C.

VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

New species and new records of Calliostomatidae

(Gastropoda: Trochoidea)
from western Pacific (II): Solomon Islands, Papua New Guinea,
Indonesia, Philippines and Taiwan
Claude VILVENS
Research Associate
Institut de Systématique, Evolution, Biodiversité (ISYEB),
Muséum national d'Histoire naturelle, CNRS, SU, EPHE, UA
CP 51, 57 rue Cuvier, 75005 Paris, France
vilvens.claude@skynet.be
urn:lsid:zoobank.org:pub:CF6D3C33-9149-4BF4-A1B0-C57FE33F904F

KEYWORDS. Gastropoda, Calliostomatidae, Solomon Islands, Papua New Guinea, Indonesia,

Philippines, Taiwan, Calliostoma, Tristichotrochus, Fautor, Akoya, Bathyfautor, Venustatrochus,
Selastele, Phenacomargarites, Thysanodonta, new species.

ABSTRACT. New records of 23 known Calliostomatidae species from an area stretching from
Solomon Islands to Taiwan are listed, extending the distribution area of some of them. The depth
ranges of these species have been completely recalculated based on all the data available for over
30 years.
14 new species are described and compared with similar species: Calliostoma pheidolon n. sp.,
C. newguineense n. sp., C. thaumaston n. sp., C. achantodes n. sp., C. zhongshaense n. sp., C.
statheron n. sp., Fautor cosmopolites n. sp., F. panteles n. sp., Akoya zhui n. sp., Selastele
solomonense n. sp., Phenacomargarites tetratropeis n. sp., Thysanodonta brucemarshalli n. sp.,
T. eumetabolos n. sp. and T. aoristos n. sp.

RESUME. De nouveaux relevés de 23 espèces connues de Calliostomatidae provenant d'une aire

allant des Iles Salomon à Taïwan sont listés, étendant ainsi l'aire de distribution d'un certain
nombre d'entre elles. Les zones de profondeurs de ces espèces ont été entièrement recalculées sur
base de l'ensemble des données disponibles pour elles depuis pus de 30 ans.
14 nouvelles espèces sont décrites et comparées avec des espèces similaires : Calliostoma
pheidolon n. sp., C. newguineense n. sp., C. thaumaston n. sp., C. achantodes n. sp.,
C. zhongshaense n. sp., C. statheron n. sp., Fautor cosmopolites n. sp., F. panteles n. sp., Akoya
zhui n. sp., Selastele solomonense n. sp., Phenacomargarites tetratropeis n. sp., Thysanodonta
brucemarshalli n. sp., T. eumetabolos n. sp. et T. aoristos n. sp..

INTRODUCTION Poppe, 2004; Poppe et al., 2006, Poppe et al., 2014;

This work is the next part of a new study of
Calliostomatidae MNHN material collected in the
western Pacific area by the French IRD-MNHN
expeditions during these last 40 years, the first part of
this study about New Caledonia area having been
published recently (Vilvens, 2023). The area
considered here start from Solomon Islands and goes
towards the West through Papua New Guinea, eastern
Indonesia, South China Sea, Philippines and as far as
Taiwan. One can wonder about the large size of this
studied area, but this can be justified by the fact that it
has appeared that some species have indeed a wide
distribution, for example from Solomon Islands to
Indonesia, or from Papua New Guinea to the
Philippines and even Taiwan.
While Calliostomatidae of some areas in the scope of
this work has been studied rather recently (Solomon
Islands: Vilvens, 2009; Philippines: Vilvens, 2000a, b;
Taiwan and Chinese seas: S.-I Huang & I.-F. Fu,
2015, 2019, 2022; Poppe & Tagaro, 2020; Thach,
2023), only poor or often very old data are available
for other areas (Papua New Guinea : Hinton, 1978
without any Calliostoma species; Indonesia :
Schepman, 1908; Knudsen, 1970; Dharma, 1988
without any Calliostoma species; Dharma, 2005 with
mostly Australian Calliostoma species; Arafura Sea,
between northern Australia and southern Indonesia :
Willan, 2002).
Although the available samples for this study are
not very numerous regarding some campaigns and
some species, this study has revealed, apart from the
extension of distribution of known species, that
numerous examined specimens belong to new species
that are described here.

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C. VILVENS New species and new records of Calliostomatidae. Part II

Material and methods
The material studied in this paper was collected by
French IRD-MNHN expeditions started up with the
goal to study the biodiversity of invertebrates,
especially molluscs, in a wide range of habitats of the
western Pacific (from Taiwan to Tonga Islands), here
especially from Solomon Islands to Taiwan through
Papua New Guinea, western Indonesia and
Philippines. More than 200 samples batches were
studied. The most numerous samples were collected
during these last 20 years, but additional samples
collected during older campaigns were also found and
therefore considered in the present work.
The following table lists the expeditions that have
collected the studied samples (data from the reference
Web site of the MNHN campaigns). The last column
is the digital object identifier (DOI) link (when
available) to websites with information about each
expedition.

Table 1. List of the MNHN expeditions in the western Pacific where samples were collected (the abbreviations
of the campaigns are used on the following map (Fig. 1)).

Campaign Sampling area Date DOI

BIOPAPUA (BP) Papua New Guinea, Bismarck Sea 8-10/2010 10.17600/10100040
CORINDON 2 (C2) Indonesia, Makassar Strait 10-11/1980 10.17600/80005712
KARUBAR (KB) Indonesia, Kai and Tanimbar Islands 10-11/1991 –
KAVALAN 2018 (KV) North-eastern Taiwan 6-7/2018 –
KAVIENG 2014 (KG) Papua New Guinea: New Ireland 6-9/2014 10.17600/14004400
MADEEP (MA) Papua New Guinea 4-5/2014 10.17600/14004000

MUSORSTOM 2 (M2) Philippines 11-12/1980 10.17600/80010211

12/2013–
NanHai 2014 (NH) South China Sea –
1/2014
PANGLAO 2004 (P4) Central Philippines 2004 –
PANGLAO 2005 (P5) Bohol and Sulu seas, Philippines 2005 –
PAPUA NIUGINI (PN) North-eastern Papua New Guinea 10-12/2012 10.17600/18000841
SALOMON 1 (S1) Solomon Islands 9/2001 10.17600/1100090
SALOMON 2 (S2) Solomon Islands 10-11/2004 10.17600/4100090
Solomon Islands: off Guadalcanal and 9-10/2007
SALOMONBOA 3 (SB3) 10.17600/7100070
Malaita
TAIWAN 2000 (T00) South-western and eastern Taiwan 7-8/2000 –
5,7–8, –
TAIWAN 2001 (T01) Southern and north-eastern Taiwan
11/2001
TAIWAN 2002 (T02) Southern and north-eastern Taiwan 5,8,9/2002 –

TAIWAN 2004 (T04) North east of Taiwan 7,8,9/2004 –

TAIWAN 2013 (T13) Southern Taiwan 5/2013 –

ZHONGSHA 2015 (ZA) South China Sea, Macclesfield Bank 7-8/2015 –

Regarding the distribution of the new species and the
extension of the distribution of known species, the
range is taken from the internal intervals of the two
extremes values: the bathymetric ranges given are the
inner values of the recorded depths, that means the
deepest minimum and the shallowest maximum of
each recorded depth range (Bouchet et al., 2008).
These ranges were computed for all the available
known samples, that means published in this work and
in the work of B.A. Marshall, 1995; Vilvens, 2005,
2009, 2012, 2014a; Vilvens & Maestrati, 2006.
Moreover, each range is provided not only for all the
available specimens but also for the only living
specimens only if applicable; when these ranges are
the same, the common range is cited once with the
"(lv)" annotation; if all the specimens are dead
collected, the range is cited with the "(dd)" annotation.

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C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

Regarding the description methodology, the main
conchological features used are (Figs 2 & 3):
♦ general shape of the shell (depressed, high spired
– conical, cyrtoconoidal, coeloconoidal - see
especially Fig. 3 below);
♦ shape of the whorls (convex, concave, straight -
with or without shoulder or keel);
♦ spiral cords of the whorls (ontogeny, number,
beads, strength);
♦ spiral cords on the base (number, beads, distance
between);
♦ shape of the aperture, the outer and the inner lip.

Figure 1. Map of locations of MNHN campaigns referenced in this paper — the abbreviations of the campaigns
are those defined in table 1; inserts from the MNHN "Réferentiel BasExp" web site
(https://expeditions.mnhn.fr/campaign/list).

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C. VILVENS New species and new records of Calliostomatidae. Part II

Figure 2. Features of Calliostomatidae shells; H : height; W : width; HA : height of the aperture; P1, P2, P3, ... :
primary cords; S1, S2, S3, … : secondary cords; T1, T2, T3, … : tertiary cords (shell : Calliostoma chlorum
Vilvens, 2005, Fiji, BORDAU 1, stn DW1454, 13.6 × 10.4 mm).

Figure 3. Shapes of Calliostomatidae shells (scale bars: 5 mm): A. Conical shape: Fautor boucheti (B.A.
Marshall, 1995), southern New Caledonia, NORFOLK 2, stn DW2081, 500–505 m, 10.1 × 9.4 mm;
B. Cyrtoconoidal shape: Laetifautor fundatus B.A. Marshall, 1995, north-eastern New Caledonia, Touho,
MONTROUZIER, stn 1269, 15–20 m, 4.2 × 3.5 mm; C. Coeloconoidal shape: Bathyfautor coriolis B.A.
Marshall, 1995, West of New Caledonia, Lansdowne Bank, EBISCO, stn CP5052, 684–715 m, 18.8 × 17.0 mm.

58
C. VILVENS
Abbreviations
Repositories
MNHN: Muséum national d'Histoire naturelle, Paris,
France.
HMNS: Houston Museum of Natural Science.
IMT: Institute of Malacology of Tokyo, Tokyo, Japan.
MM: Manchester Museum, University of Manchester.
NHMUK: Natural History Museum, London, United
Kingdom.
NIWA: National Institute of Water and Atmospheric
Sciences, Wellington, New Zealand.
NMMBA: National Museum of Marine Biology and
Aquarium, Pingtung, Taiwan.
NSMT: National Museum of Science, Tokyo, Japan.
SMF: Senckenberg Museum, Frankfurt, Germany.
ZMA: Zoologisch Museum, Amsterdam, The
Netherlands.
coll.: private collection.
Other abbreviations
H: height;
W: width;
HA: height of the aperture;
TW (tw on plates captions): number of teleoconch
whorls;
P1, P2, P3, ... : primary cords = spiral cords appearing
on the first whorls (P1 is the most adapical; the last
one is the most abapical, usually peripheral);
Pi: all the primary cords (usually i=1,…,4);
S1, S2, S3, ... : secondary cords = spiral cords
appearing after the Pi (S1 is the most adapical);
Si: all the secondary cords (usually i=1,…,4);
T1, T2, T3, … : tertiary cords = spiral cords appearing
on the last whorls (numbered following appearing
order);
Figure 4. Map of the eastern part of the area referenced in this paper (from p. 60).
NOVAPEX 25(2): 55–110, 10 juin 2024
Ti: all the tertiary cords (i=1…);
P'x, S'x: second part of a primary (secondary) cord Px
(Sx) splitting into two cords;
P"x, S"x: third part of a primary (secondary) cord Px
(Sx) already split into two cords;
X, Y: in ontogeny description, the spiral cord X
appears a little earlier than Y;
X; Y: in ontogeny description, the spiral cord X
appears clearly much earlier than Y;
(X): in ontogeny description, the spiral cord X
disappears;
stn: station;
lv: live-taken specimens present in sample;
dd: no live-taken specimens present in sample;
sub: subadult specimen;
juv: juvenile specimen.
SYSTEMATICS
We follow here the classification of WoRMS (which
itself follows Bouchet et al., 2017) where
Calliostomatidae are ranked as a family of superfamily
Trochoidea (besides e. g. Trochidae or Solariellidae),
with the five subfamilies Calliostomatinae,
Fautricinae, Margarellinae, Thysanodontinae and
Xeniostomatinae.
The different species are browsed trying to follow a
path starting in the Solomon Islands and ending at
Taiwan.
Superfamily TROCHOIDEA Rafinesque, 1815
Family CALLIOSTOMATIDAE Thiele, 1924 (1847)
Subfamily CALLIOSTOMATINAE Thiele, 1924
(1847)
Genus Calliostoma Swainson, 1840
Type species : Trochus conulus Linnaeus, 1758 (by
s.d. Herrmannsen, 1846) – Recent, Mediterranean Sea.
Grammatical gender: neuter.
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C. VILVENS New species and new records of Calliostomatidae. Part II

Calliostoma hexalyssion Vilvens, 2009 nearly straight to weakly concave with a suprasutural
Fig. 6A–H keel, at least 8 spiral cords (P2, P3; P1; P4, S3; S1, S2;
S'3; possible T1 subsutural, possible S"3 above S3)
Calliostoma (Benthastelena) hexalyssion Vilvens, with small, acute pointed beads and distance between
2009: 142–146, figs A1–A2, 73–78. Type locality: all Pi and Si much larger than their size, S3 making
Solomon Islands, Vella Gulf, between Vella Lavella keel, a subquadrangular aperture, an oblique columella
Is. and Kalombangara Is., 460–487 m. without tooth, an almost flat base with about 15 spiral
Calliostoma hexalyssion —Vilvens, 2014a: 44, 46, cords of which only the two most external and the
figs 32–33. three most internal are clearly visible and rather
strong, anomphalous, a pinkish white colour.
Material examined. Solomon Islands. SALOMON
2: stn CP2213, 07°39'S, 157°43'E, 495–650 m, 2 lv Remarks. This species was originally described from
(MNHN-IM-2007-34090, MNHN-IM-2007-34091); Solomon Islands and the new records from Papua
stn CP2214, 07°42'S, 157°44'E, 550–682 m, 1 lv New Guinea extend rather logically its distribution
(MNHN-IM-2007-34089); stn CP2243, 07°43'S, area.
156°27'E, 518–527 m, 1 juv lv (MNHN-IM-2007-
34150); stn CP2245, 07°43'S, 156°26'E, 582–609 m, 1
sub lv (MNHN-IM-2007-34149); stn CP2247,
07°45'S, 156°25'E, 686–690 m, 2 lv (MNHN-IM-
2007-34139, MNHN-IM-2007-34140); stn CP2248,
07°43'S, 156°25'E, 650–673 m, 2 lv (MNHN-IM-
2007-34137, MNHN-IM-2007-34138).
SALOMONBOA 3: stn CP2773, 09°25'S, 160°32'E,
537–619 m, 2 lv (MNHN-IM-2009-6260, MNHN-IM-
2009-6261); stn CP2849, 09°36'S, 160°46'E, 448–523
m, 1 juv lv (MNHN-IM-2009-6898).
Papua New Guinea. BIOPAPUA: stn CP3653,
02°13'S, 150°23'E, 680–700 m, 1 lv (MNHN-IM-
2013-57653), 1 lv; stn CP3654, 02°14'S, 150°16'E,
490–505 m, 1 lv (MNHN-IM-2013-57648); stn
CP3722, 06°01'S, 147°35'E, 608–625 m, 1 lv
(MNHN-IM-2013-57652); stn CP3737, 08°15'S,
150°45'E, 587 m, 1 lv (MNHN-IM-2013-57647,
MNHN-IM-2013-57654); stn CP3739, 09°09'S,
152°15'E, 503–546 m, 1 dd; stn CP3740, 09°12'S,
152°16'E, 556–645 m, 5 lv (MNHN-IM 2013-57656,
MNHN-IM-2013-57657, MNHN-IM-2013-57658,
MNHN-IM2013-57659, MNHN-IM-2013-576601 sub
lv (MNHN-IM-2013-57655), 3 lv; stn CP3742,
09°08'S, 152°19'E, 448–470 m, 1 dd; stn DW3749,
05°39'S, 153°59'E, 620–663 m, 1 dd; stn DW3757,
04°01'S, 153°36'E, 214 m, 1 dd. KAVIENG 2014: stn
CP4421, 02°23'S, 150°37'E, 470–525 m, 1 sub dd; stn
CP4422, 02°21'S, 150°38'E, 496–609 m, 3 dd; stn
CP4423, 02°20'S, 150°38'E, 550–649 m, 1 dd, 3 sub Figure 5. Map of the western part of the area
dd; stn CP4437, 02°23'S, 150°37'E, 416–535 m, 1 lv; referenced in this paper (from p. 68).
stn CP4439, 02°23'S, 150°35'E, 534–650 m, 2 lv, 1
sub dd; stn CP4440, 02°19'S, 150°37'E, 583–658 m, 1 Calliostoma simplex Schepman, 1908
lv; stn CP4448, 02°13'S, 150°12'E, 564–743 m, 2 dd, Fig. 6I–M
1 juv dd.
Calliostoma simplex Schepman, 1908: 64, pl. V, fig.
Distribution. Solomon Islands, 77–728 m, living at 5. Type locality: Indonesia, Arafura Sea, 05°48'S,
397–686 m (range computed using also data of 123°13'E, 304 m.
Vilvens, 2009; 2014a); Papua New Guinea, 214–680 Calliostoma simplex — Poppe et al., 2006: 123, pl.
m, living at 505–680 m. 64, fig. 5; Vilvens, 2009: 140, 151, figs 57–58.

Diagnosis. The main characteristics of this species are
a height up to 22.1 mm, a width up to 18.2 mm, a very
elevated, conical to slightly coeloconoidal shape, an
angulate periphery, a teleoconch of up to 9.1 whorls
60
Material examined. Papua New Guinea. PAPUA
NIUGINI: stn DW3974, 04°34'S, 146°16'E, 710 m, 1
juv dd; stn DW3976, 04°33'S, 146°17'E, 680 m, 1 dd;
stn DW3997, 06°03'S, 147°35'E, 352 m, 1 sub dd, 1
juv dd.
C. VILVENS
Indonesia, Kai Islands. KARUBAR: stn DW14,
05°18'S, 132°38'E, 245–246 m, 1 sub dd.
Distribution. Eastern Indonesia, 246–304 m (range
computed using also data of Schepman, 1908); Papua
New Guinea, 352–680 m (dd); Philippines, 191–192
m (Poppe et al., 2006).
Diagnosis. The main characteristics of this species are
a height about 14 mm, a width about 11.5 mm, an
elevated, conical shape, an angulate periphery, a
teleoconch of 7.5 straight whorls, 7 spiral cords (P2,
P3, P1; S2; S3; S1; P4), no cord making keel, a
subelliptical aperture, a slightly oblique columella
without tooth, a nearly flat base with 10 beaded, rather
thin spiral cords with a distance between them at least
1.5× their width, anomphalous, a yellowish colour.
Remarks. The few specimens here recorded and
collected in Papua New Guinea are in a rather poor
state (incrusted, broken) and/or subadult. Some of
them lack indeed S1 cord, but this seems too slight to
consider a potential different species.
Calliostoma aporia Vilvens, 2009
Fig. 6N–O
Calliostoma (Ampullotrochus) aporia Vilvens, 2009:
150–151, figs F1–F2, 107–110. Type locality:
Solomon Islands, south coast of Tetepare Is., New
Georgia Group, 105–128 m.
Calliostoma aporia — Vilvens, 2014a: 46, figs 28–29.
Material examined. Solomon Islands. SALOMON
2: stn CP2207, 07°44'S, 158°30'E, 336–341 m , 2 sub
dd, 3 juv dd; stn DW2237, 06°53'S, 156°22'E, 400 m,
1 juv dd.
Papua New Guinea. BIOPAPUA: stn DW3763,
04°40'S, 153°03'E, 160–170 m, 4 dd. KAVIENG
2014: stn DW4506, 02°35'S, 150°43'E, 94–95 m, 1
dd, 1 sub dd.
Distribution. Solomon Islands, 128–400 m (dd)
(range computed using also data of Vilvens, 2009);
Papua New Guinea, 95–160 m (dd).
Diagnosis. The main characteristics of this species are
a height up to 11.2 mm, a width up to 8.8 mm, an
elevated, conical shape, an angulate periphery, a
teleoconch of up to 7.7 straight whorls with a
suprasutural keel, at least 6 spiral cords (P1, P2, P3;
S3; P4; S1; Ti; no S2), S3 making a weak keel, a
roundly subquadrangular aperture, a weakly oblique
columella without tooth, a weakly convex to almost
flat base with 9–10 strong, granular spiral cords,
anomphalous, an off-white to light grey colour with
possible brownish flames or shades.
Remarks. As already pointed out in the original
description, C. simplex Schepman, 1908 differs from
NOVAPEX 25(2): 55–110, 10 juin 2024
C. aporia in being slightly larger, in having a S2
(commencing before S3), a keel on S3 and much
thinner spiral cords on the base.
This species was originally described from Solomon
Islands and these new records from Papua New
Guinea extend rather logically its distribution area.
Calliostoma quadricolor Schepman, 1908
Fig. 7A–D
Calliostoma quadricolor Schepman, 1908: 65, pl.V,
fig. 7. Type locality: Indonesia, from Sape Strait to
Kei Dulat Lau, 52–69 m.
Material examined. Indonesia, Kai Islands.
KARUBAR: stn DW32, 05°47'S, 132°51'E, 170–206
m, 1 sub dd.
Indonesia, Tanimbar Islands. KARUBAR: stn
DW44, 07°52'S, 132°48'E, 291–295 m, 1 sub dd.
Distribution. Indonesia, 69–291 m (dead) (range
computed using also data of Schepman, 1908).
Diagnosis. The main characteristics of this species are
a height up to 8.5 mm, a width up to 7.0 mm, an
elevated, conical shape, an angulate periphery, a
teleoconch of 7 straight whorls, up to 7 spiral cords
(P1, P2, P3; S3; S1; P4; ?no S2), S3 making keel, a
subquadrangular aperture, a slightly oblique columella
without tooth, an almost flat base with up to 13
beaded spiral cords, anomphalous, a yellowish to
greyish colour, S3 with white and reddish brown
segments, first whorls purplish.
Remarks. This is a very poorly known species. The
available syntype is clearly not the one that was used
for the writing of the original description (Marshall,
1995: 401, 415). When comparing nevertheless this
syntype to the original description (text and drawing),
it appears that a question is if there is a S2 cord
(description) or not (syntype). The two specimens here
studied from Indonesia are both in a poor state
(especially the early whorls), one being subadult
(about 5.5-6 whorls). All that is a bit short to provide
more accurate features about this species.
Calliostoma pheidolon n. sp.
Fig. 7E–F
urn:lsid:zoobank.org:act:7E7FB1AB-0829-4C61-
8DD0-806084F41B1C
Type material. Holotype (8.8 × 7.7 mm) MNHN-IM-
2000-39420.
Type locality. Papua New Guinea, off New Ireland,
BIOPAPUA, stn DW3764, 04°39'S, 153°03'E, 220 m.
Material examined. Papua New Guinea.
BIOPAPUA: stn DW3764, 04°39'S, 153°03'E, 220 m,
1 dd (holotype MNHN-IM-2000-39420).
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C. VILVENS New species and new records of Calliostomatidae. Part II

Distribution. Only known from the type locality. Colour of teleoconch cream; protoconch whitish.

