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Study Material 3- Patterns of Behaviour_240603_110557
Study Material 3- Patterns of Behaviour_240603_110557
Study Material 3- Patterns of Behaviour_240603_110557
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Orientation:
Orientation is the positioning of the body, or parts of the body, in relation to the environment.
Primary orientation is the establishment of body’s basic posture in space. The secondary
orientation in the other hand is the positioning and movement of body in response to various
stimuli.
Kinesis:
The simplest form of spatial orientation in kinesis. In kinesis, animal’s response is proportional
to the intensity of stimulation but is independent of the spatial properties of the stimulus. For
example, common woodlice (Porcellion scaber) tend to aggregate in damp places beneath rocks
and fallen logs. They move about actively at low humidity levels but are less active when the
humidity is high. They consequently spend more time in damp conditions but their enhanced
activity in dry conditions increases the chance of discoveries of a damp place. It may be inferred
that when the speed of movement and the frequency of turning of animals are affected by the
stimulus then such orientation is called kinesis. Kinesis is directed movement of animals in
relation to the sense of stimulus.
There are two types of Kinesis: A. Orthokinesis B. Klinokinesis
Klinokinesis:
In this type of kinesis the rate of change of direction increases with the increased light intensity
(According to Fraenkel and Gunn, 1940, Hinde, 1970.) The flatworm Dendrocoelum lacteum
does not move in a linear fashion but shows a twisted course of movement which includes
turning in various direction of linear movement are random and take place even in uniform
environmental condition. The rate of random turning or angular velocity is called as ‘rate of
change in direction’ (R.C.D.) and is represented in degress/Unit time. The protozoans usually
swim in a corkscrew manner and when in favourable location, the random deviations from the
line of movement are increased keeping the animal in favourable environment longer. This can
also be seen in higher crustacea and grazing mammals.
Orthokinesis:
When a relationship exists between the speed of locomotion and the intensity of stimulation
then it is called Orthokinesis. Such kinesis may be seen in Porcellion scaber (Woodlice), in
which the linear velocity of its movement is affected by the intensity of stimulus. As referred
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above, the woodlice move faster in dry condition than the humid condition. The result is that
when the woodlice are kept in a chamber with graded humidity, then they aggregate in the
humid zone. Actually, this is not because they prefer humidity but because of the fact that they
move quickly through the non-humid areas (i.e. dry areas) to humid ones. The ammocoet larvae
of lamprey Lampetra planeri are found buried in the bottom of water bodies like lakes and
ponds, with their head pointing below. If the light intensity is increased and they are exposed
to it, they exhibit a vey active phase of swimming and their head pointing below which may
help them burrow again and move away from light.
Taxis:
When the animals either move towards or away from the stimuli and their general course of
movement is the straight line joining them with the source of stimulus, such orientation is called
as ‘taxis’. If the movement of Animal towards the source of stimulus than this is called as
positive taxis and if it is opposite to this direction than it is known as negative taxis.
Klinotaxis:- This is the simplest form of orientation & undirected. Orientation by successive
comparison of stimulus intensity requires turning movements. Usually, it is called klinotaxis.
Many animals show Klinotaxis in response to gradients of chemical stimulation. This can be
seen in animals when they are confronted with environments which have, for example,
gradients of chemical stimulation.
Tropotaxis: Tropotaxis is shown by those animals which possess receptors. With the help of
receptors, they can compare the intensity of stimulus in all sites simultaneously. Tropotaxis
enables the animal to steer a course directly towards or away from the source of stimulation.
Usually the receptors are arranged in a bilaterally symmetrical fashion. It has been observed
that if the receptors of one side are put out of action then the animals are unable to decide the
course and move almost aimlessly. Such type of movement of the animal is called as “Circus
movement”.
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Telotaxis: In this type of taxis the balance does not exist and orientation of animal to light is
not altered. The animal at one time may orientate to one light source and at the other time to
the other light source. Thus it is capable of ignoring one or more light sources at a particular
time. It shows that the bilateral arrangement of the stimulus is not a necessary factor for
orientation of this type. In other words, eyes that are capable of providing information about
the direction of light by virtue of their structure are capable of Telotaxis. It can be called as a
goal directed orientation. Hermit crabs have two well developed eyes and when it is presented
with two light sources, one dim and other bright, it always moves towards the brighter one.
Male grayling butterflies, after being blinded in one eye would still fly towards the passing
females showing the goal directed orientation. When we see a sprinter doing a 100-meter dash
at an atheletic meet, we tum our eyes to get the image of the athelete on fovia centralis, the area
of retina where the vision is sharpest. From then onwards even though we may turn our eye,
head or body to follow the movement the image is kept locked on that specific location on
retina. So in spite of the image being fIxed at a place, the movement of sprinter is perceived
and the motion appreciated by our brain. This appreciation comes through the motor
organization involved in moving the eye, head or body. All the messages coming from these
different regions are integrated into continuous perception of the movement.
Menotaxis. The orienting mechanisms which help the animal to maintain a constant direction
by steering a course at constant angle to the incidence of the stimulus are called menotaxis
(Greek word meno = I remain). In this type of orientation generally the principle of negative
feedback is involved, where the output of regulating cycle acts as the input. One of the common
example of this type of orienting behaviour is the light compass reaction of the insects. The
insects use sun for guidance in homing. The ants Lasius niger use sun as a reference for homing.
