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Chapter 7—Biotechnology
MULTIPLE CHOICE
1. Restriction enzymes
a. are used to extract DNA from human cells.
b. hold the two strands together in DNA molecules.
c. help initiate transcription.
d. cut DNA at specific sites.
e. splice DNA fragments together.
ANS: D PTS: 1 DIF: Easy REF: p. 149
4. Plasmids
a. are circular DNA molecules present in many bacteria.
b. are a large linear DNA molecule isolated from bacteria.
c. are bacterial cells containing recombinant DNA molecules.
d. are transgenic animals.
e. cannot be cut with restriction enzymes.
ANS: A PTS: 1 DIF: Easy REF: p. 149
5. "Sticky ends"
a. result from DNA replication.
b. are only present in circular molecules.
c. are regions of double-stranded DNA..
d. form when recombinant DNA molecules are formed.
e. are the single-stranded regions at the ends of DNA fragments.
ANS: E PTS: 1 DIF: Easy REF: p. 149
70
Chapter 7 - Biotechnology
8. Restriction enzymes
a. are proteins isolated from certain strains of bacteria that can repair DNA.
b. are enzymes that attach to and cut DNA at specific sequences.
c. recognize and bind to any random sequence in the DNA.
d. are molecules that can help DNA move across the cell wall and enter the bacterial cells.
e. only act on DNA isolated from bacteria.
ANS: B PTS: 1 DIF: Easy REF: p. 149
10. Currently, the benefits of transgenic crops include all of the following EXCEPT
a. resistance to insects.
b. production of human proteins in large quantities.
c. raising the increased nutritional value of crops.
d. resistance to viral or fungal diseases.
e. turning modification of weeds into crops.
ANS: E PTS: 1 DIF: Easy REF: p. 150-153
71
Chapter 7 - Biotechnology
14. Before recombinant DNA techniques became available, human growth hormone was
a. isolated from children who produced excessive amounts of the hormone.
b. recovered from pituitary glands removed from human cadavers.
c. recovered from cow blood.
d. collected from many sources, such as slaughterhouses.
e. isolated from chicken eggs.
ANS: B PTS: 1 DIF: Moderate REF: p. 148
72
Chapter 7 - Biotechnology
16. In the United States, during the period between 1996 and 2006,
a. the use of herbicide tolerant transgenic crops showed the greatest increase.
b. there was a dramatic decrease in the use of transgenic crops.
c. the use of insect resistant transgenic crops showed the greatest increase.
d. the use of some types of transgenic crops increased while the use of others decreased.
e. the planting of transgenic crops was banned.
ANS: A PTS: 1 DIF: Moderate REF: p. 150-153
17. In the United States, during the period between 1996 and 2006, the most common typess of transgenic
crops
a. were tolerant of chemical weed killers.
b. were resistant to insects.
c. were resistant to bacterial infection.
d. were resistant to viral diseases.
e. had increased nutritional value.
ANS: A PTS: 1 DIF: Moderate REF: p. 150-153
21. Stem cells have been used to treat certain leukemias for decades. These stem cells are
a. isolated from donated blood.
b. pluripotent.
c. isolated from developing embryos.
d. capable of becoming many different types of cells.
e. multipotent.
ANS: E PTS: 1 DIF: Difficult REF: p. 156
73
Chapter 7 - Biotechnology
26. Before biotechnology was used, the main problem with clotting factor isolated from donated blood
was
a. there was too little clotting factor present in each sample.
b. contamination by HIV.
c. clotting factor from one individual would not work in another individual.
d. contamination by bacteria.
e. the process led to destruction of the factor.
ANS: B PTS: 1 DIF: Easy REF: p. 148
TRUE/FALSE
1. Recombinant DNA is created when DNA from two or more different organisms is joined together.
2. A transgenic organism is an animal or plant that carries a gene from another species.
3. Transgenic plants have been developed with resistance to herbicides, insects, and viral, bacterial, and
fungal diseases.