Diagnosis. A Calliostoma species of rather small size,
with a rather elevated, conical spire, 6 granular spiral
cords (P1, P2, P3; P4; S1; S4; no S2, no S3), P4 the
strongest with blunt sharp beads, a flat base with 8
rather thin granular spiral cords with interspaces larger
than cords, anomphalous, a cream colour.
Description. Shell rather small for the genus (H up to
8.8 mm, W up to 7.7 mm), slightly higher than wide,
conical in shape; spire rather elevated, height 1.1×
width, 3.8× aperture height, apical angle 60°; angulate
periphery; anomphalous.
Protoconch about 350 μm wide, 1.25 whorls, rounded,
with tiny spits, without terminal varix.
Teleoconch of 7.5 flat whorls; early whorls with 3
spiral cords, last whorl with 6 cords.
Suture weakly impressed, not canaliculated.
First whorl weakly convex, sculptured by about 15
rather thick, axial, prosocline ribs and 3 granular spiral
cords appearing almost immediately and
simultaneously; beads produced by intersection of
cords with ribs; P1 slightly thinner than the other
cords; distance between ribs similar in size to their
width, distance between cords similar to their width.
Second whorl flat, beads of cords stronger; P1 and P2
similar in size, P3 slightly stronger than the other
cords with rather thick, blunt sharp beads. On third
whorl, P4 emerging from suture, partially hidden by
next whorl. On fourth whorl, P4 completely visible,
similar in size to P3, both with blunt sharp beads; P1
and P2 slightly weaker. On fifth whorl, P4 the
strongest. On sixth whorl, S1 appearing, much thinner
than the other cords; neither S2 nor S3. On seventh
whorl, S1 similar in size to P1 and P2. On last whorl,
S4 peripheral, thin; P4 still the strongest, distance
between P3 and P4 larger than those between the other
cords that are similar to the size of cords.
Aperture subquadrate. Columella straight, slightly
oblique.
Base flat, with 8 granular spiral cords, beads produced
by intersection of cords with thin radial ridges;
distance between cords slightly larger than cords.
Discussion. Although there is a single specimen (but
fully mature) of this new species, it can be
distinguished rather easily from the similar existing
species known in the area where it has been collected.
Calliostoma pheidolon n. sp. is rather close to C.
quadricolor Schepman, 1908 from Indonesia (Fig.
7A–D) but this more or less similar in size species has
a less elevated spire, a different ontogeny of the spiral
cords (especially with S3 present but no S4), rounded
beads on all the spiral cords on the spire and more
numerous spiral cords on the base.
The new species can be compared to C. simplex
Schepman, 1908 from Indonesia (Fig. 6I–M), but this
larger species with a smaller ratio H/W has a different
ontogeny of spiral cords (especially with S2) and no
cord making keel.
The new species is also comparable to C. aporia
Vilvens, 2009 from Solomon Islands and Papua New
Guinea (Fig. 6N–O), but this more or less similar in
size species has a slightly larger H/W ratio and a
different ontogeny of cords (especially with S3 and
some Ti)
Calliostoma pheidolon n. sp. slightly remembers C.
maekawai Poppe, Tagaro & Goto, 2018 from the
Philippines (Fig. 8O), but this similar in size species
has a weakly coeloconoidal shape and much thinner
spiral cords (the ontogeny of these cords is not
provided in the original description) without an
obvious suprasutural keel on an abapical cord.
Etymology. Thrifty (Ancient Greek : φειδωλóς, -ή, -
όν) - with reference to reduced number of spiral cords
on the whorls.
Calliostoma philippei Poppe, 2004
Fig. 7G–L
Calliostoma philippei Poppe, 2004: 5–6, pl. 1. Type
locality: Philippines, off Aliguay Island, 80–200 m.
Calliostoma philippei — Poppe, Tagaro & Dekker,
2006: 122, pl. 63, fig. 6; Vilvens, 2009: 151, figs K1–
K2, 111–114; Vilvens, 2023: 54, figs 5N–O.

Figure 6 (scale bars: 5 mm)

A–H. Calliostoma hexalyssion Vilvens, 2009. A–B. Solomon Islands, SALOMON 2, stn CP2214, 550–682 m,
20.7 × 12.5 mm, 8.7 tw. C–H. Papua New Guinea, BIOPAPUA. C–D. MNHN (MNHN-IM2013-57652), stn
CP3722, 608–625 m, 21.2 × 17.2 mm, 9.0 tw. E–H. MNHN (MNHN-IM2013-57653), stn CP3653, 680–700 m,
17.7 × 14.9 mm, 8.0 tw
I–M. Calliostoma simplex Schepman, 1908. I–J. Holotype ZMA, Indonesia, Arafura Sea, SIBOGA, stn 253,
304 m, 14 × 11.5 mm, 7.5 tw. K–L. Papua New Guinea, PAPUA NIUGINI, stn DW3997, 352 m, 11.0 × 9.6 mm,
6.0 tw. M. MNHN (MNHN-IM-2014-10802), Philippines, Sulu Sea, North of Lubang, MUSORSTOM 2, stn
CP21, 191–192 m, 17.1 × 14.9 mm, 8.5 tw.
N–O. Calliostoma aporia Vilvens, 2009, Papua New Guinea, BIOPAPUA, stn DW3763, 160–170 m, 9.4 × 7.9
mm, 7.6 tw.

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C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

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C. VILVENS New species and new records of Calliostomatidae. Part II

Material examined. Solomon Islands.
SALOMONBOA 3: stn DW2818, 10°10'S, 161°55'E,
250–258 m, 1 lv (MNHN IM-2009-6079); stn
CP2819, 10°11’S, 161°54’E, 250-296 m, 1 lv
(MNHN-IM-2009-6663).
Papua New Guinea. BIOPAPUA: stn DW3640,
06°46'S, 148°00'E, 322 m, 1 dd; stn DW3719,
06°03'S, 147°36'E, 410 m, 2 dd, 2 sub dd. PAPUA
NIUGINI: stn stn DW3992, 06°03'S, 147°36'E, 450–
480 m, 4 dd, 3 juv dd; stn CP4017, 05°39'S, 148°14'E,
280–315 m, 1 dd.
Distribution. Philippines, Aliguay Island, 80–150 m
(range computed using material of Poppe et al., 2006);
Solomon Islands, 277–283 m, lv at 283–305 m (also
using data of Vilvens, 2009); West of New Caledonia,
Chesterfield plateau, 386–428 m (dd) (Vilvens, 2023);
Papua New Guinea, 315–450 m (dd).
Remarks. Calliostoma philippei Poppe, 2004 was
originally described from the Philippines based upon a
single specimen. Two other specimens were recorded
from the Philippines (Poppe et al., 2006), 5 additional
specimens were recorded from Solomon Islands
(Vilvens, 2009) and another additional one has been
recorded from Chesterfield plateau, West of New
Caledonia (Vilvens, 2023).
The material here studied, collected in Solomon
Islands and Papua New Guinea, gives some additional
features about this species: two specimens are indeed
much greater than the holotype and known samples,
with a measure of 16.0 × 10.6 mm and 8.1 whorls for
the highest one, 14.4 × 11.5 mm and 7.4 whorls for
the widest one. Moreover, the two additional whorls
are convex (instead of straight), changing the general
shape of the shell.
Because the original description is a bit vague and the
holotype is apparently not available, we propose here
only a new diagnosis taking in account the new
features, instead of a redescription that would need to
have the type in hand.
New diagnosis. The main characteristics of this
species are a height up to 16.0 mm, a width up to 11.5
mm, a very elevated shape with a conical adapical part
and a weakly cyrtoconoidal abapical part, an angulate
periphery, a teleoconch of 8.1 whorls, 7 spiral cords
(P1, P2, P3; S3; S2, P4; S1), a subquadrangular
aperture, a weakly oblique columella without tooth, an
almost flat base with 10–13 granular spiral cords and
strong axial spiral grooves, anomphalous, a
porcellaneous white colour with P1, P2, S1 and S2
brown.
Additional remarks. All the adult specimens from
Papua New Guinea are in a rather poor state, with
especially the early whorls systematically missing.
The ontogeny of cords has been described based upon
the shells of some subadult specimens: this leads to
(P1, P2, P3; S2; S3; P4; possible S1), meaning the
only difference could be that S2 and S3 appear in the
reverse order comparing to C. philippei in good state
from Solomon Islands.
Calliostoma newguineense n. sp.
Fig. 8A–D
urn:lsid:zoobank.org:act:07ADF6A4-90F2-47E7-
8B08-0FC00490C290
Type material. Holotype (21.4 × 21.3 mm) MNHN-
IM-2000-39421.
Type locality. Papua New Guinea, New Ireland,
KAVIENG 2014, stn CP4444, 02°15'S, 150°14'E,
417–421 m.
Material examined. Papua New Guinea, Solomon
Sea, Vitiaz Strait. BIOPAPUA: stn DW3719,
06°03'S, 147°36'E, 410 m, 1 dd. PAPUA NIUGINI:
stn DW3992, 06°03'S, 147°36'E, 450–480 m, 1 juv
dd.
Papua New Guinea, Bougainville seamounts.
BIOPAPUA: stn DW3745, 05°33'S, 154°00'E, 369–
377 m, 1 dd, 1 sub dd, 1 juv dd.
Papua New Guinea, New Ireland. KAVIENG 2014:
stn CP4444, 02°15'S, 150°14'E, 417–421 m, 1 dd
(holotype MNHN-IM-2000-39421).

Figure 7 (scale bars: 5 mm)

A–D. Calliostoma quadricolor Schepman, 1908, Indonesia. A–B. Syntype ZMA, SIBOGA, stn 49A, Sape
Strait, Flores, Indonesia, 69 m , 6.6 × 6.1 mm, 6.8 tw. C–D. Indonesia, Kai Islands, KARUBAR, stn DW32,
170–206 m, 8.6 × 8.8 mm, 6.0 tw.
E–F. Calliostoma pheidolon n. sp., holotype MNHN-IM-2000-39420, Papua New Guinea, off New Ireland,
BIOPAPUA, stn DW3764, 220 m, 8.8 × 7.7 mm, 7.5 tw.
G–L. Calliostoma philippei Poppe, 2004. G–H. MNHN-IM-2009-6079, Solomon Islands, SALOMONBOA 3,
stn DW2818, 250–258 m, 16.0 × 10.6 mm, 8.1 tw. I–J. MNHN-IM-2009-6663, Solomon Islands,
SALOMONBOA 3, stn unknown, 14.4 × 11.5 mm, 7.4 tw. K–L. Papua New Guinea, Salomon Sea, Vitiaz Strait.
K–L. PAPUA NIUGINI, stn DW3992, 450–480 m, 12.4 × 10.8 mm. M–N. BIOPAPUA, stn DW3719, 410 m,
15.0 × 10.5 mm.
M–P. Calliostoma altena Knudsen, 1970, Indonesia, Tanimbar Islands, KARUBAR. M–N. Stn DW61, 235–
236 m, 14.5 × 14.0 mm, 7.0 tw. O–P. Stn CP84, 246–275 m, 11.9 × 11.0 mm, 6.8 tw.

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C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

65
C. VILVENS
Distribution. Papua New Guinea, 377–450 m (dd).
Diagnosis. A rather large Calliostoma species, with a
rather elevated, coeloconoidal spire, a subangular
periphery, a teleoconch of 8.0 whorls, up to 13
granular spiral cords (P1, P2, P3, P4; S1; S2, S3; T1;
T2; T3, T4; additional Ti) with sharp beads, P3 the
strongest, making a moderate keel, a weakly convex
base with 15–18 subgranular to granular spiral cords,
distance between cords more or less similar to their
width, anomphalous, a yellowish to orangey cream
colour.
Description. Shell rather large for the genus (H up to
21.4 mm, W up to 21.3 mm), more or less as high as
wide, slightly coeloconoidal in shape; spire rather
elevated, height similar to width, 2.5× aperture
height, apical angle 65°; subangulate periphery;
anomphalous.
Protoconch about 400 μm wide, 1.25 whorls, rounded,
with tiny spits, with a terminal varix.
Teleoconch up to 8.0 whorls, early whorls flat to
slightly concave, last whorls with a slightly convex
abapical part; early whorls with 3 spiral cords, last
whorl with up to 14 cords of uneven thickness.
Suture weakly impressed, not canaliculated.
First whorl weakly convex, sculptured by about 16
rather weak prosocline ribs and 4 granular spiral cords
appearing almost immediately and simultaneously;
small beads produced by intersection of cords with
ribs; distance between ribs similar in size to their
width, interspaces between cords twice their width.
Second whorl still weakly convex, cords, ribs and
beads of cords stronger; P4 sunkening under suture at
beginning of whorl and becoming invisible; P3
stronger than the other cords with rather thick beads.
Third whorl almost flat, P3 the strongest with blunt
sharp beads, P2 the thinnest; axial ribs much stronger;
S1 appearing, very thin. On fourth whorl, S1 still
much thinner than the other cords; P3 the strongest
and producing keel; P4 slightly emerging from suture
but still almost completely hidden by next whorl. Fifth
whorl slightly concave, S2 appearing, S3 appearing a
bit later; P3 and P1 stronger than the other cords, P3
the strongest, Si thinner than P2, S3 the thinnest. On
sixth whorl, T1 appearing between P1 and S1, T2 later
between S1 and P2; beads of all Pi and Si pointed. On
seventh whorl, T3 appearing between P2 and S2 and
T4 between S2 and P3. On last whorl, P4 emerging,
additional Ti appearing on the large specimens;
moderate keel on P3; beads of all cords rather close
and sharp, making a prickly appearance to the surface.
Aperture subquadrate with a rounded outer lip.
Columella straight, slightly oblique.
Base weakly convex, with 15–18 subgranular to
granular spiral cords, innermost ones stronger, beads
produced by intersection of cords with thin radial
ridges, distance between cords more or less similar to
their width.
66
New species and new records of Calliostomatidae. Part II
Colour of teleoconch yellowish to orangey cream;
protoconch whitish.
Discussion. Calliostoma newguineense n. sp. is rather
close to Tristichotrochus tosaensis Kuroda & Habe,
1961 from southern Japan (Fig. 8E), but this similar in
size species has a more elevated spire with a ratio
H/W greater than 1, a different ontogeny regarding the
order of appearance of the Si with (S2; S3; S1) instead
of (S1, S2; S3) and a double keel made by P3 et S3
instead a single keel produced by P3.
The new species is also rather close to Calliostoma
rufomaculatum Schepman, 1908 from Indonesia (Fig.
8G–H), but the holotype is smaller for the same
number of whorls, has a more elevated spire with
slightly concave last whorls and a different ontogeny
of teleoconch cords (P1, P2, P3; S3; S1, S2; Ti).
C. newguineense n. sp. may also be compared to C.
sakashitai (Sakurai, 1994) from Japan and East China
Sea (Fig. 8F), but this species has a smaller size for
the same number of whorls, a more elevated spire with
a higher H/W ratio and a different ontogeny of spiral
cords (S2; S3; S1) instead of (S1, S2; S3).
The general shape of the new species remembers the
one of C. crassicostatum Schepman, 1908 from
Indonesia, Celebes Sea (Fig. 8M–N) but this species is
much smaller (H of 9 mm) for a similar number of
whorls, has a peripheral suprasutural spiral cord and
has a partially open umbilicus.
Remarks. The holotype is the single specimen fully
mature and in good state, the other samples being
subadult and/or with broken parts, making them
inappropriate to be designated as paratypes.
Etymology. Named after the country of type locality.
Calliostoma thaumaston n. sp.
Fig. 8I–L, Table 2
urn:lsid:zoobank.org:act:F1D202EC-3D9A-49FA-
AB3B-7E81E7EBAE2A
Type material. Holotype (15.0 × 14.1 mm) MNHN-
IM-2000-39422. Paratype: 1 MNHN (MNHN-IM-
2000-39423, as listed below).
Type locality. Indonesia, Kai Islands, KARUBAR,
stn CP25, 05°30'S, 132°52'E, 336–346 m.
Material examined. Indonesia, Kai Islands.
KARUBAR: stn CP25, 05°30'S, 132°52'E, 336–346
m, 1 lv (holotype MNHN-IM-2000-39422).
Papua New Guinea, Salomon Sea, Vitiaz Strait.
PAPUA NIUGINI: stn DW3992, 06°03'S, 147°36'E,
450–480 m, 1 dd (paratype MNHN-IM-2000-39423).
Distribution. Indonesia, Kai Islands, 336–346 m (lv);
Papua New Guinea, Salomon Sea, Vitiaz Strait, 450–
480 m (dd).
C. VILVENS
Diagnosis. A Calliostoma species of large size, with a
moderately elevated, conical to slightly cyrtoconoidal
spire, a subangulate periphery, up to 7.2 whorls flat
except on the last whorls weakly convex, at least 10
granular spiral cords (P1, P2, P3; S2, S1; S3; P4; T1,
T2; T3, possible T4), a weakly convex to almost flat
base with 14 rather thin subgranular to granular spiral
cords with interspaces less or equal to their width,
anomphalous, a light orange to brown colour.
Description. Shell rather large for the genus (H up to
19.7 mm, W up to 18.0 mm), slightly higher than
wide, conical to slightly cyrtoconoidal; spire rather
elevated, height 1.1× width, 2.5× to 3.0× aperture
height, apical angle 65° to 70°; subangulate periphery;
anomphalous.
Protoconch 390 μm wide (holotype, missing on
paratype), 1.25 whorls, rounded, with a few tiny spits,
without terminal varix.
Teleoconch up to 7.2 whorls, early whorls flat, last
whorls weakly convex; early whorls with 3 spiral
cords, last whorl with at least 10 cords.
Suture visible, neither impressed nor canaliculated.
First whorl weakly convex, sculptured by about 16
weak prosocline ribs and 3 granular spiral cords
appearing almost immediately and simultaneously;
beads produced by intersection of cords with ribs; P1
slightly thinner than the other cords; distance between
ribs similar in size to width of ribs, interspaces
between cords about 2× their width. Second whorl
Table 2. Calliostoma thaumaston n. sp.: Shells measurements in mm for types.
TW H W
Holotype KARUBAR CP25
6.5 15.0
MNHN-IM-2000-39422
Paratype PAPUA NIUGINI:
DW3992 MNHN-IM-2000- 7.2 19.7
39423
Remarks. At first glance, the two specimens of the
new species may look a bit different. But a more
careful examination, especially of the ontogeny of the
spiral cords, while taking in account the difference of
size and the number of whorls, leads to the conclusion
that there is no objective reason to separate them.
Discussion. Calliostoma thaumaston n. sp., especially
the paratype, could remind of C. newguineense n. sp.
from Papua New Guinea (Fig. 8A–D), but this similar
in size species has a coeloconoidal shape (not conical
or weakly cyrtoconoidal), a different ontogeny of the
Si (S2, S1; S3 while S1; S2, S3; for C. newguineense)
and rounded spiral cords beads instead of acute sharp
beads.
The new species is weakly similar to some samples of
C. emmanueli Vilvens, 2000 from the Philippines and
Solomon Islands (Fig. 9K–R), but this similar species
has a different ontogeny regarding the Si (S2, S3; S1;
P4), slightly more convex whorls and more numerous
whorls for a same size.
NOVAPEX 25(2): 55–110, 10 juin 2024
flat, beads of cords stronger; P1 and P2 similar in size,
P3 slightly stronger than the other cords with blunt
sharp beads; axial ribs stronger. On third whorl, P3
much stronger than other cords with thicker beads; S2
appearing, very thin; S1 appearing a bit later; S3
appearing near the end of the whorl while S1 and S2
already rather strong. On fourth whorl, axial sculpture
vanishing; P3 still the strongest, P1 stronger than all
the other cords; P4 slightly emerging from suture,
partially hidden by next whorl. On fifth whorl, P4
more visible. On sixth whorl, T1 appearing between
P1 and S1, T2 a bit later between S1 and P2; T3
appearing near the end of the whorl between S2 and
P3 while T1 and T2 already rather strong; possible T4
between P2 and S2. On last whorl, P4 peripheral,
thinner than all other Pi and Si; all cords rather close,
distance between smaller than their width.
Aperture subelliptical. Columella slightly curved,
slightly oblique.
Base weakly convex to almost flat, with 14 spiral
cords, innermost cords stronger and granular,
innermost ones thinner and subgranular, beads
produced by intersection of cords with thin radial
ridges; distance between cords slightly equal or
smaller than their width.
Colour of teleoconch light orange to brown;
protoconch light orange (holotype).
Operculum circular, multispiral, corneous, light
brown.
HA H/W H/HA
14.1 6.1 1.06 2.46
18.0 6.5 1.09 3.03
C. thaumaston n. sp. may be compared to Fautor
cosmopolites n. sp. from Papua New Guinea to
Taiwan (Fig. 18A–H), but this species is much larger
for a same number of whorls (with a ratio H/TW of
1.3–1.7 instead of 2.3–2.7), has a H/W ratio less than
1.1 (instead of 1.1–1.2 for C. cosmopolites) and a
different ontogeny of the Si (S2, S1; S3 while S2; S3;
S1 for C. cosmopolites)
The general shape of the new species (especially the
holotype) remembers the one of C. crassicostatum
Schepman, 1908 from Indonesia (Celebes Sea) (Fig.
8M–N) but this species is much smaller (H of 9 mm)
for a similar number of whorls, has a peripheral
suprasutural spiral cord and has a partially open
umbilicus.
Etymology. Surprising, unexpected (Greek:
θαυμαστόσ, -ή, -όν) - with reference to the apparent
difference between the two types of the new species.
67
C. VILVENS New species and new records of Calliostomatidae. Part II

Calliostoma altena Knudsen, 1970
Fig. 7M–P
Calliostoma altena Knudsen, 1970: 18, pl. 1A–B, 2A–
B. Type locality: Pulu Tonijn, Makassar Strait, S.W.
of Celebes, Indonesia, 900 m.
Material examined. Indonesia, Tanimbar Islands.
KARUBAR: stn DW61, 09°05'S, 132°44'E, 235–236
m, 2 dd; stn CP84, 09°23'S, 131°09'E, 246–275 m, 2
sub dd.
Distribution. Indonesia, Makassar Strait, 900 m;
Tanimbar Islands, 236–246 m (dd).
Diagnosis. The main characteristics of this species are
a height up to 32.0 mm, a width up to 29.0 mm, an
elevated, conical to slightly cyrtoconoidal shape, a
subangulate periphery, a teleoconch of 9 straight
whorls, at least 6 granular spiral cords (P2, P3; P4; S2;
P1; S3; T1 between P2–S2, T2 between S2–P3; T3
between P3–S3, T4 between P1–P2), P2 and P3 with
strong beads, other cords with sharp small pointed
beads, P3 the strongest, making keel, P2 making a
secondary carina, a subquadrangular aperture, a
vertical arcuate columella without tooth, a weakly
convex base with at least 10 flat, smooth spiral cords
(30 in the original description), 2–3 innermost cords
stronger, anomphalous, a whitish colour with some
darker flames.
Remarks. The peculiar ontogeny of the spiral cords
and shape of the teleoconch makes it difficult to
mistake this species for another species of the
considered area. It seems that the present records are
the first Recent ones since the description.
Calliostoma scobinatum (Reeve, 1863)
Fig. 9A–F
Zizyphinus scobinatus A.Adams in Reeve, 1863: pl.V,
fig. 29. Type locality: Bombay, depth unknown.
Ziziphinus scobinatus — Brazier, 1878: 44.
Calliostoma scobinatum — Kaicher, 1986: card TR1-
2153.
Calliostoma (Fautor) scobinatum — B. A. Marshall,
1995: 399-401.
Calliostoma scobinatus — Subba Rao, 2003: 86, pl. 8,
Fig. 7.
Calliostoma scobinatum — Poppe, Tagaro & Dekker,
2006: 123, pl. 64, figs 1-3; Vilvens, 2009: 140, figs
45–50, M1–M2.
Material examined. Philippines, Bohol Sea.
PANGLAO 2004: stn P3, 09°31'N, 123°42'E, 100 m,
1 dd; stn T27, 09°33'N, 123°51'E, 106–137 m, 1 dd.
Distribution. India (Bombay area), depth unknown;
Torres Straits, 22 m (Brazier, 1878); Philippines, 81–
106 m (range computed using material of Poppe et al,
2006); Solomon Islands, 82–83 m (Vilvens, 2009).

