Evolution II

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30 Evolution II 30.

1 Mechanism of evolution

30 Evolution II

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30 Evolution II

Think about…
30.1Mechanism of evolution
30.2Speciation
Recall Think about…
Concept map

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30 Evolution II

The largest living tortoise


Galapagos
Islands

South
America
native to

Galapagos tortoise

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30 Evolution II

The largest living tortoise


On different islands
 different kinds of Galapagos tortoises

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30 Evolution II

The largest living tortoise


saddle-shaped shell

limited ground
vegetation
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30 Evolution II

The largest living tortoise


saddle-shaped shell

allows neck to move


upwards

tortoise can feed on limited ground


tall vegetation vegetation
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30 Evolution II

The largest living tortoise


dome-shaped shell

plentiful ground
vegetation
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30 Evolution II

The largest living tortoise


dome-shaped shell

no need to move the


neck upwards to feed plentiful ground
on tall vegetation vegetation
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30 Evolution II

The largest living tortoise


Online video:Evo
lution of the gian
t tortoises

Some scientists believe


that they have evolved
from a common ancestor South
living in the mainland of America
South America.

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30 Evolution II

How might different kinds of


Galapagos tortoises arise from a
common ancestor?

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30 Evolution II

When do scientists consider


different groups of organisms to
be different species?

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30 Evolution II 30.1 Mechanism of evolution

30.1 Mechanism of evolution


How does evolution take place?
Lamarck’s theory
(Lamarckism 拉馬克學說 )
Darwin’s theory
(Darwinism 達爾文主義 )

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
• proposed in 1809 by:
Jean Baptiste
Lamarck

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
He suggested that:
1 environmental changes created needs that
caused organisms to modify their existing
structures to meet these needs.

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
He suggested that:
existing structures
repeatedly not used
used
well
degenerate
developed

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
He suggested that:
2 acquired characters ( 後天獲得的性狀 )
would be passed on to later generations.

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
Evolution of the giraffe’s neck:

short-necked
ancestors

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
Environmental changes created needs for
organisms to develop new features.

stretched their
necks to get
leaves on trees no grass

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
Environmental changes created needs for
organisms to develop new features.
neck became
longer due to
repeated use

acquired character
passed on to offspring
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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
After many generations…

extremely
long neck

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
• August Weismann did an experiment
to test Lamarck’s idea
tails cut

× ×
mating offspring
with tails

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
• August Weismann did an experiment
to test Lamarck’s idea
tails cut again

× ×
offspring
with tails

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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
• August Weismann did an experiment
to test Lamarck’s idea
process repeated

× ×
new generations were
born with complete tails
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30 Evolution II 30.1 Mechanism of evolution

A Lamarckism
• showed that acquired characters were
NOT passed on to later generations

disproved Lamarckism

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism Online video:D


arwin’s story

• proposed by Charles Darwin and


Alfred Russel Wallace in 1858
• also known as the theory
of natural selection
( 自然選擇 )

Charles Darwin
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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
• Darwin’s ideas came from a visit to
the Galapagos Islands

South
America
• finches on the islands
were very similar to
those on the mainland
of South America
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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
• but the beaks were different
 adapted to different food sources

nuts large flying cacti insect


insects insects larvae
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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
• Darwin suggested that:

all originated from a single species


on the mainland of South America

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
• after the voyage, Darwin worked on his idea
of natural selection for 20 years

published a book titled


On the Origin of Species
in 1859

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
• came from deductions which were made
from a series of observations

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
Observation 1:
Organisms can
produce large
numbers of
offspring, but the
population remains
fairly stable.

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
Observation 1: Deduction:
Organisms can Many fail to survive
produce large and reproduce due to
numbers of competition for limited
offspring, but the resources
population remains  struggle for
fairly stable. existence
( 生存競爭 )

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
Observation 2:
There is variation
among individuals of
the same species.

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
Observation 2: Deduction:
There is variation Individuals with
among individuals of favourable characters
the same species. are better adapted to
the environment
 higher chance of
survival
 survival of the
fittest ( 適者生存 )
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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
Observation 3:
Some favourable
characters are
passed on to
offspring.

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
Observation 3: Deduction:
Some favourable Better adapted
characters are individuals pass their
passed on to favourable characters
offspring. on to offspring.