74
Chapter 7 - Biotechnology
4. As gene transfer technology becomes more sophisticated, novel combinations of traits may be
developed, requiring specific management plans.
5. Humans have only been able to genetically modify agricultural plants and animals since recombinant
DNA technology was developed.
6. Animal models can be used to screen drugs to identify those that improve symptoms of different
diseases.
7. Embryonic stem cells can only form a limited number of cell types.
8. Currently, adult stem cells can only form a limited number of cell types.
9. Currently, the most common form of stem cell therapy uses adult stem cells.
10. Currently, gene therapy is a common form of treatment for many genetic disorders.
MATCHING
75
Chapter 7 - Biotechnology
ESSAY
1. What are restriction enzymes, plasmids, and recombinant DNA molecules? Describe the steps
involved in the production of recombinant DNA molecules.
ANS:
A restriction enzyme is a protein that cuts the DNA into fragments at specific sites. Plasmids are small,
circular DNA molecules found in many species of bacteria. Researchers have genetically modified
plasmids to create carrier molecules called vectors that are used in recombinant DNA technology.
Recombinant DNA can be defined as the combination of DNA from two or more different organisms.
The steps involved in producing recombinant DNA molecules:
• Identify the gene of interest.
• Extract DNA from cells containing the gene.
• Treat the DNA with a restriction enzyme. Often these cuts leave an overhanging single-stranded
region called a “sticky end”.
• Cut plasmid DNA with the same restriction enzyme.
• The DNA fragments from the cells containing the gene and fragments of plasmid DNA are mixed
and spliced together to form recombinant DNA molecules).
2. What are GM crops? Describe the advantages and concerns with the use of these crops.
ANS:
GM (genetically-modified) crops contain genes from another organism. In agriculture, transgenic
plants are used directly as crops with nutritional enhancements or herbicide resistance. Answers
dealing with the advantages and concerns with the use of GM crops will vary but should note that
genetic modifications allow us to make crop plants that are resistant to herbicides, pests, and disease,
and to change nutritional value, chemical content, growing season, and crop yields. Concerns have
been raised about the safety and environmental impact of transgenic crops, safety and labeling, the
rights of farmers, loss of traditional crops, and biodiversity, among others.
76
Chapter 7 - Biotechnology
SHORT ANSWER
1. What advantages has biotechnology provided over previous methods to produce genetically modified
crops?
ANS:
Humans have been genetically modifying agricultural plants and animals for more than 10,000 years
using selective breeding and crossbreeding to produce the diversity of domesticated plants and animals
we depend on for our food. Biotechnology has changed only the way and the rate at which these
changes are made. It has increased the specificity and predictability of changes that can be made and
also has expanded the range of species that can donate genes.
2. Distinguish between embryonic stem cells and adult stem cells. Make sure to include information on
the source of each cell, potential to form different cell types, and their current and potential uses.
ANS:
Embryonic stem cells (ESCs) are isolated from a developing embryo, and are called pluripotent
because they are able to form so many different cell types, including muscle, bone, skin, and nerve
tissue. Embryonic stem cells are being studied to understand the early stages of the disease process and
to develop therapies to treat diseases. Adult stem cells are found in and isolated from many tissues of
the adult body. These stem cells can also generate different cell types to replace those that are worn out
or damaged. However, adult stem cells are called multipotent because they can form only a limited
number of cell types. Adult stem cells have been used for several decades to treat diseases. One type of
adult stem found in bone marrow divides to form new blood cells to replace those that die, such as in
leukemia. Many biotech companies are developing therapies based on the use of embryonic and adult
stem cells. Many of these therapies may lead to treatment for conditions including spinal cord injury,
regrowth of burned tissue, and damaged heart tissue. Several clinical trials are now under way to
determine whether embryonic stem cells can be used to treat diseases.
3. What is gene therapy? What are technical problems that must be overcome before gene therapy
becomes a common treatment for genetic diseases?