Figure 8 (scale bars: 5 mm)

A–D. Calliostoma newguineense n. sp., Papua New Guinea. A–B. Holotype MNHN-IM-2000-39421, New
Ireland, KAVIENG 2014, stn CP4444, 417–421 m, 21.4 × 21.3 mm, 8.0 tw. C–D. Bougainville seamounts,
BIOPAPUA, stn DW3745, 369–377 m, 16.6 × 21.3 mm.
E. Tristichotrochus tosaensis Kuroda & Habe, 1961, New Caledonia, eastern Lifou, Cap des Pins,
MUSORSTOM 6, CP465, 22.4 × 20.4, 8 tw.
F. Calliostoma sakashitai (Sakurai, 1994), Taiwan, Dali district, 15.2 × 12.6 mm, Fu coll.
G–H. Calliostoma rufomaculatum Schepman, 1908, holotype ZMA, SIBOGA, stn 204, Indonesia, peninsula of
Sulawesi, Buton strait, 22 m, 14.5 × 12.5, 8 tw.
I–L. Calliostoma thaumaston n. sp. I–J. Holotype MNHN-IM-2000-39422, Indonesia, Kai Islands, KARUBAR,
336–346 m, 15.0 × 14.1 mm, 6.5 tw. K–L. Paratype MNHN-IM-2000-39423, Papua New Guinea, Salomon Sea,
Vitiaz Strait, PAPUA NIUGINI, stn DW3992, 450–480 m, 19.7 × 18.0 mm, 7.2 tw.
M–N. Calliostoma crassicostatum Schepman, 1908, Indonesia, Celebes Sea, SIBOGA, stn 133, Lirung,
Salibabu Island, 9 × 8 mm, 36 m, 7.5 tw, drawings from the original description.
O. Calliostoma maekawai Poppe, Tagaro & Goto, 2018, holotype HMNS 597890, Philippines, Balicasag Island,
50–150 m, 10.9 × 10.0 mm (© www.conchology.be).

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C. VILVENS New species and new records of Calliostomatidae. Part II

Diagnosis. The main characteristics of this species are
a height up to 15.9 mm, a width up to 12.0 mm, an
elevated, conical shape, an angulate periphery, a
teleoconch of 8 straight whorls, at least 7 rather thin
spiral cords (P2, P3; P1; P4; S3; S2; S1; some Ti first
on the abapical part) with weakly pointed beads,
distance between cords similar in size to cords, P3 the
strongest on first whorls and S3 the strongest on next
whorls making a weak keel, a subelliptical to
subquadrangular aperture, a slightly oblique columella
without tooth, a moderately convex base (only for
fully mature specimens, weakly convex to almost flat
on subadult specimens) with 8–12 beaded spiral cords
(interspaces at most similar to their width),
anomphalous, a pinkish to yellowish cream colour
with axial light brown flames, S3 with white and light
brown segments for 2–3 beads.
Remarks. Originally described from India, this
species was reported from the Philippines by Poppe et
al. (2006) with a list of 4 stations. The specimens
figured in this work (pl. 64, figs. 1–3) are all smaller
than the types. Among the other available specimens
cited but not figured in the same study, one show
clearly an almost flat base with an angulate periphery
(stn L65-68) whereas a specimen with about 8 whorls
(stn T9) shows a more convex base with a subangulate
base, both with a number of whorls less or equal to 6.
Calliostoma trotini Poppe, Tagaro & H. Dekker, 2006
Fig. 9G–J
Calliostoma trotini Poppe, Tagaro & H. Dekker, 2006:
126, pl. 67, fig. 2. Type locality: Balabac, Palawan
Island, Philippines, depth unknown.
Material examined. Papua New Guinea. KAVIENG
2014: stn DW4453, 02°36'S, 150°38'E, 64–74 m, 1
dd.
South-western Taiwan. TAIWAN 2002: stn CP168,
22°27'N, 120°27'E, 53 m, 1 lv.
Distribution. Philippines, depth unknown; Papua
New Guinea, 64–74 m (dd); Taiwan, lv at 53 m.
Remarks and diagnosis attempt. The original
description of Calliostoma trotini Poppe, Tagaro & H.
Dekker, 2006 from Philippines (unavailable NMP
holotype: 11.8 × 11.3 mm, 6.5 whorls, 8 spiral cords
on the base) is very unclear regarding the ontogeny of
the spiral cords. Also the pictures coming with it don't
permit to establish this ontogeny. So all that can be
said here is that these two specimens respectively
from Papua New Guinea and Taiwan look very like
this species with the following features:
♦ 7.0 × 6.1 mm (KAVIENG 2014 - Papua New
Guinea) and 10.1 × 8.6 mm (TAIWAN 2002 -
Taiwan);
♦ conical, moderately elevated spire;
♦ angulate periphery;
♦ 6.6 to 7.9 straight whorls with the ontogeny (P1,
P2, P3; S1, P4), P3 the strongest, making keel;
♦ base with 3 poorly defined outer spiral cords and
1 granular inner one;
♦ narrow umbilicus.
The single specimen from Taiwan was first thought to
be maybe a different unknown species, because of its
higher number of whorls (7.9 tw) for a similar height
and its smaller ratio H/W. But all the other features
are the same as the known specimens of C. trotini,
except that the Taiwanese specimen has additional Ti
on the last whorls. Only additional material from
Taiwan could enlighten about the possible variations
of this species.
Calliostoma emmanueli Vilvens, 2000
Fig. 9K–R
Calliostoma emmanueli Vilvens, 2000: 4, figs 6–8.
Type locality: Balicasag Island, Bohol, Philippines,
180-240 m.

Figure 9 (scale bars: 5 mm)

A–F. Calliostoma scobinatum (Reeve, 1863). A–B. Lectotype NHMUK (BMNH ) 19688277, India, Bombay,
depth unknown, 15.5 × 11.8 mm, 9.0 tw. C–F. Philippines, PANGLAO 2004. C–D. Panglao Island, stn T9, 97–
120 m, 11.3 × 9.4 mm, 8.1 tw. E–F. Pamilacan Island, stn L65–68, 55–81 m, 6.3 × 5.6 mm, 6.5 tw.
G–J. Calliostoma trotini Poppe, Tagaro & H. Dekker, 2006. G–H. Papua New Guinea, New Ireland,
KAVIENG, stn DW4453, 64–74 m, 7.0 × 6.1 mm, 6.7 tw. I–J. South-western Taiwan, TAIWAN 2002, stn
CP168, 53 m, 10.1× 8.6 mm, 7.9 tw.
K–R. Calliostoma emmanueli Vilvens, 2000. K–N. Philippines, Philippines, Bohol, Balicasag Island. K–L.
Holotype MNHN-IM-2000-30357, 180–240 m, 12.0 × 11.5 mm, 7.0 tw. M–N. 120 m, 10.5 × 9.0 mm, 6.8 tw,
Vilvens coll.. O–R. Indonesia, KARUBAR. O–P. Tanimbar Islands, stn DW49, 206–210 m, 8.5 × 8.2 mm, 6.6
tw. Q–R. Kai Islands, stn DW30, 111–118 m, 15.4 × 13.3 mm, 7.8 tw.

70
C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

71
C. VILVENS
Calliostoma emmanueli — Poppe et al., 2006: 118, pl.
61, figs 5, 7.
Material examined. Indonesia, Kai Islands.
KARUBAR: stn DW30, 05°39'S, 132°56'E, 111–118
m, 1 dd; stn DW49, 08°00'S, 132°59'E, 206–210 m, 1
lv, 1 sub lv, 1 juv.
Solomon Islands. SALOMON 1: stn CP1859,
09°33'S, 160°37'E, 283–305 m, 1 juv lv.
Papua New Guinea, Bismarck Sea. PAPUA
NIUGINI: stn DW3973, 04°34'S, 146°17'E, 411–430
m, 2 juv dd.
Philippines. Fishermen, Balicasag Is., 7/2004, 1 sub
dd.
Distribution. Philippines, Visayas, 100–215 m (range
computed using also data from Poppe et al., 2006);
Indonesia, Malaku (Moluccas), 118–206 m, lv at 206–
210 m; Papua New Guinea, Bismarck Sea, 411–430 m
(dd); Solomon Islands, 411–430 m (lv).
Remarks. These new records extend significantly the
distribution of this species. Originally described from
the Philippines with a single specimen (holotype 12.0
× 11.5 mm, 7 tw), this species was characterized
among other features by a moderately elevated conical
spire with a large apical angle. Poppe et al. (2006)
reported new specimens from Philippines, giving
frontal view of two shells: one of them matching
perfectly with the holotype (pl. 61, fig. 5 : 9 × 8.3
mm) and another one (pl. 61, fig. 7 : 11.7 × 9.4 mm)
more unusual regarding its apical angle and its more
elevated spire, but apparently matching all the other
features of C. emmanueli. So it seems that this species
has some variations about this apical angle.
This last form leads then to compare this species with
C. simplex Schepman, 1908 (Fig. 6I–M): this species
similar in size and in ontogeny is strictly conical with
flat whorls, thinner spiral cords on these whorls and
granular (not nearly smooth) spiral cords on the base
The specimens of C. emmanueli from Indonesia have
a S3 spiral cord appearing a bit later than on the
specimens from Philippines. One of them is especially
large (15.4 × 13.3 for 7.7 whorls), revealing, among
others, for the first time some additional Ti (T1
between P1–S1, T2 between P1–T1, T3 between P3–
S3. The diagnosis has then to be adjusted.
New diagnosis. The main characteristics of this
species are a height up to 15.4.0 mm, a width up to
13.3 mm, a moderately to rather elevated, more or less
conical shape, an angulate periphery, a teleoconch of
7.7 whorls (early whorls almost flat, last whorls
weakly convex), 7 spiral cords (P1, P2, P3; S2, S3;
S1; P4) with additional Ti on very large specimens, no
cord making keel, P1 slightly stronger, a
subquadrangular aperture, a slightly arcuate columella
without tooth, an almost flat base with 10–12 nearly
smooth to weakly subgranular flat spiral cords,
72
New species and new records of Calliostomatidae. Part II
anomphalous, a light brown colour with large reddish
brown patches on first whorls.
Calliostoma paucicostatum Kosuge, 1984
Fig. 10A–I
Calliostoma (Tristichotrochus) paucicostatum
Kosuge, 1984: 5-6, pl. 2, figs 5–7. Type locality:
Punta Engano, Mactan, Philippines, depth unknown.
Calliostoma paucicostatum — Vilvens, 2000: 88, figs
10-11; Poppe et al., 2006: 121–122, pl. 60, fig 3.
Material examined. Indonesia, Kai Islands.
KARUBAR: stn CP05, 05°49'S, 132°18'E, 296–299
m, 3 lv; stn CP09, 05°23'S, 132°29'E, 368–389 m, 1
dd; stn DW13, 05°26'S, 132°38'E, 417–425 m, 1 dd.
Philippines, Bohol Sea. PANGLAO 2004: stn T9,
09°34'N, 123°50–51'E, 97–120 m, 1 juv dd.
PANGLAO 2005: stn CP2350, 09°31'N, 124°01'E,
602–738 m; 1 dd, 1 juv dd; stn CP2358, 08°52'N,
123°37'E, 569–583 m, 1dd, 1 juv dd; stn CP2363,
09°06'N, 123°25'E, 437–439 m, 1 dd; stn CP2381,
08°43'N, 123°19'E, 259–280 m, 1 lv, 1 sub dd; stn
CP2384, 08°46'N, 123°16'E, 624–647 m, 1 lv; stn
CP2388, 09°27'N, 123°35'E, 762–786 m, 1 dd; stn
CP2389, 09°28'N, 123°38'E, 784–786 m, 3 dd; stn
CP2394, 09°29'N, 123°40'E, 470–566 m, 1 lv; stn
CP2397, 09°35'N, 123°42'E, 642–669 m, 2 dd.
Distribution. Philippines, 100–784 m, lv at 280–624
m (also using data of Poppe et al., 2006); Indonesia,
299–417 m, lv at 296–299 m; Taiwan, depth unknown
(Poppe et al., 2006).
Diagnosis. The main characteristics of this species are
a height up to 19.2 mm, a width up to 18.0 mm, an
elevated, conical to slightly coleonidal shape, an
angulate periphery, a teleoconch of 8 straight whorls,
at least 5 granular spiral cords (P2, P3; P1; S1, S3;
possible S2, Ti between P1-S1, S1-P2, P2-P3; P4)
with sharp pointed beads, S3 the strongest, making
keel, P3 making a secondary carina, a
subquadrangular aperture, a slightly oblique columella
without tooth, a weakly convex base with about 10
beaded spiral cords, anomphalous, a yellowish brown
colour.
Remarks. The Indonesian records here reported
extend considerably the distribution area towards the
South. This leaves however a large gap between the
two parts of this area.
Calliostoma suduirauti Bozzetti, 1997
Fig. 10J–Q
Calliostoma suduirauti Bozzetti, 1997: 43, figs 1–4.
Type locality: Balicasag Island, Bohol, Philippines,
140 m.
Calliostoma suduirauti —Vilvens, 2000: 88, figs 4–5;
Poppe et al., 2006: 124, pl. 65, figs 5–6.
C. VILVENS
Material examined. Solomon Islands.
SALOMONBOA 3: stn DW2827, 10°26'S, 161°51'E,
134–272 m, 1 lv.
Papua New Guinea, Solomon Sea. PAPUA
NIUGINI: stn CP4017, 05°39'S, 148°14'E, 280–315
m, 1 dd.
Papua New Guinea, New Ireland. BIOPAPUA: stn
CP3759, 04°00'S, 153°36'E, 287–352 m, 1 sub dd.
KAVIENG 2014: stn DW4505, 02°33'S, 150°47'E,
84–92 m, 1 dd.
Indonesia, Kai Islands. KARUBAR: stn DW18,
05°18'S, 133°01'E, 205–212 m, 1 dd; stn DW24,
05°32'S, 132°51'E, 230–243 m, 1 dd.
Distribution. Philippines, 140 m (range computed
using also data of Poppe et al., 2006); Solomon
Islands, 134–272 m; Papua New Guinea, 92–287 m
(dd); Indonesia, 212–230 m (dd).
Diagnosis. The main characteristics of this species are
a height up to 13.8 mm, a width up to 12.2 mm, an
elevated, conical to slightly coeloconoidal shape, an
angulate periphery, a teleoconch of 7 straight whorls,
at least 6 spiral cords (P1, P2, P3; S2; S1; T1 between
P2 and S2; possible T2 between P1-S1 et S1-P2; P4;
no S3), P3 the strongest with sharp nodules, S2
thinner also with smaller sharp nodules, a
subquadrangular aperture, a slightly oblique columella
without tooth, an almost flat base with up to 12
beaded spiral cords, a narrow umbilicus partially
covered by a columellar expansion, a light brown
colour with darker axial flames, P3 segmented into
alternate white and brown zones.
Remarks. Calliostoma paucicostatum Kosuge, 1984
and C. suduirauti Bozzetti, 1997, both described from
the Philippines, are clearly two close species. This was
already pointed out in the original description, with as
justification of the difference "a stronger peripheral
keel, brighter coloration and smaller size", and also by
Poppe (2006), without more argumentation. Based
upon the study of the available material of the two
species (especially their holotype), one can use mainly
as distinction criteria:
♦ a different ontogeny of the Si and Ti, with
respectively (S1, S3; possible S2, Ti between P1-S1,
S1-P2, P2-P3) for C. paucicostatum and (S2; S1; T1
between P2 and S2; possible T2 between P1-S1 et S1-
P2; P4; no S3) for C. suduirauti;
♦ the double keel on P3 and S3 especially visible
on penultimate whorls with S3 the strongest on the
last whorl for C. paucicostatum and a single very
strong keel on P3 for C. suduirauti.
On the other hand, the Indonesian, Papua New
Guinean and Solomon Sea samples here reported
extend surprisingly the distribution area of this species
towards the South and the Southeast of its original
locality (the Philippines). This leaves however some
large gaps between these areas.
NOVAPEX 25(2): 55–110, 10 juin 2024
Calliostoma ticaonicum (A. Adams, 1851)
Fig. 11A–H
Ziziphinus ticaonicus A. Adams, 1853: 167. Type
locality: Ticao Island, Masbate, Philippines, 11 m (6
fathoms).
Calliostoma ticaonicum — Kaicher, 1986: card TR1-
2158; Vilvens, 2000: figs 4–5; Poppe et al., 2006:
125, pl. 95, figs 2-4,7.
Material examined. Indonesia, Makassar strait.
CORINDON 2: stn CH294, 02°38'S, 117°50'E, 46–57
m, 3 lv, 3 sub lv.
Philippines. Fishermen, Balicasag Is., 07/2004, 2 sub
dd.
Distribution. Philippines, 25–140 m (also using data
of Poppe et al., 2006); Indonesia, Makassar straits,
46–57 m.
Diagnosis. The main characteristics of this species are
a height up to 17.0 mm, a width up to 15.0 mm, an
elevated, slightly cyrtoconoidal shape, an subangulate
periphery, a teleoconch of 8.5 convex whorls, up to 15
thin spiral cords (P1, P2, P3, P4; S1, S2, S3; Ti),
granular on the first whorls, subgranular on the next
whorls, P4 the strongest, a subelliptical aperture, a
arcuate columella without tooth, a moderately convex
base with about 20 beaded spiral cords, an open
narrow umbilicus, an ivory background colour with
brownish flames, first whorls possibly darker.
Remarks. This is a rather variable species, as pointed
out by Poppe et al. (2006), especially regarding the
height of the spire, the shape of the whorls and the
presence of not of a more or less marked keel on the
last whorl: the two Indonesian living specimens here
figured (Fig. 11A–D) render this variability very well.
Calliostoma poppei Vilvens, 2000
Fig. 11I–J
Calliostoma poppei Vilvens, 2000: 3–4, figs 1–3.
Type locality: Balicasag Island, Bohol, Philippines,
180 m.
Calliostoma poppei — Poppe et al., 2006: 122, pl. 63,
fig. 2.
Material examined. Philippines, Bohol. PANGLAO
2004: stn P1, 09°36'N, 123°45'E, 90–200 m, 1 lv.
Distribution. Philippines, 150–180 m (range
computed using also data of Poppe et al., 2006).
Diagnosis. The main characteristics of this species are
a height up to 15.0 mm, a width up to 12.0 mm, an
elevated, conical shape, an angulate periphery, a
teleoconch of up to 8 flat whorls, 6 spiral cords (P2,
P3, P1; P4; S1; S2), P3 the strongest with coarse thick
beads and making keel, P1 slightly thinner with
73
C. VILVENS New species and new records of Calliostomatidae. Part II

rounded beads, other cords subgranular to almost
smooth, a subquadrangular aperture, a vertical, arcuate
columella, an almost flat base with about 10 spiral
cords (innermost cords granular, outermost thinner
and almost smooth), a narrow umbilicus partly
covered by the expansion of the columella, a pink or
reddish-brown background colour with brownish
flammules or blotches, P3 and P4 with alternate white
and brown areas.
Calliostoma dedonderi Vilvens, 2000
Fig. 11K–O
Calliostoma dedonderi Vilvens, 2000: 87–88, figs 1–
3. Type locality: Philippine Islands, Bohol, Balicasag
Island, 140 m.
Calliostoma dedonderi — Poppe et al., 2006: 118, pl.
61, figs 1–4, 6.
Material examined. Philippines, Bohol. PANGLAO
2004: stn P1, 09°36'N, 123°45'E, 90–200 m, 1 juv dd;
stn L35, 09°36'N, 123°46'E, 90 m, 1 juv dd; stn L69–
73, 09°31'N, 123°41'E, 90–98 m, 5 juv lv; stn L79,
09°37'N, 123°46'E, 50 m, 3 dd.
Philippines, Aliguay Island. By fisherman: 140–160
m, 1 dd, 160 m, 1 dd.
Distribution. Philippines, 50–140 m, lv at 50 m
(range computed using also data of Poppe et al.,
2006).
Diagnosis. The main characteristics of this species are
a height up to 13.8 mm, a width up to 8.6 mm, an
elevated, conical shape, an angulate periphery, a
teleoconch of up to 8 flat to weakly concave (big
specimens) whorls, up to 7 spiral cords (P2, P3, P1;
S3; P4; S1; S2), P3 the strongest with sharp beads, one
bead for two to five on last whorls stronger and
sharper, a subquadrangular aperture, a weakly curved
and slightly oblique columella, a weakly convex to
almost flat base with 9-10 subgranular to nearly
smooth spiral cords, anomphalous, a nacreous white
colour with reddish-brow axial flammules, P3, S3, P4
and cords of the base with alternate regular orange
blotches.
Remarks. Since the original description of 2000,
greater specimens were found in Philippines and
reported by Poppe et al. (2006). One of the figured
specimens is unusually great (13.8 × 8.6 m) and have
about 8 whorls.
Calliostoma vilvensi Poppe, 2004
Fig. 12A–B
Calliostoma vilvensi Poppe, 2004: 6–8, pl. 2, figs 1a–
2b. Type locality: Philippines, offshore Aliguay
Island, depth unknown.
Calliostoma vilvensi — Poppe et al., 2006: 127, pl.
66, figs 1–2.
Material examined. Philippines, Bohol. PANGLAO
2004: stn L51-60, 09°38'N, 123°48'E, 62 m, 2 dd, 1
sub dd, 2 juv dd.
Distribution. Philippines, 50–150 m (range computed
using also data of Poppe et al., 2006).
Diagnosis. The main characteristics of this species are
a height up to 22.0 mm, a width up to 20.6 mm, a
moderately elevated, conical shape, an angulate
periphery, a teleoconch of up to 9 flat whorls, up to 7
spiral cords (P1, P2, P3; P4; S1; T1 between P1–S1;
S4), P1 the strongest, a subquadrangular aperture, a
weakly arcuate and slightly oblique columella, a
weakly convex to almost flat base with about 12
granular to subgranular spiral cords, anomphalous, a
yellowish brown colour with brownish streaks or
flames.
Calliostoma formosense E. A. Smith, 1907
Fig. 12G–N
Calliostoma formosensis E. A. Smith, 1907: 205, text
fig. Type locality: South Formosa, depth unknown.
Calliostoma (Calliostoma) formosensis — Higo et al.,
1999a, 61.
Calliostoma formosensis — Dong, 2002, 111, fig.
115.

