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Living Physical Geography
Instructor’s Manual
PART II
The Biosphere and the Geography of Life
PART SUMMARY
Like Part I, solar energy underpins all chapters in Part II. All life on Earth uses solar energy either
directly, as in the case of plants and herbivores, or indirectly, as in the case of carnivores. Part II
consists of four chapters, each exploring the biosphere from a geographer’s perspective:
Chapter 7 Patterns of Life: Biogeography
Chapter 8 Climate and Life: Biomes
Chapter 9 Soil and Water Resources
Chapter 10 The Living Hydrosphere: Ocean Ecosystems
Chapter 7 introduces the basic elements of the discipline of biogeography and explores
geographic patterns of life. Chapter 8 couples biomes with global climate types because biomes
are a result of climate. Chapter 9 explores soil and water as resources for people. The
biogeography of the oceans is discussed in Chapter 10.
TEACHING CONTEXT
Covering the biosphere immediately after the atmosphere makes sense for two reasons. First,
the theme of solar energy doing work on Earth’s physical systems continues from the
atmosphere. Living organisms use solar radiation, either directly or indirectly, for their energy
source. Second, Chapter 8 discusses biomes in the context of global climate types. Teaching
biomes and climate jointly is an efficient and logical use of classroom time. Combining them
also avoids the redundancy caused by covering climate types in the atmosphere then covering
them again in the biosphere, as is commonly done.
Similarly, soil and water resources are topics that are naturally combined. The two
resources relate to one another in the context of agriculture and feeding people.
The importance of life in the oceans has been overlooked in physical geography textbooks.
This chapter provides a portrait of the biogeography of the oceans. The oceans are important
for several reasons, particularly because they cover a significant portion of the planet and much
of humanity relies on them for food.
TEACHING TIPS
• Biogeography is an important subdiscipline within geography. Students should be exposed
to the main ideas in chapters 7 and 8, including basic geographic patterns of life, geographic
1
patterns resulting from evolution, factors that control the geographic ranges of organisms,
organizing schemes of the biosphere, and the climate-biomes concept.
• Chapter 9’s soil and water resources is most effectively taught and learned if it is couched in
the context of providing resources for people, including food and water resources.
• Marine biogeography can be explored as sequenced in the chapter, or the material can be
tailored to suit one’s preferences and time schedule.
CHAPTER 7
Patterns of Life: Biogeography
CHAPTER SUMMARY
Three major patterns of biodiversity are that the tropics are more biodiverse than high
latitudes, large islands are more biodiverse than small islands, and organisms migrate and can
change biodiversity on a seasonal basis. Many biogeographic patterns reveal evolutionary
histories of regions and the organisms living in them. Most species’ geographic ranges are the
result of limiting factors. Organisms disperse to obtain resources. Ecological disturbance often
reduces biodiversity in an area, and organisms return to areas of disturbance through ecological
succession. The biosphere can be organized by energy flows, genetic similarities, and the
geographic space a given unit of life occupies. Coconut dispersal exemplifies the concepts of
dispersal, as well as domestication and evolution.
Three global biodiversity patterns include the latitudinal biodiversity gradient, which refers
to the tropics as being more biodiverse than high latitudes. Large islands are more biodiverse
than small islands. Biogeographic islands can include mountaintops or lakes, and so on.
Migration can also affect biodiversity for any given area.
Biogeographic patterns sometimes reveal evolutionary events for a given region. Where
two regions share similar physical characteristics but are geographically separate, such as
occurs in geographically separate deserts, genetically unrelated organisms will evolve similar
adaptations. Patterns of evolutionary divergence occur among genetically similar organisms
after they become reproductively isolated from the population from which they came.
Evolutionary divergence commonly occurs after organisms disperse to remote geographic
islands. All species arose through the process of evolution by natural selection.
Limiting factors play an important role in determining the geographic distribution of
organisms. Biological limiting factors include competition, predation, and mutualism. Physical
limiting factors include light, water, and temperature. All organisms are subject to limiting
factors.
Dispersal is the permanent relocation of individuals within a population. Dispersal is
important for both an individual and for a species. An individual may disperse to reduce limiting
factors and increase resources. A species can expand or shift its geographic range through time
as individuals disperse. When people disperse an organism to a geographic region where it did
not previously live, the organism is considered non-native, and invasive if it escapes into the
2
wild and competes with native organisms. When organisms disperse on their own to a new
location they are considered colonizers. Invasive species are a problem in most areas worldwide
today.