ANS:
Gene therapy involves the placement of normal, functioning genes in a person who carries one or two
mutant alleles of that gene and having the normal allele or alleles make the correct protein. However,
two major problems exist. First, after a gene is isolated, it must be delivered to target cells and inserted
into a chromosome. Then the gene must be switched on to direct the synthesis of enough of the normal
protein to correct the genetic disorder. Unfortunately, although some gene therapy clinical trials have
been partially successful, others have been tragic.
77
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IMPERIAL GAGE
Prunus domestica
1. Am. Gard. Cal. 588. 1806. 2. Prince Treat. Hort. 25, 26. 1828. 3.
Lond. Hort. Soc. Cat. 147, 148. 1831. 4. Prince Pom. Man. 2:56. 1832. 5.
Kenrick Am. Orch. 209. 1835. 6. Mag. Hort. 6:123. 1840. 7. Cultivator
10:167 fig. 1843. 8. Downing Fr. Trees Am. 278 fig. 107. 1845. 9. Floy-
Lindley Guide Orch. Gard. 302, 383. 1846. 10. N. Y. Agr. Soc. Rpt. 343 fig.
1847. 11. Thomas Am. Fruit Cult. 325, 326 fig. 254. 1849. 12. Mag. Hort.
16:454. 1850. 13. Am. Pom. Soc. Cat. 54. 1852. 14. Elliott Fr. Book 411.
1854. 15. U. S. Pat. Off. Rpt. 148, Pl. 5 fig. 1. 1864. 16. Barry Fr. Garden
413. 1883. 17. Mathieu Nom. Pom. 443. 1889. 18. Wickson Cal. Fruits
355. 1891. 19. Mich. Sta. Bul. 103:34. 1894. 20. Guide Prat. 154, 364.
1895. 21. Cornell Sta. Bul. 131:187. 1897. 22. Va. Sta. Bul. 134:42. 1902.
23. Ohio Sta. Bul. 162:239, 254, 255. 1905.
Flushing Gage 7, 8, 11, 12, 14, 15, 17, 20. Flushing Gage 3, 9. Harper
22. Imperial Gage 17, 20. Imperial Green Gage 7. Jenkinson’s Imperial 6,
14, 15. Prince’s Gage 4, 12, 17. Prince’s Imperial Gage 4, 5, 6, 10.
Prinzens Kaiser Reine Claude 20. Prince’s Kaiser Reine-Claude 17.
Prince’s Imperial Gage 8, 11, 14, 15, 17, 20. Prince’s White Gage 4, 12,
17. Reine-Claude de Flushings 20. Reine-Claude Imperiale 20. Reine-
Claude Imperiale 17. Reine-Claude Verte Imperiale 17. Reine-Claude
Imperiale de Prince 17, 20. Reine-Claude Blanche de Boston 17, 20.
Reine-Claude Verte Superieure 20. Superior Gage 9. Superior Green
Gage 12, 14, 15, 17, 20. Superior Green Gage? 3. Superiour Green Gage
8. White Gage? 1, 2, 20. White Gage 14, 15. White Gage of Boston 7, 8,
11, 17.
ITALIAN PRUNE
ITALIAN PRUNE
Prunus domestica
1. Lond. Hort. Soc. Cat. 152. 1831. 2. Prince Pom. Man. 2:78. 1832. 3.
Kenrick Am. Orch. 262. 1832. 4. Manning Book of Fruits 106. 1838. 5. Am.
Pom. Soc. Cat. 214, 220. 1836. 6. Downing Fr. Trees Am. 381. 1857. 7.
Cultivator 8:52 fig. 1860. 8. Hogg Fruit Man. 366. 1866. 9. Downing Fr.
Trees Am. 924. 1869. 10. Pom. France 7: No. 22, fig. 1871. 11. Mas Le
Verger 6:69, fig. 35. 1866-73. 12. Oberdieck Deut. Obst. Sort. 442. 1881.
13. Lauche Deut. Pom. No. 2, Pl. 4, 22. 1882. 14. Barry Fr. Garden 412.
1883. 15. Cat. Cong. Pom. France 360. 1887. 16. Mathieu Nom. Pom.