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
Observation 3: Deduction:
Some favourable Accumulation of
characters are favourable characters
passed on to over many generations
offspring.  individuals quite
different from their
ancestor
 evolution
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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
Animation 30.1

Evolution of the giraffe’s neck:

ancestor of
giraffes

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
Variation in neck length:

long neck
short neck

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism

when weather
became dry

no grass grew

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism

short-necked giraffes
could not eat leaves
from treetops
died

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism

long-necked giraffes could


eat leaves from treetops

survived

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism

long-necked

 favourable
character for survival
 passed on to offspring

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism
After many generations…
long-necked giraffes
predominated in the
population

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism

Different

ancestor modern giraffes

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30 Evolution II 30.1 Mechanism of evolution

B Darwinism

Evolution

ancestor modern giraffes

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30 Evolution II 30.1 Mechanism of evolution

Natural selection in action:


i) Peppered moths ( 斑點蛾 )
ii) Insecticide-resistant insect pests
iii)Antibiotic-resistant bacteria

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30 Evolution II 30.1 Mechanism of evolution

i) Peppered moths
Variations exist among moths:

black peppered
moth (mutant 突變
型)
white peppered moth
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30 Evolution II 30.1 Mechanism of evolution

i) Peppered moths
Before industrialization
easily seen by birds

camouflaged ( 偽
裝)
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30 Evolution II 30.1 Mechanism of evolution

i) Peppered moths
Before industrialization
easily seen by birds

eaten by birds

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30 Evolution II 30.1 Mechanism of evolution

i) Peppered moths
During industrialization

tree trunks were


darkened

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30 Evolution II 30.1 Mechanism of evolution

i) Peppered moths
During industrialization
easily seen by birds

camouflaged

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30 Evolution II 30.1 Mechanism of evolution

i) Peppered moths
During industrialization
easily seen by birds

eaten by birds

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30 Evolution II 30.1 Mechanism of evolution

i) Peppered moths
During industrialization
higher chance of
survival and
reproduction

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30 Evolution II 30.1 Mechanism of evolution

i) Peppered moths
During industrialization
higher chance of
survival and
reproduction

increase in
population in
polluted areas
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30 Evolution II 30.1 Mechanism of evolution

ii) Insecticide-resistant insect pests


• DDT, an insecticide, was used in the
1940s to kill mosquitoes
• but many mosquitoes became
resistant to DDT within a few years

Why?
Online video:
Historical clips
on DDT

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30 Evolution II 30.1 Mechanism of evolution

ii) Insecticide-resistant insect pests


Genetic variations exist among
mosquitoes:

resistant
to DDT
non-resistant
to DDT
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30 Evolution II 30.1 Mechanism of evolution

ii) Insecticide-resistant insect pests


When DDT was used:

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30 Evolution II 30.1 Mechanism of evolution

ii) Insecticide-resistant insect pests


When DDT was used:
most non-resistant
mosquitoes died

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30 Evolution II 30.1 Mechanism of evolution

ii) Insecticide-resistant insect pests


When DDT was used:
higher chance of
survival and
reproduction

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30 Evolution II 30.1 Mechanism of evolution

ii) Insecticide-resistant insect pests


When DDT was used:

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30 Evolution II 30.1 Mechanism of evolution

ii) Insecticide-resistant insect pests


Repeated use of DDT:
proportion of
DDT-resistant
mosquitoes 

DDT becomes
less effective
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30 Evolution II 30.1 Mechanism of evolution

iii) Antibiotic-resistant bacteria


Genetic variations exist among bacteria:
resistant
non-resistant

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30 Evolution II 30.1 Mechanism of evolution

iii) Antibiotic-resistant bacteria


When antibiotic is used:
died

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30 Evolution II 30.1 Mechanism of evolution

iii) Antibiotic-resistant bacteria


When antibiotic is used:
higher chance of survival
and reproduction

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30 Evolution II 30.1 Mechanism of evolution

iii) Antibiotic-resistant bacteria


Indiscriminate use of antibiotic:
proportion of resistant bacteria 

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30 Evolution II 30.1 Mechanism of evolution

iii) Antibiotic-resistant bacteria


Example: Staphylococcus aureus
( 金黃葡萄球菌 )
• some strains resistant
to penicillin and
methicillin

Online video:
Safe use of
antibiotics (×10 000)
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30 Evolution II 30.1 Mechanism of evolution

C Neo-Darwinism
• with more knowledge in genetics and
related areas of biology,
e.g. mutations cause variations
• Darwinism is refined as
neo-Darwinism ( 新達爾文主
義)

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30 Evolution II 30.1 Mechanism of evolution

1 Charles Darwin put forward the


theory of natural selection
to explain how evolution takes
place.