Most ecosystems experience disturbance in various forms, ranging from damage caused by
strong winds to devastating volcanic eruptions. All disturbance events are followed by
ecological succession, in which species return to the area that was disturbed. Fire is a type of
disturbance that was removed from many forested areas in North America. The ecological
importance of fire is now recognized by forest managers, and in many areas fire is being
allowed to return.
The biosphere is very complex. Three schemes of categorization, or hierarchies, are used to
organize the biosphere’s complexity. The trophic hierarchy is based on energy flows.
Photosynthetic organisms convert solar energy to chemical energy. This chemical energy is then
transferred to other organisms as they consume one another. In the trophic hierarchy, only
10% of the energy of one organism is transferred to the next organism. The taxonomic
hierarchy uses genetic similarities to form nested groups of organisms. The spatial hierarchy
organizes the biosphere geographically by the space that is occupied, ranging from the
individual to biomes to the entire biosphere.
The Geographic Perspectives at the end of the chapter explores the biogeographic history of
the coconut palm and its means of dispersal to illustrate the concepts of dispersal,
domestication, and mutualism. Coconuts and people are mutualist partners. The coconut palm
has benefited from people cultivating and dispersing it. People have benefited from the
coconut palm because it provides food and material resources.
TEACHING CONTEXT
Biogeography is an important subdiscipline within geography. The material in this chapter
provides a good introduction to many foundational concepts in biogeography. This chapter is
also a prerequisite for Chapter 8. Much of Chapter 8 assumes that the student has read this
chapter first.
TEACHING TIPS
• This chapter presents an abrupt departure from Part I, the atmosphere. Create a sense of
continuity by emphasizing that physical geography is ultimately about the study of energy’s
effects on Earth’s physical systems. Part II continues with the study of solar energy in the
biosphere.
• Biogeography is different from biology in that biogeography emphasizes spatial patterns.
Analysis of those patterns reveals process. Emphasize the spatial aspects of topics as much
as possible when teaching this unit.
• Students usually enjoy the Yellowstone wolf story (Figure 7.14) and mutualist interactions
among organisms. Find other examples of mutualism, but have the students determine how
3
the relationship is an example of mutualism. Other examples include corals and their algae,
acacia and ant guardians, and squirrels and oaks/acorns.
_____________________________________________________________________________
1. Determine how many trophic levels the owl is above the primary producer level (see Figure
7.25), and what percentage of the original energy is available at the owl’s trophic level.
2. Convert the percentage of energy captured at the owl’s trophic level to a decimal (by
dividing it by 100).
3. Divide the owl’s biomass (0.5 kg) by the decimal you obtained in step 2.
Answer:
Step 1: The owl is three trophic levels up. Each level passes on 10%. Thus, the owl receives
0.1% of the energy originally available from the primary producers.
Step 2: 0.1%/100 = 0.001.
Step 3: 0.5 kg/0.001 = 500 kg of primary producer biomass are required to support one
tertiary-level consumer owl that weighs 0.5 kg.
Consider This
1. How do you think the sooty shearwater knows where it is in such vast geographic
space? What “map” is the bird using? Birds navigate using an internal compass that
4
allows them to detect Earth’s magnetic field. This compass allows them to navigate
across the featureless oceans on a constant course.
2. What specific “push–pull” factors can you think of that would compel the sooty
shearwater to migrate such great distances? The most important push factor is winter.
The pull factors are nesting and mating opportunities, as well as increased food
resources.
Consider This
1. What trophic level(s) does the eastern emerald elysia occupy? It is a primary producer
because it makes its own energy directly from the Sun. It is also a primary consumer
because it eats algae.
2. How would photosynthesis be an advantage to this animal? Photosynthesis in animals
should be a significant advantage because animals can supplement their diet with
energy obtained from the Sun. The Sun may be a reliable source of energy during times
when other types of food are not.
Problem 7.1
This placemark highlights a November 2012 eruption of Tolbachik volcano on the Kamchatka
Peninsula. Activate the folder for this problem to answer the questions below.