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If these ants are confirmed to a dark box for few hours while in the middle of their homeward
journey, after their release they would maintain the same angle to the sun as the one when they
were kept in the box. They would not compensate for the movement of sun during these hours
which changed it's angle. This would then lead them to a different path then the one taking
them home.
Mnemotaxis. Interestingly mnemotaxis was postulated as a possibility by Kuhn in 1919. But
his critics argued against it and he succumbed to their pressure and retracted his theory as no
proper example could be found then. However, he has been vindicated now as many examples
have been discovered. Mnemotaxis is a form of orientation based on memory. The water shrew
(Neomys fodiens) exhibits this memory resident orientation when it follows the same path
every time. If its memory map is disturbed by changing a part of its path, it will stop and start
exploring the surroundings, it will then retrace its steps till it locates a known land mark and
then will resume the earlier course and would try to overcome the difficulty posed by the
changed path. It can also be seen in children reciting poetry, who repeat the earlier lines if they
forget anyone line in the middle of recitation.
Learning:
Irreversible change in response to particular stimuli as opposed to the reversible changes that
result from changes in motivation.
Kinds of learning behaviour:
The various levels of learning found in the animal kingdom have been classified ethologically in several
ways which are recognized by animal psychologists. 1. Habituation 2. Imprinting 3. Conditioned
Reflexes or Associative learning 4. Trial-and-error learning 5. Latent learning 6. Insight learning 7.
Reasoning.
1. Habituation:
2. Imprinting
Imprinting is actually a specialized form of learning that is seen clearly in many kinds of birds during
their early period of life (Hess, 1958.) When the early period over’s, the birds are unable to learn. For
example, young ducks will normally follow their mother around soon after they hatch. This following
behaviour is the result of hatching. Apparently imprinting will occur only for a short time after the bird
is hatched. If imprinting is prevented from occurring by hatching the egg in an incubator and not
exposing the young bird to an adult bird for some days, it never occurs. Experiments have been
performed with ducks and other birds hatched in incubators. In the case of many, the first large moving
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object seen by the newly hatched animal (bird) will be the stimulus for imprinting. If the first such
object seen is a man, the young birds will follow the man about. In simple form, if the young duckling
are hatched artificially and then exposed early to some moving object other than their true mother,
they will behave towards the object as they normally would to their mother duck, following the object
wherever it goes. Such a “substitute mother” might be a ball bicycle, an electric train engine, or even
a human.
The sight and smell of food can cause salivation. Similar a dinner gong sound can produce salivation.
Associative learning or learned reflex is of two types: A-classical conditioning and BInstrumental
conditioning.
Trial and error learning results when instrumental conditioning is made more complex by introducing
several variables. Various types of mazes i.e. problem apparatus as weel as other multiple choice
situations have been developed to measure learning ability in various animals. Attempts have been
made to show that various protozoans, expecially paramaecia, can lern. Even experiments designed to
illustrate habitation learning have not been entriley successful. In fact, learning has not yet been
successfully demonstrated in the simple multicellular animals or in any of the radially symmetrical
animals (Coelenterata, Ctenophora, and Echinodermeta).
5. Latent learning Latent learning is acquired even when there is no particular reward or
punishment associated with an animal’s activity. Many animals appear to be naturally “curious”,
exploring their surroundings in great detail. When finches are relased into an aviary for example, they
will investigate every nook and cranny until they are thoroughly familiar with their new home. When
a hive of honeybees is closed up, moved to a new location, and then reopened, many of the emerging
bees appear to perform short orientation flights, lasting only a minute or two. They emerge and fly in
ever widening circles around the hive, apparently learning to recognize the surroundings sufficiently
to find there, way back from a longer flight.
6. Insight learning The most sophisticated kind of learning is called insight learning, the ability
to use knowledge obtained in one context to solve a problem in a different context. The chimpanzee,
who pokes a stick into an ant nest, allows the ants to climb on the stick and then licks them off, is using
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insight learning if it has “reasoned” beforehand that the stick would make a suitable tool for capturing
the ants. Insight learning may also be termed as “intelligence.” Insight learning differs from trial-and-
error leaning because it involves the sudden production of anew response. In the human species
insight learning is most highly developed. However, birds, such as crow also possess this type of
learning. Actually human behaviour contains many learned responses and depends especially on
insight learning.
7. Reasoning With increasing complexity of the nervous system in the higher vertebrates’, for
instance, mammals and especially the primates, the behavioural pattern many not always be innate or
acquired from past experiences but a complex of one more ability of reasoning. Reasoning is of the
top of all modes of adaptive behaviour. Reasoning capacity includes the ability to solve complex
problems with something more than simple trail-and-error habit or stimulus-response modification.
In other words, reasoning is the ability to use past experience and logical deductions to solve some
new complex problem.
Animals are often used as points of comparison when talking about AI and the possibility of
its consciousness. Animals used to be thought of as non-conscious, pre-programmed "machines", and
had few rights and protections. Science has since revealed that many species, from insects to whales,
have the hallmarks of consciousness. We must be careful with the language we use when comparing
animals and AI, to avoid equating the two. A popular topic of conversation is whether AI will ever
become conscious. But what exactly do we mean by consciousness? Consciousness is hard to define.
To put it loosely, it’s a state of awareness about the “self". It allows us to solve problems by thinking
(rather than trial and error), and to create a narrative about our lives through memory and language.
But the problem with defining consciousness in a broad way is that we can only ever understand it
through the lens of our own experience. We don’t really know what it means to be a conscious non-
human because we simply cannot imagine the way non-humans experience the world.