436. 1889. 17. Wickson Cal. Fruits 358. 1891. 18. Guide Prat. 155, 362.
1895. 19. Oregon Sta. Bul. 45:23 fig. 1897. 20. Cornell Sta. Bul. 131:187,
fig. 44. 1897. 21. Wash. Sta. Bul. 38:7, 8. 1899. 22. W. N. Y. Hort. Soc.
Rpt. 44:92. 1899. 23. Waugh Plum Cult. 111 fig. 1901. 24. U. S. D. A. Div.
Pom. Bul. 10:6. 1901. 25. Mass. Sta. An. Rpt. 17:158. 1905.
Altesse Double 8, 9, 10, 15, 18. August Zwetsche 16. Auguste
Zwetsche 10, 18. Blaue Riesenzwetsche 16, 18. Bleue d’Italie 15.
Couetsche d’Italie 18. Couetsche Fellenberg 10, 18. D’Italie 18. Double
Blackpruim 16, 18. Fausse Altesse 16, 18. Fellemberg 14. Fellemberg 8,
16, 18. Fellenberg 5, 9, 11, 16, 18, 19, 20, 23. Fellenburg 22. Fellenberg
5, 6, 7, 9, 17. Fellenburg 25. Feltemberg 10, 18. Fellenberg Quetsche 16,
18. Fellenberger Zwetsche 12, 13, 16, 18. Grosse Früh Zwetsche? 16.
German Prune 19, 22. Italienische Blaue Zwetsche 11, 16. Italianische
blaue zwetsche 18. Italian Guetsche 10, 18. Italian Prune 6, 7, 10, 11, 16,
17, 18, 25. Italian Quetsche 8, 9, 15, 16. Italianische Zwetsche 18.
Italienische Pflaumen Zwetsche 16. Italienische Zwetsche 11, 13.
Italianische Zwetsche 10. Italianische blanc Zwetsche 10. Italienische
Zwetsche 12, 13, 16. Large German Prune 17. Prune d’Italie 8, 9, 10, 16.
Pflaume mit dem Pfirschenblatt 18. Pflaume Mit dem Pfirsichblatt 16.
Prune Suisse 6. Quetsche 18. Quetsche Bleue d’Italie 10, 11, 16, 18.
Quetsche d’Italie 1, 10, 11, 15, 18. Quetsche d’Italie 3, 8, 9, 11, 16.
Schweizer Zwetsche 12, 13, 16. Schweizerzwetsche 18. Swiss Prune 17,
19, 22. Semiana 8, 10, 16, 18. Turkish Prune 22. Zwetsche von Dätlikon
16, 18.
The Italian Prune is one of the most widely grown of all plums. Its
home is Italy and it is grown in all of the plum regions of continental
Europe; is well known in England; is third or fourth in popularity in
the Atlantic States of America; is by long odds the leading plum in
the Pacific Northwest where it is chiefly used in prune-making and is
grown somewhat for prunes and for shipping green in California.
There are several reasons why this plum is so popular. To begin
with, it is finely flavored whether eaten out of hand, stewed or
otherwise prepared for the table or cured as a prune. The fruit is a
little too tart to be ranked as a first-rate dessert plum and yet it is one
of the best of the prunes for this purpose, though it must be fully ripe
to be fit for dessert; in cooking it changes to a dark, wine color, very
attractive in appearance, with a most pleasant, sprightly flavor; as a
cured prune the flesh is firm and meaty, yet elastic, of good color and
a perfect freestone, making when cooked the same attractive
looking, fine-flavored, sprightly sauce to be had from the green fruits;
the prunes from this variety, too, are noted for long-keeping. In the
uncured state the variety keeps and ships well. The trees are usually
large, hardy, productive, well formed and bear regularly; yet they are
not ideal and the variety fails chiefly in tree-characters. The trees are
often capricious to soil and climate, do not always bear well, seem to
be susceptible to diseases, are preyed upon by insects and suffer in
particular from dry or hot weather. Were all of these troubles of the
tree to befall the variety at one time it would of necessity give way to
better sorts, but happily they are to be found for most part in illy-
adapted conditions or in certain seasons; the Italian Prune well cared
for in locations to which it is suited must long remain one of the
leading plums despite the faults of the trees.