1. What kind of ecological succession will follow this eruption?
a. Primary succession
b. Secondary succession
(answer: B)
2. How long is this lava flow?
a. 2 km
b. 11 km
c. 18 km
d. 25 km
(answer: B)
3. Zoom out. Where is this eruption occurring?
a. Japan
b. Alaska
c. Russia
d. North Korea
(answer: C)
5
Problem 7.2
This placemark highlights the Ponderosa fire, which burned 15,000 acres in Northern California
in August 2012. Activate the folder for this problem to see a natural color satellite image of the
smoke from the Ponderosa fire burning on August 19. The red outlines show hot spots where
fires were still burning. If you zoom out, you will see the smoke column from the Chips fire to
the south and east.
1. Based on the photo accompanying this placemark, what kind of fire was this at the location
of the placemark?
a. A surface fire
b. A canopy fire
(answer: A)
2. What kind of ecological succession will occur after this fire?
a. Primary succession
b. Secondary succession
(answer: B)
3. What is the length of the burning area shown within the red outline?
a. 7 km
b. 13 km
c. 20 km
d. 100 km
(answer: A)
4. Deactivate the fires layer by clicking the folder for this problem and zooming in to the
placemark. This image was made before the fire, so no burn scar is visible. You can see a
number of bare patches made by clear-cut logging of the forest here. What kind of ecological
succession will take place in these patches?
a. Primary succession
b. Secondary succession
(answer: B)
Problem 7.3
This placemark visits the Aleutian Islands of Alaska in the North Pacific Ocean.
1. What kind of biological dispersal do island chains such as this provide?
a. Stepping-stone dispersal
b. Waif dispersal
c. Sweepstakes dispersal
(answer: A)
Problem 7.4
This placemark visits Ascension Island.
1. How far is this island from the nearest continent?
a. 500 mi
6
b. 900 mi
c. 1,400 mi
d. 2,000 mi
(answer: B)
2. How did native terrestrial organisms reach Ascension?
a. Stepping-stone dispersal
b. Waif dispersal
c. Sweepstakes dispersal
(answer: C)
Problem 7.5
California’s Channel Islands are featured here.
1. About how far from the mainland is the island nearest to the mainland?
a. 2 mi
b. 19 mi
c. 45 mi
d. 90 mi
(answer: B)
2. Which of the Channel Islands would you expect to have the highest plant and animal
diversity?
a. San Nicolas
b. San Clemente
c. San Miguel
d. Santa Cruz
(answer: D)
3. The Channel Island fox (Urocyon littoralis) dispersed to the islands about 16,000 years ago.
By what means did it reach the islands?
a. Stepping-stone dispersal
b. Waif dispersal
c. Sweepstakes dispersal
(answer: B)
4. The fox is found on six of the islands. Assuming the fox reached the Channel Islands only
once, by what means did it disperse to the other five islands?
a. Stepping-stone dispersal
b. Waif dispersal
c. Sweepstakes dispersal
(answer: A)
5. There are six subspecies of the Channel Island fox, none of which looks like the ancestral
gray fox on the mainland. What process produced these changes?
a. Convergent evolution
b. Divergent evolution
7
c. Ecological succession
d. Ecological disturbance
(answer: B)
Concept Review
8
8. What is a population? Provide biogeographic examples that result from evolution once
populations are geographically isolated. A population is a group of organisms that interact and
interbreed in the same geographic area. The example used in this book is warm-water
tolerance among salmon. Other examples include penicillin-resistant strains of tuberculosis or
strains of bacteria that are resistant to antibiotics.
9. What is convergent evolution? Convergent evolution is the evolutionary process in which
two or more unrelated organisms evolve to look alike.
10. What is divergent evolution? Divergent evolution is the process in which individuals in one
population evolve to look different from closely related individuals in another population.
11. What are biogeographic regions and how do they form? Biogeographic regions are
continental-scale regions that contain genetically similar groups of plants and animals. Physical
barriers, like deserts, mountains, and oceans, isolate groups of organisms in one region from
groups in another region.
9
17. What is a keystone species? Give an example. How does the concept of a keystone species
relate to limiting factors? The gray wolf in Yellowstone was used as an example in the text. Its
removal led to increased elk populations, decreased riparian vegetation, and more stream
erosion. Beaver and its pond ecosystems also diminished without the riparian vegetation on
which it depends. Wolf kill also supports many scavenger species. The removal of the wolf from
the park was a limiting factor for many other organisms that depend on the wolf indirectly.