Figure 10 (scale bars: 5 mm)

A–I. Calliostoma paucicostatum Kosuge, 1984. A–B. Holotype IMT-84-46, Philipines, Cebu, Mactan, Punta
Engano, 19.2 × 18.0 mm, 8 tw. C–G. Philippines, Bohol Sea, PANGLAO 2005. C–D. Stn CP2384, 624–647 m,
19.0 × 18.5 mm, 7.5 tw. E–G. Specimen with repairs, stn CP2389, 784–786 m, 16.2 × 15.1 mm, 7.1 tw. H–I.
Indonesia, Kai Islands, KARUBAR, stn CP05, 296–299 m, 12.0 × 11.8 mm, 6.9 tw.
J–Q. Calliostoma suduirauti Bozzetti, 1997. J–K. Holotype MNHN-IM-2000-30355, Philippines, Bohol,
Balicasag Island, 140 m, 14.1 × 12.4 mm, 7.0 tw. L–M. Solomon Islands, SALOMONBOA 3, stn DW2827,
134–272 m, 11.2 × 11.1 m, 7.1 tw. N–O. Papua New Guinea, Salomon Sea, PAPUA NIUGINI, stn CP4017,
280–315 m, 11.4 × 10.9 mm, 7.2 tw. P–Q. Indonesia, Kai Islands, KARUBAR, stn DW18, 205–212 m, 11.8 ×
11.0 mm, 7.3 tw.

74
C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

75
C. VILVENS
Material examined. Southern Taiwan. TAIWAN
2002; stn CP171, 22°14'N, 120°30'E, 195 m, 1 lv.
North-eastern Taiwan. TAIWAN 2000: off Tashi,
depth unknown (fishermen), 2 lv, 3 sub lv. TAIWAN
2001: stn CP75, 24°57'N, 122°02'E, 139 m, 1 juv dd;
stn CP79, 24°50'N, 122°00'E, 145–200 m, 1 lv, 8 sub
lv; stn CP80, 24°51'N, 121°59'E, 194–214 m, 5 lv, 2
sub lv, 2 juv lv; stn CP114, 24°51'N, 121°58'E, 128–
250 m, 4 lv, 5 sub lv, 6 juv lv. TAIWAN 2004: stn
CP243, 24°52'N, 121°55'E, 117–120 m, 5 lv; stn
CP244, 24°53'N, 121°56'E, 122–123 m, 14 lv, 1 juv
dd; stn CP249, 24°57'N, 122°05'E, 212–220 m, 1 lv.
Off Tashi, dredged, 5/2001, 3 dd.
Distribution. Taiwan, 50–300 m, lv at 120–212 m
(range computed using also Higo et al., 1999a);
southern Japan (Kyūshū), 190–210 m (Higo et al.,
1999a).
Diagnosis. The main characteristics of this species are
a height up to about 55 mm, a width up to about 55
mm, a moderately elevated, coeloconoidal shape, an
angulate periphery, a teleoconch of up to 9 whorls,
more or less flat except the last ones convex, up to 25
spiral cords (P1, P3; P4; P2; S1; S2; about 20 Ti
appearing by successive intercalations), P4 much
stronger on early and median whorls, but only a bit
stronger on the last whorls, a subquadrangular
aperture, an arcuate, oblique columella, a weakly
convex to almost flat base with up to about 35 thin
spiral cords, innermost granular and outermost
subgranular only, a large umbilicus, a light brow
colour with darker flames and patches.
Remarks. The protoconch of all the large specimens
studied here is systematically broken. But this
protoconch is safe on smaller specimens (as for
example the protoconch of 300 µm of a specimen
Figure 11 (scale bars: 5 mm)
A–H. Calliostoma ticaonicum (A.Adams, 1851). A–D. Indonesia, Makassar straits, CORINDON 2, stn CH294,
46–57 m. A–B. 15.8 × 14.2 mm, 8.0 tw. C–D. 14.3 × 14.3 mm, 7.6 tw. E–H. Philippines, Bohol, Balicasag
Island, by fishermen, 140 m, Vilvens coll.. E–F.
18.2 × 15.6 mm, 8.6 tw. G–H. 16.9 × 14.7 mm, 8.5 tw.
I–J. Calliostoma poppei Vilvens, 2000, Philippines, Bohol, PANGLAO 2004, stn P1, 90–200 m, 10.7 × 8.8 mm,
7.5 tw.
K–O. Calliostoma dedonderi Vilvens, 2000, Philippines, Bohol, PANGLAO 2004. K–N. stn L79, 50 m. K–L.
7.9 × 6.3 mm, 6.9 tw. M–N. 8.3 × 6.3 mm, 6.9 tw. O. (scale bar: 1 mm) Juvenils specimen, stn L69–73, 90–98
m, 2.3 × 1.8 mm, 4.0 tw.
76
New species and new records of Calliostomatidae. Part II
collected at stn CP79 of TAIWAN 2001, being 17.3 ×
18.4 mm for 7.9 tw).
Calliostoma achantodes n. sp.
Fig. 12C–E
urn:lsid:zoobank.org:act:85CB3F68-8162-49EE-
A301-02B876905D75
Type material. Holotype (12.0 × 10.7 mm) MNHN-
IM-2000-39424.
Type locality. South-western Taiwan, TAIWAN
2002, stn CP162, 22°10'N, 120°38'E, 190–200 m.
Material examined. South-western Taiwan.
TAIWAN 2002: stn CP162, 22°10'N, 120°38'E, 190–
200 m, 1 dd (holotype MNHN-IM-2000-39424).
Distribution. Only known from the type locality.
Diagnosis. A Calliostoma species of rather small size,
with a moderately elevated, conical spire, an angulate
periphery, a teleoconch of 6.9 whorls, 7 granular to
finally spiny spiral cords (P1, P2, P3; S3, P4; S2; S1),
P3 the strongest with sharp beads, making a moderate
keel, a flat base with 10 granular spiral cords, distance
between cords similar to width of cords, anomphalous,
a pinkish grey colour.
Description. Shell rather small for the genus (H 12.0
mm, W 10.7 mm), higher than wide, conical in shape;
spire rather elevated, 3.5× aperture height, apical
angle 60°; angulate periphery; anomphalous.
Protoconch about 420 μm wide (eroded on the single
type), of 1.25 whorls.
Teleoconch up to 6.9 straight whorls; early whorls
with 3 spiral cords, last whorl with 7 cords of uneven
width.
Suture weakly impressed, not canaliculated.
C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

77
C. VILVENS New species and new records of Calliostomatidae. Part II

First whorl weakly convex, sculptured by rather wide,
prosocline ribs and 3 granular spiral cords appearing
immediately and simultaneously; beads produced by
intersection of cords with ribs; distance between ribs
similar in size to width of ribs, distance between cords
smaller than their width. Second whorl flat, cords, ribs
and beads of cords stronger; distance between Pi
similar to their width; beads of P3 pointed beyond the
half of the whorl. On third whorl, P3 stronger than the
other cords with pointed, rather thick beads; S3
appearing, P4 partially appearing but covered by next
whorl. On fourth whorl, P4 sinking under suture, S2
appearing, S3 almost as strong as P3 with similar
pointed beads, axial sculpture vanishing. On fifth
whorl, beads of all the Pi and Si pointed; P3 making a
moderate keel. On last whorl, S1 appearing, very thin
and P4 emerging from suture.
Aperture subquadrate with a rounded outer lip.
Columella straight, slightly oblique.
Base almost flat, with 10 granular spiral cords,
innermost ones slightly stronger, beads produced by
intersection of cords with thin radial ridges, distance
between cords to width of cords.
Colour of teleoconch pinkish grey.
Discussion. Considering the keel on P3 on
Calliostoma achantodes n. sp., two other species seem
rather similar at first glance.
The new species is rather close to C. paucicostatum
Kosuge, 1984 described from the Philippines (Fig.
10A–I), but this slightly larger species has a slightly
coeloconoidal shape, thinner spiral cords with a
different ontogeny (especially with S1 and S3
appearing simultaneously) and a main keel on S3 with
P3 making a secondary carina.
C. achantodes n. sp. may also be compared to C.
suduirauti Bozzetti, 1997 from the Philippines to
Solomon Islands (Fig. 10J–Q), but this similar in size
species has thinner spiral cords except P3 with a
different ontogeny (especially with S1 appearing
much sooner and no S3).
The new species may also be compared to C. hungi S.-
I Huang & I.-F. Fu, 2019 from Taiwan Strait (Fig.
12F), but this smaller species has a less elevated spire
and compact rows of rounded beads with interspaces
much narrow than width of cords. The ontogeny
described in the original description of this species
(P1, P2, P3; S3; S2; S1) could be in fact (P1, P2, P3;
P4; S2; S1; no S3) because S3 could be likely P4.
Etymology. Thorny (Ancient Greek: άχανθώδης, -ες)
- with reference to the pointed beads on all the spiral
cords of the teleoconch.
Calliostoma zhongshaense n. sp.
Fig. 13A–F, Table 3
urn:lsid:zoobank.org:act:04D3128E-B145-45A6-
8C84-ACB5BB2941E2
Type material. Holotype (16.6 × 15.0 mm) MNHN-
IM-2013-59731. Paratypes: 2 MNHN (MNHN-IM-
2013-59733, MNHN-IM-2000-39425), 1 NMMBA
(as listed below).
Type locality. South China Sea, Macclesfield Bank,
ZHONGSHA 2015, stn CP4146, 16°09'N, 114°16'E,
232–314 m.
Material examined. South China Sea, Macclesfield
Bank. ZHONGSHA 2015: stn CP4146, 16°09'N,
114°16'E, 232–314 m, 3 lv (holotype MNHN-IM-
2013-59731, paratype IM-2013-59733, paratype
NMMBA NMMB-M011800), 2 sub lv (MNHN-IM-
2013-59732, MNHN-IM-2013-61909), 3 sub lv, 5 juv
lv; stn CP4148, 16°07'N, 114°19'E, 218–281 m, 1 lv;
stn CP4153, 16°14'N, 114°30'E, 318 m, 1 lv (paratype
MNHN-IM-2000-39425), 4 sub lv; stn CP4154,
16°15'N, 114°34'E, 321–326 m, 6 lv.
Distribution. South China Sea, Macclesfield Bank,
281–321 m (lv).

Figure 12 (scale bars: 5 mm)

A–B. Calliostoma vilvensi Poppe, 2004, Philippines, Bohol, PANGLAO 2004, stn L51–60, 62 m, 13.1 × 12.7
mm, 6.6 tw.
C–E. Calliostoma achantodes n. sp., holotype MNHN-IM-2000-39424, TAIWAN 2002, stn CP162, 190–200
m, 12.0 × 10.7 mm, 7.0 tw.
F. Calliostoma hungi S.-I Huang & I-F. Fu, 2019, holotype NMNS-007817-00004, Taiwan Strait, 6.8 × 6.7 mm,
6.8 tw (original description).
G–N. Calliostoma formosense E. A. Smith, 1907, Taiwan. G–H. Depressed form, TAIWAN 2004, stn CP244,
24°53'N, 122–123 m, 47.7 × 63.1 mm, 8.5 tw. I–J. More elevated form, TAIWAN 2000, by fishermen, 44.0 ×
54.1 mm, 8.8 tw. K–N. Subadult with elevated form, TAIWAN 2001, stn CP79, 145–200 m, 17.3 × 18.4 mm,
7.9 tw. N. (scale bar: 1 mm) Details of the protoconch and early whorls.

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C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

79
C. VILVENS
Diagnosis. A Calliostoma species of medium size,
with an elevated, conical spire, a weakly subangular
periphery, a teleoconch up to 7.5 whorls, 10 to 13
spiral cords (P1, P2, P3; S2; S3, S1; T1, T2; T3; 3
possible additional Ti; P4) with sharp beads, P3 the
strongest, making a strong keel, a weakly convex base
with 9–10 granular spiral cords, outermost cords
closer than inner ones, a narrow and deep umbilicus, a
pinkish white colour with light brown patches on
abapical cords.
Description. Shell of medium size for the genus (H up
to 17.2 mm, W to 15.2 mm), conical (or very weakly
coeloconoidal) in shape, slightly higher than wide
(ratio H/W of 1.1); spire rather elevated, 3.0× to 3.6×
aperture height, apical angle 60°; weakly subangulate
periphery; narrow and deep umbilicus.
Protoconch about 400 μm wide, 1.25 whorls, rounded,
with reticulate pattern and with a thin straight terminal
varix.
Teleoconch up to 7.5 slightly convex whorls, with a
strong abapical keel at the two thirds of the height
except on the two earliest whorls; early whorls with 3
spiral cords, last whorl with 10–13 spiral cords of
uneven width.
Suture weakly impressed, not canaliculated.
First whorl weakly convex, sculptured by about 20
low prosocline ribs and 3 granular spiral cords P1, P2
and P3 appearing immediately and simultaneously;
small beads produced by intersection of cords with
Table 3. Calliostoma zhongshaense n. sp.: Shells measurements in mm for types.
TW H W
Holotype ZHONGSHA 2015
CP4146 MNHN-IM-2013- 7.5
59731
Paratype 1 ZHONGSHA
2015 CP4146 MNHN-IM- 7.4
2013-59733
Paratype 2 ZHONGSHA
2015: CP4153 MNHN-IM- 7.5
2000-39425
Paratype 3 ZHONGSHA
2015 CP4146 NMMBA 7.3
NMMB-M011800
Discussion. Calliostoma zhongshaense n. sp. is rather
close to C. thachi Alf & Stratmann, 2007 from South
China Sea (off Central Viet Nam) (Fig. 13G–H), but
this slightly larger species with a similar ontogeny of
spiral cords has clearly a coeloconoidal shape with a
shape of the whorls concave under the keel, a less
elevated spire (ratio H/W of 1.0 instead of 1.1) and 2
keels on P3 and P4 (instead of a single one on P3 with
P4 thin on the new species).
The new species is also rather close to C. nanshaense
Z.-Z. Dong, 2002 from Nansha Islands (Fig. 13I–J),
but this species is smaller for more whorls (8), has a
coeloconoidal shape with concave whorls, spiral cords
80
New species and new records of Calliostomatidae. Part II
ribs; distance between ribs about 1.5× their width,
interspaces between cords 1.5× their width. On second
whorl cords, ribs and beads of cords stronger, P3 the
strongest, interspaces between cords 2× their width.
On third whorl, P3 making a moderate keel with beads
becoming blunt sharp. On fourth whorl, S2 appearing,
keel on P3 much stronger with beads strongly pointed;
beads of P2 blunt sharp. On fifth whorl, S3 appearing
close to P3, S1 appearing half a whorl later; beads of
all cords pointed; axial ribs vanishing, except under
S3. On seventh whorl, T1 appearing between P3 and
S3, T2 a bit later between P2 and S2. On last whorl,
T3 appearing between S2 and P3, possible additional
Ti between P1 and S1, S1 and P2 and above P1; P4
emerging, thin.
Aperture subquadrate with a rounded outer lip.
Columella straight, slightly oblique.
Base weakly convex, with 9–10 granular spiral cords,
innermost cords stronger and outermost cords closer
than inner ones; beads produced by intersection of
cords with thin radial ridges.
Umbilicus open on adult specimen (more than 7
whorls), narrow and deep, partially bordered by an
expansion of the columella.
Colour of teleoconch pinkish white, light brown
patches on P3, possibly also on S3 and P4; protoconch
white.
Operculum circular, multispiral, corneous, light
brown.
HA H/W H/HA
16.6 15.0 5.20 1.11 3.19
17.0 15.2 4.70 1.12 3.62
16.6 15.2 5.50 1.09 3.02
17.2 15.0 5.10 1.15 3.37
similar in size and a large open umbilicus with a spiny
spiral cord at rim.
One can compare the new species to C. thrincoma
Melvill & Standen, 1903 from the Gulf of Oman (Fig.
13K–L), but this species is smaller, has two keels (the
strongest one on P3, the other one, double, on S3 and
P4) and lacks an umbilicus.
Calliostoma zhongshaense n. sp. remembers a little
Tristichotrochus aculeatus (G. B. Sowerby III, 1912)
from Japan (Figs 14K–N, 15A–N, 16A–N), but this
species has 3 keels on P2, P3, P4 and lacks an
umbilicus.
C. VILVENS
The new species is slightly similar to Calliostoma
katoi (Sakurai, 1994) from Japan (Fig. 13M–N), but
this species is smaller for more whorls (8), has a
weakly coeloconoidal shape, a more elevated spire,
and the beads of the spiral cords are thinner and not
pointed.
Etymology. With reference to the type locality: the
Macclesfield Bank is the main part of what China
calls the Zhongsha Islands, grouping skerries, entirely
submerged banks, seamounts and shoals in the South
China Sea.
Calliostoma statheron n. sp.
Fig. 14A–B
urn:lsid:zoobank.org:act:0F95E94F-7B2C-424F-
9AB1-B693A3B49D52
Type material. Holotype (17.5 × 17.6 mm) MNHN-
IM-2013-44341.
Type locality. South China Sea, off An-Da Chiao,
NanHai 2014, stn DW4111, 10°25'N, 114°46'E, 462–
1039 m.
Material examined. South China Sea, off An-Da
Chiao. NanHai 2014: stn DW4111, 10°25'N,
114°46'E, 462–1039 m, 1 lv (holotype MNHN-IM-
2013-44341).
Distribution. Only known from the type locality.
Diagnosis. A Calliostoma species of medium size,
with a moderatly elevated, cyrtoconoidal spire, a
weakly subangular periphery, a teleoconch with 6.3
whorls with depressed suprastural area except on the
last one, 11 spiral cords (P1, P2, P3; S2; S1; S3, P4;
T1 between P1 and S1, T2 between S1 and P2; T3 P2
and S2; T4 between S2 and P3) with sharp beads, P3
the strongest), making a strong keel, a weakly convex
base with 15 granular spiral cords, outermost cords
closer and thinner than inner ones, no umbilicus, a
nacreous white colour.
Description. Shell of medium size for the genus (H =
17.5 mm, W = 17.6 mm), slightly cyrtoconoidal in
shape, as high as wide; spire moderately elevated,
2.7× aperture height, apical angle 70°; very weakly
subangulate periphery; anomphalous.
Protoconch of about 600 µm, almost completely
broken on the single available specimen.
Teleoconch up to 6.3 whorls, all more or less straight
except penultimate one slightly convex and last one
convex; early whorls with 3 spiral cords, last whorl
with 11 spiral cords of uneven thickness.
Suture weakly impressed, slightly canaliculated.
First whorl almost straight, sculptured by about 25
strong, very prosocline ribs and 3 granular spiral cords
P1, P2 and P3 appearing immediately and
simultaneously; small beads produced by intersection
NOVAPEX 25(2): 55–110, 10 juin 2024
of cords with ribs; distance between ribs about 1.5×
their width, interspaces between cords 2× their width;
depressed suprasutural area. On second whorl cords,
ribs and beads of cords much stronger, interspaces
between ribs 2× their width; S2 appearing. On third
whorl, beads of Pi becoming blunt sharp, S1
appearing. On fourth whorl, axial sculpture vanishing,
all 5 cords similar in size, interspaces between them
similar to their width. On fifth whorl, S3 appearing;
P4 slightly emerging from suture. On sixth whorl, T1
appearing between P1 and S1 simultaneously with T2
between S1 and P2; T3 appearing between P2 and S2
a bit later. On last whorl, T4 appearing between S2
and P3; P1 slightly stronger than other cords;
suprasutural area no more depressed, filled by S3
becoming stronger.
Aperture subrounded. Columella weakly arcuate,
slightly oblique.
Base weakly convex, with 15 spiral cords, innermost
cords strong with interspaces wider than their width,
outermost ones thin with interspaces narrower than
their width; beads produced by intersection of cords
with thin radial ridges.
Umbilicus completely closed, covered by an
expansion of the columella.
Colour of teleoconch nacreous white.
Operculum circular, multispiral, corneous, light
brown.
Discussion. Calliostoma statheron n. sp. seems at first
glance rather close to C. thaumaston n. sp. from
Indonesia and Papua New Guinea (Fig. 8I–L), but this
species, although similar in size, ontogeny of the
spiral cords and number of whorls, has a conical
shape, no depressed suprasutural area, a more angulate
periphery and spiral cords P2, S2 and P3 thicker.
C. statheron n. sp. seems also rather close to C.
shawni Poppe & Tagaro, 2020 from Japan to Malaysia
(Fig. 14C–D), but this species is much smaller for a
similar number of whorls (holotype: 7.0 × 6.4 with
about 6–6.5 tw), has a more elevated spire and only 7
spiral cords Pi and Si without Ti.
The new species may remember C. lui S.-I Huang & I-
F. Fu, 2019 from north-eastern Taiwan (Fig. 14E–F),
but this species is much smaller for a higher number
of whorls (holotype: 9.2 × 9.7 7.3 tw) and has strong
keels on P2, P3 and P4.
One can compare the new species to C. basulense
Poppe, Tagaro & Vilvens, 2014 from the Philippines
(Fig. 14G–H), but this smaller species (11.5–14.6 ×
11.0-12.6 mm for about 7 tw) has a much more
elevated spire with P3 stronger and making a weak
keel on the median whorls.
One can finally also compare the new species to C.
sakashitai (Sakurai, 1994) from Japan and East China
Sea (Fig. 8F), but this species is smaller for a larger
number of whorls, a much more elevated spire (with a
larger H/W ratio) and an ontogeny of cords different,
especially regarding S3 and S1.
81
C. VILVENS New species and new records of Calliostomatidae. Part II