The Italian Prune originated in Italy at least a century ago and has
long been common in northern Italy, especially in the vicinity of
Milan. The London Horticultural Society catalog for 1831 first
mentions it in England and the following year it was described in
America by Prince as an excellent prune recently introduced from
Europe. The American Pomological Society recommended it in 1856
as worthy of further testing and in 1862 it was added to the fruit
catalog list of this society. The origin of the name Fellenberg, a very
common synonym, is explained by Lauche[218] who says: “It came to
Germany through a Mr. Fellenberg and is therefore spread under his
name and also under the names Schweizerzwetsche and
Fellenberger Zwetsche.” He further adds that the variety “is still not
known in Germany as it deserves, considering its quality, size and
productiveness.”
JAPEX
Triflora ×
JEFFERSON
JEFFERSON
Prunus domestica
1. Downing Fr. Trees Am. 279, 280 fig. 108. 1845, 2. Horticulturist 1:11,
93. 1846. 3. Floy-Lindley Guide Orch. Gard. 420. 1846. 4. Thomas Am.
Fruit Cult. 325, 326, fig. 251. 1849. 5. Mag. Hort. 16:453 fig. 25. 1850. 6.
Hovey Fr. Am. 2:1, Pl. 1851. 7. Am. Pom. Soc. Cat. 54. 1852. 8. Elliott Fr.
Book 411. 1854. 9. Thompson Gard. Ass’t 518, Pl. 1. 1859. 10. Mas Le
Verger 6:17, Pl. 9. 1866-73. 11. Pom. France 7: No. 28. 1871. 12. Hogg
Fruit Man. 707. 1884. 13. Gaucher Pom. Prak. Obst. No. 95, Col. Pl. 1894.
14. Cornell Sta. Bul. 131:188. 1897.
Bingham incor. 2, 8. Prune Jefferson 11.
JUICY
JUICY
1. Burbank Cat. 20. 1893. 2. Cal. State Bd. Hort. 53. 1897. 3. Vt. Sta.
Bul. 67:15. 1898. 4. Ohio Sta. Bul. 113:161. 1899. 5. Conn. Pom. Soc.
Rpt. 155. 1900. 6. Ohio Sta. Bul. 162:256. 257. 1905. 7. Mass. Sta. An.
Rpt. 17:161. 1905.
KELSEY
Prunus triflora
KERR
Prunus triflora
1. Ga. Hort. Soc. Rpt. 52. 1889. 2. Am. Gard. 12:307, 501. 1891. 3. Ibid.
13:700. 1892. 4. Kerr Cat. 1894. 5. Cornell Sta. Bul. 62:25. 1894. 6. Ga.
Hort. Soc. Rpt. 95. 1895. 7. Ala. Sta. Bul. 85:443. 1897. 8. Am. Pom. Soc.
Cat. 41. 1899. 9. Cornell Sta. Bul. 175:136. 1899. 10. Waugh Plum Cult.
137. 1901. 11. Ga. Hort. Soc. Rpt. XIII. 1904. 12. Ohio Sta. Bul. 162:256,
257. 1905.
Hattankio 1. Hattankio 7. Hattankin No. 2. 2, 3. Hattonkin No. 2. 4, 5, 10.
Hattonkin 9. Hattankio No. 2. 6, 11. Hattankio Oblong 9, 11. Hattankio 10.
Hattan 10. Hytankio 10. Hytan-Kayo 10.
Kerr is about the best of the yellow Trifloras and is one of the best
of all early plums of its species. It is very productive, sometimes
over-bearing, and should always be thinned. The quality of the plums
is good and the fruits are attractive in appearance. The faults of the
variety are that the fruits drop as they ripen, though they color if
picked green, and in some localities the tree-characters are poor.