Beaver, for example, depend on the wolf to control elk populations. Without that control, the
elk limit the beaver by taking its food.
18. What is mutualism? Give examples of it. How can it be a limiting factor for a given
organism? Mutualism refers to reciprocal actions that benefit two or more different species.
Examples include flowers and their pollinators, and clownfish and sea anemones. Mutualism is
a limiting factor when the mutualist partner is absent. This limits the potential of both
organisms.
10
24. Review the importance of fire to western North American forests. What problem has
developed? What is being done to address it? Fire is an integral and necessary element in
many ecosystems worldwide, including North American forests. In areas where fire naturally
occurred with frequency, the vegetation and organisms living there were adapted and
coevolved with fire. In the 1930s the U.S. Forest Service implemented a policy of fire
suppression. This caused fuels to build up. Now with the increased fuel loads in the forests, fires
burn hotter and uncontrollably. Prescribed burns and mechanical thinning are being used to
reduce fuels loads.
25. What is ecological succession? What causes it? What are seres? Succession is the step-by-
step changes in vegetation in an ecosystem that follow a disturbance event. Ecological
disturbance causes succession. Seres are stages of succession. For example, after a disturbance
event grasses and ferns may be the first sere. This may be followed by shrubs, then small trees,
then mature oak forest. Each of these are examples of seres.
26. What is a climax community? Is this model always accurate? The climax model idea states
that if left undisturbed, vegetation will return to its original state. The model is not accurate.
The problem with the climax community model is that it ignores anthropogenic influences,
climate change, non-native species and the occurrence of repeated disturbance events that
continually reset the clock of succession. The modern view sees geographic regions as
composed of mosaics of different seres.
11
31. What types of consumers recycle dead organisms? Scavengers (such as a coyote) eat
carrion, detritivores (such as an earthworm) eat rotting organic material, decomposers (like
fungi and bacteria) break down organic remains into simpler compounds.
32. What are homologous traits? How are they used to create taxonomic groups? These are
traits that originate from a single ancestral trait. In other words, they share a genetic
relationship. Organisms that share a common ancestor have homologous traits. Taxonomists
group organisms by shared genetic similarities.
33. Compare common names and scientific names. On what is each based? How is each type
of name useful? There are often several common names for a single species. Scientific names
assign only one genus and species name for each species. Common names are in the local
language, but scientific names are always in Latin. Common names are generally based on
superficial appearance. Scientific names are based on genetic similarities among organisms.
Common names are easy to use and remember, but they can be misleading because there are
many of them. Scientific names can be unfriendly to the public, but they are efficient and
provide useful information about the organism’s genetic relationships to other organisms.
34. What is a community? How is the term defined? Give examples of a few communities. A
community consists of different populations of organisms interacting in a geographic area. Tide
pool community, grazing community in grasslands, ponderosa pine community, songbird
community, and so on. There are no hard rules in referencing a community.
35. What is the carbon cycle? What is the nitrogen cycle? How do they work? Why are they
important? The carbon cycle refers to how the carbon atom moves within and between Earth’s
physical systems. The nitrogen cycle refers to how the nitrogen moves within and between
Earth’s physical systems. Both carbon and nitrogen are essential elements for living beings.
Both have different reservoirs and means of movement. Generally, nitrogen travels from the
air, into soil, then into plants, then into other living organisms, and eventually back to the
atmosphere. Nitrogen from the atmosphere must be fixed so plants can use it. Carbon cycles in
much the same way as nitrogen. One important difference is that carbon is stored in the
lithosphere in fossil fuels. When people burn these fuels the carbon becomes carbon dioxide in
the atmosphere. This contributes to climate change.
12
40. What are the two primary limiting factors controlling the coconut’s geographic range?
Low humidity and low temperature.
41. What is artificial selection? How does it relate to domestication? How does the coconut
illustrate the process of artificial selection? Artificial selection occurs when people select
desirable traits from organisms. Organisms like the coconut are domesticated through artificial
selection. Through time, people create versions of these organisms that suit the needs of
people. The modern Cocos nucifera is a product of artificial selection through many years.
Compared to its wild ancestors, as studied in the fossil record, modern coconut seeds are much
larger.
42. Describe the ecological relationship between humans and the coconut. Coconut and
people are an example of mutualism—both parties have benefited in the relationship.