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30 Evolution II 30.1 Mechanism of evolution

1 The theory suggests that


variations exist among individuals
within a species. The individuals
compete with one another for
limited resources (i.e. struggle
for existence ).
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30 Evolution II 30.1 Mechanism of evolution

1 When the environment changes,


individuals with favourable
characters have a higher chance
of survival and (i.e.
reproduction
survival
). of the fittest

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30 Evolution II 30.1 Mechanism of evolution

1 They pass the favourable


characters on to their offspring .
Over many generations, the action
of natural selection leads to the
evolution of new species.

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30 Evolution II 30.1 Mechanism of evolution

2 Lamarck’s theory Darwin’s theory


1 Environmental 1 Variations
change creates already exist
a needfor within species.
organisms to
change.

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30 Evolution II 30.1 Mechanism of evolution

2 Lamarck’s theory Darwin’s theory


2 In order to survive, 2 The environment
individuals can selects those
develop new individuals with
features in
favourable
response to the
characters to
environmental
change. survive, and
eliminates others.
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30 Evolution II 30.1 Mechanism of evolution

2 Lamarck’s theory Darwin’s theory


3 Acquired 3 Individuals with
characters are favourable
passed on to later characters have a
generations, higher chance of
making them quite survival and
different from their reproduction .
ancestor.

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30 Evolution II 30.1 Mechanism of evolution

2 Lamarck’s theory Darwin’s theory


3 Acquired 3 The offspring
characters are inherit these
passed on to later characters, making
generations, them quite different
making them quite from their ancestor.
different from their
ancestor.

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30 Evolution II 30.1 Mechanism of evolution

3 Genetic variations exist among


insect pests in their resistance to the
insecticide. When the insecticide is
applied, the insecticide-resistant insect
pests have a higher chance of survival
and reproduction.

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30 Evolution II 30.1 Mechanism of evolution

3 When the insecticide is used


repeatedly, the proportion of the
resistant insect pests in the population
increases in subsequent
generations.

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30 Evolution II 30.1 Mechanism of evolution

3 As a result, the insecticide becomes


less effective in killing insect pests.

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30 Evolution II 30.1 Mechanism of evolution

4 Genetic variations exist among


bacteria in their resistance to a certain
antibiotic. When the antibiotic is used,
the antibiotic-resistant bacteria have
a higher chance of survival and
reproduction.

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30 Evolution II 30.1 Mechanism of evolution

4 With the indiscriminate use of the


antibiotic, the proportion of the
resistant bacteria in the population
increases in subsequent generations.

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30 Evolution II 30.2 Speciation

30.2 Speciation
• the process of forming new species
• in the 1940s…
I put forward
a hypothesis to
explain speciation.

Ernst Mayr
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30 Evolution II 30.2 Speciation

1 A single species of tortoise that can


interbreed freely. Animation 30.2

same
species
forest
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30 Evolution II 30.2 Speciation

2
mountain range forms

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30 Evolution II 30.2 Speciation

2 Two groups are isolated ( 隔離 ).

group A group B

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30 Evolution II 30.2 Speciation

2 They cannot meet and interbreed.

gene flow stops

group A
 group B

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30 Evolution II 30.2 Speciation

2 Mutations within each group produce


different genetic variations.

group A
 group B

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30 Evolution II 30.2 Speciation

2 Climate starts to change.

wetter drier

group A group B

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30 Evolution II 30.2 Speciation

2 Environmental conditions become different.

wetter drier

group A group B

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30 Evolution II 30.2 Speciation

3 Natural selection acts on the two groups


in different ways.

evolve independently

group A group B

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30 Evolution II 30.2 Speciation

4 After many generations…

large genetic differences

group A group B

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30 Evolution II 30.2 Speciation

4 After many generations…

cannot interbreed to
large genetic
produce differences
fertile offspring

group A group B

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30 Evolution II 30.2 Speciation

4 They become two different species.

species
group AA group BB
species

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30 Evolution II 30.2 Speciation

Isolation mechanisms
Geographic isolation ( 地理隔離 )
is one of the mechanisms.
Organisms are separated by
physical barriers.

mountain
ranges deserts
oceans rivers
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30 Evolution II 30.2 Speciation

Geographic isolation
• example: squirrels in Grand Canyon

with a
white tail

without a the Colorado


white tail River

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30 Evolution II 30.2 Speciation

Geographic isolation
• example: squirrels in Grand Canyon

a river formed ancestral population


of the squirrels

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30 Evolution II 30.2 Speciation

Geographic isolation
• example: squirrels in Grand Canyon
two groups of squirrels could not
meet and interbreed
 evolved independently

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30 Evolution II 30.2 Speciation

Geographic isolation
• example: squirrels in Grand Canyon

two distinct species formed

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30 Evolution II 30.2 Speciation