Etymology. Stable, steady, firm (Ancient Greek
σταθερός, -ά, - όν) - with reference to the similar
height and width of the shell making a steady shape.
Genus Tristichotrochus Ikebe, 1942
Type species: Calliostoma aculeatum G. B. Sowerby
III, 1912 (by o.d.) – Recent, Japan.
Grammatical gender: masculine.
Tristichotrochus aculeatus (G. B. Sowerby III, 1912)
Figs 14K–N, 15A–N, 16A–N
Calliostoma aculeatum G.B.Sowerby III, 1912: p.
473, text-fig. 3. Type locality: Japan.
Calliostoma (Tristichotrochus) aculeatus — Ikebe,
1942: 257–258, pl. XXVII, figs 1–3.
Tristichotrochus aculeatus — Kira, 1962: 11, pl. 8,
fig. 12; Kuroda et al., 1971: 23–24, pl. 10, figs 11–12.
Calliostoma aculeatum — Kaicher, 1986: card TR1-
2078; Higo et al., 1999a, b: 61; Sasaki, 2000: 71–72,
fig. 99; Dong, 2002, 102, fig. 103; Qi Zhongyan,
2004: 20, pl. 009, fig. A.
Calliostoma aculeatum aliguayensis — Poppe et al.,
2006: 115, pl. 60, figs 1–2. Syn. nov.
Material examined. Indonesia, Kai and Tanimbar
Islands. KARUBAR: stn DW02, 05°47'S, 132°13'E,
209–240 m, 1 dd, 1 sub dd, 1 juv dd; stn CP05,
05°49'S, 132°18'E, 296–299 m, 2 lv; stn DW15,
05°17'S, 132°41'E, 212–221 m, 1 dd, 1 juv dd; stn
CP16, 05°17'S, 132°50'E, 315–349 m, 1 dd; stn CP69,
08°42'S, 131°53'E, 356–368 m, 3 dd, 1 sub dd; stn
CP78, 09°06'S, 131°24'E, 284–295 m, 1 lv; stn CP77,
08°57'S, 131°27'E, 346–352 m, 1 lv; stn CP84,
09°23'S, 131°09'E, 246–275 m, 3 lv.
North east of Taiwan. TAIWAN 2000: stn CP58,
24°33'N, 122°06'E, 221–254 m, 1 dd; stn DW36,
21°54'N, 120°36'E, 300–331 m, 1 sub dd. TAIWAN
2001: stn CP79, 24°50'N, 122°00'E, 145–200 m, 1 lv;
stn CP114, 24°51'N, 121°58'E, 128–250 m, 1 dd.
TAIWAN 2004: stn CP247, 24°52'N, 122°02'E, 487–
540 m, 1 dd.
East China Sea, North east of Taiwan. KAVALAN
2018: stn CP4173, 25°29’N, 122°23’E, 261–330 m, 1
dd; stn CP4175, 25°25’N, 122°28’E, 434–535 m, 1
dd.
South west of Taiwan. TAIWAN 2000: stn CP11,
22°16'N, 119°14'E, 263–276 m, 1 dd; stn DW34,
22°01'N, 120°36'E, 240–246 m, 1 dd; stn DW36,
21°54'N, 120°36'E, 300–331 m, 2 dd, 1 sub dd.
North of the South China Sea. ZHONGSHA 2015:
stn CP4135, 19°59'N, 114°38'E, 211–218 m, 2 sub dd.
Philippines, Bohol Sea. PANGLAO 2004: stn P1,
09°36'N, 123°45'E, 90–200 m, 1 sub lv, 2 juv dd; stn
P3, 09°31'N, 123°42'E, 100 m, 1 lv. PANGLAO 2005:
stn DW2339, 09°32'N, 123°44'E, 164–176 m, 1 dd;
stn CP2340, 09°29'N, 123°44'E, 271–318 m, 3 lv, 2
sub dd; stn CP2343, 09°27'N, 123°49'E, 273–356 m, 9
lv, I juv dd; stn CP2344, 09°28'N, 123°50'E, 128–142
m, 2 lv; stn CP2349, 09°32'N, 123°56'E, 219–240 m,
1 lv; stn CP2360, 08°49'N, 123°38'E, 357–372 m, 1
lv; stn CP2361, 08°53'N, 123°34'E, 516–543 m, 1 dd;
stn CP2362, 08°57'N, 123°33'E, 679–740 m, 1 dd; stn
CP2381, 08°43'N, 123°19'E, 259–280 m, 2 dd; stn
CP2395, 09°36'N, 123°44'E, 382–434 m, 1 dd, 1 juv
lv; stn CP2407, 09°41'N, 123°49'E, 256–268 m, 2 dd;
stn CP2409, 09°45'N, 123°45'E, 220–257 m, 1dd.
Distribution. Japan, East China sea and Korea, 50–
100 m (Higo et al., 1999a); Philippines (as subspecies
aliguayensis), 100–679 m, lv at 100–382 m (range
computed using also data of Poppe et al., 2006);
Indonesia, 221–356 m, lv at 275–346 m; Taiwan,
200–487 m, lv at 145–200 m.
Diagnosis. The main characteristics of this species are
a height up to 18.3 mm, a width up to 16.7 mm, an
elevated, conical to slightly cyrtoconoidal shape, an
angulate periphery, a teleoconch of up to 9 weakly
convex whorls, up to 11 granular spiral cords (P2, P3,
P1; P4; S2; S1, S3; T1 between P2 and S2 the earliest
of the Ti; other Ti possible between P1 and S1, S1 and
P2, S2 and P3, P3 and S3; ) with pointed beads, P3 the
strongest, P2 and P4 almost as strong as P3, the three
cords making keel, a subquadrangular aperture, an
oblique columella without tooth, a slightly convex
base with up to 13 subgranular spiral cords,
anomphalous, an ivory to hazel background colour
with brownish flames.

Figure 13 (scale bars: 5 mm)

A–F. Calliostoma zhongshaense n. sp., South China Sea, Macclesfield Bank, ZHONGSHA 2015. A–D. Stn
CP4146, 232–314 m. A–B. Holotype MNHN-IM-2013-59731, 16.6 × 15.0 mm, 7.5 tw. C–D. Paratype MNHN-
IM-2013-59733, 17.0 × 15.2 mm, 7.5 tw. E–F. Paratype MNHN-IM-2000-39425, stn CP4153, 318–318 m, 16.6
× 15.2 mm, 7.5 tw.
G–H. Calliostoma thachi Alf & Stratmann, 2007, South China Sea, Central Viet Nam, off Nha Trang (from the
original description). G. Holotype SMF 328943, 16.7 × 16.9 mm, 8 tw. H. Paratype D.Stratmann coll., 19.8 ×
20.2 mm.
I–J. Calliostoma nanshaense Z.-Z. Dong, 2002, drawings from the original description Nansha Islands, 14.0 ×
14.0 mm, 8 tw.
K–L. Calliostoma thrincoma Melvill & Standen, 1903, syntype NHMUK 1903.12.15.122, Gulf of Oman, 10.0
× 8.8 mm.
M–N. Calliostoma katoi (Sakurai, 1994), holotype NSMT, Japan, 13.1 × 11.2 mm, 8 tw.
82
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83
C. VILVENS New species and new records of Calliostomatidae. Part II

Remarks. The subspecies Tristichotrochus aculeatus
aliguayensis was described by Poppe et al. from the
Philippines (2006), the differences pointed out being
"its much thicker and heavier shell, the more
pronounced sculpture and in general its lower spire" -
likely a rather light criteria to separate the nominal
species and the supposed subspecies, the both ones
sharing particularly the same ontogeny of spiral cords
on the teleoconch. One can just add that the original
description of C. aculeatus aliguayensis points out an
open umbilicus, while the specimens from Japan are
usually anomphalous.
Qi Zhongyan (2004) had already illustrated a
specimen of C. aculeatum that has a moderately (not
rather) elevated spire. But the new material here
reported from Indonesia, East China Sea and Taiwan
bring specimens
♦ with moderately (KARUBAR DW15 and CP77,
TAIWAN 2001 CP79 and CP114, Kii (Japan)
specimen) to rather elevated spire (KARUBAR CP84
and CP69, KAVALAN 2018 CP4175, TAIWAN 2000
CP58, Minabe (Japan) specimens)
♦ with closed (TAIWAN 2001 CP79, KARUBAR
DW15, CP84 and CP77, Minabe (Japan) and Kii
(Japan) specimens) to partially or even almost
completely open umbilicus (TAIWAN 2001 CP114,
KAVALAN 2018 CP4175, KARUBAR CP69),
Most of the specimens from the Philippines have
indeed a moderately elevated spire and a (usually
partially) open umbilicus. PANGLAO 2005, stn
CP2395 sample brings a large specimen with an open
umbilicus (Fig. 16G–H) and a much smaller specimen
with a closed umbilicus (Fig. 16I–J). There is also a
completely anomphalous specimen in the material
recorded by Poppe et al. in their original description
(PANGLAO 2005, stn CP2380 - Fig. 16E–F).
Finally, it then seems likely that C. aculeatus is a
widespread species with variations regarding the
height of the spire (moderately elevated to rather
elevated) and the umbilicus (closed, partially closed or
open), without subspecies.
Additional remark. The close species Calliostoma
soyoae Ikebe, 1942 from Japan (Fig. 14I–J) is rather
easy to distinguish at a glance from C. aculeatus by its
more convex whorls and a much thinner sculpture
with weaker keels.
Genus Fautor Iredale, 1924
Type species: Ziziphinus comptus A. Adams, 1855 (by
o.d.) – Recent, southern Australia.
Grammatical gender: masculine
Remarks. This genus, of which the types species is
the Australian Fautor comptus (A. Adams, 1855) (Fig.
17M–N), was early considered as a subgenus of the
genus Calliostoma. B. A. Marshall (1995) has
characterized it by having narrower and more finely
beaded spiral cords, opposed to those of Calliostoma.
One can add to this feature the presence of axial ribs
on the early whorls whereas this sculpture is usually
very weak or absent on Calliostoma species (B. A.
Marshall, 1995) and also a rather typical more or less
conical shape. There are by now 21 accepted Fautor
species (WoRMS, 2023). But other species has been
described as belonging to the subgenus Fautor and
should be added to this list: Calliostoma (Fautor)
aprosceptum Vilvens, 2009, C. (F.) diaphoros
Vilvens, 2009.
Fautor paradigmatus (B. A. Marshall, 1995)
Fig. 17A–L
Calliostoma (Fautor) paradigmatum B. A. Marshall,
1995: 395–397, figs 13–15, 119, 155. Type locality:
southern New Caledonia, 505–550 m.
Calliostoma (Fautor) paradigmatum — Vilvens,
2005: 2; Vilvens, 2009: 132, figs 25–26; Vilvens,
2012: 18, figs 51–54.
Calliostoma paradigmatum — Vilvens, 2014a: 44,
figs 26–27; Vilvens, 2014b: figs 55–56.
Fautor paradigmatus — Vilvens, 2023: 54–55, figs
6A–H.

Figure 14 (scale bars: 5 mm)

A–B. Calliostoma statheron n. sp., holotype MNHN- IM-2013-44341, South China Sea, off An-Da Chiao,
NanHai 2014, stn DW4111, 462–1039 m, 17.5 × 17.6 mm, 6.3 tw.
C–D. Calliostoma shawni Poppe & Tagaro, 2020, holotype MNHN, Japan, Okinawa Island, 12–13 m, 7.0 × 6.4
mm, about 6.5 tw (from the original description).
E–F. Calliostoma lui S.-I Huang & I-F. Fu, 2019, holotype NMNS-007817-00003, north-eastern Taiwan,
"shallow water", 9.2 × 9.8 mm, 7.3 tw (from the original description).
G–H. Calliostoma basulense Poppe, Tagaro & Vilvens, 2014, holotype MNHN-IM-2012-18895, Philippines,
Mindanao, Surigao, Basul Island, 14.6 × 12.5 mm, about 7 tw (from the original description).
I–J. Calliostoma soyoae Ikebe, 1942, Nada, pref. Wakayama, Japan, 14.2 × 13.7 mm, 7.2 tw, Vilvens coll..
K–N. Tristichotrochus aculeatus (G. B. Sowerby III, 1912). K–L. Japan, Minabe,
17.8 × 16.0 mm, 7.6 tw. M–N. Japan, Kii, 36 m, 17.6 × 17.2 mm, 7.5 tw.

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85
C. VILVENS New species and new records of Calliostomatidae. Part II

Material examined. Solomon Islands.
SALOMONBOA 3: stn CP2823, 10°27'S, 162°02'E,
240–572 m, 1 juv dd.
Papua New Guinea. PAPUA NIUGINI: stn
DW4076, 04°01'S, 144°55'E, 860 m, 1 juv dd.
Indonesia, Kai Islands. KARUBAR: stn CP09,
05°23'S, 132°29'E, 368–389 m, 1 lv; stn DW13,
05°26'S, 132°38'E, 417–425 m, 2 sub dd, 2 juv dd; stn
CP20, 05°15'S, 132°59'E, 769–809 m, 1 lv.
Philippines, Bohol. PANGLAO 2004: stn P1,
09°36'N, 123°45'E, 90–200 m, 1 lv juv.
PANGLAO 2005: stn CP2361, 08°53'N, 123°34'E,
516–543 m, 1 juv dd.
Taiwan. TAIWAN 2000: stn DW37, 21°52'N,
120°35'E, 420–503 m, 1 dd; stn FP16, 22°18'N,
119°15'E, 351–360 m, 1 juv dd.
South China Sea. ZHONGSHA 2015: stn DW4161,
20°33'N, 116°39'E, 538–560 m, 1 dd.
Distribution. Southern New Caledonia, 410–795 m,
lv at 550–795 m (B. A. Marshall, 1995; Vilvens,
2009; Vilvens, 2023); northern New Caledonia, 585–
622 m, lv at 585 mm (B. A. Marshall, 1995; Vilvens,
2023); West of New Caledonia, 300–900 m, lv at
900–950 m (Vilvens, 2023); Solomon Islands, 240–
572 m (dd); Papua New Guinea, 860 m (dd);
Indonesia, Kai Islands, 389–769 m (lv); Philippines,
Bohol, 200–516 m (lv); Taiwan, 360–420 m(dd);
South China Sea, 538–560 m (dd); Tonga, 342–500 m
(dd) (Vilvens, 2005); Austral Archipelago, 570–620 m
(dd) (Vilvens, 2014a); Tarava Seamounts, 660–670 m
(dd) (Vilvens, 2012).
Diagnosis. The main characteristics of this species are
a height up to 13.9 mm, a width up to 11.4 mm, an
elevated, conical shape, a subangulate to slightly
rounded periphery, a teleoconch of up to 7.6 weakly
convex whorls, at least 6 granular spiral cords (P2, P3;
P1; P4; S2; S1; S3 possibly missing or weak if
present; possible Ti between Pi and Si), a depressed
suprasutural area above the impressed suture, S3 if
present and P4 thinner than the other cords, a
subelliptical aperture, an oblique columella without
tooth, a weakly convex base with 12-17 subgranular
spiral cords, anomphalous, a nacreous white colour.
Remarks. It's a wonder to record here this originally
New Caledonian species so far from its known range,
although a few specimens were already recorded in
the Entrecasteaux Reefs in the north east of New
Caledonia. More overall, it's also amazing that this
species has been recorded along a so large area, from
Tonga and even French Polynesia to South China Sea
and southern Taiwan. But the ontogeny and the shape
of the spiral cords, coupled with the peculiar shape of
the whorls with a depressed suprasutural area, of all
the specimens studied match perfectly those of the
holotype from New Caledonia. More far to the North,
Calliostoma uranipponense (Okutani, 1969) from
Japan (Honshu) (Fig. 18I–J) is rather close: it has
similar spiral cords on the teleoconch but without S3,
a larger suprasutural area, all cords except P1 and S1
thinner and almost smooth.
Fautor cosmopolites n. sp.
Fig. 18A–H, Table 4
urn:lsid:zoobank.org:act:9D9FC92E-E390-4C05-
9D2B-923FDD2F9F88
Type material. Holotype (8.8 × 7.7 mm) MNHN-
IM-2000-39426. Paratypes: 3 MNHN (MNHN-IM-
2000-39427, MNHN-IM-2000-39428, MNHN-IM-
2000-39429 (as listed below).
Type locality. Indonesia, Kai Islands, KARUBAR,
stn CP16, 05°17'S, 132°50'E, 315–349 m.
Material examined. Papua New Guinea, Salomon
Sea. MADEEP: stn DW4268, 05°33'S, 153°59'E,
383–720 m, 1 dd (paratype 1 MNHN-IM-2000-
39427).
Papua New Guinea, seamounts near Bougainville.
BIOPAPUA: stn DW3748, 05°37'S, 154°01'E, 398–
399 m, 1 dd (paratype 2 MNHN-IM-2000-39428).
Papua New Guinea, Bismarck Sea, South-east of
Vokeo Island. PAPUA NIUGINI: stn DW4074,
03°16'S, 144°05'E, 460–605 m, 1 juv dd; stn CP4079,
04°34'S, 145°52'E, 960 m, 1 juv lv.
Indonesia, Kai Islands. KARUBAR: stn CP16,
05°17'S, 132°50'E, 315–349 m, 1 lv (holotype
MNHN-IM-2000-39426).

Figure 15 (scale bar: 5 mm)

A–N. Tristichotrochus aculeatus (G. B. Sowerby III, 1912). A–B. Indonesia, Kai Islands, KARUBAR, stn
DW15, 212–221 m, 14.9 × 15.7, 6.8 tw. C–D. Indonesia, Tanimbar Islands, KARUBAR, stn CP77, 346–352 m,
17.0 × 17.0 mm, 6.9 tw. E–F. Specimen without Si, North east of Taiwan, TAIWAN 2000, stn CP58, stn CP79,
145–200 m, 13.6 × 13.5 mm, 6.5 tw. G–H. Indonesia, Tanimbar Islands, KARUBAR, stn CP84, 246–275 m,
13.7 × 12.6 mm, 7.0 tw. I–J. Specimen with umbilicus partially covered, East China Sea, North east of Taiwan,
KAVALAN 2018, stn CP4175, 434–535 m, 14.9 × 14.1 mm, 7.1 tw. K–L. North east of Taiwan, TAIWAN
2000, stn CP58, 221–254 m, 15.9 × 15.4 mm, 7.0 tw. M–N. Specimen with umbilicus partially covered, north
east of Taiwan, TAIWAN 2001, stn CP114, 128–250 m, 14.5 × 15.5 mm, 5.9 tw.

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C. VILVENS New species and new records of Calliostomatidae. Part II

Southern Taiwan. TAIWAN 2000: stn DW5,
22°41'N, 119°56'E, 213–236 m, 4 dd (including
paratype 3 MNHN-IM-2000-39429); stn DW36,
21°54'N, 120°36'E, 300–331 m, 1 dd.
North-eastern Taiwan. TAIWAN 2001: stn CP74,
24°51'N, 121°59'E, 220 m, 1 dd .
Distribution. Papua New Guinea, 399–960 m, lv at
960 m; Indonesia, Kai Islands, 315–349 m (lv);
Taiwan, 220–300 m (dd).
Diagnosis. A Fautor species of rather small size, with
a rather elevated, conical spire, a subangulate to
slightly angulate periphery, a teleoconch of up to 6.4
flat to weakly convex whorls, at least 7 granular spiral
cords (P1, P2, P3; S2; S3; S1; P4; possible Ti), a
nearly flat to weakly convex base with 10-12 spiral
cords, innermost ones granular, outermost ones
thinner and subgranular, anomphalous, a yellowish to
whitish ivory colour.
their width; S2 appearing; axial sculpture vanishing.
On fourth whorl, P1 the strongest with strong beads
axially elongated on some specimens (Indonesia); S3
appearing neat the end of the whorl. On fifth whorl,
S1 appearing; S3 still rather thin, depressed
suprasutural area still visible. Additional Ti may
appear on some specimens (especially those collected
near Taiwan): T1 between P2 and S2, T2 between S2
and P3, T3 between S1 and P2. On last whorl, P4
peripheral, thin.
Aperture subquadrate with rounded angles. Columella
straight, slightly oblique.
Base nearly flat to weakly convex, with 10–12 spiral
cords, inner cords granular with distance between
them similar to their width, outer cords thinner and
subgranular with distance between them slightly
smaller than their width.
Colour of teleoconch yellowish to whitish ivory;
protoconch white.
Operculum circular, multispiral, corneous, brown.