Critical-Thinking Questions
1. The theory of evolution has been a source of controversy between scientists and some
religious groups since it was first developed. Do you find it controversial? Explain. Answers
will vary. Religiously minded students may take offense or they may not. The instructor may
wish to mention that science explains “how” things come about but it does not explain nor
attempt to explain “why” things come about. Framing the issue this way can be very productive
and stimulate class discussion.
2. What trophic level(s) do you occupy? Why is it more efficient for people to eat as primary
consumers rather than as secondary consumers? People with plant-based diets exclusively
(called vegan) are primary consumers. Everybody else is omnivorous and primary, secondary,
tertiary consumers and so on, depending on what the meal is. Because of the loss of 90% of
energy with each higher trophic level, it’s more efficient to feed plants or grain directly to
people rather than to animals first.
3. By definition, when people move an organism to a new place, that organism is non-native.
People brought the coconut to Hawai’i over one thousand years ago. Because that was so
long ago, do you think enough time has passed that we can consider this plant native to
Hawai’i now? Explain. Answers will vary. There is no right answer. Students may stick to the
strict definition that any species brought outside its native geographic range by people is non-
native. They may also argue that enough time has elapsed, several thousand years in some
cases, for the organism to be considered native.
4. Does evolutionary convergence (Section 7.1) form from analogous traits or homologous
traits (Section 7.5)? Explain. It forms from analogous traits. Convergence, by definition, occurs
when two unrelated organisms come to look alike. This can only happen in an analogous
context.
5. Compare natural selection (Section 7.1) with artificial selection (Section 7.6). How are they
the same and how are they different? They are the same in that characteristics or traits are
13
being selected. With natural selection, nature does the selecting. With artificial selection,
people do the selecting.
Test Yourself
1. True or false? When a species disperses on its own to a new location outside its geographic
range, it is considered a native species. (answer: true)
2. True or false? Allen’s rule states that there are more species in the tropics than at higher
latitudes. (answer: false)
3. True or false? After ecological disturbance, ecosystems do not always return to the same
climax community that existed prior to the disturbance. (answer: true)
4. True or false? Many species have more than one common name. (answer: true)
5. Multiple choice: What an organism does to obtain food is called its
a. niche.
b. habitat.
c. prey.
d. community.
(answer: niche)
6. Multiple choice: Which of the following is not an evolutionary adaptation to fire by plants?
a. Serotinous cones
b. Crown sprouting
c. Deep roots
d. Fire-proof leaves
(answer: D)
7. Multiple choice: Which of the following does fire suppression often cause?
a. A small fire
b. A crown fire
c. A surface fire
d. A prescribed burn
(answer: B)
8. Multiple choice. Which of following is not an example of a limiting factor?
a. Temperature
b. Light
c. Predation
d. A scavenger
(answer: D)
9. Fill in the blank: A __________ is a stage of ecological succession following ecological
disturbance. (answer: sere)
10. Fill in the blank: __________ is the term used to describe an interaction between two
species from which both species benefit. (answer: Mutualism)
14
Picture This. Your Turn
1. What trophic level do the copper sharks occupy? The sharks occupy the fourth trophic level
or the tertiary consumer level.
2. Given the 10% rule, how much energy from the primary producers do they receive? They
receive 0.1% of the energy from the primary producers, phytoplankton.
15
Living Physical Geography Questions
1. Why do animals migrate? Animals migrate from areas where resources are in short supply to
areas where they are more abundant.
2. Why do cacti have spines? Cacti have spines to ward off animals trying to take their stores of
water.
3. How are wolves important to ecosystems? Wolves function as keystone species in
ecosystems, and many unrelated species benefit from their presence.
4. Why do coconuts float? Coconuts float because they have evolved to disperse long distances
on ocean currents.
16
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The threidy knit he’s woven wi’,
’Ud fain destroy what sicht he has
O’ this puir transitory stage,
Yet tho’ he kens the fragment is
O’ little worth he e’er can view,
Jalousin’ it’s a cheatrie weed,
He tyauves wi’ a’ his micht and main
To keep his sicht despite his kind
Conspirin’ as their nature is
’Gainst ocht wi’ better sicht than theirs.
O little Life
In which Daith guises and deceives itsel’,
Joy that mak’s Grief a Janus,
Hope that is Despair’s fause-face,
And Guid and Ill that are the same,
Save as the chance licht fa’s!