Limpets live on rocky shores. They attach


themselves firmly to rocks to avoid being washed
away by wave action. Limpets have a cone-shaped
shell.
limpet

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30 Evolution II 30.2 Speciation

The height-length ratio of the shell varies with the


degree of exposure to wave action.

limpet

heigh
t
length

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30 Evolution II 30.2 Speciation

A student measures the height-length ratios of the


shells of two groups of limpets. These two groups
are living on two different shores, X and Y, in the
same area.
limpet

heigh
t
length

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30 Evolution II 30.2 Speciation

limpet
The mean values of the
ratios are shown in the
table below. heigh
length t

Mean value of the height-length


Shore
ratio of the shell
X 0.42
Y 0.24
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30 Evolution II 30.2 Speciation

a Which shore (X or Y) is more likely to be


exposed to stronger wave action? Explain with
reference to the concept of natural selection.
(7
marks)
Mean value of the height-length
Shore
ratio of the shell
X 0.42
Y 0.24
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30 Evolution II 30.2 Speciation

Answer

a Shore Y (1)
Compared with the limpets on shore X, the
limpets on shore Y have a smaller height-length
ratio, showing that their shells are flatter. (1)

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30 Evolution II 30.2 Speciation

Answer

a Limpets with a flatter shell are better adapted to


the environment (1)
as this character reduces the chance of them
being washed away by wave action. (1)

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30 Evolution II 30.2 Speciation

Answer

a Thus, limpets with a flatter shell have a higher


chance of survival. (1)
They breed and pass this character on to their
offspring. (1)
Over many generations, they predominate in the
population on shore Y. (1)

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30 Evolution II 30.2 Speciation

b Limpets are slow-moving. Those living on


shore X are unlikely to mate with those living
on shore Y. However, they are not considered
as geographically isolated. Suggest one
possible reason for this.
(Hint: Limpets spend a short period of their life
as free-swimming larvae before settling on
rock.) (3
marks)
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30 Evolution II 30.2 Speciation

Answer

b The limpet larvae may be carried by ocean


currents from shore X to shore Y, or the other
way round. (1)
The larvae settle on the shores and grow to
maturity and interbreed. (1)
Thus, there is still gene flow between the two
groups of limpets. (1)
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30 Evolution II 30.2 Speciation

Answer

b The limpet larvae may be carried by ocean


Different groups of organisms in
currents from shore X to shore Y, or the other
a
way round. population
(1) must be reproductively
isolated (i.e. they can no longer
The larvaeinterbreed)
settle on thebefore
shores evolving
and grow to
into
maturity and interbreed. (1)
new species.
Thus, there is still gene flow between the two
groups of limpets. (1)
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30 Evolution II 30.2 Speciation

1 Speciation is the process by


which new species form.

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30 Evolution II 30.2 Speciation

2 Speciation occurs when a


population separates into small
groups. The groups are isolated
from one another (e.g. by a
physical barrier) so that they
can / cannot interbreed.
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30 Evolution II 30.2 Speciation

2 Each group may then evolve


independently by
natural selection.

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30 Evolution II 30.2 Speciation

2 After many generations, genetic


variations may accumulate to a
point where individuals of different
isolated groups can no longer
interbreed to produce fertile
offspring.
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30 Evolution II 30.2 Speciation

2 The isolated groups become


different species.

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30 Evolution II

1 How might different kinds of


Galapagos tortoises arise from a
common ancestor?

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30 Evolution II

Individuals from the mainland of


South America arrived in the different
islands of Galapagos.
They were geographically isolated
from one another by the ocean.

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30 Evolution II

Mutations took place in different


groups of tortoises which produced
offspring with different genetic
variations.

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30 Evolution II

Individuals, which were better adapted


to the new environment, had a higher
chance of survival and reproduction.
They passed the favourable characters
on to their offspring.
Over many generations, different kinds
of tortoises with different characters
formed.
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30 Evolution II

2 When do scientists consider different


groups of organisms to be different
species?

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30 Evolution II

When different groups of organisms


cannot interbreed to produce fertile
offspring, they are considered as
different species.

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30 Evolution II

Evolution
can be explained by
theory of natural
proposed selection
acts on
by
Darwin and individuals
Wallace with variations

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30 Evolution II

individuals with
variations
may be

individuals with individuals without


favourable characters favourable characters

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30 Evolution II

individuals with individuals without


favourable characters favourable characters
have have
higher chance of survival lower chance of survival
and reproduction and reproduction
leads to leads to
population of individuals population of individuals
with favourable without favourable
characters increases characters decreases
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30 Evolution II

individuals with
variations
may become
isolated
populations
may give rise to
new species

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