks before those of the
foothills palo verde. The staggered blooming times help to
prevent hybridization between the two species, both of which
are pollinated by native bees and the introduced honeybee.
Other trees that flower at this time include the honey
mesquite (Prosopis glandulosa). The mesquite’s leguminous
pods will be an important food source for much of the desert’s
fauna in the coming months. Meanwhile, most desert cacti
are putting on flower buds and may be already blooming. The
prickly pear (Opuntia spp.) (center) provides pollen for bees
and migrating birds.

Photo credits: (bottom left and right) Steve Amesbury; (bottom center) Aaryn Olsson.
The orchid-like flowers of the desert willow (Chilopsis linearis; right)
are a beautiful pink or purple color. They first begin to appear in April
and last until August. Desert willows are most often found in arroyos
or near streams. During the autumn and winter, they lose their
leaves.
The velvet mesquite (Prosopis velutina) is the common
species of mesquite in the Sonoran Desert. It
generally begins to bloom in April. The flowers are
small and form an elongated cluster called a catkin.
Groves of mesquite, known as bosques (below), often
line arroyos.

April
Sun Mon Tue Wed Thu Fri Sat
1 2 3 4

5 6 7 8 9 10 11

12 13 14 15 16 17 18

19 20 21 22 23 24 25

26 27 28 29 30

Record Low Max Monthly Precipitation (1905): 3.53 inches Record High
(1901): 28F (1965): 104F

Photo credits: (Top right, bottom left, and background) Steve Amesbury; (Top middle, Top left) Mark Dimmitt, Arizona-
Sonora Desert Museum
May
Saguaros bloom in April and May, providing a
valuable nectar source for the endangered Lesser
Long-nosed bat (Leptonycteris curasoae). This bat
undertakes long migratory flights, relying on the
nectar from columnar cacti such as the saguaro and
organ pipe cactus (Stenecereus thurberi). Their
dependence on nectar necessitates a tight coupling
of their migration with the timing of flowering of the
saguaro and other cacti. Therefore, the paths these
bats take are often called “nectar corridors.”
The white-winged dove (Zenadia asiatica) is another
migrant that utilizes the saguaro.

Photo credits: (Top left) Merlin D. Tuttle, Bat Conservation International; (Background).
May
Sun Mon Tue Wed Thu Fri Sat
1 2

3 4 5 6 7 8 9

Ironwoods (Olneya tesota) break buds in


10 11 12 13 14 15 16
late April and early May, flowering for 10-
18 days before shedding their flowers,
leaving a pink blanket on the desert floor.
17 18 19 20 21 22 23
Flower production is variable, with large
production occurring every 2-5 years.
Insect pollinators, including many species
of bees, visit the ironwood flowers, whose 24 25 26 27 28 29 30

flowers persist beyond the flowering


phase of palo verdes (Cercidium spp.).
The anatomy of the pea-shaped flower 31

(upper right) requires the pollinator to bury


itself between the petals, ensuring it picks
up and deposits pollen during its visit. Record Low Max Monthly Precipitation (1931): 1.34 inches Record High
(1899): 32F (1910): 111F
Photo credits: (Top left and background) Mark Dimmit, Arizona-
Arizona-Sonora Desert Museum.
Museum.
June
One of the main economic activities in the Sonoran
Desert region is cattle ranching, these activities are
strongly related to phenological responses from
the desert to climatic pulses. Rain will affect the
timing and quantity of plant production, directly
impacting cattle ranching.
Other important activities are tourism and hunting.
Hunting is regulated by specific cyclic events
intrinsic to all species such as mating or birth of
wildlife. Tourism is conducted at different times of
the year in the presence of events such
as flowering, fruit harvesting and wildlife spotting.

Photo credits: (Top right) Jacinto Mayboca Esparza ; (Bottom left) Jose Raul Romo Leon; (Background) Don Knight.
Some of the traditional activities
related to harvest the native flora
happen during June:
The traditional saguaro
(Carnegiea gigantea) fruit harvest
of the Tohono O’odham (Desert
People) Nation takes place
primarily during this month.

June
Sun Mon Tue Wed Thu Fri Sat
1 2 3 4 5 6

7 8 9 10 11 12 13

14 15 16 17 18 19 20
Summer
Solstice

21 22 23 24 25 26 27

28 29 30

Record Low Record High


Max Monthly Precipitation (1938): 2.07 inches
(1908): 43F (1960): 115F

Photo credits: T. Beth Kinsey , the Firefly Forest.


July Average high temperature: 105
Average low temperature: 80
Prior to 2008 the Arizona Monsoon was Warmest ever: 121
defined as beginning after the third Coldest ever: 61
consecutive day of dew points above 55oF. Average precipitation: .8
On average this concurred around July 7
with the monsoon continuing for the next two
months. Starting in 2008 the National
Weather Service has defined the Arizona
monsoon as beginning on June 15 and
ending on September 30

Photo credits: USGS, Arizona Water Science Center.