Description. Shell rather small for the genus (H up to
10.6 mm, W up to 8.7 mm), slightly higher than wide,
conical in shape; spire rather elevated, height 1.1× to
1.2× width, 3.0× to 3.5× aperture height, apical angle
65°; subangulate to slightly angulate periphery;
anomphalous.
Protoconch of 360–400 μm wide, 1.25 whorls,
rounded, with tiny spits, without terminal varix.
Teleoconch up to 6.4 flat to weakly convex whorls;
early whorls with 3 spiral cords, last whorl with 7
cords.
Suture impressed, not canaliculated.
First whorl weakly convex, sculptured by about 20
moderately thick prosocline ribs and 3 granular spiral
cords P1, P2, P3 appearing almost immediately and
simultaneously; beads produced by intersection of
cords with ribs; P1 slightly thinner than the other
cords; distance between ribs similar in size to width of
ribs, distance between cords about 2× their width. On
second whorl, flat beads of cords stronger; P1, P2 and
P3 becoming similar in size; axial ribs more
prosocline, distance between them 1.5× their width;
P3 making a moderate keel, producing a depressed
area above the suture. On third whorl, beads of cords
strong and separate; distance between cords similar to
Discussion. The new species is rather close to Fautor
paradigmatus (B. A. Marshall, 1995) widespread in
the Indo-Pacific (Fig. 17A–L), but is different mainly
by having S1 and S3 cords appearing systematically in
the opposite order (S1 first on F. paradigmatum, S3
first on F. cosmopolites); moreover, S3 is always
present on the new species, producing a much less
depressed suprasutural area.
F. cosmopolites n. sp. remembers also Calliostoma
diaphoros Vilvens, 2009 from Solomon Islands (Fig.
19A–B), but this similar in size species has a S3
appearing the first (before the other Si) with spiral
cords different in size, P1 being much stronger than
the other spirals cords that are relatively thin.
The new species may also be compared to C.
emmanueli Vilvens, 2000 from the Philippines (Fig.
9K–R), but this species lacks the suprasutural
depressed area with P3 making a moderate keel and
has nearly smooth to weakly subgranular flat spiral
cords on the base, instead of (sub) granular ones.
Etymology. Cosmopolitan (Ancient Greek:
χόσμουπολίτης, -ου) - with reference to the large
distribution area of the new species (from Papua New
Guinea to north-eastern Taiwan).

Figure 16 (scale bar: 5 mm)

A–D. Tristichotrochus aculeatus (G. B. Sowerby III, 1912). A–B. Indonesia, Tanimbar Islands, KARUBAR,
stn CP69, 356–368 m, 19.2 × 17.7 mm, 7.2 tw. C–D. Southern Philippines, Davao, Talikud Island, 13.7 × 13.8
mm, 6.5 tw.
E–N. Tristichotrochus aculeatus aliguayensis (Poppe, Tagaro & H. Dekker, 2006), Philippines, Bohol Sea,
PANGLAO 2005, determination by Poppe, Tagaro & H. Dekker, 2006. E–F. Off Balicasag IslanStn CP2380,
150–163 m, 14.9 × 14.4 mm, 6.9 tw. G–J. Maribojoc Bay, stn CP2395, 382–434 m. G–H. 19.0 × 17.9 mm, 7.1
tw. I–J. 10.4 × 10.0 mm, 5.9 tw. K–L. Off Pamilacan Island, stn CP2343, 273–356 m, 14.2 × 14.4 mm, 6.9 tw.
M–N. Off Balicasag Island, stn CP2340, 271–318 m, 18.3 × 18.0 mm, 7.3 tw.

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C. VILVENS New species and new records of Calliostomatidae. Part II

Table 4. Fautor cosmopolites n. sp.: Shells measurements in mm for types.

TW H W HA H/W H/HA
Holotype KARUBAR CP16
6.2 10.6 8.7 3.1 1.22 3.42
MNHN-IM-2000-39426
Paratype 1 MADEEP
DW4268 MNHN-IM-2000- 6.1 9.1 8 3 1.14 3.03
39427
Paratype 2 BIOPAPUA
DW3748 MNHN-IM-2000- 6.2 8.1 7.3 2.7 1.11 3.00
39428
Paratype 3 TAIWAN2000
DW5 MNHN-IM-2000- 6.4 9.5 8.4 2.7 1.13 3.52
39429

Fautor panteles n. sp. base with 8 weakly subgranular to granular spiral

Fig. 18K–O, Table 5 cords with distance smaller than cords, anomphalous,
urn:lsid:zoobank.org:act:E35C17E9-2329-4B4F- a white background colour with spiral cords orange
B035-7280D5556BA8 except S3 white.

Type material. Holotype (9.9 × 8.3 mm) MNHN-
IM-2000-39430. Paratype: (MNHN-IM-2000-39431,
as listed below).
Type locality. Papua New Guinea, off New Ireland,
BIOPAPUA, stn DW3745, 369–377 m.
Material examined. Papua New Guinea, off New
Ireland. BIOPAPUA: stn DW3745, 05°33'S,
154°00'E, 369–377 m, 1 dd (holotype MNHN-IM-
2000-39430).
Solomon Islands, Sealark channel. SALOMONBOA
3: stn DW2860, 09°20'S, 160°13'E, 328–342 m, 1 sub
dd (paratype MNHN-IM-2000-39431), 1 juv dd.
Indonesia, Tanimbar Islands. KARUBAR: stn
DW50, 07°59'S, 133°02'E, 184–186 m, 1 juv dd.
Distribution. Papua New Guinea, 369–377 m (dd);
Solomon Islands, 328–342 m (dd); Indonesia,
Tanimbar Islands, 184–186 m (dd).
Diagnosis. A Calliostoma species of rather small size,
with a rather elevated, conical spire, 7 granular spiral
cords (P1, P2, P3; S2, S3; S1, P4), S3 the strongest
with blunt sharp beads and making keel, a almost flat
Description. Shell rather small for the genus (H up to
9.5 mm, W up to 8.3 mm), higher than wide, conical
in shape; spire rather elevated, height 1.2× width, 3.7×
to 3.8× aperture height, apical angle 60°; angulate
periphery; anomphalous.
Protoconch about 450 μm wide, 1.25 whorls, rounded,
with a reticulate pattern of tiny spaces, without
terminal varix.
Teleoconch up to 6.5 whorls; early whorls weakly
convex with 3 spiral cords, next whorls flat, last whorl
with 7 granular spiral cords.
Suture poorly visible, neither impressed nor
canaliculated.
First whorl weakly convex, sculptured by about 15
rather weak prosocline ribs and 3 granular spiral cords
P1, P2, P3 appearing almost immediately and
simultaneously; beads produced by intersection of
cords with ribs; P1 very thinner than the other cords;
distance between ribs about 1.5× to 2× their width,
distance between cords more or less the same as their
width. On second whorl flat, P3 much stronger than
the other Pi, with blunt sharp beads; P1 becoming
similar in size to P2 only near the end of the whorl.

Figure 17 (scale bars: 5 mm)

A–L. Fautor paradigmatus (B.A.Marshall, 1995). A–D. Indonesia, Kai Islands, KARUBAR. A–B. Stn CP20,
769–809 m, 12.5 × 11.6 mm, 6.1 tw. C–D. Stn CP09, 368–389 m, 10.1 × 9.0 mm, 6.2 tw. E–F. Philippines,
Bohol, PANGLAO 2004, stn P1, 90–200 m, 7.5 × 6.4 mm, 5.4 tw. G–H. Southern Taiwan, Bashi channel,
TAIWAN 2000, stn DW37, 420–503 m, 10.9 × 9.8 mm, 6.9 tw. I–J. Holotype MNHN-IM-2000-31531, New
Caledonia, MUSORSTOM 4, stn DW220, 505–550 m, 11.4 × 10.0 mm, 6.7 tw, goldened for SEM. K–L.
MNHN-IM-2013-65483, West of New Caledonia, KANADEEP 1, stn CP5055, 900–950 m, 13.9 × 11.2 mm, 7.2
tw.
M–N. Fautor comptus (A. Adams, 1855), lectotype NHMUK, 11.1 × 7.9 mm.

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C. VILVENS New species and new records of Calliostomatidae. Part II

Near the middle of the third whorl, S2 and S3
appearing, quickly as strong as P1; P3 the strongest
with strong pointed beads and making keel. On fourth
whorl, S3 with thick beads and stronger than the other
cord except P3 still the strongest. On fifth whorl, S3
definitively the strongest with blunt sharp beads and
making keel; P3 similar in size to P1, other cords
slightly thinner. On last whorl, S1 appearing, much
thinner than the other cords; P4 emerging from suture,
as thin as S1; distance between cords (except S1)
about 1.5× their width.
Aperture subquadrate. Columella straight, slightly
oblique.
Base almost flat, with 8 spiral cords, innermost ones
stronger and granular, outermost ones subgranular to
nearly smooth; distance between cords slightly smaller
than their width.
Colour of teleoconch white, with spiral cords orange
except S3 white; protoconch white.

Table 5. Fautor panteles n. sp.: Shells measurements in mm for types.

TW H W HA H/W H/HA
Holotype BIOPAPUA
DW3745 MNHN-IM-2000- 6.5 9.9 8.3 2.6 1.19 3.81
39430
Paratype SALOMONBOA3
DW2860 MNHN -IM-2000- 6 8.5 7.2 2.3 1.18 3.70
39431

Discussion. Fautor panteles n. sp. is close to
Calliostoma quadricolor Schepman, 1908 from
Indonesia (Fig. 7A–D) but this similar in size species
has a less elevated spire (with a ratio H/W of 1.0–1.1
instead of 1.2 for the new species), a different
ontogeny of spiral cords with S2 absent (instead of
present) and a smaller H/TW ratio (1.4–1.5 instead of
1.4–1.5).
The new species is comparable to Calliostoma simplex
Schepman, 1908 from Indonesia (Fig. 6I–M) but this
species has all its spiral cords similar in size, whereas
S3 is significantly the strongest making keel on the
new species with P1 and P3 stronger than the other
cords.
The new species may be compared to C. soloensis van
Regteren Altena, 1938 from Java (Pliocene) (Fig.
19C–D) but this much larger species (20 × 19 mm, 10
tw) has a double peripheral keel (apparently on S3 and
P4, although the accurate description of the ontogeny
of cords can't be found in the original description)
instead of a single keel on S3.
Fautor panteles n. sp. slightly remembers C.
maekawai Poppe, Tagaro & Goto, 2018 from the
Philippines (Fig. 8O), but this slightly larger species
has a weakly coeloconoidal shape and much thinner
spiral cords (the ontogeny of these cords is not
provided in the original description) without an
obvious suprasutural keel on an abapical cord.
Remarks. Again, one can wonder about the large
distribution of this species, in the same way as Fautor
paradigmatus or Calliostoma emmanueli.
Etymology. Perfect (Greek: παντελής, -ές) - with
reference to the perfectly conical shape of the shell.

Figure 18 (scale bar: 5 mm)

A–H. Fautor cosmopolites n. sp.. A–B. Holotype MNHN-IM-2000-39426, Indonesia, Kai Islands, KARUBAR,
stn CP16, 315–349 m, 8.8 × 7.7 mm, 6.2 tw. C–D. Paratype 1 MNHN-IM-2000-39427, Papua New Guinea,
Salomon Sea, MADEEP, stn DW4268, 383–720 m, 9.1 × 8.0 mm, 6.1 tw. E–F. Paratype 2 MNHN-IM-2000-
39428, Papua New Guinea, BIOPAPUA, stn DW3748, 398–399 m, 8.1 × 7.3 mm, 6.2 tw. G–H. Paratype 3
MNHN-IM-2000-39429, Southern Taiwan, TAIWAN 2000, stn DW5, 213–236 m, 9.5 × 8.4 mm, 6.4 tw.
I–J. Calliostoma uranipponense (Okutani, 1969), holotype NSMT-Mo 69544, off Akita, North-east of Honshu,
10.1 × 8.8 mm,
K–O. Fautor panteles n. sp. K–M. Holotype MNHN-IM-2000-39430, Papua New Guinea, off New Ireland,
BIOPAPUA, stn DW3745, 369–377 m, 9.9 × 8.3 mm, 6.5 tw. N–O. Paratype MNHN-IM-2000-39431, Solomon
Islands, Sealark channel, SALOMONBOA 3, stn DW2860, 328–342 m, 8.5 × 7.2 mm, 6.0 tw.

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C. VILVENS
Genus Akoya Habe, 1961
Type species: Calliostoma akoya Kuroda, 1942 (by
o.d.) – Recent, Japan.
Grammatical gender: feminine.
Diagnosis. Based upon the type species Akoya akoya
(Kuroda, 1942), the features of the genus are a rather
elevated spire, an angulate periphery, numerous thin
spiral cords finely beaded on the early whorls and
nearly smooth except the most adapical ones on the
last whorls, a weakly convex base with thin, low spiral
cord, a subquadrate aperture, a nearly closed
umbilicus covered more or less completely by the
expansion of the columella.
Akoya haliarchus (Melvill, 1889)
Fig. 19G–L
Zizyphinus haliarchus Melvill, 1889: 32–33, pl. 2, fig.
3. Type locality: Australia, depth unknown.
Tristichotrochus haliarchus — Kira, 1961, 12, pl.8,
fig. 20; Kuroda, 1971, 22, pl. 10, fig. 15.
Calliostoma (Calliostoma) haliarchus — Higo et al.,
1999a, 61.
Calliostoma haliarchus — Sasaki, 2000, 73, fig. 97;
Dong, 2002, 108–109, fig. 102; Poppe et al., 2006:
119, pl. 62, fig 4.
Material examined. Solomon Islands. SALOMON
2: stn CP2187, 08°17'S, 160°00'E, 482–604 m, 1 dd.
Philippines, Bohol Sea. PANGLAO 2005: stn
CP2368, 08°56'N, 123°17'E, 318–322 m, 1 dd.
Taiwan. Southern Taiwan. TAIWAN 2000: stn
DW36, 21°54'N, 120°36'E, 300–331 m, 1 dd.
East of Taiwan. TAIWAN 2000: stn DW60, 24°40'N,
122°12'E, 418–532 m, 1 sub dd.
Distribution. Japan, 50–200 m (Kuroda et al., 1971);
Philippines, 150–318 m (range computed also using
data from Poppe, 2006); Taiwan, East China Sea,
100–418 m (range computed also using data from
Dong, 2002); Solomon Islands, 482–604 m (dd).
Diagnosis. The main characteristics of this species are
a height up to 50 mm, a width up to 50 mm, a
moderately elevated, conical shape, an angulate
periphery, a teleoconch of up about 9 straight whorls,
more or less 20 spiral cords (P1, P2, P3; P4; S2; S1,
P4; T1 between P2 and S2 the earliest of the Ti; T2
between S1 and P2, T3 between S2 and P3; other
numerous Ti by intercalation between the existing
cords), all granular on earliest whorls, adapical cords
granular and abapical cords subgranular on the last
whorls, all very thin except P4 much stronger, a
subquadrangular aperture, a arcuate columella without
tooth, a weakly convex base with at least 30 thin,
smooth spiral cords, anomphalous, a beige
background colour with light orange brown flames
and P4 white with regular brown wide dots.
94
New species and new records of Calliostomatidae. Part II
Remarks. The specimens here studied are very
crumbly and hardly broken. On the specimen from
Solomon Islands, the first and the last whorls are
missing, while the remaining whorls have huge holes.
On the specimen of Philippines, the first and the last
whorls are also missing while the teleoconch surface
is so encrusted that it is very difficult to distinguish
the numerous and thin spiral cords.
As already pointed out by Poppe et al. (2006), this
species has been found in Japan, Taiwan area and
Shanghai area. It seems very scarce in the Philippines
area.
Genus changing. Considering the diagnosis of the
genus Akoya and the one of Tristichotrochus
haliarchus (Melvill, 1889), it seems likely that this
species should be placed into the genus Akoya. The
name Akoya haliarchus (Melvill, 1889) new comb. is
therefore proposed for it.
Akoya zhui n. sp.
Fig. 19E–F, Table 6
urn:lsid:zoobank.org:act:93241ADD-A576-4E4F-
8E68-4C09498D3E6A
Type material. Holotype (17.8 × 17.8 mm) MNHN-
IM-2000-39432. Paratype: 1 MNHN (MNHN-IM-
2000-39433, as listed below).
Type locality. South west of Taiwan, TAIWAN 2000,
stn DW34, 22°01'N, 120°36'E, 246–240 m.
Material examined. South west of Taiwan.
TAIWAN 2000: stn DW34, 22°01'N, 120°36'E, 240–
246 m, 1 dd (holotype MNHN-IM-2000-39432 ); stn
DW36, 21°54'N, 120°36'E, 300–331 m, 1 sub dd
(paratype MNHN-IM-2000-39433 ).
East of Taiwan. TAIWAN 2000: stn DW60, 24°40'N,
122°12'E, 418–532 m, 1 juv dd.
Distribution. South west of Taiwan, 246–300 m (dd);
East of Taiwan, 418–532 m (dd).
Diagnosis. A calliostomatid species of medium size,
with a rather elevated, conical spire, an angulate
periphery, a teleoconch of up about 7.6 straight
whorls, 16–18 thin, smooth spiral cords (P1, P2, P3;
S2; P4, S1, P4; T1 between P2 and S2; T2 between S1
and P2, T3 between S2 and P3; other numerous Ti by
intercalation between the existing cords), all very thin
and similar in size except P4 slightly stronger, a
subquadrangular aperture, a straight columella without
tooth, a weakly convex base with about 25 thin,
smooth spiral cords, anomphalous, a whitish beige
background colour with one spiral cord in two light
brown, P4 uniformly beige.
C. VILVENS
Description. Shell of medium size for the family (H
up 17.8 mm, W to 17.8 mm), as high as wide, conical
in shape; spire rather elevated, height 1.0× to 1.3×
width, 2.8× to 2.9× aperture height, apical angle 65°;
angulate periphery; anomphalous.
Protoconch about 350–360 μm wide, 1.25 whorls,
rounded, with a reticulate pattern of tiny spaces,
without terminal varix.
Teleoconch up to 7.6 whorls; two first whorls weakly
convex with 3 spiral cords, next whorls flat, last whorl
with 16–18 smooth spiral cords.
Suture poorly visible, impressed, not canaliculated.
First whorl weakly convex, sculptured by about 20
thin prosocline ribs and 3 granular spiral cords
appearing almost immediately and simultaneously;
beads produced by intersection of cords with ribs; P1
thinner than the two other cords; distance between ribs
similar to their width, distance between cords more or
less the same as their width. On second whorl flat, P1
almost as strong as the other Pi, S2 appearing near the
end of the whorl. On third whorl, P4 emerging from
Table 6. Akoya zhui n. sp.: Shells measurements in mm for types.
TW H W
Holotype TAIWAN 2000
DW34 MNHN-IM-2000- 7.7 17.8
39432
Paratype TAIWAN 2000
DW36 MNHN-IM-2000- 6.7 11.6
39433
Discussion. Akoya zhui n. sp. is clearly rather close to
Akoya haliarchus (Melvill, 1889) originally described
from Japan (Fig. 19G–L), but this species, similar in
shape and with a close ontogeny of spiral cords, is
very much larger for a similar number of whorls, has a
smaller protoconch (about 500 µm instead of about
350 µm) and has a peripheral spiral cord P4 much
stronger than the other cords, a white colour with
regular brown wide dots (for the n. sp., P4 is only very
weakly stronger than the other cords and has the same
colour as the other ones).
The new species may be compared to Akoya akoya
(Kuroda, 1942) from Japan (Fig. 19O–P), but this
larger species (25 × 29, mm, 8 tw) has much more
convex whorls, a different ontogeny of spiral cords
(P1, P2, P3; S2; S1; T1 between P1 and S1; T2
between P2 and S2; T3 between S1 and P2; P4;
numerous additional Ti) and adapical spiral cords of
the last whorls granular, not smooth.
Considering the straight whorls and the numerous thin
spiral cords, the new species may be compared to
Calliostoma connyae Poppe, Tagaro & Vilvens, 2014
from the Philippines (Fig. 19M–N), but this much
larger species (34.5 × 29.5 mm, 8 tw) has a much
more elevated spire for a similar number of whrols,
granular (not smooth) spiral cords on the last whorls
and two suprasutural peripheral cords making
moderate keels.
NOVAPEX 25(2): 55–110, 10 juin 2024
suture, smooth, partially covered by next whorl; S1
appearing; axial sculpture vanishing; no S3. On fourth
whorl, P1 the strongest cord; P4 completely visible;
T1 appearing between S2 and P2 and a bit later T2
and T3 respectively between S1 and P2, P2 and P3.
On next whorls, other numerous Ti appearing by
intercalation between the existing cords; abapical
cords becoming smooth, adapical cords smooth about
two whorls later; all cords thin and similar in width
except P4 weakly stronger. On last whorl, at least 16
thin smooth spiral cords except P4 peripheral slightly
stronger; distance between cords less or equal to their
width.
Aperture subquadrate. Columella more or less straight,
slightly oblique.
Base weakly convex, with 25 very thin, smooth spiral
cords; distance between cords greater or equal to their
width.
Colour of teleoconch whitish beige, one in two spiral
cord light brown, P4 uniformly beige; protoconch
white.
HA H/W H/HA
17.8 6.3 1.00 2.83
9.1 4.0 1.27 2.90
Etymology. Conical (Chinese: 锥 or錐 - Zhuī) - with
reference to the perfect conical shape of the shell.
Genus: Bathyfautor Marshall, 1995
Type species: Bathyfautor rapuhia B.A. Marshall,
1995 (by o.d.) – Recent, New Zealand.
Grammatical gender: masculine
Bathyfautor multispinosus (Schepman, 1908)
Fig. 20A–B
Calliostoma multispinosum Schepman, 1908: 64-65,
pl. V, fig. 7. Type locality: Indonesia, Makassar Strait,
00°32'S, 119°40'E, 655 m.
Bathyfautor multispinosus — B. A. Marshall, 1995:
420.
Calliostoma multispinosum — Poppe et al., 2006:
121, pl. 60, fig. 5.
Material examined. Indonesia, Tanimbar Islands.
KARUBAR: stn CP39, 07°47'S, 132°26'E, 466–477
m, 3 lv; stn CC56, 08°16'S, 131°59'E, 549–552 m, 4
lv; stn CP72, 08°36'S, 131°33'E, 676–699 m, 3 dd, 4
sub dd, 1 juv dd; stn CP75, 08°46'S, 131°36'E, 451–
452 m, 3 dd, 2 sub dd.
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C. VILVENS New species and new records of Calliostomatidae. Part II