This variety was imported from Japan by Frost and Burgess,
Riverside, California, and was distributed under the group name
Hattankio No. 2 or Hattonkin No. 2. As Georgeson was also
distributed under the same name, though under a different number,
confusion resulted. To better distinguish between the two, L. H.
Bailey, in 1894, named Hattonkin No. 2 Kerr, in honor of J. W. Kerr,
the noted plum specialist, of Denton, Maryland. In 1899 the variety
was placed on the fruit catalog list of the American Pomological
Society. The following description is compiled.
KING DAMSON
Prunus insititia
1. Watkins Nur. Cat. 48. 1892?. 2. Am. Gard. 14:146, 147. 1893. 3.
Garden 53:265. 1898. 4. Can. Exp. Farm Bul. 2nd Ser. 3:51. 1900. 5.
Thompson Gard. Ass’t 4:161. 1901.
Bradley’s King 5. Bradley’s King of Damsons 3. King of Damsons 1, 2.
The fruit of King Damson runs large for a Damson and the flavor is
agreeable, so agreeable that the variety is really a very good dessert
fruit late in the season. This Damson is little grown in America and
deserves much wider cultivation. A peculiarity of the plum is that
there is always more or less doubling of the petals. Very little is
known regarding the history of this excellent variety, but it seems
probable that it originated in Kent, England, where it is much grown.
KIRKE
Prunus domestica
1. Pom. Mag. 3:111, Pl. 1830. 2. Lond. Hort. Soc. Cat. 149. 1831. 3.
Kenrick Am. Orch. 263. 1832. 4. Downing Fr. Trees Am. 306. 1845. 5.
Floy-Lindley Guide Orch. Gard. 281, 382. 1846. 6. Mag. Hort. 15:488 fig.
43. 1849. 7. Thompson Gard. Ass’t 518, Pl. 1. 1859. 8. Mas Le Verger
6:15, fig. 8. 1866-73. 9. Am. Pom. Soc. Cat. 36. 1875. 10. Pom. France 7:
No. 26. 1871. 11. Flor. & Pom. 47. 1876. 12. Oberdieck Deut. Obst. Sort.
430. 1881. 13. Lauche Deut. Pom. 16, Pl. IV. 1882. 14. Hogg Fruit Man.
708. 1884. 15. Guide Prat. 154, 358. 1895. 16. Gard. Chron. 24:19. 1898.
17. Gaucher Pom. Prak. Obst. No. 96, Col. Pl. 1894. 18. Rev. Hort. 500.
1898. 19. Soc. Nat. Hort. France Pom. 536. 1904.
De Kirke 15. Kirke’s 2, 4, 6, 7, 9, 14, 17. Kirke’s 8, 10, 15, 17. Kirke’s
Pflaume 12, 13. Kirke’s Pflaume 8, 10, 15, 17. Kirke’s Plum 1, 5, 8, 10, 11,
16, 18. Kirk’s Plum 3, 5. Kirke 17. Kirke’s Plum 15, 17, 19. Prune de Kirke
18. Prune de Kirke 8, 10, 17. Prune Kirke 19.
LAFAYETTE
Prunus domestica
1. Prince Pom. Man. 2:96. 1832. 2. Tucker’s Gen. Farmer 3:153. 1839.
3. Elliott Fr. Book 427. 1854. 4. Am. Pom. Soc. Rpt. 222, 244. 1858. 5.
Hogg Fruit Man. 368. 1866. 6. Downing Fr. Trees Am. 916. 1869. 7. Guide
Prat. 160, 359. 1895.
Gifford’s Lafayette 1, 4, 6. Gifford’s La Fayette 2, 3.
LAIRE
Prunus orthosepala?
Laire is cultivated locally in Rooks and neighboring counties in
Kansas and is highly spoken of by those who grow it. The
description of the variety is made from information sent from the
United States Department of Agriculture. For a further account of this
plum the reader is referred to the discussion of Prunus orthosepala,
page 97. The name is derived from that of the man who first brought
the plums under cultivation some twenty or twenty-five years ago.