Vegetation
greenness is With the arrive of the monsoon the Sonoran
measured here desert landscape changes dramatically from
by the MODIS arid and dry scenery to a green humid-warm
sensor, on the and vivid ecosystem.
TERRA satellite. From a phenological point of view, many
Satellites are processes occur during this month in an
routinely used to accelerated way, influenced by the excessive
measure sun hours (photoperiod), precipitation, and
landscape-scale higher temperatures. The result is an increase
phenology. in vegetation greenness, meaning more food
for animals and humans.

July
Sun Mon Tue Wed Thu Fri Sat
1 2 3 4

5 6 7 8 9 10 11

12 13 14 15 16 17 18

19 20 21 22 23 24 25

* 1972,
Landsat 1

26 27 28 29 30 31
Drought Levels Based on
Monthly Streamflow discharge,
July 2008 Record Low Record High
Max Monthly Precipitation (1984): 7.56 inches
(1911): 49F (1995): 115F

Photo credits: USGS, Arizona Water Science Center. * Satellite launch dates.
August
August is the peak of the monsoon season and
month with the greatest rainfall (~3 in) in the
Sonoran Desert, which only averages 3-15 inches
of rain per year. Monsoon is defined as a seasonal
change in the wind direction with strong and brief
thunderstorms that bring heavy rain. The summer
monsoon in the Sonoran Desert brings moisture
from the Gulfs of Mexico and California. The winter
and summer rain seasons and mild winters of the
Sonoran Desert enable its unique biodiversity.
Populat household plants such as Caesalpinia
(bottom left) and Tecoma stans bloom throughout
the monsoon season.

Photo credits: (Top) T. Beth Kinsey, the Firefly Forest; (Bottom) Willem J. D. van Leeuwen; (Background) Jahan Kariyeva.
Sonoran desert toads (Bufo alvarius) emerge following the
rains to feed and breed in large temporary rain pools. During
the rest of the year this species hibernates underground.
Sonoran desert toads occur in a number of habitats ranging
from creosote bush desert scrub and grasslands to oak-pine
woodlands and tropical deciduous forests. These toads
consume a large variety of insects throughout their lives.

August
Sun Mon Tue Wed Thu Fri Sat
1

August is the month of the distinctive


buzzing sound in the melodies of the
Sonoran Desert. These signature sounds 2 3 4 5 6 7 8

are the mating tunes of the male cicada.


Sonoran Desert or Apache cicadas 9 10 11 12 13 14 15

(Diceroprocta apache) are the only


insects known to be able to cool
themselves by sweating during summer 16 17 18 19 20 21 22

heat. Adult cicadas feed on the watery


liquids inside trees and
23 24 25 26 27 28 29
nymphs feed from tree
roots underground
during their few years 30 31

of development.

Record Low Max Monthly Precipitation (1935): 5.61 inches Record High
(1917): 55F (1944): 112F

Photo credits: (Top and bottom) T. Beth Kinsey, the Firefly Forest; (Background) Jahan Kariyeva.
September
September is a transition
month climatologically and
biologically. The monsoon
rains recede rapidly and
drier conditions increase.
Temperatures are still hot
but may cool towards the
end of the month as
summer ends and fall
begins.

Plants and flowers like the turpentine bush


(Ericameria laricifolia) are still blooming from the
monsoon rains. The prickly pear's fruit (Opuntia
spp.) become fully ripe and ready to eat.

September is the breeding season for big horn


sheep (Ovis canadensis).

Photo credits: (Top and Background) T. Beth Kinsey, the Firefly Forest; (Bottom Left) Paul Berquist, Tucson Sonora Desert
Museum.
September is a time of insect
activity due to standing water
from the monsoon rains. The
risk of West Nile Virus
transmission from mosquitoes
is high. Brilliant butterflies like
the gray hairstreak (Strymon
melinus) will decorate and
pollinate the landscape.

September is the peak time of bird


migration. A variety of birds can be seen September
passing through or settling in for the Sun Mon Tue Wed Thu Fri Sat
winter. Rufous hummingbirds 1 2 3 4 5

(Selasphorus rufus) arrive for the winter


while Western Kingbirds (Tyrannus
6 7 8 9 10 11 12
verticalis) gather to migrate south.

13 14 15 16 17 18 19

20 21 22 23 24 25 26
Autumnal
Equinox

27 28 29 30

Record Low Record High


Max Monthly Precipitation (1996): 4.41 inches
(1965): 42F (1950): 112F

Photo credits: (Top left) Stephen Minter, Tucson Sonora Desert Museum; (Top right) T. Beth Kinsey, the Firefly Forest; (Bottom
left) Paul and Joyce Berquist, Tucson Sonora Desert Museum; (Background) T. Beth Kinsey, the Firefly Forest.
October
October brings a noticeable change in temperatures and
precipitation, as the days shorten and the monsoon season
officially ends. Precipitation decreases markedly and
temperatures begin to vary more between day and night, both
climatic shifts providing signals to plants and animals to begin
their next phenophase. Many plants start to turn color in
October, due to an interruption in chlorophyll production and
subsequent increase in the reds and yellows reflected by other
chemicals, e.g., anthocyanins. A leafless coralbean (Erythrina
flabelliformis) at right and a summer annual (below) show that
autumn colors can be quite dramatic in the Sonoran Desert.