Distribution. Indonesia, 550–650 m (range computed
using also data of Schepman, 1908), lv at 477–549 m;
Philippines, 552–592 m (Poppe et al., 2006).
Diagnosis. The main characteristics of this species are
a height up to 21.8 mm, a width up to 21.0 mm, an
elevated, conical to slightly coeloconoidal shape, an
angulate periphery, a teleoconch of up to 8.5 more or
less straight whorls, 11 spiral cords (P1, P2, P3; P4;
S2, S3; S1; T1, T2, T3 respectively between P1 and
S1, P2 and S2, S2 and P3; S4), beads of P1 sharp,
beads of S3 sharp and finally fused into a strong keel,
a subquadrangular aperture, an oblique columella
without tooth, a moderately convex base with a large
median smooth area bordered by 4 smooth external
and 4 strongly beaded internal spiral cords,
anomphalous, an ecru to off-white colour.
Genus: Venustatrochus Powell, 1951
Type species: Venustatrochus georgianus Powell,
1951 (by o.d.) – Recent, Antarctica.
Grammatical gender: masculine
Venustatrochus malaita (Vilvens, 2009)
Fig. 20C–N
Calliostoma (Benthastelena) malaita Vilvens, 2009:
146–147, figs C1–C2, 87–90. Type locality: Solomon
Islands, Indispensable Strait between Santa Isabel and
Malaita, SALOMON 2, stn CP2186, 08°17'S,
160°00'E, 487–541 m.
Venustatrochus malaita — B. A. Marshall, 2016: 123,
figs 2E–F.
Material examined. Solomon Islands. SALOMON
2: stn CP2184, 08°17'S, 160°00'E, 464–523 m, 1 lv
(MNHN-IM-2007-18521).
Papua New Guinea, Solomon Sea. MADEEP: stn
DW4305, 10°46'S, 151°10'E, 666–680 m, 1 dd.
Papua New Guinea, New Ireland. KAVIENG 2014:
stn CP4420, 02°24'S, 150°36'E, 425–442 m, 1 sub dd;
stn CP4422, 02°21'S, 150°38'E, 496–609 m, 4 dd; stn
CP4439, 02°23'S, 150°35'E, 534–650 m, 1 dd.
BIOPAPUA: stn CP3653, 02°13'S, 150°23'E, 680–
700 m, 1 sub lv.
West of New Caledonia, Lansdowne. EBISCO: stn
CP2623, 20°06'S, 160°19'E, 691–886 m, 1 dd.
South of New Caledonia, Banc Athos. NORFOLK
2: stn DW2064, 25°17'S, 168°56'E, 609–691 m, 1 lv
(MNHN-IM-2007-18334).
North-eastern Taiwan. TAIWAN 2004: stn CP248,
24°52'N, 122°02'E, 516–557 m, 1 sub dd.
Distribution. Solomon Islands, 487–523 m, lv at 523
m (range computed using also data of Vilvens, 2009);
Papua New Guinea, 442–680 m, lv at 680–700 m;
West of New Caledonia, Lansdowne, 609–691 m (lv);
South of New Caledonia, Athos & Porthos Bank, 691–
795 m, lv at 609–691 m (range using also data of
Vilvens, 2009); North-eastern Taiwan, 516–557 m
(dd).
Diagnosis. The main characteristics of this species are
a height up to 20.7 mm, a width up to 22.7 mm, a
rather elevated, conical shape, an angulate periphery, a
teleoconch of up to 6.3 convex whorls with a median
shoulder, up to 12 beaded spiral cords (P1, P2, P3; P4;
S1, S2; T1 between P2 and S2, possible T2 between
S1 and P2, P1 and S1; possible additional Ti), P2 the
strongest making a moderate keel on the median
shoulder, a subquadrangular aperture, a slightly
oblique columella without tooth, a moderately base
with about 20 smooth spiral cords; 10 innermost cords
slightly thicker than 10 outermost cords,
anomphalous, a nacreous white colour.

Figure 19 (scale bars: 5 mm)

A–B. Calliostoma diaphoros Vilvens, 2009, holotype MNHN-IM-2000-22076, Solomon Islands, New Georgia
Sound, off Santa "isabel, SALOMON 2, stn CP2201, 307–310 m,
11.2 × 9.1 mm, 7.2 tw.
C–D. Calliostoma soloensis van Regteren Altena, 1938, Java (Pliocene), 20 × 19 mm, 10 tw, pictures of the
original description.
E–F. Akoya zhui n. sp., south west of Taiwan, holotype MNHN-IM-2000-39432, TAIWAN 2000, stn DW34,
246–240 m, 17.8 × 17.8 mm, 7.7 tw.
G–L. Akoya haliarchus (Melvill, 1889). G–H. Solomon Islands, SALOMON 2, stn CP2187, 482–604 m, 32.5 ×
28.4 mm, 5 tw. I–J. Holotype MM MANCH FF 7778, Australia, 40 × 36 (in the original description), 39 × 40
mm (estimated from the photograph). K–L. Japan, 90 m, 42.7 × 41.6 mm, 8.8 tw, Vilvens coll..
M–N. Calliostoma connyae Poppe, Tagaro & Vilvens, 2014, holotype, Philippines, Mindanao, Balut Islands,
34.4 × 29.5 mm, 8 tw.
O–P. Akoya akoya (Kuroda, 1942), Japan, 17.2 × 18.6 mm, 6.5 tw, Vilvens coll.

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97
C. VILVENS New species and new records of Calliostomatidae. Part II

Remarks. These new records enlarge significantly the
distribution area of this species. It appears clearly that,
whereas the number of specimens in the various
stations of the collects is small and that this species is
found only in deep waters (range of 523–691 m), this
species is widespread in the western Indo-Pacific area.
One can also observe some variations among the
various specimens. Larger specimens have a more
elevated spire. The only specific feature of the
specimens of Papua New Guinea is the tertiary cord
between P1 and S1, appearing more or less at the
same time as the T2 between S1 and P2. The
specimen from Lansdowne (EBISCO stn CP2623, east
of New Caledonia) has less numerous spiral cords on
the base, but this is the only difference with the
specimens from Solomon Islands. Finally, the
specimen from Taiwan is a bit more atypical regarding
his smaller size for a number of whorls (5.9) almost
similar to the one of the other specimens.
Subfamily FAUTRICINAE B.A. Marshall, 1995
Genus Selastele B.A. Marshall, 1995
Type species: Calliostoma onustum Odhner (by o.d.) –
Recent, northern New Zealand.
Grammatical gender: neuter.
Selastele solomonense n. sp.
Fig. 21F–I
urn:lsid:zoobank.org:act:3174AD82-4930-4F04-
9C21-F45C6382D17C
Type material. Holotype (6.2 × 5.2 mm) MNHN-IM-
2000-39434 Paratype : 2 MNHN (MNHN-IM-2000-
39435, MNHN -IM-2000-39436, as listed below).
Type locality. Solomon Islands, New Georgia,
SALOMON 2, stn CP2219, 07°58'S, 157°34'E, 650–
836 m.
Material examined. Solomon Islands, New Georgia.
SALOMON 2: stn CP2219, 07°58'S, 157°34'E, 650–
836 m, 1 dd (holotype MNHN-IM-2000-39434), 1 sub
dd.
Solomon Islands, Vela Gulf. SALOMON 2: stn
CP2263, 07°55'S, 156°51'E, 485–520 m, 1 dd.
Papua New Guinea, Bismack Sea. PAPUA
NIUGINI: stn DW4031, 04°53'S, 145°49'E, 370 m, 1
dd (paratype 1 MNHN-IM-2000-39435).
Papua New Guinea, Solomon Sea. MADEEP: stn
DW4327, 08°17'S, 149°45'E, 666–695 m, 1 dd
(paratype 2 MNHN-IM-2000-39436).
Distribution. Solomon Islands, 520–650 m (dd);
Papua New Guinea, 370–666 m (dd).
Diagnosis. A Selastele species of rather large size,
with an elevated, conical spire, 7 granular spiral cords
(P1, P2, P3; S3; P4; S2; S1), all cords with beads
separate and more or less pointed, P3, S3 and P1 the
strongest, an almost flat base with 8 spiral cords of
which intermediate ones possibly indistinct, no
umbilicus, a yellowish white colour.
Description. Shell of large size for the genus (H up to
7.5 mm, W up to 5.5 mm), higher than wide, conical
in shape; spire elevated, height 1.2× to 1.4× width,
2.5× to 3.9× aperture height, apical angle about 60°;
angulate periphery; anomphalous.
Protoconch about 360–450 μm wide, 1.25 whorls,
rounded, sculptured by a network of hexagonal spaces
clearly visible, without terminal varix.
Teleoconch of 5.9 whorls, flat except the first one
convex; first whorl with 3 spiral cords, last whorl with
7 cords.
Suture impressed, not canaliculated.
First whorl convex, sculptured by 3 spiral cords with
rounded beads; P1, P2 and P3 appearing immediately,
P3 the strongest, P1 much thinner than the two other
ones; prosocline threads very close on the first half of
whorl, distance between threads becoming 2× their
size on the second half; threads connecting beads of
cords. On second whorl, S3 emerging from suture,
almost completely covered by next whorl; P2 slightly
thinner than P1; distance between threads at least 3×
their width; beads of cords largely separate, becoming
more or less sharp. S2 appearing near the end of
second whorl (paratype 1) to the beginning of fourth
whorl (paratype 2); S1 appearing half a whorl to one
whorl later. On third whorl, S3 fully visible; P4
partially emerging from suture; axial threads still
present and rather strong with interspaces 3× their
width. On next whorls, P3, S3 and P1 the strongest
cords; axial threads weakening and widening. On last
whorl, P4 completely visible, peripheral, thinner than
other cords.

Figure 20 (scale bars: 5 mm)

A–B. Bathyfautor multispinosus (Schepman, 1908), Indonesia, Tanimbar Islands, KARUBAR, stn CC56, 549–
552 m, 21.8 × 21.0 mm, 8.6 tw.
C–N. Venustatrochus malaita (Vilvens, 2009). C–D. MNHN-IM-2007-18521, Solomon Islands, SALOMON 2,
stn CP2184, 464–523 m, 21.2 × 23.3 mm, 6.2 tw. E–F. Papua New Guinea, New Ireland, KAVIENG 2014, stn
CP4439, 534–650 m, 23.7 × 18.4 mm, 6.5 tw. G–H. Papua New Guinea, Solomon Sea, MADEEP, stn DW4305,
666–680 m, 28.2 × 26.9 mm, 6.6 tw. I–J. West of New Caledonia, Lansdowne, EBISCO, stn CP2623, 691–886
m, 18.8 × 20.8 mm, 6.1 tw. K–L. South of New Caledonia, Banc Athos, NORFOLK 2, stn DW2064, 25°17'S,
168°56'E, 609–691 m, 23.5 × 23.6 mm, 6.5 tw. M–N. North-eastern Taiwan, TAIWAN 2004, stn CP248, 516–
557 m, 12.5 × 13.1 mm, 5.9 tw.
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99
C. VILVENS
Aperture subquadrate. Columella slightly curved,
vertical.
Base almost flat, with about smooth to subgranular 8
spiral cords, distance between them similar to their
size; innermost cord the strongest, penultimate
outermost cord stronger than all other cords, cords in
the middle area possibly weak to almost completely
eroded; no umbilicus.
Colour of teleoconch and base yellowish white.
Discussion. Among the known Selastele species, the
new species is only rather close to Selastele pictum B.
A. Marshall, 1995 from the Loyalty Islands (Fig. 21J),
but this similar in size species has a very peculiar
ontogeny of cords with S1 absent and P2 vanishing
and replaced par two Ti.
Remarks. There are some variations among the
available specimens of Selastele solomonense n. sp.,
regarding
♦ the shape of beads of the cords on the whorls:
almost rounded and blunt sharp (paratype 1) to prickly
pointed (paratype 2)
♦ the spiral cords on the base that can be
subgranular (paratype 1) to completely smooth
(holotype, paratype 2), even completely indistinct in
the middle area (holotype, paratype 2)
♦ as already mentioned, the position of the
beginning of the Si.
Etymology. From the Solomon area - with reference
to the Solomon Islands and the Solomon Sea (and
neighbouring Bismarck Sea) where the specimens
have been collected.
Genus Phenacomargarites B. A. Marshall, 2016
Type species: Phenacomargarites williamsae B. A.
Marshall, 2016 (by o.d.) – Recent, Solomon Islands,
SW of Santa Isobel, 8°24′S, 159°22′E, 897–1057 m.
Grammatical gender: masculine.
Phenacomargarites tetratropeis n. sp.
Fig. 21A–B
urn:lsid:zoobank.org:act:B53A804B-60C4-4354-
928A-F848E79993D0
Type material. Holotype (17.2 × 18.1 mm) MNHN-
IM-2000-39437.
Type locality. Solomon Islands, SALOMON 1, stn
CP1807, 09°42'S, 160°53'E, 1077–1135 m.
Material examined. Solomon Islands. SALOMON
1: stn CP1807, 09°42'S, 160°53'E, 1077–1135 m, 1 dd
(holotype MNHN-IM-2000-39437).
Distribution. Only known from the type locality.
Diagnosis. A Phenacomargarites species of rather
small size, with a moderately elevated, cyrtoconoidal
100
New species and new records of Calliostomatidae. Part II
spire, a rounded periphery, 19 spiral cords (P1, P3, P4;
P2; S4; T1, T2; T3; T4, T5; other Ti) and 4 keels on
P3, P4, S4 and P5, a convex base with 25 smooth,
rather thin, close spiral cords, a very narrow umbilicus
partially covered by an expansion of the columella, a
white colour.
Description. Shell rather small for the genus (H of
17.2 mm, W of 18.1 mm), wider than high,
cyrtoconoidal in shape; spire moderately elevated,
height 0.95× width, 1.8× aperture height, apical angle
90°; rounded periphery; narrow umbilicus.
Protoconch about 440 μm wide, 1.2 whorls, rounded,
with a reticulate pattern, without terminal varix.
Teleoconch of 5.0 convex whorls; early whorls with 5
spiral cords, last whorl with 19 spiral cords.
Suture weakly impressed, not canaliculated.
First whorl convex, sculptured by 3 smooth spiral
cords P1, P3 and P4 appearing immediately and
simultaneously; P2 appearing after the half of the
whorl; P5 appearing near the end of the whorl, almost
completely covered by suture with the next whorl;
distance between cords about 2× their width, except
P1 and P2 closer. On second whorl, P4 the strongest,
P3 stronger than the other cords, both making weak
keels; weak, flat prosocline ribs appearing after the
half of the whorl between cords, stronger in the area
between suture and P3 and making P1, P2 and P3
weakly subgranular, beads produced by intersection of
cords with ribs; interspaces between ribs about 2×
their width; S4 appearing near the end of the whorl,
subgranular. On third whorl, P3 and P4 making
stronger keel, S4 stronger; surface between suture and
P3 with a reticulate pattern; T1 and T2 appearing near
the end of the whorl, respectively between suture and
P1, P1 and P2. On fourth whorl, T3 appearing
between P3 and P4, T4 a bit later between P2 and P3,
T5 between P4 and P5; S4 making an additional keel.
On last whorl, P5 emerging from suture; new
additional thin spiral cords appearing on the abapical
half, giving a total number of cords of 19; P2 and P3
still making strong keels, S4 and P5 making two
slightly weaker keels.
Aperture subcircular. Columella slightly arcuate,
vertical.
Base convex, with 25 smooth, rather thin, close spiral
cords.
Umbilicus very narrow (about 5% of the width of the
base), with almost vertical wall, partially covered by
an expansion of the columella.
Colour of teleoconch and protoconch white.
Discussion. Phenacomargarites tetratropeis n. sp.
slightly remembers P. incomptus B. A. Marshall, 2016
from Chatham Rise (East of Chatham Islands, New
Zealand) (Fig. 21E), but this larger species (28.9 ×
33.0 mm, 5.3 tw) is rather different, having only a
single angulation on P4.
In the same geographical area of Solomon Islands,
P. williamsae B. A. Marshall, 2016 (Fig. 21C–D) is
C. VILVENS
slightly larger than tne new species (20.2 × 22.1 mm,
5.3 tw) with only two keels and a weak angulation.
Etymology. Carinated with four keels (Ancient greek:
τρόπίς, -εως (carina) with τετρα (four times) as prefix)
- with reference to the four carinas on the last whorl of
the shell.
Subfamily THYSANODONTINAE B. A. Marshall,
1988
Genus Thysanodonta B. A.Marshall, 1988
Type species: Thysanodonta aucklandica B. A.
Marshall, 1988 (by o.d..) – Recent, New Zealand.
Grammatical gender: feminine.
Thysanodonta boucheti B. A. Marshall, 1988
Fig. 21K–L
Thysanodonta boucheti B. A. Marshall, 1988: 219–
220, figs 3 G–I. Type locality: northern New
Caledonia, 300–350 m.
Thysanodonta boucheti — B. A. Marshall, 1995: 434,
fig. 168; Vilvens & Maestrati, 2006: 3, figs 1–2;
Vilvens, 2023: 72, fig. 13E–F.
Material examined. Solomon Islands.
SALOMONBOA 3: stn CP2823, 10°27'S, 162°02'E,
240–572 m, 1 sub dd.
Distribution. North-west of New Caledonia, 350–613
m, lv at 502–516 m; east of New Caledonia, 435–510
m (dd); west of New Caledonia, 443–569 m (dd);
southern New Caledonia, 365–685 m, lv at 361–365 m
(all using data of New Caledonia - see Vilvens, 2023);
Wallis et Futuna, 640–730 m (dd) (Vilvens &
Maestrati, 2006); Solomon Islands, 240–572 m (dd).
Diagnosis. The main characteristics of this species are
a height up to 8.9 mm, a width up to 6.9 mm, an
elevated, conical shape, an angulate periphery, a
teleoconch of up to 6.0 whorls flat except the last one
convex, 5 granular spiral cords on the last whorl (4 on
penultimate whorl) (P3; P1, P2; S3, P4; neither S1 nor
S2), all broad except P4 very thin and only visible on
the last whorl, a subquadrate aperture, a vertical
columella without tooth, a convex base with 7–8
subgranular to granular (innermost one only), rather
strong rounded spiral cords, no umbilicus, a white
nacreous colour.
Thysanodonta eucosmia B. A. Marshall, 1995
Fig. 21M–P
Thysanodonta eucosmia B. A. Marshall, 1995:437–
439, figs 113–115, 148–149, 158. Type locality:
southern New Caledonia, 675–680 m.
Thysanodonta eucosmia — Vilvens & Maestrati,
2006: 3, figs 3–4; Vilvens, 2023: 72–73, fig. 13A–D.
NOVAPEX 25(2): 55–110, 10 juin 2024
Material examined. Solomon Islands.
SALOMONBOA 3: stn CP2823, 10°27'S, 162°02'E,
240–572 m, 1 juv dd.
Distribution. Southern New Caledonia to Norfolk
Ridge, 360–805 m, lv at 365–680 m (range computed
using also data of B. A. Marshall, 1995; Vilvens &
Maestrati, 2006); Solomon Islands, 240–572 m (dd).
Diagnosis. The main characteristics of this species are
a height up to 12.0 mm, a width up to 9.0 mm, an very
elevated, conical shape, a rounded periphery, a
teleoconch of up to 7.0 whorls flat except the last ones
convex, 9–11 thin, granular spiral cords on the last
whorls (P3, P4; P1, P2, S1, S2; S3; up to 4 Ti on the 4
last whorls); P3 stronger and flange-like on first
whorls, all cords similar in size, a subcircular aperture,
a slightly curved, vertical columella without tooth, a
convex base with 9–13 nearly smooth, rather strong,
rounded spiral cords, anomphalous, a white colour.
Remarks. This single, and moreover juvenile,
specimen extends the distribution area of this species
towards the North. Additional material is clearly
needed to confirm is presence in the Solomon Islands.
Thysanodonta brucemarshalli n. sp.
Fig. 21Q–S
urn:lsid:zoobank.org:act:484A82B4-E923-4C40-
8FC8-F80FE2B23DED
Type material. Holotype (9.2 × 7.5 mm) MNHN-IM-
2000-39438.
Type locality. Papua New Guinea, Bismarck Sea,
PAPUA NIUGINI, stn DW3973, 04°34'S, 146°17'E,
411–430 m.
Material examined. Papua New Guinea, Bismarck
Sea. PAPUA NIUGINI: stn DW3973, 04°34'S,
146°17'E, 411–430 m, 1 dd (holotype MNHN-IM-
2000-39438).
Distribution. Only known from the type locality.
Diagnosis. A Thysanodonta species of medium size,
with a moderately elevated, slightly cyrtoconoidal
spire, 8 granular spiral cords on the last whorl (P3; P1,
P2; S3; S1, S2; P4; P'1), S3 peripheral with beads
looking as opisthocline narrow ribs, a subquadrate
aperture, a straight, slightly oblique columella, an
almost flat base with 10 smooth spiral cords, without
umbilicus, a nacreous yellowish white colour.
Description. Shell of medium size for the genus (H of
9.2 mm, W of 7.5 mm), higher than wide, slightly
cyrtoconoidal in shape; spire moderately elevated,
height 1.2× width, 3.0× aperture height, apical angle
62°; angulate periphery; anomphalous.
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Protoconch 350 μm wide, 1.25 whorls, rounded,
sculptured by a network of hexagonal spaces
detectable even at low magnification and without
terminal varix.
Teleoconch up to 6.1 weakly convex to almost flat
whorls; early whorls with 1 strong spiral cord, last
whorl with 7 cords.
Suture weakly impressed and not canaliculated on first
whorls, harder to detect on last whorls.
First whorl convex, sculptured by about 20 thin, weak,
slightly prosocline threads, four weak indistinct spiral
threads on the adapical part and one strong, making
keel, weakly subgranular spiral cord P3; interspaces
between threads similar to their size. On second
whorl, P1 and P2 resolving, granular; axial threads
thicker and more prosocline, connecting beads of
cords; P3 still stronger than the other cords; S3
emerging from suture at half of whorl; S1 and S2
appearing near end of whorl. On third whorl, all cords
more or less similar in size; axial ribs and keel on P3
vanishing, making an almost flat shape to the whorl.
On fourth whorl, P4 appearing, narrower than other
cords. On fifth whorl, P1 dividing into two cords of
the same width; beads of all the other cords
lengthening axially. On last whorls, beads of all cords
axially lengthened except those of P4 emerging,
subperipheral; beads getting narrower and more
numerous from P1 to S3; S3 peripheral with its beads
transforming into opisthocline narrow ribs.
Aperture subquadrate. Columella straight, slightly
oblique, ending abruptly at meeting with outer lip.
Base almost flat, with 10 smooth spiral cords, distance
between them similar to their size; outermost cords
dividing into two similar parts.
Colour of teleoconch nacreous yellowish white;
protoconque white.
Discussion. Thysanodonta brucemarshalli n. sp. has
highly distinct features, especially regarding S3 and
P1. It is rather close to T. boucheti B. A. Marshall,
1988 from New Caledonia (Fig. 21K–L) but this
similar in size species lacks S1 and S2 and none of the
peripheral cords has opisthocline ribs.
The new species may also be compared to T.
aucklandica Marshall, 1988 from New Zealand (Fig.
22S), but this similar in size species has a strictly
conical shape, a different number and ontogeny of
cords (P3; P1, P2, S2; S3; P4; no S1) and the
peripheral cord lacks opisthocline ribs.
Etymology. After Bruce Anders Marshall, thorough
malacologist, expert on New Zealand Mollusca of the
Museum of New Zealand Te Papa Tongarewa and
founder, among numerous other taxa, of the genus
Thysanodonta and of the Thysanodontinae family.
Thysanodonta eumetabolos n. sp.
Fig. 22A–D, Table 7
urn:lsid:zoobank.org:act:69379648-6102-4011-8FF2-
982228FAA301
Type material. Holotype (5.9 × 4.5 mm) MNHN-IM-
2000-39439. Paratype: 1 MNHN (MNHN-IM-2000-
39440, as listed below).
Type locality. Papua New Guinea, Bismarck Sea,
PAPUA NIUGINI, stn DW3973, 04°34'S, 146°17'E,
411–430 m.

Figure 21 (scale bars: 5 mm)

A–B. Phenacomargarites tetratropeis n. sp., holotype MNHN-IM-2000-39437, Solomon Islands, SALOMON

1, stn CP1807, 1077–1135 m, 17.2 × 18.1 mm, 5.0 tw.
C–D. Phenacomargarites williamsae B.A. Marshall, 2016, holotype MNHN IM-2007-18543, Solomon Islands,
South-West of Santa Isobel, 897–1057 m, 20.2 × 22.1 mm, 5.3 tw.
E. Phenacomargarites incomptus B.A. Marshall, 2016, holotype NIWA 54346, Aloha Seamount, East of
Chatham Islands, New Zealand, 801–823 m, 28.9 × 33.0 mm, 5.3 tw.
F–I. Selastele solomonense n. sp. F–G. Holotype MNHN-IM-2000-39434, Solomon Islands, New Georgia,
SALOMON 2, stn CP2219, 650–836 m, 6.2 × 5.2 mm, 5.7 tw. H–I. Paratype MNHN-IM-2000-39435, Papua
New Guinea, Bismack Sea, PAPUA NIUGINI, stn DW4031, 370 m, 5.5 × 4.2 mm, 5.7 tw.
J. Selastele pictum B.A. Marshall, 1995, holotype MNHN-IM-2000-31679, south-eastern New Caledonia,
Loyalty Islands, SMIB 5, stn DW 87, 370 m, 5.5 × 4.2 mm (goldened for Scanning Electron Microscopy).
K–L. Thysanodonta boucheti B. A. Marshall, 1988, Solomon Islands, SALOMONBOA 3, stn CP2823, 240–
572 m, 6.2 × 4.8 mm, 5.6 tw.
M–P. Thysanodonta eucosmia B. A. Marshall, 1995. M–N. Juvenile, Solomon Islands, SALOMONBOA 3, stn
CP2823, 240–572 m, 3.8 × 3.2 mm, 4.2 tw. O–P. MNHN-IM-2013-57605, south-eastern New Caledonia,
EXBODI, stn DW3077, 420–540 m, 10.3 × 7.1 mm, 7.0 tw
Q–S. Thysanodonta brucemarshalli n. sp., holotype MNHN-IM-2000-39438, Papua New Guinea, Bismarck
Sea, PAPUA NIUGINI, stn DW3973, 9.2 × 7.5 m, 6.1 tw.

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C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

103
C. VILVENS New species and new records of Calliostomatidae. Part II

Material examined. Papua New Guinea, Bismarck
Sea. PAPUA NIUGINI: stn DW3973, 04°34'S,
146°17'E, 411–430 m, 1 dd (holotype MNHN-IM-
2000-39439), 2 sub dd, 1 juv dd.
Solomon Sea. MADEEP: stn DW4287, 09°12'S,
153°56'E, 340–375 m, 2 juv dd; stn DW4290,
09°13'S, 153°54'E, 593 m, 1 sub dd (paratype MNHN-
IM-2000-39440).
Distribution. Papua New Guinea, Bismarck Sea,
411–430 m (dd); Solomon Sea, 375–593 m (dd).
Diagnosis. A small Thysanodonta species, with a
moderately elevated, slightly cyrtoconoidal spire, 6
(possibly 7) granular spiral cords on the last whorl
(P3; P1, P2; S3; S2; P4, possible T1), all cords similar
in size except P4, a subcircular to subelliptical
aperture, a straight, slightly oblique columella, a
weakly convex base with 10 subgranular to smooth
spiral cords, without umbilicus, a light brown to white
colour.
Description. Shell small for the genus (H up to 6.1
mm, W up to 4.9 mm), higher than wide,
cyrtoconoidal in shape; spire moderately elevated,
height 1.2× to 1.3× width, 2.7× to 2.8× aperture
height, apical angle 65° to 75°; subangulate periphery;
anomphalous.
Protoconch about 350-400 μm wide, 1.25 whorls,
rounded, sculptured by a network of hexagonal spaces
detectable even at low magnification (especially on
the paratype) and with a thin terminal varix.
Teleoconch up to 6.2 weakly convex whorls; early
whorls with 3 spiral cords, last whorl with 6 (possibly
7) ones.
Suture weakly impressed, not canaliculated, harder to
detect on last whorls.
First whorl convex, sculptured by about 25 very thin,
close, strongly prosocline threads and with P1, P2 and
P3 appearing immediately; P1 and P2 very thin and
hard to detect, P3 much stronger, subgranular, making
keel. On second whorl, P1 and P2 stronger but thinner
than P3, granular; axial threads thicker, connecting
beads of cords, interspaces 1.5×–2× their width; S3
and then S2 appearing near half of whorl. On third
whorl, P1 and S3 similar in size, stronger than P2 and
S2, weaker than P3; axial ribs vanishing. On next
whorls, all cords gradually similar in size; distance
between them similar in size to their width. On last
whorl, P4 appearing, much narrower than other cords;
T1 possibly appearing between S2 and P3 (holotype).
Aperture subcircular (holotype) to subelliptical
(paratype). Columella straight, slightly oblique,
ending abruptly at meeting with outer lip.
Base weakly convex, with 10 spiral cords, subgranular
(holotype) to smooth (paratype), distance between
them similar to their size, innermost cords slightly
stronger.
Colour of teleoconch light brown to white;
protoconque white.

Figure 22 (scale bars: 5 mm)

A–D. Thysanodonta eumetabolos n. sp. A–B. Holotype MNHN-IM-2000-39439, Papua New Guinea, Bismarck
Sea, PAPUA NIUGINI, stn DW3973, 411–430 m, 5.9 × 4.5 mm, 6.2 tw. C–D. Paratype MNHN-IM-2000-
39440, Solomon Sea,. MADEEP, stn DW4290, 593 m, 6.1 × 4.9 mm, 5.5 tw.
E–F. Thysanodonta cassis Vilvens & Maestrati, western New Caledonia, Coral Sea, EBISCO, stn DW2634,
342–347 m, 7.2 × 5.6 mm, 6.0 tw.
G–N. Thysanodonta aoristos n. sp., Papua New Guinea. G–H. Holotype MNHN-IM-2000-39441, Solomon Sea,
MADEEP, stn DW4295, 516 m, 9.6 × 6.9 mm, 6.7 tw. I–N. Vitiaz strait, PAPUA NIUGINI, stn DW3992, 450–
480 m. I–J. Paratype 1 MNHN-IM-2000-39442, 8.2 × 6.7 mm, 6.9 tw. K–L. Paratype 4 MNHN-IM-2000-39445,
8.1 × 6.0 mm, 6.7 tw. M–N. Paratype 5 MNHN-IM-2000-39446, 7.9 × 6.0 mm, 6.0 tw.
O–P. Thysanodonta festiva Marshall, 1995, MNHN, southern New Caledonia, BERYX 11, stn DW18, 250–270
m, 6.5 × 4.4 mm.
Q–R. Thysanodonta pileum Vilvens & Maestrati, 2006, holotype MNHN-IM-2000-5957, New Caledonia,
SMIB 8, stn DW152–154, 305–367 m, 5.0 × 3.3 mm, 6.0 tw.
S. Thysanodonta aucklandica B.A. Marshall, 1988, holotype, 10.2 × 8.5 mm, 6.3 tw (from the original
description)

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C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

105
C. VILVENS
Table 7. Thysanodonta eumetabolos n. sp.: Shells measurements in mm for types.
TW H W
Holotype PAPUA NIUGINI
DW3973 MNHN-IM-2000- 6.2
39439
Paratype MADEEP DW4290
5.5
MNHN -IM-2000-39440
Discussion. Thysanodonta eumetabolos n. sp. is rather
close to T. cassis Vilvens & Maestrati, 2006 from
New Caledonia (Fig. 22E–F), but this slightly larger
species has a different ontogeny of the spiral cords
(P3; P1, P2; P4; S2; S3) with moreover P1 and S3 not
similar to the other ones.
The new species may be compared to T. pileum Vilvens
& Maestrati, 2006 from New Caledonia (Fig. 22Q–R),
but this species similar in size and shape has a more
elevated spire and a very different ontogeny of spiral
cords ((P3, P4; P1, P2; S2, S1; no S3).
Etymology. Variable (Ancient Greek: εύμετάϐολος, -
ον)- with reference to the variability of the new
species, regarding the small differences between the
holotype and the paratype, although they share the
main typical features.
Thysanodonta aoristos n. sp.
Fig. 22G–N, Table 8
urn:lsid:zoobank.org:act:2994BC2A-181F-414E-
A688-E6EFF412936C
Type material. Holotype (9.6 × 6.9 mm) MNHN-IM-
2000-39441. Paratypes : 5 MNHN (MNHN-IM-2000-
39442, MNHN-IM-2000-39443,MNHN-IM-2000-
39444, MNHN-IM-2000-39445, MNHN-IM-2000-
39446, as listed below).
Type locality. Papua New Guinea, Solomon Sea,
MADEEP, stn DW4295, 09°36'S, 152°54'E, 516 m.
Material examined. Solomon Sea. MADEEP: stn
DW4295, 09°36'S, 152°54'E, 516 m, 1 dd (holotype
MNHN-IM-2000-39441).
Papua New Guinea, Vitiaz strait. BIOPAPUA: stn
DW3719, 06°03'S, 147°36'E, 410 m, 1 dd (paratype 3
MNHN-IM-2000-39444), 1 sub dd, 3 juv dd.
PAPUA NIUGINI: stn DW3992, 06°03'S, 147°36'E,
450–480 m, 4 dd (paratype 1MNHN-IM-2000-39442,
paratype 2 MNHN-IM-2000-39443, paratype 4
MNHN-IM-2000-39445, paratype 5 MNHN-IM-
2000-39446), 1 juv dd.
Solomon Islands. SALOMON 1: stn DW1825,
09°51'S, 160°58'E, 340–391 m, 1 dd.
Distribution. Solomon Sea, 516 m (dd); Papua New
Guinea, Vitiaz strait, 410–450 m (dd); Solomon
Islands, 340–391 m (dd).
106
New species and new records of Calliostomatidae. Part II
HA H/W H/HA
5.9 4.5 2.20 1.31 2.68
6.1 4.9 2.20 1.24 2.77
Diagnosis. A Thysanodonta species of medium size,
with a rather elevated, conical to very slightly
cyrtoconoidal spire, a more or less angulate periphery,
at least 6 (possibly up to 10) granular spiral cords on
the last whorl (P3; P1, P2; S3; S2; P4; possible T1;
possible T2, T3, T4), all cords similar in size except
P4 a bit thinner, a weakly convex to almost flat base
with 8–10 smooth to subgranular spiral cords, without
umbilicus, a ivory colour, possibly with light brown
hue.
Description. Shell of medium size for the genus (H up
to 9.6 mm, W up to 7.1 mm), higher than wide, more
or less conical to slightly cyrtoconoidal in shape; spire
rather elevated, height 1.2× to 1.4× width, 3.0× to
4.6× aperture height, apical angle 60°; subangulate to
angulate periphery; anomphalous.
Protoconch about 360-420 μm wide, 1.25 whorls,
rounded, sculptured by a network of hexagonal spaces
(visible only on paratypes 2 and 3) and without
terminal varix.
Teleoconch up to 6.9 flat whorls, except the first one
weakly convex and possibly the last one very convex
(paratypes 4 and 5); early whorls with 4 spiral cords,
last whorl with at least 6 (possibly up to 10) ones.
Suture weakly impressed, not canaliculated, harder to
detect on last whorls.
First whorl convex, sculptured by about 15–20 rather
thin, rather close, prosocline threads (features a bit
variable on the types) and P3 appearing immediately,
subgranular, making keel; P1 and P2 appearing a bit
later, S3 near end of whorl. On second whorl, P1 and
P2 stronger but thinner than P3 and S3, granular; axial
threads thicker with interspaces wider (1.5–2× their
width); S2 appearing near end of whorl. On third
whorl, keel on P3 and axial threads vanishing; all 4
cords gradually similar in size. P4 partially emerging
from suture near end of third whorl or on fourth
whorl. On next whorls, all cords similar in size;
distance between them similar in size to their width.
On last whorl, P4 appearing, narrower than other
cords; T1 possibly appearing between P1 and S1
(holotype); T2, T3 and T4 possibly appearing
respectively between P2 and S2, S2 and P3, P3 and S3
(paratypes 4 and 5).
Aperture subquadrate. Columella straight, weakly
oblique, ending abruptly at meeting with outer lip.
Base weakly convex to almost flat, with 8–10 smooth
(holotype, paratype 1) to subgranular (paratypes 3, 4
C. VILVENS NOVAPEX 25(2): 55–110, 10 juin 2024

and 5) spiral cords, distance between them similar to Colour of teleoconch ivory, possibly with light brown
their size, innermost cords slightly stronger. hue; protoconch white.

Table 8. Thysanodonta aoristos n. sp.: Shells measurements in mm for types.

TW H W HA H/W H/HA
Holotype MADEEP DW4295
6.7 9.6 6.9 2.9 1.39 3.31
MNHN-IM-2000-39441
Paratype 1 PAPUA NIUGINI
DW3992 MNHN-IM-2000- 6.9 8.2 6.7 2.7 1.22 3.04
39442
Paratype 2 PAPUA NIUGINI
DW3992 MNHN-IM-2000- 6.6 9.6 7.1 2.3 1.35 4.17
39443
Paratype 3 BIOPAPUA: stn
DW3719 MNHN-IM-2000- 6.3 7.4 5.9 1.6 1.25 4.63
39444
Paratype 4 PAPUA NIUGINI
DW3992 MNHN-IM-2000- 6.7 8.1 6.0 2.2 1.35 3.68
39445
Paratype 5 PAPUA NIUGINI
DW3992 MNHN-IM-2000- 6.0 7.9 6.0 2.2 1.32 3.59
39446

Discussion. Thysanodonta aoristos n. sp. is close to T. ACKNOWLEDGEMENTS

festiva B. A. Marshall, 1995 from southern New
Caledonia (Fig. 22O–P), but this smaller species (H up
to 7.4 mm for 7.3 tw) is brightly coloured, has a
narrower apical angle (42°–44°) and has a peculiar,
more complicated ontogeny of spiral cords (P3; (P4);
P1, P2, S2, S3; S'3) with P4 fused with S3 and S3
dividing later into two cords.
The new species is also rather close to T. boucheti B.
A. Marshall, 1988 from New Caledonia (Fig. 21K–L)
but this species is smaller for a similar number of
whorls and has only 5 spiral cords, lacking S2.
Remarks. One could at first glance consider that
paratypes 4 and 5 do not belong to the same species as
the other types of Thysanodonta aoristos n. sp.,
mainly because they have multiple Ti and a last whorl
very convex. But simultaneous careful examination of
the specimens collected at the station DW3992 of the
PAPUA NIUGINI campaign, from which almost all
types do come from, shows that they share the same Pi
and Si cords ontogeny while the holotype is
intermediate between the "no Ti" state and the
"numerous Ti" state. Also, regarding the convexity of
the last whorl, the paratype 5 is intermediate between
strong convexity (paratype 4) and almost flat shape
(holotype and paratype 1) for this last whorl.
Etymology. Hesitant, indecisive (Ancient Greek :
άόριστος, -ον) - as if the personified new species
could not choose between Ti of no Ti spiral cords,
convex or flat last whorl.
This paper is based on material originating from many
cruises carried between 1980 and 2015 on board
French, Taiwanese, Indonesian and Philippines
research vessels, mostly as part of the "Tropical Deep-
Sea Benthos" programme (formerly "Campagnes
Musorstom"). Scientific partners included National
Taiwan Ocean University (Keelung), National Taiwan
University (Taipei), University of Papua New Guinea
(Port Moresby), Pusat Penelitian dan Pengembangan
Oseanologi LIPI (Jakarta), University of San Carlos
(Cebu City) and the Bureau of Fisheries and Aquatic
Resources (Manila). Specifically, material from
Taiwan and the South China Sea was collected as part
of the French–Taiwanese deep-sea project TF-
DeepEvo funded by ANR and NSC (ANR 12-ISV7-
0005-01, PIs Sarah Samadi,Wei-Jen Chen). Access to
French ship time was made possible through Institut
de Recherche pour le Développement (IRD, formerly
ORSTOM), IFREMER (formerly CNEXO), and the
Flotte Océanographique Française. Additional
expedition funding was provided by the Total
Foundation, Prince Albert II of Monaco Foundation,
Stavros Niarchos Foundation. All expeditions
operated under the regulations then in force in the
countries in question.
I thank Philippe Bouchet, Barbara Buge, Virginie
Héros, Pierre Lozouet and Philippe Maestrati
(MNHN) for assistance throughout my work, from
curation of expedition material, access to the
malacological resources of the MNHN and issuing
loans, to cataloguing relevant specimens, and
especially Virginie Héros for reading and completing
the manuscript.

107
C. VILVENS
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C. VILVENS New species and new records of Calliostomatidae. Part II

Appendix : list of the Calliostomatidae species studied in this work

Calliostoma hexalyssion Vilvens, 2009 - p.60

Calliostoma simplex Schepman, 1908 - p.60
Calliostoma aporia Vilvens, 2009 - p.61
Calliostoma quadricolor Schepman, 1908 - p.61
Calliostoma pheidolon n. sp. - p.61
Calliostoma philippei Poppe, 2004 - p.61
Calliostoma newguineense n. sp. - p.64
Calliostoma thaumaston n. sp. - p.66
Calliostoma altena Knudsen, 1970 - p.68
Calliostoma scobinatum (Reeve, 1863) - p.68
Calliostoma trotini Poppe, Tagaro & H. Dekker, 2006 - p.70
Calliostoma emmanueli Vilvens, 2000 - p.70
Calliostoma paucicostatum Kosuge, 1984 - p.72
Calliostoma suduirauti Bozzetti, 1997 - p.72
Calliostoma ticaonicum (A.Adams, 1851) - p.73
Calliostoma poppei Vilvens, 2000 - p.73
Calliostoma dedonderi Vilvens, 2000 - p.74
Calliostoma vilvensi Poppe, 2004 - p.74
Calliostoma formosense E. A. Smith, 1907 - p.74
Calliostoma achantodes n. sp. - p.76
Calliostoma zhongshaense n. sp. - p.78
Calliostoma statheron n. sp. - p.81
Tristichotrochus aculeatus (G. B. Sowerby III, 1912) - p.82
Fautor paradigmatus (B.A. Marshall, 1995) - p.84
Fautor cosmopolites n. sp. - p.86
Fautor panteles n. sp. - p.90
Akoya haliarchus (Melvill, 1889) - p.94
Akoya zhui n. sp. - p.94
Bathyfautor multispinosus (Schepman, 1908) - p.95
Venustatrochus malaita (Vilvens, 2009) - p.96
Selastele solomonense n. sp. - p.98
Phenacomargarites tetratropeis n. sp. - p.100
Thysanodonta boucheti B. A. Marshall, 1988 - p.101
Thysanodonta eucosmia B.A. Marshall, 1995 - p.101
Thysanodonta brucemarshalli n. sp. - p.101
Thysanodonta eumetabolos n. sp. - p.102
Thysanodonta aoristos n. sp. - p.106

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