Photo credits: (Bottom) T. Beth Kinsey, the Firefly Forest; (Top) Willem J. D. van Leeuwen; (Background) Jahan Kariyeva.
As the autumn progresses resident birds briefly sing again,
and migrating birds make a south-bound pass through the
Sonoran Desert. Wintering hawks and hummingbirds arrive,
and summering shore birds and water fowl migrate south.
Anna’s hummingbirds (Calypte anna; male shown at left) are
back in the Sonoran Desert to breed, starting in late
September. Both male and female Anna's Hummingbirds
have dark bills, green backs, grayish-green undersides, and
lack of prominent eye stripes.

Elf owls (Micrathene whitneyi; below), October


who have spent the summer in the
Sun Mon Tue Wed Thu Fri Sat
Sonoran Desert eating bugs, prepare 1 2 3
to migrate south before their prey is
killed off by colder temperatures.
4 5 6 7 8 9 10

11 12 13 14 15 16 17
* 1978,
AVHRR

18 19 20 21 22 23 24

25 26 27 28 29 30 31

Record Low Record High


Max Monthly Precipitation (1983): 5.78 inches
(1897): 29F (1950): 104F

Photo credits: (Top) Will Elder, National Park Service; (Bottom) T. Beth Kinsey, the Firefly Forest; (Background) Jahan Kariyeva.
* Satellite launch dates.
November
Although November is the beginning of the winter precipitation season, the
precipitation that typically occurs is inadequate to satiate the parched
landscape. Temperatures continue to decline and the Sonoran Desert
generally experiences its first freezing temperatures of the season by the last
week of the month. The dry conditions combined with winds generated by
storms moving across the Great Basin to the north can result in dust storms.
Besides particles of soil, the wind can also disperse microscopic biota across
the desert, which is an important part of desert ecology. In the Sonoran
Desert, the soil dwelling fungi Coccidioides immitis and Coccidioides
posadasii are suspected of releasing their spores during this period. If these
spores are inhaled by humans they can cause a
usually mild respiratory disease called “Valley Fever”. Deciduous trees that inhabit the canyons of the
Sonoran Desert lose their leaves after a brief
flash of color.

In the foothills surrounding the Sonoran Desert


grasses that prospered during the monsoonal rains
turn a golden brown.
Photo credits: (Top) Ted Myers, Arizona-Sonora Desert Museum; (Bottom) James Tamerius.
Young male mule
deer (Odocoileus
hemionus) are active,
rubbing the velvet
from their antlers in
preparation for
sparring in the winter.

Desert-mistletoe (Phoradendron spp.) -- a November


hemiparasite -- begins to grow on
Ironwood, Mesquite, Palo Verde and
Acacia trees during November.

The phainopepla (Phainopepla nitens) is a


wintering bird in the Sonoran Desert.
Mistletoe berries are their primary food Record Low Record High
Max Monthly Precipitation (1905): 4.61 inches
source, although they eat insects as well. (1919): 19F (1924): 94F

Photo credits: (Top left ) Mark A. Dimmit, Arizona-Sonora Desert Museum; (Top right) Paul Berquist, Arizona-Sonora Desert
Museum; (Bottom left) T. Beth Kinsey, Firefly Forest
December
December is the month when the hunting activity
begins for species like mule deer, white tailed deer
(Odocoileus virginianus) and the desert big horn
sheep (Ovis canadensis), in many areas of the
Sonoran Desert.

Photo credits: (Top right and background) Rogelio Lizarraga; (Top left) Stephen Minter.
Winter crops abound in the Sonoran Desert, one of Agricultural produce unique to this region is sent
the few places within the U.S. where you can grow to the whole country where the weather
crops the whole year due to its mild temperatures conditions are not as favorable as in the Sonoran
and abundant sunshine. Desert in December.

Pistachio Lettuce Chili pepper Date palm

December
Sun Mon Tue Wed Thu Fri Sat
1 2 3 4 5

6 7 8 9 10 11 12

13 14 15 16 17 18 19

20 21 22 23 24 * 1999, 25 26
Winter TERRA
Solstice

27 28 29 30 31

Drought Levels Based on


Monthly Streamflow
discharge, December Record Low Record High
Max Monthly Precipitation (1914): 5.85 inches
(1901): 10F (1970): 88F
2007
Photo credits: USGS, Arizona Water Science Center. * Satellite launch dates.
Photo credits: Steve Amesbury.

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